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404 lines
92 KiB
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<document id="D87C18A4F2DD82A4CB02D6FA0F243338" ID-CLB-Dataset="3587" ID-DOI="10.5252/geodiversitas2021v43a20" ID-GBIF-Dataset="8e41f2cf-1620-45db-847c-54fc788f51ad" ID-ISSN="1638-9395" ID-Zenodo-Dep="5679113" ID-ZooBank="urn:lsid:zoobank.org:pub:04F82471-9F26-4935-97CE-1A286E08C958" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1636557091778" checkinUser="felipe" docAuthor="Luján, Àngel H., Čerňanský, Andrej, Bonilla-Salomón, Isaac, Březina, Jakub & Ivanov, Martin" docDate="2021" docId="523F87B8FF9EDC55FF09F9A4E31CF8D0" docLanguage="en" docName="Geodiversitas.43.20.691-707.pdf" docOrigin="Geodiversitas 43 (20)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Testudo (Chersine) kalksburgensis Toula 1896" docType="treatment" docVersion="8" lastPageNumber="700" masterDocId="AE06FFC0FF94DC5EFFA7FFFCE63EFF9C" masterDocTitle="Fossil turtles from the early Miocene localities of Mokrá-Quarry (Burdigalian, MN 4), South Moravian Region, Czech Republic" masterLastPageNumber="707" masterPageNumber="691" pageNumber="699" updateTime="1699469340771" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="178266E26C38FD2DA28BBE001DAAB92E">Fossil turtles from the early Miocene localities of Mokrá-Quarry (Burdigalian, MN 4), South Moravian Region, Czech Republic</mods:title>
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<mods:name id="382D9D85E39A79EEDC96F2865149AC3A" type="personal">
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<mods:namePart id="0EDFF30A2B9E9EF59E9AAE74609DAD3F">Luján, Àngel H.</mods:namePart>
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<mods:affiliation id="8AC48E217F69722C35E2025C3D266F1E">Masaryk University, Faculty of Science, Department of Geological Sciences, 611 37 Brno (Czechia) and Comenius University in Bratislava, Faculty of Natural Sciences, Department of Ecology, 842 15 Bratislava (Slovakia) and Institut Català de Paleontologia Miquel Crusafont, Universitat Autònoma de Barcelona, Edifici ICTA, 08193 Cerdanyola del Vallès (Spain)</mods:affiliation>
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<mods:nameIdentifier id="1A3981ABCF9821BD37E45196C9E35033" type="email">angel.lujan@icp.cat</mods:nameIdentifier>
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<mods:namePart id="3AEDE76BCD874B053ED80F816248EA95">Čerňanský, Andrej</mods:namePart>
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<mods:affiliation id="00A710E34ABD9CF8BC802BB759CA5384">Comenius University in Bratislava, Faculty of Natural Sciences, Department of Ecology, 842 15 Bratislava (Slovakia)</mods:affiliation>
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<mods:nameIdentifier id="9FADE3EAB61031E326C3EFB03EAF16BB" type="email">cernansky.paleontology@gmail.com</mods:nameIdentifier>
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<mods:namePart id="293123DFAD755BE0AF0A953C4FF0C67E">Bonilla-Salomón, Isaac</mods:namePart>
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<mods:affiliation id="CB9AC38DB3290DF87C085B7F7D60D72F">Department of Geology and Paleontology, Faculty of Natural Sciences, Comenius University, Mlynská dolina, Ilkovičova, 842 15 Bratislava (Slovakia)</mods:affiliation>
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<mods:nameIdentifier id="F8E394841F20E374B9C3F883E70643A3" type="email">salomon1@uniba.sk</mods:nameIdentifier>
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<mods:namePart id="B379D95120903C71FCE48D4D39ABA882">Březina, Jakub</mods:namePart>
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<mods:affiliation id="12B7FF7BB4B863A040158925C2A733A0">Moravian Museum, Department of Geology and Paleontology, 659 37 Brno (Czechia)</mods:affiliation>
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<mods:nameIdentifier id="26B825774B40ECB8332221CB9C0AD285" type="email">jbrezina@mzm.cz</mods:nameIdentifier>
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<mods:namePart id="2095DADA4BC9BD06D7F467D676583C48">Ivanov, Martin</mods:namePart>
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<mods:affiliation id="222A661F060E83B226CFB735081F9982">Masaryk University, Faculty of Science, Department of Geological Sciences, 611 37 Brno (Czechia)</mods:affiliation>
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<mods:nameIdentifier id="0A7E799EF9DDAF1F184AA133C823FCF5" type="email">mivanov@sci.muni.cz</mods:nameIdentifier>
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<mods:title id="AD5911010DEB943D6300EECF1C57EC80">Geodiversitas</mods:title>
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<mods:part id="C81F97F58E6D511DCFA02C4F0CFE816A">
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<mods:date id="DFA03923C1FD91CDD29ABCAF2AF70CCC">2021</mods:date>
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<treatment id="523F87B8FF9EDC55FF09F9A4E31CF8D0" ID-DOI="http://doi.org/10.5281/zenodo.5700005" ID-GBIF-Taxon="190303793" ID-Zenodo-Dep="5700005" LSID="urn:lsid:plazi:treatment:523F87B8FF9EDC55FF09F9A4E31CF8D0" httpUri="http://treatment.plazi.org/id/523F87B8FF9EDC55FF09F9A4E31CF8D0" lastPageId="11" lastPageNumber="700" pageId="10" pageNumber="699">
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<subSubSection id="928C6525FF9EDC54FF09F9A4E7C1F90F" pageId="10" pageNumber="699" type="materials_examined">
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<subSubSection id="928C6525FF9EDC54FF09F9A4E4E3F9EF" box="[174,733,1624,1651]" pageId="10" pageNumber="699" type="nomenclature">
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<taxonomicName id="1D964D2DFF9EDC54FF09F9A4E4E3F9EF" authority="Toula, 1896" authorityName="Toula" authorityYear="1896" box="[174,733,1624,1651]" class="Reptilia" family="Testudinidae" genus="Testudo" isUncertain="true" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="699" phylum="Chordata" rank="species" species="kalksburgensis" subGenus="Chersine">
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<emphasis id="E8E2EABCFF9EDC54FF09F9A4E739F9EF" bold="true" box="[174,263,1624,1651]" italics="true" pageId="10" pageNumber="699">Testudo</emphasis>
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(
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<emphasis id="E8E2EABCFF9EDC54FEBFF9A4E744F9EF" bold="true" box="[280,378,1624,1651]" italics="true" pageId="10" pageNumber="699">Chersine</emphasis>
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) cf.
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<emphasis id="E8E2EABCFF9EDC54FE09F9A4E471F9EF" bold="true" box="[430,591,1624,1651]" italics="true" pageId="10" pageNumber="699">kalksburgensis</emphasis>
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<bibRefCitation id="BE074B5FFF9EDC54FDF2F9A5E4E3F9EF" author="TOULA F." box="[597,733,1625,1651]" pageId="10" pageNumber="699" pagination="915 - 920" refId="ref15838" refString="TOULA F. 1896. - Ueber neue Wirbelthierreste aus dem Tertiar Oesterreichs und Rumeliens. I. Neue Landschildkrotenreste (Testudo kalksburgensis n. sp.) aus den mediterranen Strandbildungen von Kalksburg bei Wien. Zeitschrift der DeutschenGeologischen Gesellschaft 48 (4): 915 - 920." type="journal article" year="1896">
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<collectingRegion id="1852F84CFF9EDC54FDF2F9A5E4A6F9EF" box="[597,664,1625,1651]" country="Russia" name="Tula" pageId="10" pageNumber="699">Toula</collectingRegion>
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, 1896
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</bibRefCitation>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="928C6525FF9EDC54FE2BF985E7C1F90F" box="[396,511,1657,1684]" pageId="10" pageNumber="699" type="description">
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<paragraph id="DA2936AEFF9EDC54FE2BF985E7C1F90F" blockId="10.[174,733,1624,1684]" box="[396,511,1657,1684]" pageId="10" pageNumber="699">
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(
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<figureCitation id="42AD2A2BFF9EDC54FE31F985E7E6F908" box="[406,472,1657,1684]" captionStart="FIG" captionStartId="11.[132,143,1940,1957]" captionTargetId="figure-19@12.[132,1455,207,1870]" captionTargetPageId="12" captionText="FIG. 5. — Carapace remains of Testudo (Testudo) cf. kalksburgensis from Mokrá-Quarry localities:A-D, Pal. 1320 (nuchal); A, B, dorsal;C, D, visceral; E, Pal. 1322 (neural 4) in dorsal; F, Pal. 1323 (neural 6) in dorsal; G, Pal. 1350 (neural 7) in dorsal; H-J, Pal. 1325 (pygal); H, external; I, J, internal; K, L, Pal. 1326 (costal 1) in dorsal; M, N, Pal. 1329 (costal 2) in dorsal; O, P, Pal. 1330 (costal 3) in dorsal; Q, R, Pal. 1331 (costal 6) in dorsal; S-V, Pal. 1333 (peripheral 1); S, T, external; U, V, internal; W-Z, Pal. 1334 (peripheral 2); W, X, external; Y, Z, internal; A’, B’, Pal. 1338 (peripheral 5) in external; C’-F’, Pal. 1340 (peripheral 7); C’, D’, exter-" figureDoi="http://doi.org/10.5281/zenodo.5679133" httpUri="https://zenodo.org/record/5679133/files/figure.png" pageId="10" pageNumber="699">Figs 5</figureCitation>
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;
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||
<figureCitation id="42AD2A2BFF9EDC54FE40F985E7CBF90F" box="[487,501,1657,1683]" captionStart="FIG" captionStartId="13.[132,142,1894,1911]" captionTargetBox="[132,1457,215,2025]" captionTargetId="figure-16@13.[131,1457,209,1875]" captionTargetPageId="13" captionText="FIG. 6. — Plastral remains ofTestudo (Testudo) cf.kalksburgensis: A-D’, from Mokrá-Quarry localities;A-F, Pal.1351A (left epiplastron);A, C, visceral;B, D, ventral;E, F, left lateral; G-I, Pal. 1354 (entoplatron); G, visceral;H, I, ventral;J-L, Pal. 1353 (entoplatron); J, visceral; K, L, ventral; M, N, Pal. 1352 (entoplatron);M, visceral; N, ventral; O-R, Pal.1351B (hyoplastron);O, P, visceral;Q, R, ventral;S-V, Pal.1355 (hyoplastron);S, T, visceral;U, V, ventral;W-Z, Pal.1356 (hypoplastron);W, X, visceral;Y, Z, ven- tral;A’-D’, Pal.1360 (xiphiplastron);A’, B’, visceral;C’, D’, ventral;E’-H’, Pal.1361 (xiphiplastron);E’, F’, visceral;G’, H’, ventral views;I’, reconstruction of plastron.Thick lines correspond to scute sulci, dotted lines denote plate sutures, and oblique lines indicate missing plate portions. Abbreviations: Ab, abdominal; An, anal; ent, ento- plastron;epi, epiplastron; hyo, hyoplastron;hyp, hypoplastron;Fe, femoral; Gu, gular; Hu, humeral;Pe, pectoral; xi, xiphiplastron. Scale bars:A-H’, 1 cm; I’, 2.5 cm." figureDoi="http://doi.org/10.5281/zenodo.5679141" httpUri="https://zenodo.org/record/5679141/files/figure.png" pageId="10" pageNumber="699">6</figureCitation>
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)
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</paragraph>
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</subSubSection>
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</subSubSection>
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<paragraph id="DA2936AEFF9EDC54FF23F948E7E8F97A" blockId="10.[132,774,1716,1766]" pageId="10" pageNumber="699">
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<materialsCitation id="6AFE3CF3FF9EDC54FF23F948E7E8F97A" ID-GBIF-Occurrence="3400720302" pageId="10" pageNumber="699">LOCALITIES. — MWQ1/2001, MWQ2/2003, MWQTC/2001, MWQ4/2018 and MCQ3/2005.</materialsCitation>
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</paragraph>
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<paragraph id="DA2936AEFF9EDC54FF23F903E28BFDB0" blockId="10.[131,776,1791,2027]" lastBlockId="10.[811,1457,215,556]" pageId="10" pageNumber="699">
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STUDIED MATERIAL. —
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<materialsCitation id="6AFE3CF3FF9EDC54FE25F8FDE28BFDB0" ID-GBIF-Occurrence="3400720304" pageId="10" pageNumber="699">
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<emphasis id="E8E2EABCFF9EDC54FE25F8FDE415F885" bold="true" box="[386,555,1793,1817]" pageId="10" pageNumber="699">
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<collectingCountry id="A281763EFF9EDC54FE25F8FDE415F885" box="[386,555,1793,1817]" name="Czech Republic" pageId="10" pageNumber="699">Czech Republic</collectingCountry>
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</emphasis>
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.
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<collectingRegion id="1852F84CFF9EDC54FD9CF903E68AF8AE" country="Czech Republic" name="Jihomoravsky" pageId="10" pageNumber="699">South Moravia Region</collectingRegion>
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, Mokrá-Quarry, carapace remains (
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<figureCitation id="42AD2A2BFF9EDC54FDB9F8E6E466F8AD" box="[542,600,1817,1842]" captionStart="FIG" captionStartId="11.[132,143,1940,1957]" captionTargetId="figure-19@12.[132,1455,207,1870]" captionTargetPageId="12" captionText="FIG. 5. — Carapace remains of Testudo (Testudo) cf. kalksburgensis from Mokrá-Quarry localities:A-D, Pal. 1320 (nuchal); A, B, dorsal;C, D, visceral; E, Pal. 1322 (neural 4) in dorsal; F, Pal. 1323 (neural 6) in dorsal; G, Pal. 1350 (neural 7) in dorsal; H-J, Pal. 1325 (pygal); H, external; I, J, internal; K, L, Pal. 1326 (costal 1) in dorsal; M, N, Pal. 1329 (costal 2) in dorsal; O, P, Pal. 1330 (costal 3) in dorsal; Q, R, Pal. 1331 (costal 6) in dorsal; S-V, Pal. 1333 (peripheral 1); S, T, external; U, V, internal; W-Z, Pal. 1334 (peripheral 2); W, X, external; Y, Z, internal; A’, B’, Pal. 1338 (peripheral 5) in external; C’-F’, Pal. 1340 (peripheral 7); C’, D’, exter-" figureDoi="http://doi.org/10.5281/zenodo.5679133" httpUri="https://zenodo.org/record/5679133/files/figure.png" pageId="10" pageNumber="699">Fig. 5</figureCitation>
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A-V’): Pal. 1319, nuchal; Pal. 1320, nuchal; Pal. 1321, nuchal; Pal. 1322, neural 4; Pal. 1323, neural 6; Pal. 1350, neural 7; Pal. 1322, neural 4; Pal. 1324, pygal; Pal. 1325, pygal; Pal. 1328, right costal 1; Pal. 1326, left costal 1; Pal. 1327, left costal 1; Pal. 1329, right costal 2; Pal. 1330, left costal 3; Pal. 1331, left costal 6; Pal. 1332, left peripheral 1; Pal. 1333, right peripheral 1; Pal. 1334, left peripheral 2; Pal. 1335, left peripheral 2; Pal. 1336, right periph- eral 2; Pal. 1337, right peripheral 2; Pal. 1338, right peripheral 5; Pal. 1339, left peripheral 7; Pal. 1340, right peripheral 7; Pal. 1341, right peripheral 8; Pal. 1342, left peripheral 9; Pal. 1343, right peripheral 9; Pal. 1344, left peripheral 10; Pal. 1345, left peripheral 10; Pal. 1346, left peripheral 10; Pal. 1347, left peripheral 11; Pal. 1348, left peripheral 11. Pal. 1349, left peripheral 11. Plastral remains (
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<figureCitation id="42AD2A2BFF9EDC54FBD8FE8AE282FE12" box="[1151,1212,374,398]" captionStart="FIG" captionStartId="13.[132,142,1894,1911]" captionTargetBox="[132,1457,215,2025]" captionTargetId="figure-16@13.[131,1457,209,1875]" captionTargetPageId="13" captionText="FIG. 6. — Plastral remains ofTestudo (Testudo) cf.kalksburgensis: A-D’, from Mokrá-Quarry localities;A-F, Pal.1351A (left epiplastron);A, C, visceral;B, D, ventral;E, F, left lateral; G-I, Pal. 1354 (entoplatron); G, visceral;H, I, ventral;J-L, Pal. 1353 (entoplatron); J, visceral; K, L, ventral; M, N, Pal. 1352 (entoplatron);M, visceral; N, ventral; O-R, Pal.1351B (hyoplastron);O, P, visceral;Q, R, ventral;S-V, Pal.1355 (hyoplastron);S, T, visceral;U, V, ventral;W-Z, Pal.1356 (hypoplastron);W, X, visceral;Y, Z, ven- tral;A’-D’, Pal.1360 (xiphiplastron);A’, B’, visceral;C’, D’, ventral;E’-H’, Pal.1361 (xiphiplastron);E’, F’, visceral;G’, H’, ventral views;I’, reconstruction of plastron.Thick lines correspond to scute sulci, dotted lines denote plate sutures, and oblique lines indicate missing plate portions. Abbreviations: Ab, abdominal; An, anal; ent, ento- plastron;epi, epiplastron; hyo, hyoplastron;hyp, hypoplastron;Fe, femoral; Gu, gular; Hu, humeral;Pe, pectoral; xi, xiphiplastron. Scale bars:A-H’, 1 cm; I’, 2.5 cm." figureDoi="http://doi.org/10.5281/zenodo.5679141" httpUri="https://zenodo.org/record/5679141/files/figure.png" pageId="10" pageNumber="699">Fig. 6</figureCitation>
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A-H’): Pal. 1351A, left epiplastron; Pal. 1352, entoplastron; Pal. 1353, entoplastron; Pal. 1354, entoplastron; Pal. 1351B, right hyoplastron; Pal. 1355, right hyoplastron; Pal. 1356, left hypoplastron; Pal. 1357, right hypoplastron; Pal. 1358, right hypoplastron; Pal. 1359, right hypoplastron; Pal. 1360, left xiphiplastron; Pal. 1361, left xiphiplastron; Pal. 1362, left xiphiplastron.
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</materialsCitation>
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</paragraph>
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<paragraph id="DA2936AEFF9EDC54FC8AFDAAE5FDFDF2" blockId="10.[811,1458,598,2028]" box="[813,963,598,624]" pageId="10" pageNumber="699">DESCRIPTION</paragraph>
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<paragraph id="DA2936AEFF9EDC54FC8AFD8AE285FC53" blockId="10.[811,1458,598,2028]" pageId="10" pageNumber="699">
|
||
<taxonomicName id="1D964D2DFF9EDC54FC8AFD8AE541FD0C" authorityName="Linnaeus" authorityYear="1758" box="[813,895,630,656]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="699" phylum="Chordata" rank="genus">
|
||
<emphasis id="E8E2EABCFF9EDC54FC8AFD8AE541FD0C" box="[813,895,630,656]" italics="true" pageId="10" pageNumber="699">Testudo</emphasis>
|
||
</taxonomicName>
|
||
material recovered from Mokrá-Quarry consists mainly of disarticulated plates (Pal. 1319-1362). However, some costals, peripherals and neurals are missing. They belong to a medium-sized
|
||
<taxonomicName id="1D964D2DFF9EDC54FB0FFD29E2C6FD73" authorityName="Linnaeus" authorityYear="1758" box="[1192,1272,725,751]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="699" phylum="Chordata" rank="genus">
|
||
<emphasis id="E8E2EABCFF9EDC54FB0FFD29E2C6FD73" box="[1192,1272,725,751]" italics="true" pageId="10" pageNumber="699">Testudo</emphasis>
|
||
</taxonomicName>
|
||
, which reached approximately
|
||
<quantity id="1D6E9B4BFF9EDC54FC47FD09E269FC8C" box="[992,1111,757,784]" metricMagnitude="-1" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="10" pageNumber="699" unit="cm" value="22.5" valueMax="25.0" valueMin="20.0">20-25 cm</quantity>
|
||
. The following description is based on the entire available material, but not all the plates are depicted (
|
||
<figureCitation id="42AD2A2BFF9EDC54FBACFCC9E272FCCC" box="[1035,1100,821,848]" captionStart="FIG" captionStartId="11.[132,143,1940,1957]" captionTargetId="figure-19@12.[132,1455,207,1870]" captionTargetPageId="12" captionText="FIG. 5. — Carapace remains of Testudo (Testudo) cf. kalksburgensis from Mokrá-Quarry localities:A-D, Pal. 1320 (nuchal); A, B, dorsal;C, D, visceral; E, Pal. 1322 (neural 4) in dorsal; F, Pal. 1323 (neural 6) in dorsal; G, Pal. 1350 (neural 7) in dorsal; H-J, Pal. 1325 (pygal); H, external; I, J, internal; K, L, Pal. 1326 (costal 1) in dorsal; M, N, Pal. 1329 (costal 2) in dorsal; O, P, Pal. 1330 (costal 3) in dorsal; Q, R, Pal. 1331 (costal 6) in dorsal; S-V, Pal. 1333 (peripheral 1); S, T, external; U, V, internal; W-Z, Pal. 1334 (peripheral 2); W, X, external; Y, Z, internal; A’, B’, Pal. 1338 (peripheral 5) in external; C’-F’, Pal. 1340 (peripheral 7); C’, D’, exter-" figureDoi="http://doi.org/10.5281/zenodo.5679133" httpUri="https://zenodo.org/record/5679133/files/figure.png" pageId="10" pageNumber="699">Figs 5</figureCitation>
|
||
;
|
||
<figureCitation id="42AD2A2BFF9EDC54FBFDFCC9E256FCD3" box="[1114,1128,821,847]" captionStart="FIG" captionStartId="13.[132,142,1894,1911]" captionTargetBox="[132,1457,215,2025]" captionTargetId="figure-16@13.[131,1457,209,1875]" captionTargetPageId="13" captionText="FIG. 6. — Plastral remains ofTestudo (Testudo) cf.kalksburgensis: A-D’, from Mokrá-Quarry localities;A-F, Pal.1351A (left epiplastron);A, C, visceral;B, D, ventral;E, F, left lateral; G-I, Pal. 1354 (entoplatron); G, visceral;H, I, ventral;J-L, Pal. 1353 (entoplatron); J, visceral; K, L, ventral; M, N, Pal. 1352 (entoplatron);M, visceral; N, ventral; O-R, Pal.1351B (hyoplastron);O, P, visceral;Q, R, ventral;S-V, Pal.1355 (hyoplastron);S, T, visceral;U, V, ventral;W-Z, Pal.1356 (hypoplastron);W, X, visceral;Y, Z, ven- tral;A’-D’, Pal.1360 (xiphiplastron);A’, B’, visceral;C’, D’, ventral;E’-H’, Pal.1361 (xiphiplastron);E’, F’, visceral;G’, H’, ventral views;I’, reconstruction of plastron.Thick lines correspond to scute sulci, dotted lines denote plate sutures, and oblique lines indicate missing plate portions. Abbreviations: Ab, abdominal; An, anal; ent, ento- plastron;epi, epiplastron; hyo, hyoplastron;hyp, hypoplastron;Fe, femoral; Gu, gular; Hu, humeral;Pe, pectoral; xi, xiphiplastron. Scale bars:A-H’, 1 cm; I’, 2.5 cm." figureDoi="http://doi.org/10.5281/zenodo.5679141" httpUri="https://zenodo.org/record/5679141/files/figure.png" pageId="10" pageNumber="699">6</figureCitation>
|
||
). Most of the dermal grooves and sutures are visible. There is no ornamentation on the carapace or plastron. Growth lines are discernible in some carapace plates, and more specifically in both peripheral and costal plates (e.g.,
|
||
<figureCitation id="42AD2A2BFF9EDC54FB97FC49E2B9FC53" box="[1072,1159,948,975]" captionStart="FIG" captionStartId="11.[132,143,1940,1957]" captionTargetId="figure-19@12.[132,1455,207,1870]" captionTargetPageId="12" captionText="FIG. 5. — Carapace remains of Testudo (Testudo) cf. kalksburgensis from Mokrá-Quarry localities:A-D, Pal. 1320 (nuchal); A, B, dorsal;C, D, visceral; E, Pal. 1322 (neural 4) in dorsal; F, Pal. 1323 (neural 6) in dorsal; G, Pal. 1350 (neural 7) in dorsal; H-J, Pal. 1325 (pygal); H, external; I, J, internal; K, L, Pal. 1326 (costal 1) in dorsal; M, N, Pal. 1329 (costal 2) in dorsal; O, P, Pal. 1330 (costal 3) in dorsal; Q, R, Pal. 1331 (costal 6) in dorsal; S-V, Pal. 1333 (peripheral 1); S, T, external; U, V, internal; W-Z, Pal. 1334 (peripheral 2); W, X, external; Y, Z, internal; A’, B’, Pal. 1338 (peripheral 5) in external; C’-F’, Pal. 1340 (peripheral 7); C’, D’, exter-" figureDoi="http://doi.org/10.5281/zenodo.5679133" httpUri="https://zenodo.org/record/5679133/files/figure.png" pageId="10" pageNumber="699">Fig. 5K</figureCitation>
|
||
, A’).
|
||
</paragraph>
|
||
<paragraph id="DA2936AEFF9EDC54FCE3FC28E3AAFAB2" blockId="10.[811,1458,598,2028]" pageId="10" pageNumber="699">
|
||
The nuchal plate is hexagonal, slightly wider than long, and its anterior edge is pointed (
|
||
<figureCitation id="42AD2A2BFF9EDC54FB0FFC08E2D4FB93" box="[1192,1258,1012,1039]" captionStart="FIG" captionStartId="11.[132,143,1940,1957]" captionTargetId="figure-19@12.[132,1455,207,1870]" captionTargetPageId="12" captionText="FIG. 5. — Carapace remains of Testudo (Testudo) cf. kalksburgensis from Mokrá-Quarry localities:A-D, Pal. 1320 (nuchal); A, B, dorsal;C, D, visceral; E, Pal. 1322 (neural 4) in dorsal; F, Pal. 1323 (neural 6) in dorsal; G, Pal. 1350 (neural 7) in dorsal; H-J, Pal. 1325 (pygal); H, external; I, J, internal; K, L, Pal. 1326 (costal 1) in dorsal; M, N, Pal. 1329 (costal 2) in dorsal; O, P, Pal. 1330 (costal 3) in dorsal; Q, R, Pal. 1331 (costal 6) in dorsal; S-V, Pal. 1333 (peripheral 1); S, T, external; U, V, internal; W-Z, Pal. 1334 (peripheral 2); W, X, external; Y, Z, internal; A’, B’, Pal. 1338 (peripheral 5) in external; C’-F’, Pal. 1340 (peripheral 7); C’, D’, exter-" figureDoi="http://doi.org/10.5281/zenodo.5679133" httpUri="https://zenodo.org/record/5679133/files/figure.png" pageId="10" pageNumber="699">Fig. 5</figureCitation>
|
||
A-D). The posterolateral edges are straight, whereas the anterior ones are curved in medial direction. A transverse thickening is recognizable on the visceral surface of the nuchal (
|
||
<figureCitation id="42AD2A2BFF9EDC54FA8CFBA8E39CFBF3" box="[1323,1442,1108,1135]" captionStart="FIG" captionStartId="11.[132,143,1940,1957]" captionTargetId="figure-19@12.[132,1455,207,1870]" captionTargetPageId="12" captionText="FIG. 5. — Carapace remains of Testudo (Testudo) cf. kalksburgensis from Mokrá-Quarry localities:A-D, Pal. 1320 (nuchal); A, B, dorsal;C, D, visceral; E, Pal. 1322 (neural 4) in dorsal; F, Pal. 1323 (neural 6) in dorsal; G, Pal. 1350 (neural 7) in dorsal; H-J, Pal. 1325 (pygal); H, external; I, J, internal; K, L, Pal. 1326 (costal 1) in dorsal; M, N, Pal. 1329 (costal 2) in dorsal; O, P, Pal. 1330 (costal 3) in dorsal; Q, R, Pal. 1331 (costal 6) in dorsal; S-V, Pal. 1333 (peripheral 1); S, T, external; U, V, internal; W-Z, Pal. 1334 (peripheral 2); W, X, external; Y, Z, internal; A’, B’, Pal. 1338 (peripheral 5) in external; C’-F’, Pal. 1340 (peripheral 7); C’, D’, exter-" figureDoi="http://doi.org/10.5281/zenodo.5679133" httpUri="https://zenodo.org/record/5679133/files/figure.png" pageId="10" pageNumber="699">Fig. 5C, D</figureCitation>
|
||
). The cervical scute is longer than wide and its total length constitutes less than a half of the nuchal plate. It is well developed both dorsally and viscerally according to the three preserved plates (Pal. 1319-21). The anterior edge is narrower than the posterior ones. The lateral sulci are almost straight and parallel to each other (
|
||
<figureCitation id="42AD2A2BFF9EDC54FAA8FAEFE3B9FAB2" box="[1295,1415,1299,1326]" captionStart="FIG" captionStartId="11.[132,143,1940,1957]" captionTargetId="figure-19@12.[132,1455,207,1870]" captionTargetPageId="12" captionText="FIG. 5. — Carapace remains of Testudo (Testudo) cf. kalksburgensis from Mokrá-Quarry localities:A-D, Pal. 1320 (nuchal); A, B, dorsal;C, D, visceral; E, Pal. 1322 (neural 4) in dorsal; F, Pal. 1323 (neural 6) in dorsal; G, Pal. 1350 (neural 7) in dorsal; H-J, Pal. 1325 (pygal); H, external; I, J, internal; K, L, Pal. 1326 (costal 1) in dorsal; M, N, Pal. 1329 (costal 2) in dorsal; O, P, Pal. 1330 (costal 3) in dorsal; Q, R, Pal. 1331 (costal 6) in dorsal; S-V, Pal. 1333 (peripheral 1); S, T, external; U, V, internal; W-Z, Pal. 1334 (peripheral 2); W, X, external; Y, Z, internal; A’, B’, Pal. 1338 (peripheral 5) in external; C’-F’, Pal. 1340 (peripheral 7); C’, D’, exter-" figureDoi="http://doi.org/10.5281/zenodo.5679133" httpUri="https://zenodo.org/record/5679133/files/figure.png" pageId="10" pageNumber="699">Fig. 5A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="DA2936AEFF9EDC54FCE3FACFE587FA71" blockId="10.[811,1458,598,2028]" pageId="10" pageNumber="699">
|
||
Only three neurals plates are preserved, which vary in shape from subsquare (i.e., neural 4) to hexagonal (i.e., neurals 6 to 7:
|
||
<figureCitation id="42AD2A2BFF9EDC54FC7DFA8FE225FA12" box="[986,1051,1395,1422]" captionStart="FIG" captionStartId="11.[132,143,1940,1957]" captionTargetId="figure-19@12.[132,1455,207,1870]" captionTargetPageId="12" captionText="FIG. 5. — Carapace remains of Testudo (Testudo) cf. kalksburgensis from Mokrá-Quarry localities:A-D, Pal. 1320 (nuchal); A, B, dorsal;C, D, visceral; E, Pal. 1322 (neural 4) in dorsal; F, Pal. 1323 (neural 6) in dorsal; G, Pal. 1350 (neural 7) in dorsal; H-J, Pal. 1325 (pygal); H, external; I, J, internal; K, L, Pal. 1326 (costal 1) in dorsal; M, N, Pal. 1329 (costal 2) in dorsal; O, P, Pal. 1330 (costal 3) in dorsal; Q, R, Pal. 1331 (costal 6) in dorsal; S-V, Pal. 1333 (peripheral 1); S, T, external; U, V, internal; W-Z, Pal. 1334 (peripheral 2); W, X, external; Y, Z, internal; A’, B’, Pal. 1338 (peripheral 5) in external; C’-F’, Pal. 1340 (peripheral 7); C’, D’, exter-" figureDoi="http://doi.org/10.5281/zenodo.5679133" httpUri="https://zenodo.org/record/5679133/files/figure.png" pageId="10" pageNumber="699">Fig. 5</figureCitation>
|
||
E-G). It is noteworthy that none of them is more than twice as wide as long, and that they are encroached transversally by the intervertebral sulci (
|
||
<figureCitation id="42AD2A2BFF9EDC54FC91FA2FE549FA72" box="[822,887,1491,1518]" captionStart="FIG" captionStartId="11.[132,143,1940,1957]" captionTargetId="figure-19@12.[132,1455,207,1870]" captionTargetPageId="12" captionText="FIG. 5. — Carapace remains of Testudo (Testudo) cf. kalksburgensis from Mokrá-Quarry localities:A-D, Pal. 1320 (nuchal); A, B, dorsal;C, D, visceral; E, Pal. 1322 (neural 4) in dorsal; F, Pal. 1323 (neural 6) in dorsal; G, Pal. 1350 (neural 7) in dorsal; H-J, Pal. 1325 (pygal); H, external; I, J, internal; K, L, Pal. 1326 (costal 1) in dorsal; M, N, Pal. 1329 (costal 2) in dorsal; O, P, Pal. 1330 (costal 3) in dorsal; Q, R, Pal. 1331 (costal 6) in dorsal; S-V, Pal. 1333 (peripheral 1); S, T, external; U, V, internal; W-Z, Pal. 1334 (peripheral 2); W, X, external; Y, Z, internal; A’, B’, Pal. 1338 (peripheral 5) in external; C’-F’, Pal. 1340 (peripheral 7); C’, D’, exter-" figureDoi="http://doi.org/10.5281/zenodo.5679133" httpUri="https://zenodo.org/record/5679133/files/figure.png" pageId="10" pageNumber="699">Fig. 5</figureCitation>
|
||
E-G).
|
||
</paragraph>
|
||
<paragraph id="DA2936AEFF9EDC54FCE4FA0FE38CF950" blockId="10.[811,1458,598,2028]" pageId="10" pageNumber="699">
|
||
The pygal plate is trapezoidal with slightly concave anterolateral margins. Its external surface is moderately convex, whereas its internal one is rather concave (
|
||
<figureCitation id="42AD2A2BFF9EDC54FAA4F9CEE37DF9D1" box="[1283,1347,1586,1613]" captionStart="FIG" captionStartId="11.[132,143,1940,1957]" captionTargetId="figure-19@12.[132,1455,207,1870]" captionTargetPageId="12" captionText="FIG. 5. — Carapace remains of Testudo (Testudo) cf. kalksburgensis from Mokrá-Quarry localities:A-D, Pal. 1320 (nuchal); A, B, dorsal;C, D, visceral; E, Pal. 1322 (neural 4) in dorsal; F, Pal. 1323 (neural 6) in dorsal; G, Pal. 1350 (neural 7) in dorsal; H-J, Pal. 1325 (pygal); H, external; I, J, internal; K, L, Pal. 1326 (costal 1) in dorsal; M, N, Pal. 1329 (costal 2) in dorsal; O, P, Pal. 1330 (costal 3) in dorsal; Q, R, Pal. 1331 (costal 6) in dorsal; S-V, Pal. 1333 (peripheral 1); S, T, external; U, V, internal; W-Z, Pal. 1334 (peripheral 2); W, X, external; Y, Z, internal; A’, B’, Pal. 1338 (peripheral 5) in external; C’-F’, Pal. 1340 (peripheral 7); C’, D’, exter-" figureDoi="http://doi.org/10.5281/zenodo.5679133" httpUri="https://zenodo.org/record/5679133/files/figure.png" pageId="10" pageNumber="699">Fig. 5</figureCitation>
|
||
H-J). The marginal scutes 12 are missing, and therefore the supracaudal scute is not divided by a sagittal groove (Pal. 1324-25:
|
||
<figureCitation id="42AD2A2BFF9EDC54FC8CF96EE551F931" box="[811,879,1682,1709]" captionStart="FIG" captionStartId="11.[132,143,1940,1957]" captionTargetId="figure-19@12.[132,1455,207,1870]" captionTargetPageId="12" captionText="FIG. 5. — Carapace remains of Testudo (Testudo) cf. kalksburgensis from Mokrá-Quarry localities:A-D, Pal. 1320 (nuchal); A, B, dorsal;C, D, visceral; E, Pal. 1322 (neural 4) in dorsal; F, Pal. 1323 (neural 6) in dorsal; G, Pal. 1350 (neural 7) in dorsal; H-J, Pal. 1325 (pygal); H, external; I, J, internal; K, L, Pal. 1326 (costal 1) in dorsal; M, N, Pal. 1329 (costal 2) in dorsal; O, P, Pal. 1330 (costal 3) in dorsal; Q, R, Pal. 1331 (costal 6) in dorsal; S-V, Pal. 1333 (peripheral 1); S, T, external; U, V, internal; W-Z, Pal. 1334 (peripheral 2); W, X, external; Y, Z, internal; A’, B’, Pal. 1338 (peripheral 5) in external; C’-F’, Pal. 1340 (peripheral 7); C’, D’, exter-" figureDoi="http://doi.org/10.5281/zenodo.5679133" httpUri="https://zenodo.org/record/5679133/files/figure.png" pageId="10" pageNumber="699">Fig. 5</figureCitation>
|
||
H-J). The shape of the vertebral scutes cannot be ascertained with a confidence because they are incomplete.
|
||
</paragraph>
|
||
<paragraph id="DA2936AEFF9EDC54FCE5F92EE21BF877" blockId="10.[811,1458,598,2028]" pageId="10" pageNumber="699">
|
||
Only five of the eight costal plates are present (i.e., costals 1-3 and 5-6), which are trapezoidal. Costals 1 and 3 host the intervertebral sulci (
|
||
<figureCitation id="42AD2A2BFF9EDC54FBEFF8EEE337F8B0" box="[1096,1289,1809,1836]" captionStart="FIG" captionStartId="11.[132,143,1940,1957]" captionTargetId="figure-19@12.[132,1455,207,1870]" captionTargetPageId="12" captionText="FIG. 5. — Carapace remains of Testudo (Testudo) cf. kalksburgensis from Mokrá-Quarry localities:A-D, Pal. 1320 (nuchal); A, B, dorsal;C, D, visceral; E, Pal. 1322 (neural 4) in dorsal; F, Pal. 1323 (neural 6) in dorsal; G, Pal. 1350 (neural 7) in dorsal; H-J, Pal. 1325 (pygal); H, external; I, J, internal; K, L, Pal. 1326 (costal 1) in dorsal; M, N, Pal. 1329 (costal 2) in dorsal; O, P, Pal. 1330 (costal 3) in dorsal; Q, R, Pal. 1331 (costal 6) in dorsal; S-V, Pal. 1333 (peripheral 1); S, T, external; U, V, internal; W-Z, Pal. 1334 (peripheral 2); W, X, external; Y, Z, internal; A’, B’, Pal. 1338 (peripheral 5) in external; C’-F’, Pal. 1340 (peripheral 7); C’, D’, exter-" figureDoi="http://doi.org/10.5281/zenodo.5679133" httpUri="https://zenodo.org/record/5679133/files/figure.png" pageId="10" pageNumber="699">Fig. 5K, L, O, P</figureCitation>
|
||
, W’), whereas the costals 2 and 6 host the interpleural sulci (
|
||
<figureCitation id="42AD2A2BFF9EDC54FAECF8CDE5B2F8F0" captionStart="FIG" captionStartId="11.[132,143,1940,1957]" captionTargetId="figure-19@12.[132,1455,207,1870]" captionTargetPageId="12" captionText="FIG. 5. — Carapace remains of Testudo (Testudo) cf. kalksburgensis from Mokrá-Quarry localities:A-D, Pal. 1320 (nuchal); A, B, dorsal;C, D, visceral; E, Pal. 1322 (neural 4) in dorsal; F, Pal. 1323 (neural 6) in dorsal; G, Pal. 1350 (neural 7) in dorsal; H-J, Pal. 1325 (pygal); H, external; I, J, internal; K, L, Pal. 1326 (costal 1) in dorsal; M, N, Pal. 1329 (costal 2) in dorsal; O, P, Pal. 1330 (costal 3) in dorsal; Q, R, Pal. 1331 (costal 6) in dorsal; S-V, Pal. 1333 (peripheral 1); S, T, external; U, V, internal; W-Z, Pal. 1334 (peripheral 2); W, X, external; Y, Z, internal; A’, B’, Pal. 1338 (peripheral 5) in external; C’-F’, Pal. 1340 (peripheral 7); C’, D’, exter-" figureDoi="http://doi.org/10.5281/zenodo.5679133" httpUri="https://zenodo.org/record/5679133/files/figure.png" pageId="10" pageNumber="699">Fig. 5M, N, Q, R</figureCitation>
|
||
, W’). The pleural scutes are poorly preserved, so no significant details can be discerned. Despite this, the pleuromarginal sulcus coincides with the costoperipheral suture all along the preserved peripheral and costal plates (e.g.,
|
||
<figureCitation id="42AD2A2BFF9EDC54FCCAF82DE5DBF870" box="[877,997,2001,2028]" captionStart="FIG" captionStartId="11.[132,143,1940,1957]" captionTargetId="figure-19@12.[132,1455,207,1870]" captionTargetPageId="12" captionText="FIG. 5. — Carapace remains of Testudo (Testudo) cf. kalksburgensis from Mokrá-Quarry localities:A-D, Pal. 1320 (nuchal); A, B, dorsal;C, D, visceral; E, Pal. 1322 (neural 4) in dorsal; F, Pal. 1323 (neural 6) in dorsal; G, Pal. 1350 (neural 7) in dorsal; H-J, Pal. 1325 (pygal); H, external; I, J, internal; K, L, Pal. 1326 (costal 1) in dorsal; M, N, Pal. 1329 (costal 2) in dorsal; O, P, Pal. 1330 (costal 3) in dorsal; Q, R, Pal. 1331 (costal 6) in dorsal; S-V, Pal. 1333 (peripheral 1); S, T, external; U, V, internal; W-Z, Pal. 1334 (peripheral 2); W, X, external; Y, Z, internal; A’, B’, Pal. 1338 (peripheral 5) in external; C’-F’, Pal. 1340 (peripheral 7); C’, D’, exter-" figureDoi="http://doi.org/10.5281/zenodo.5679133" httpUri="https://zenodo.org/record/5679133/files/figure.png" pageId="10" pageNumber="699">Fig. 5K, L</figureCitation>
|
||
, W’).
|
||
</paragraph>
|
||
<paragraph id="DA2936AEFF9FDC55FF3CFF2BE72BFBF3" blockId="11.[130,778,215,1868]" pageId="11" pageNumber="700">
|
||
Peripheral plates 1-3, together with the nuchal, make up the anterior opening of the shell. Peripheral 1 is heptagonal, while the peripheral 2 is subtriangular. Peripheral 1 displays a moderately developed spike at about the middle of the anterior edge. The presence of protrusions on the remaining peripherals (i.e., 2 and 3) cannot be evaluated. Peripheral 5 (Pal. 1338:
|
||
<figureCitation id="42AD2A2BFF9FDC55FEA1FE6AE778FE2D" box="[262,326,406,433]" captionStart="FIG" captionStartId="11.[132,143,1940,1957]" captionTargetId="figure-19@12.[132,1455,207,1870]" captionTargetPageId="12" captionText="FIG. 5. — Carapace remains of Testudo (Testudo) cf. kalksburgensis from Mokrá-Quarry localities:A-D, Pal. 1320 (nuchal); A, B, dorsal;C, D, visceral; E, Pal. 1322 (neural 4) in dorsal; F, Pal. 1323 (neural 6) in dorsal; G, Pal. 1350 (neural 7) in dorsal; H-J, Pal. 1325 (pygal); H, external; I, J, internal; K, L, Pal. 1326 (costal 1) in dorsal; M, N, Pal. 1329 (costal 2) in dorsal; O, P, Pal. 1330 (costal 3) in dorsal; Q, R, Pal. 1331 (costal 6) in dorsal; S-V, Pal. 1333 (peripheral 1); S, T, external; U, V, internal; W-Z, Pal. 1334 (peripheral 2); W, X, external; Y, Z, internal; A’, B’, Pal. 1338 (peripheral 5) in external; C’-F’, Pal. 1340 (peripheral 7); C’, D’, exter-" figureDoi="http://doi.org/10.5281/zenodo.5679133" httpUri="https://zenodo.org/record/5679133/files/figure.png" pageId="11" pageNumber="700">Fig. 5</figureCitation>
|
||
A’, B’), which is the only preserved plate involved in the shell bridge, is rectangular and rather flat, unlike posterior peripherals. Pal. 1338 displays a very weak longitudinal lateral ridge that is placed slightly above of the marginoabdominal sulcus. Peripherals 7
|
||
<emphasis id="E8E2EABCFF9FDC55FD9AFDEAE478FDAC" bold="true" box="[573,582,534,560]" pageId="11" pageNumber="700">-</emphasis>
|
||
11 (
|
||
<figureCitation id="42AD2A2BFF9FDC55FDD5FDEAE48EFDAD" box="[626,688,534,561]" captionStart="FIG" captionStartId="11.[132,143,1940,1957]" captionTargetId="figure-19@12.[132,1455,207,1870]" captionTargetPageId="12" captionText="FIG. 5. — Carapace remains of Testudo (Testudo) cf. kalksburgensis from Mokrá-Quarry localities:A-D, Pal. 1320 (nuchal); A, B, dorsal;C, D, visceral; E, Pal. 1322 (neural 4) in dorsal; F, Pal. 1323 (neural 6) in dorsal; G, Pal. 1350 (neural 7) in dorsal; H-J, Pal. 1325 (pygal); H, external; I, J, internal; K, L, Pal. 1326 (costal 1) in dorsal; M, N, Pal. 1329 (costal 2) in dorsal; O, P, Pal. 1330 (costal 3) in dorsal; Q, R, Pal. 1331 (costal 6) in dorsal; S-V, Pal. 1333 (peripheral 1); S, T, external; U, V, internal; W-Z, Pal. 1334 (peripheral 2); W, X, external; Y, Z, internal; A’, B’, Pal. 1338 (peripheral 5) in external; C’-F’, Pal. 1340 (peripheral 7); C’, D’, exter-" figureDoi="http://doi.org/10.5281/zenodo.5679133" httpUri="https://zenodo.org/record/5679133/files/figure.png" pageId="11" pageNumber="700">Fig. 5</figureCitation>
|
||
C’-W’), together with the pygal plate (
|
||
<figureCitation id="42AD2A2BFF9FDC55FE61FDCAE43DFDCD" box="[454,515,566,593]" captionStart="FIG" captionStartId="11.[132,143,1940,1957]" captionTargetId="figure-19@12.[132,1455,207,1870]" captionTargetPageId="12" captionText="FIG. 5. — Carapace remains of Testudo (Testudo) cf. kalksburgensis from Mokrá-Quarry localities:A-D, Pal. 1320 (nuchal); A, B, dorsal;C, D, visceral; E, Pal. 1322 (neural 4) in dorsal; F, Pal. 1323 (neural 6) in dorsal; G, Pal. 1350 (neural 7) in dorsal; H-J, Pal. 1325 (pygal); H, external; I, J, internal; K, L, Pal. 1326 (costal 1) in dorsal; M, N, Pal. 1329 (costal 2) in dorsal; O, P, Pal. 1330 (costal 3) in dorsal; Q, R, Pal. 1331 (costal 6) in dorsal; S-V, Pal. 1333 (peripheral 1); S, T, external; U, V, internal; W-Z, Pal. 1334 (peripheral 2); W, X, external; Y, Z, internal; A’, B’, Pal. 1338 (peripheral 5) in external; C’-F’, Pal. 1340 (peripheral 7); C’, D’, exter-" figureDoi="http://doi.org/10.5281/zenodo.5679133" httpUri="https://zenodo.org/record/5679133/files/figure.png" pageId="11" pageNumber="700">Fig. 5</figureCitation>
|
||
H-J), form the posterior opening of the shell. An elongated and subvertical scar of the dorsal projection of the hypoplastron is discernible in peripheral 7 internally (
|
||
<figureCitation id="42AD2A2BFF9FDC55FE2BFD6AE7F4FD2C" box="[396,458,661,688]" captionStart="FIG" captionStartId="11.[132,143,1940,1957]" captionTargetId="figure-19@12.[132,1455,207,1870]" captionTargetPageId="12" captionText="FIG. 5. — Carapace remains of Testudo (Testudo) cf. kalksburgensis from Mokrá-Quarry localities:A-D, Pal. 1320 (nuchal); A, B, dorsal;C, D, visceral; E, Pal. 1322 (neural 4) in dorsal; F, Pal. 1323 (neural 6) in dorsal; G, Pal. 1350 (neural 7) in dorsal; H-J, Pal. 1325 (pygal); H, external; I, J, internal; K, L, Pal. 1326 (costal 1) in dorsal; M, N, Pal. 1329 (costal 2) in dorsal; O, P, Pal. 1330 (costal 3) in dorsal; Q, R, Pal. 1331 (costal 6) in dorsal; S-V, Pal. 1333 (peripheral 1); S, T, external; U, V, internal; W-Z, Pal. 1334 (peripheral 2); W, X, external; Y, Z, internal; A’, B’, Pal. 1338 (peripheral 5) in external; C’-F’, Pal. 1340 (peripheral 7); C’, D’, exter-" figureDoi="http://doi.org/10.5281/zenodo.5679133" httpUri="https://zenodo.org/record/5679133/files/figure.png" pageId="11" pageNumber="700">Fig. 5</figureCitation>
|
||
E’, F’). As a rule, the peripherals
|
||
<quantity id="1D6E9B4BFF9FDC55FF18FD4AE72DFD53" box="[191,275,693,720]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" metricValueMax="2.54" metricValueMin="2.032" pageId="11" pageNumber="700" unit="in" value="9.0" valueMax="10.0" valueMin="8.0">8-10 in</quantity>
|
||
<taxonomicName id="1D964D2DFF9FDC55FE87FD49E74EFD53" authorityName="Linnaeus" authorityYear="1758" box="[288,368,693,719]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="700" phylum="Chordata" rank="genus">
|
||
<emphasis id="E8E2EABCFF9FDC55FE87FD49E74EFD53" box="[288,368,693,719]" italics="true" pageId="11" pageNumber="700">Testudo</emphasis>
|
||
</taxonomicName>
|
||
are rectangular with dorsal surfaces slightly concave, whereas in
|
||
<emphasis id="E8E2EABCFF9FDC55FE68FD29E466FD73" box="[463,600,725,751]" italics="true" pageId="11" pageNumber="700">
|
||
<taxonomicName id="1D964D2DFF9FDC55FE68FD29E46AFD73" authorityName="Pomel" authorityYear="1847" box="[463,596,725,751]" class="Agaricomycetes" family="Fomitopsidaceae" genus="Ptychogaster" kingdom="Fungi" order="Polyporales" pageId="11" pageNumber="700" phylum="Basidiomycota" rank="genus">Ptychogaster</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
they are rather subrectangular and the dorsal concavity is well-developed. Peripheral 11 hosts the lateral edge of the supracaudal scute (
|
||
<figureCitation id="42AD2A2BFF9FDC55FF29FCC9E6F2FCCC" box="[142,204,821,848]" captionStart="FIG" captionStartId="11.[132,143,1940,1957]" captionTargetId="figure-19@12.[132,1455,207,1870]" captionTargetPageId="12" captionText="FIG. 5. — Carapace remains of Testudo (Testudo) cf. kalksburgensis from Mokrá-Quarry localities:A-D, Pal. 1320 (nuchal); A, B, dorsal;C, D, visceral; E, Pal. 1322 (neural 4) in dorsal; F, Pal. 1323 (neural 6) in dorsal; G, Pal. 1350 (neural 7) in dorsal; H-J, Pal. 1325 (pygal); H, external; I, J, internal; K, L, Pal. 1326 (costal 1) in dorsal; M, N, Pal. 1329 (costal 2) in dorsal; O, P, Pal. 1330 (costal 3) in dorsal; Q, R, Pal. 1331 (costal 6) in dorsal; S-V, Pal. 1333 (peripheral 1); S, T, external; U, V, internal; W-Z, Pal. 1334 (peripheral 2); W, X, external; Y, Z, internal; A’, B’, Pal. 1338 (peripheral 5) in external; C’-F’, Pal. 1340 (peripheral 7); C’, D’, exter-" figureDoi="http://doi.org/10.5281/zenodo.5679133" httpUri="https://zenodo.org/record/5679133/files/figure.png" pageId="11" pageNumber="700">Fig. 5</figureCitation>
|
||
S’-W’). The marginal scute 1 is trapezoidal, whereas the remaining posterior marginals are either subsquare or subrectangular (
|
||
<figureCitation id="42AD2A2BFF9FDC55FE90FC89E74BFC0C" box="[311,373,885,912]" captionStart="FIG" captionStartId="11.[132,143,1940,1957]" captionTargetId="figure-19@12.[132,1455,207,1870]" captionTargetPageId="12" captionText="FIG. 5. — Carapace remains of Testudo (Testudo) cf. kalksburgensis from Mokrá-Quarry localities:A-D, Pal. 1320 (nuchal); A, B, dorsal;C, D, visceral; E, Pal. 1322 (neural 4) in dorsal; F, Pal. 1323 (neural 6) in dorsal; G, Pal. 1350 (neural 7) in dorsal; H-J, Pal. 1325 (pygal); H, external; I, J, internal; K, L, Pal. 1326 (costal 1) in dorsal; M, N, Pal. 1329 (costal 2) in dorsal; O, P, Pal. 1330 (costal 3) in dorsal; Q, R, Pal. 1331 (costal 6) in dorsal; S-V, Pal. 1333 (peripheral 1); S, T, external; U, V, internal; W-Z, Pal. 1334 (peripheral 2); W, X, external; Y, Z, internal; A’, B’, Pal. 1338 (peripheral 5) in external; C’-F’, Pal. 1340 (peripheral 7); C’, D’, exter-" figureDoi="http://doi.org/10.5281/zenodo.5679133" httpUri="https://zenodo.org/record/5679133/files/figure.png" pageId="11" pageNumber="700">Fig. 5</figureCitation>
|
||
A-D, S-V, W’). Based on two nuchal plates (Pal. 1319-21), two peripherals 1 (Pal. 1332-33) and three costals 1 (Pal. 1326-28), the triple junction amongst the pleural 1, vertebral 1, and marginal 1 is located outside the nuchal plate. Noticeably, the posterior border of the marginal 5 (Pal. 1338;
|
||
<figureCitation id="42AD2A2BFF9FDC55FE23FBE8E7FDFBB3" box="[388,451,1044,1071]" captionStart="FIG" captionStartId="11.[132,143,1940,1957]" captionTargetId="figure-19@12.[132,1455,207,1870]" captionTargetPageId="12" captionText="FIG. 5. — Carapace remains of Testudo (Testudo) cf. kalksburgensis from Mokrá-Quarry localities:A-D, Pal. 1320 (nuchal); A, B, dorsal;C, D, visceral; E, Pal. 1322 (neural 4) in dorsal; F, Pal. 1323 (neural 6) in dorsal; G, Pal. 1350 (neural 7) in dorsal; H-J, Pal. 1325 (pygal); H, external; I, J, internal; K, L, Pal. 1326 (costal 1) in dorsal; M, N, Pal. 1329 (costal 2) in dorsal; O, P, Pal. 1330 (costal 3) in dorsal; Q, R, Pal. 1331 (costal 6) in dorsal; S-V, Pal. 1333 (peripheral 1); S, T, external; U, V, internal; W-Z, Pal. 1334 (peripheral 2); W, X, external; Y, Z, internal; A’, B’, Pal. 1338 (peripheral 5) in external; C’-F’, Pal. 1340 (peripheral 7); C’, D’, exter-" figureDoi="http://doi.org/10.5281/zenodo.5679133" httpUri="https://zenodo.org/record/5679133/files/figure.png" pageId="11" pageNumber="700">Fig. 5</figureCitation>
|
||
A’, B’) is parallel (i.e., instead of oblique) relative to those anteroposterior edges of the peripheral 5.
|
||
</paragraph>
|
||
<paragraph id="DA2936AEFF9FDC55FF3CFB88E49AFA12" blockId="11.[130,778,215,1868]" pageId="11" pageNumber="700">
|
||
A partial epiplastron, together with the hyoplastron (
|
||
<figureCitation id="42AD2A2BFF9FDC55FF29FB68E6F1FB32" box="[142,207,1172,1198]" captionStart="FIG" captionStartId="13.[132,142,1894,1911]" captionTargetBox="[132,1457,215,2025]" captionTargetId="figure-16@13.[131,1457,209,1875]" captionTargetPageId="13" captionText="FIG. 6. — Plastral remains ofTestudo (Testudo) cf.kalksburgensis: A-D’, from Mokrá-Quarry localities;A-F, Pal.1351A (left epiplastron);A, C, visceral;B, D, ventral;E, F, left lateral; G-I, Pal. 1354 (entoplatron); G, visceral;H, I, ventral;J-L, Pal. 1353 (entoplatron); J, visceral; K, L, ventral; M, N, Pal. 1352 (entoplatron);M, visceral; N, ventral; O-R, Pal.1351B (hyoplastron);O, P, visceral;Q, R, ventral;S-V, Pal.1355 (hyoplastron);S, T, visceral;U, V, ventral;W-Z, Pal.1356 (hypoplastron);W, X, visceral;Y, Z, ven- tral;A’-D’, Pal.1360 (xiphiplastron);A’, B’, visceral;C’, D’, ventral;E’-H’, Pal.1361 (xiphiplastron);E’, F’, visceral;G’, H’, ventral views;I’, reconstruction of plastron.Thick lines correspond to scute sulci, dotted lines denote plate sutures, and oblique lines indicate missing plate portions. Abbreviations: Ab, abdominal; An, anal; ent, ento- plastron;epi, epiplastron; hyo, hyoplastron;hyp, hypoplastron;Fe, femoral; Gu, gular; Hu, humeral;Pe, pectoral; xi, xiphiplastron. Scale bars:A-H’, 1 cm; I’, 2.5 cm." figureDoi="http://doi.org/10.5281/zenodo.5679141" httpUri="https://zenodo.org/record/5679141/files/figure.png" pageId="11" pageNumber="700">Fig. 6</figureCitation>
|
||
A-F, O-R), forms the anterior plastral lobe, which is rather trapezoidal (
|
||
<figureCitation id="42AD2A2BFF9FDC55FED2FB48E788FB52" box="[373,438,1204,1230]" captionStart="FIG" captionStartId="13.[132,142,1894,1911]" captionTargetBox="[132,1457,215,2025]" captionTargetId="figure-16@13.[131,1457,209,1875]" captionTargetPageId="13" captionText="FIG. 6. — Plastral remains ofTestudo (Testudo) cf.kalksburgensis: A-D’, from Mokrá-Quarry localities;A-F, Pal.1351A (left epiplastron);A, C, visceral;B, D, ventral;E, F, left lateral; G-I, Pal. 1354 (entoplatron); G, visceral;H, I, ventral;J-L, Pal. 1353 (entoplatron); J, visceral; K, L, ventral; M, N, Pal. 1352 (entoplatron);M, visceral; N, ventral; O-R, Pal.1351B (hyoplastron);O, P, visceral;Q, R, ventral;S-V, Pal.1355 (hyoplastron);S, T, visceral;U, V, ventral;W-Z, Pal.1356 (hypoplastron);W, X, visceral;Y, Z, ven- tral;A’-D’, Pal.1360 (xiphiplastron);A’, B’, visceral;C’, D’, ventral;E’-H’, Pal.1361 (xiphiplastron);E’, F’, visceral;G’, H’, ventral views;I’, reconstruction of plastron.Thick lines correspond to scute sulci, dotted lines denote plate sutures, and oblique lines indicate missing plate portions. Abbreviations: Ab, abdominal; An, anal; ent, ento- plastron;epi, epiplastron; hyo, hyoplastron;hyp, hypoplastron;Fe, femoral; Gu, gular; Hu, humeral;Pe, pectoral; xi, xiphiplastron. Scale bars:A-H’, 1 cm; I’, 2.5 cm." figureDoi="http://doi.org/10.5281/zenodo.5679141" httpUri="https://zenodo.org/record/5679141/files/figure.png" pageId="11" pageNumber="700">Fig. 6</figureCitation>
|
||
I’). Despite not being entirely preserved, the epiplastral dorsal pad seems rectangular and longer than wide. It is moderately developed posteroventrally and overhangs it slightly. Consequently, a small gular pocket is present (
|
||
<figureCitation id="42AD2A2BFF9FDC55FF5CFACFE707FAD1" box="[251,313,1331,1357]" captionStart="FIG" captionStartId="13.[132,142,1894,1911]" captionTargetBox="[132,1457,215,2025]" captionTargetId="figure-16@13.[131,1457,209,1875]" captionTargetPageId="13" captionText="FIG. 6. — Plastral remains ofTestudo (Testudo) cf.kalksburgensis: A-D’, from Mokrá-Quarry localities;A-F, Pal.1351A (left epiplastron);A, C, visceral;B, D, ventral;E, F, left lateral; G-I, Pal. 1354 (entoplatron); G, visceral;H, I, ventral;J-L, Pal. 1353 (entoplatron); J, visceral; K, L, ventral; M, N, Pal. 1352 (entoplatron);M, visceral; N, ventral; O-R, Pal.1351B (hyoplastron);O, P, visceral;Q, R, ventral;S-V, Pal.1355 (hyoplastron);S, T, visceral;U, V, ventral;W-Z, Pal.1356 (hypoplastron);W, X, visceral;Y, Z, ven- tral;A’-D’, Pal.1360 (xiphiplastron);A’, B’, visceral;C’, D’, ventral;E’-H’, Pal.1361 (xiphiplastron);E’, F’, visceral;G’, H’, ventral views;I’, reconstruction of plastron.Thick lines correspond to scute sulci, dotted lines denote plate sutures, and oblique lines indicate missing plate portions. Abbreviations: Ab, abdominal; An, anal; ent, ento- plastron;epi, epiplastron; hyo, hyoplastron;hyp, hypoplastron;Fe, femoral; Gu, gular; Hu, humeral;Pe, pectoral; xi, xiphiplastron. Scale bars:A-H’, 1 cm; I’, 2.5 cm." figureDoi="http://doi.org/10.5281/zenodo.5679141" httpUri="https://zenodo.org/record/5679141/files/figure.png" pageId="11" pageNumber="700">Fig. 6</figureCitation>
|
||
A-C). The gular scutes are triangular, with slightly sinuous lateral margins, and form an angle of less than 45° relative to the sagittal axis (
|
||
<figureCitation id="42AD2A2BFF9FDC55FD86FA8FE45CFA11" box="[545,610,1395,1421]" captionStart="FIG" captionStartId="13.[132,142,1894,1911]" captionTargetBox="[132,1457,215,2025]" captionTargetId="figure-16@13.[131,1457,209,1875]" captionTargetPageId="13" captionText="FIG. 6. — Plastral remains ofTestudo (Testudo) cf.kalksburgensis: A-D’, from Mokrá-Quarry localities;A-F, Pal.1351A (left epiplastron);A, C, visceral;B, D, ventral;E, F, left lateral; G-I, Pal. 1354 (entoplatron); G, visceral;H, I, ventral;J-L, Pal. 1353 (entoplatron); J, visceral; K, L, ventral; M, N, Pal. 1352 (entoplatron);M, visceral; N, ventral; O-R, Pal.1351B (hyoplastron);O, P, visceral;Q, R, ventral;S-V, Pal.1355 (hyoplastron);S, T, visceral;U, V, ventral;W-Z, Pal.1356 (hypoplastron);W, X, visceral;Y, Z, ven- tral;A’-D’, Pal.1360 (xiphiplastron);A’, B’, visceral;C’, D’, ventral;E’-H’, Pal.1361 (xiphiplastron);E’, F’, visceral;G’, H’, ventral views;I’, reconstruction of plastron.Thick lines correspond to scute sulci, dotted lines denote plate sutures, and oblique lines indicate missing plate portions. Abbreviations: Ab, abdominal; An, anal; ent, ento- plastron;epi, epiplastron; hyo, hyoplastron;hyp, hypoplastron;Fe, femoral; Gu, gular; Hu, humeral;Pe, pectoral; xi, xiphiplastron. Scale bars:A-H’, 1 cm; I’, 2.5 cm." figureDoi="http://doi.org/10.5281/zenodo.5679141" httpUri="https://zenodo.org/record/5679141/files/figure.png" pageId="11" pageNumber="700">Fig. 6</figureCitation>
|
||
B-D).
|
||
</paragraph>
|
||
<paragraph id="DA2936AEFF9FDC55FF3CFA6FE539F9F1" blockId="11.[130,778,215,1868]" pageId="11" pageNumber="700">
|
||
The three available entoplastra (Pal. 1352-54:
|
||
<figureCitation id="42AD2A2BFF9FDC55FD2DFA6FE4F9FA31" box="[650,711,1427,1453]" captionStart="FIG" captionStartId="13.[132,142,1894,1911]" captionTargetBox="[132,1457,215,2025]" captionTargetId="figure-16@13.[131,1457,209,1875]" captionTargetPageId="13" captionText="FIG. 6. — Plastral remains ofTestudo (Testudo) cf.kalksburgensis: A-D’, from Mokrá-Quarry localities;A-F, Pal.1351A (left epiplastron);A, C, visceral;B, D, ventral;E, F, left lateral; G-I, Pal. 1354 (entoplatron); G, visceral;H, I, ventral;J-L, Pal. 1353 (entoplatron); J, visceral; K, L, ventral; M, N, Pal. 1352 (entoplatron);M, visceral; N, ventral; O-R, Pal.1351B (hyoplastron);O, P, visceral;Q, R, ventral;S-V, Pal.1355 (hyoplastron);S, T, visceral;U, V, ventral;W-Z, Pal.1356 (hypoplastron);W, X, visceral;Y, Z, ven- tral;A’-D’, Pal.1360 (xiphiplastron);A’, B’, visceral;C’, D’, ventral;E’-H’, Pal.1361 (xiphiplastron);E’, F’, visceral;G’, H’, ventral views;I’, reconstruction of plastron.Thick lines correspond to scute sulci, dotted lines denote plate sutures, and oblique lines indicate missing plate portions. Abbreviations: Ab, abdominal; An, anal; ent, ento- plastron;epi, epiplastron; hyo, hyoplastron;hyp, hypoplastron;Fe, femoral; Gu, gular; Hu, humeral;Pe, pectoral; xi, xiphiplastron. Scale bars:A-H’, 1 cm; I’, 2.5 cm." figureDoi="http://doi.org/10.5281/zenodo.5679141" httpUri="https://zenodo.org/record/5679141/files/figure.png" pageId="11" pageNumber="700">Fig. 6</figureCitation>
|
||
G-N) are hexagonal both ventrally and viscerally. They are partially covered by the gular scutes, which generally extend up to the middle of the entoplastron. The ventral surface of the epiplastra, covered by the gular scutes, is not in relief. In all specimens, the gular scutes are not crossed by the humeropectoral sulcus transversally (
|
||
<figureCitation id="42AD2A2BFF9FDC55FDEDF9AEE4C4F9F1" box="[586,762,1618,1645]" captionStart="FIG" captionStartId="13.[132,142,1894,1911]" captionTargetBox="[132,1457,215,2025]" captionTargetId="figure-16@13.[131,1457,209,1875]" captionTargetPageId="13" captionText="FIG. 6. — Plastral remains ofTestudo (Testudo) cf.kalksburgensis: A-D’, from Mokrá-Quarry localities;A-F, Pal.1351A (left epiplastron);A, C, visceral;B, D, ventral;E, F, left lateral; G-I, Pal. 1354 (entoplatron); G, visceral;H, I, ventral;J-L, Pal. 1353 (entoplatron); J, visceral; K, L, ventral; M, N, Pal. 1352 (entoplatron);M, visceral; N, ventral; O-R, Pal.1351B (hyoplastron);O, P, visceral;Q, R, ventral;S-V, Pal.1355 (hyoplastron);S, T, visceral;U, V, ventral;W-Z, Pal.1356 (hypoplastron);W, X, visceral;Y, Z, ven- tral;A’-D’, Pal.1360 (xiphiplastron);A’, B’, visceral;C’, D’, ventral;E’-H’, Pal.1361 (xiphiplastron);E’, F’, visceral;G’, H’, ventral views;I’, reconstruction of plastron.Thick lines correspond to scute sulci, dotted lines denote plate sutures, and oblique lines indicate missing plate portions. Abbreviations: Ab, abdominal; An, anal; ent, ento- plastron;epi, epiplastron; hyo, hyoplastron;hyp, hypoplastron;Fe, femoral; Gu, gular; Hu, humeral;Pe, pectoral; xi, xiphiplastron. Scale bars:A-H’, 1 cm; I’, 2.5 cm." figureDoi="http://doi.org/10.5281/zenodo.5679141" httpUri="https://zenodo.org/record/5679141/files/figure.png" pageId="11" pageNumber="700">Fig. 6H, I, K, L</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="DA2936AEFF9FDC55FF3CF98EE32FFD4C" blockId="11.[130,778,215,1868]" lastBlockId="11.[811,1457,215,720]" pageId="11" pageNumber="700">
|
||
Pal. 1351B is the best preserved hyoplastron (
|
||
<figureCitation id="42AD2A2BFF9FDC55FD20F98EE4FAF910" box="[647,708,1650,1676]" captionStart="FIG" captionStartId="13.[132,142,1894,1911]" captionTargetBox="[132,1457,215,2025]" captionTargetId="figure-16@13.[131,1457,209,1875]" captionTargetPageId="13" captionText="FIG. 6. — Plastral remains ofTestudo (Testudo) cf.kalksburgensis: A-D’, from Mokrá-Quarry localities;A-F, Pal.1351A (left epiplastron);A, C, visceral;B, D, ventral;E, F, left lateral; G-I, Pal. 1354 (entoplatron); G, visceral;H, I, ventral;J-L, Pal. 1353 (entoplatron); J, visceral; K, L, ventral; M, N, Pal. 1352 (entoplatron);M, visceral; N, ventral; O-R, Pal.1351B (hyoplastron);O, P, visceral;Q, R, ventral;S-V, Pal.1355 (hyoplastron);S, T, visceral;U, V, ventral;W-Z, Pal.1356 (hypoplastron);W, X, visceral;Y, Z, ven- tral;A’-D’, Pal.1360 (xiphiplastron);A’, B’, visceral;C’, D’, ventral;E’-H’, Pal.1361 (xiphiplastron);E’, F’, visceral;G’, H’, ventral views;I’, reconstruction of plastron.Thick lines correspond to scute sulci, dotted lines denote plate sutures, and oblique lines indicate missing plate portions. Abbreviations: Ab, abdominal; An, anal; ent, ento- plastron;epi, epiplastron; hyo, hyoplastron;hyp, hypoplastron;Fe, femoral; Gu, gular; Hu, humeral;Pe, pectoral; xi, xiphiplastron. Scale bars:A-H’, 1 cm; I’, 2.5 cm." figureDoi="http://doi.org/10.5281/zenodo.5679141" httpUri="https://zenodo.org/record/5679141/files/figure.png" pageId="11" pageNumber="700">Fig. 6</figureCitation>
|
||
O-R), which hosts entirely both the humeropectoral and pectoroabdominal scutes: the former is nearly straight and obliquely oriented relative to the sagittal plane (i.e., only slightly sinuous:
|
||
<figureCitation id="42AD2A2BFF9FDC55FEEEF90EE79EF890" box="[329,416,1778,1804]" captionStart="FIG" captionStartId="13.[132,142,1894,1911]" captionTargetBox="[132,1457,215,2025]" captionTargetId="figure-16@13.[131,1457,209,1875]" captionTargetPageId="13" captionText="FIG. 6. — Plastral remains ofTestudo (Testudo) cf.kalksburgensis: A-D’, from Mokrá-Quarry localities;A-F, Pal.1351A (left epiplastron);A, C, visceral;B, D, ventral;E, F, left lateral; G-I, Pal. 1354 (entoplatron); G, visceral;H, I, ventral;J-L, Pal. 1353 (entoplatron); J, visceral; K, L, ventral; M, N, Pal. 1352 (entoplatron);M, visceral; N, ventral; O-R, Pal.1351B (hyoplastron);O, P, visceral;Q, R, ventral;S-V, Pal.1355 (hyoplastron);S, T, visceral;U, V, ventral;W-Z, Pal.1356 (hypoplastron);W, X, visceral;Y, Z, ven- tral;A’-D’, Pal.1360 (xiphiplastron);A’, B’, visceral;C’, D’, ventral;E’-H’, Pal.1361 (xiphiplastron);E’, F’, visceral;G’, H’, ventral views;I’, reconstruction of plastron.Thick lines correspond to scute sulci, dotted lines denote plate sutures, and oblique lines indicate missing plate portions. Abbreviations: Ab, abdominal; An, anal; ent, ento- plastron;epi, epiplastron; hyo, hyoplastron;hyp, hypoplastron;Fe, femoral; Gu, gular; Hu, humeral;Pe, pectoral; xi, xiphiplastron. Scale bars:A-H’, 1 cm; I’, 2.5 cm." figureDoi="http://doi.org/10.5281/zenodo.5679141" httpUri="https://zenodo.org/record/5679141/files/figure.png" pageId="11" pageNumber="700">Fig. 6R</figureCitation>
|
||
), while the latter is curved and transversally oriented relative to the sagittal plane (
|
||
<figureCitation id="42AD2A2BFF9FDC55FD08F8EEE6A2F8D0" captionStart="FIG" captionStartId="13.[132,142,1894,1911]" captionTargetBox="[132,1457,215,2025]" captionTargetId="figure-16@13.[131,1457,209,1875]" captionTargetPageId="13" captionText="FIG. 6. — Plastral remains ofTestudo (Testudo) cf.kalksburgensis: A-D’, from Mokrá-Quarry localities;A-F, Pal.1351A (left epiplastron);A, C, visceral;B, D, ventral;E, F, left lateral; G-I, Pal. 1354 (entoplatron); G, visceral;H, I, ventral;J-L, Pal. 1353 (entoplatron); J, visceral; K, L, ventral; M, N, Pal. 1352 (entoplatron);M, visceral; N, ventral; O-R, Pal.1351B (hyoplastron);O, P, visceral;Q, R, ventral;S-V, Pal.1355 (hyoplastron);S, T, visceral;U, V, ventral;W-Z, Pal.1356 (hypoplastron);W, X, visceral;Y, Z, ven- tral;A’-D’, Pal.1360 (xiphiplastron);A’, B’, visceral;C’, D’, ventral;E’-H’, Pal.1361 (xiphiplastron);E’, F’, visceral;G’, H’, ventral views;I’, reconstruction of plastron.Thick lines correspond to scute sulci, dotted lines denote plate sutures, and oblique lines indicate missing plate portions. Abbreviations: Ab, abdominal; An, anal; ent, ento- plastron;epi, epiplastron; hyo, hyoplastron;hyp, hypoplastron;Fe, femoral; Gu, gular; Hu, humeral;Pe, pectoral; xi, xiphiplastron. Scale bars:A-H’, 1 cm; I’, 2.5 cm." figureDoi="http://doi.org/10.5281/zenodo.5679141" httpUri="https://zenodo.org/record/5679141/files/figure.png" pageId="11" pageNumber="700">Fig. 6R, V</figureCitation>
|
||
). The preserved portion of the humeral scute indicates that this was trapezoidal. The abdominal seems to be the largest scute of the plastron. Moreover, its medial sector is slightly oblique (
|
||
<figureCitation id="42AD2A2BFF9FDC55FBB2FEEBE264FEAD" box="[1045,1114,279,305]" captionStart="FIG" captionStartId="13.[132,142,1894,1911]" captionTargetBox="[132,1457,215,2025]" captionTargetId="figure-16@13.[131,1457,209,1875]" captionTargetPageId="13" captionText="FIG. 6. — Plastral remains ofTestudo (Testudo) cf.kalksburgensis: A-D’, from Mokrá-Quarry localities;A-F, Pal.1351A (left epiplastron);A, C, visceral;B, D, ventral;E, F, left lateral; G-I, Pal. 1354 (entoplatron); G, visceral;H, I, ventral;J-L, Pal. 1353 (entoplatron); J, visceral; K, L, ventral; M, N, Pal. 1352 (entoplatron);M, visceral; N, ventral; O-R, Pal.1351B (hyoplastron);O, P, visceral;Q, R, ventral;S-V, Pal.1355 (hyoplastron);S, T, visceral;U, V, ventral;W-Z, Pal.1356 (hypoplastron);W, X, visceral;Y, Z, ven- tral;A’-D’, Pal.1360 (xiphiplastron);A’, B’, visceral;C’, D’, ventral;E’-H’, Pal.1361 (xiphiplastron);E’, F’, visceral;G’, H’, ventral views;I’, reconstruction of plastron.Thick lines correspond to scute sulci, dotted lines denote plate sutures, and oblique lines indicate missing plate portions. Abbreviations: Ab, abdominal; An, anal; ent, ento- plastron;epi, epiplastron; hyo, hyoplastron;hyp, hypoplastron;Fe, femoral; Gu, gular; Hu, humeral;Pe, pectoral; xi, xiphiplastron. Scale bars:A-H’, 1 cm; I’, 2.5 cm." figureDoi="http://doi.org/10.5281/zenodo.5679141" httpUri="https://zenodo.org/record/5679141/files/figure.png" pageId="11" pageNumber="700">Fig. 6</figureCitation>
|
||
Y-Z). The hypo-xiphiplastral suture is roughly straight and well developed, which means that a plastral hinge is absent. The xiphiplastron is trapezoidal and its ventral side is very flat and crossed by the femoroanal sulcus (
|
||
<figureCitation id="42AD2A2BFF9FDC55FBF9FE6AE29EFE2C" box="[1118,1184,406,432]" captionStart="FIG" captionStartId="13.[132,142,1894,1911]" captionTargetBox="[132,1457,215,2025]" captionTargetId="figure-16@13.[131,1457,209,1875]" captionTargetPageId="13" captionText="FIG. 6. — Plastral remains ofTestudo (Testudo) cf.kalksburgensis: A-D’, from Mokrá-Quarry localities;A-F, Pal.1351A (left epiplastron);A, C, visceral;B, D, ventral;E, F, left lateral; G-I, Pal. 1354 (entoplatron); G, visceral;H, I, ventral;J-L, Pal. 1353 (entoplatron); J, visceral; K, L, ventral; M, N, Pal. 1352 (entoplatron);M, visceral; N, ventral; O-R, Pal.1351B (hyoplastron);O, P, visceral;Q, R, ventral;S-V, Pal.1355 (hyoplastron);S, T, visceral;U, V, ventral;W-Z, Pal.1356 (hypoplastron);W, X, visceral;Y, Z, ven- tral;A’-D’, Pal.1360 (xiphiplastron);A’, B’, visceral;C’, D’, ventral;E’-H’, Pal.1361 (xiphiplastron);E’, F’, visceral;G’, H’, ventral views;I’, reconstruction of plastron.Thick lines correspond to scute sulci, dotted lines denote plate sutures, and oblique lines indicate missing plate portions. Abbreviations: Ab, abdominal; An, anal; ent, ento- plastron;epi, epiplastron; hyo, hyoplastron;hyp, hypoplastron;Fe, femoral; Gu, gular; Hu, humeral;Pe, pectoral; xi, xiphiplastron. Scale bars:A-H’, 1 cm; I’, 2.5 cm." figureDoi="http://doi.org/10.5281/zenodo.5679141" httpUri="https://zenodo.org/record/5679141/files/figure.png" pageId="11" pageNumber="700">Fig. 6</figureCitation>
|
||
A’-C’). It is noteworthy that the distinct notch in their lateral margins, between the anal and femoral scutes, is missing(
|
||
<figureCitation id="42AD2A2BFF9FDC55FB4BFE2AE312FE6C" box="[1260,1324,470,496]" captionStart="FIG" captionStartId="13.[132,142,1894,1911]" captionTargetBox="[132,1457,215,2025]" captionTargetId="figure-16@13.[131,1457,209,1875]" captionTargetPageId="13" captionText="FIG. 6. — Plastral remains ofTestudo (Testudo) cf.kalksburgensis: A-D’, from Mokrá-Quarry localities;A-F, Pal.1351A (left epiplastron);A, C, visceral;B, D, ventral;E, F, left lateral; G-I, Pal. 1354 (entoplatron); G, visceral;H, I, ventral;J-L, Pal. 1353 (entoplatron); J, visceral; K, L, ventral; M, N, Pal. 1352 (entoplatron);M, visceral; N, ventral; O-R, Pal.1351B (hyoplastron);O, P, visceral;Q, R, ventral;S-V, Pal.1355 (hyoplastron);S, T, visceral;U, V, ventral;W-Z, Pal.1356 (hypoplastron);W, X, visceral;Y, Z, ven- tral;A’-D’, Pal.1360 (xiphiplastron);A’, B’, visceral;C’, D’, ventral;E’-H’, Pal.1361 (xiphiplastron);E’, F’, visceral;G’, H’, ventral views;I’, reconstruction of plastron.Thick lines correspond to scute sulci, dotted lines denote plate sutures, and oblique lines indicate missing plate portions. Abbreviations: Ab, abdominal; An, anal; ent, ento- plastron;epi, epiplastron; hyo, hyoplastron;hyp, hypoplastron;Fe, femoral; Gu, gular; Hu, humeral;Pe, pectoral; xi, xiphiplastron. Scale bars:A-H’, 1 cm; I’, 2.5 cm." figureDoi="http://doi.org/10.5281/zenodo.5679141" httpUri="https://zenodo.org/record/5679141/files/figure.png" pageId="11" pageNumber="700">Fig. 6</figureCitation>
|
||
A’, C’). The femoral is trapezoidal and much longer medially than the anal scutes, which are subrectangular and wider than long. The femoroanal sulcus is slightly sinuous and obliquely oriented relative to the sagittal plane. The anal notch is wider than long, and its visceral area covered by the anal scutes is variable, from moderately (Pal. 1361:
|
||
<figureCitation id="42AD2A2BFF9FDC55FA8DFD6AE356FD2C" box="[1322,1384,662,688]" captionStart="FIG" captionStartId="13.[132,142,1894,1911]" captionTargetBox="[132,1457,215,2025]" captionTargetId="figure-16@13.[131,1457,209,1875]" captionTargetPageId="13" captionText="FIG. 6. — Plastral remains ofTestudo (Testudo) cf.kalksburgensis: A-D’, from Mokrá-Quarry localities;A-F, Pal.1351A (left epiplastron);A, C, visceral;B, D, ventral;E, F, left lateral; G-I, Pal. 1354 (entoplatron); G, visceral;H, I, ventral;J-L, Pal. 1353 (entoplatron); J, visceral; K, L, ventral; M, N, Pal. 1352 (entoplatron);M, visceral; N, ventral; O-R, Pal.1351B (hyoplastron);O, P, visceral;Q, R, ventral;S-V, Pal.1355 (hyoplastron);S, T, visceral;U, V, ventral;W-Z, Pal.1356 (hypoplastron);W, X, visceral;Y, Z, ven- tral;A’-D’, Pal.1360 (xiphiplastron);A’, B’, visceral;C’, D’, ventral;E’-H’, Pal.1361 (xiphiplastron);E’, F’, visceral;G’, H’, ventral views;I’, reconstruction of plastron.Thick lines correspond to scute sulci, dotted lines denote plate sutures, and oblique lines indicate missing plate portions. Abbreviations: Ab, abdominal; An, anal; ent, ento- plastron;epi, epiplastron; hyo, hyoplastron;hyp, hypoplastron;Fe, femoral; Gu, gular; Hu, humeral;Pe, pectoral; xi, xiphiplastron. Scale bars:A-H’, 1 cm; I’, 2.5 cm." figureDoi="http://doi.org/10.5281/zenodo.5679141" httpUri="https://zenodo.org/record/5679141/files/figure.png" pageId="11" pageNumber="700">Fig. 6</figureCitation>
|
||
A’, B’) to well developed (Pal. 1362:
|
||
<figureCitation id="42AD2A2BFF9FDC55FB26FD49E2FFFD53" box="[1153,1217,693,719]" captionStart="FIG" captionStartId="13.[132,142,1894,1911]" captionTargetBox="[132,1457,215,2025]" captionTargetId="figure-16@13.[131,1457,209,1875]" captionTargetPageId="13" captionText="FIG. 6. — Plastral remains ofTestudo (Testudo) cf.kalksburgensis: A-D’, from Mokrá-Quarry localities;A-F, Pal.1351A (left epiplastron);A, C, visceral;B, D, ventral;E, F, left lateral; G-I, Pal. 1354 (entoplatron); G, visceral;H, I, ventral;J-L, Pal. 1353 (entoplatron); J, visceral; K, L, ventral; M, N, Pal. 1352 (entoplatron);M, visceral; N, ventral; O-R, Pal.1351B (hyoplastron);O, P, visceral;Q, R, ventral;S-V, Pal.1355 (hyoplastron);S, T, visceral;U, V, ventral;W-Z, Pal.1356 (hypoplastron);W, X, visceral;Y, Z, ven- tral;A’-D’, Pal.1360 (xiphiplastron);A’, B’, visceral;C’, D’, ventral;E’-H’, Pal.1361 (xiphiplastron);E’, F’, visceral;G’, H’, ventral views;I’, reconstruction of plastron.Thick lines correspond to scute sulci, dotted lines denote plate sutures, and oblique lines indicate missing plate portions. Abbreviations: Ab, abdominal; An, anal; ent, ento- plastron;epi, epiplastron; hyo, hyoplastron;hyp, hypoplastron;Fe, femoral; Gu, gular; Hu, humeral;Pe, pectoral; xi, xiphiplastron. Scale bars:A-H’, 1 cm; I’, 2.5 cm." figureDoi="http://doi.org/10.5281/zenodo.5679141" httpUri="https://zenodo.org/record/5679141/files/figure.png" pageId="11" pageNumber="700">Fig. 6</figureCitation>
|
||
E’, F’).
|
||
</paragraph>
|
||
<subSubSection id="928C6525FF9FDC55FC8AFD0AE31CF8D0" pageId="11" pageNumber="700" type="discussion">
|
||
<paragraph id="DA2936AEFF9FDC55FC8AFD0AE5ABFC92" blockId="11.[811,1457,758,1868]" box="[813,917,758,784]" pageId="11" pageNumber="700">REMARKS</paragraph>
|
||
<paragraph id="DA2936AEFF9FDC55FC8AFCE9E5B1FAB2" blockId="11.[811,1457,758,1868]" pageId="11" pageNumber="700">
|
||
The genus
|
||
<taxonomicName id="1D964D2DFF9FDC55FC3AFCE9E5D2FCB3" authorityName="Linnaeus" authorityYear="1758" box="[925,1004,789,815]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="700" phylum="Chordata" rank="genus" sensu="lato">
|
||
<emphasis id="E8E2EABCFF9FDC55FC3AFCE9E5D2FCB3" box="[925,1004,789,815]" italics="true" pageId="11" pageNumber="700">Testudo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F3D157C7FF9FDC55FC54FCE9E22FFCB3" box="[1011,1041,789,815]" pageId="11" pageNumber="700" sensu="lato">s.l.</taxonomicNameLabel>
|
||
is a clade with five extant and multiple extinct species of terrestrial tortoises of western Palearctic distribution (
|
||
<bibRefCitation id="BE074B5FFF9FDC55FC64FCAAE313FCF3" author="LAPPARENT DE BROIN F. & BOUR R. & PARHAM J. F. & PERALA J." box="[963,1325,853,880]" pageId="11" pageNumber="700" pagination="803 - 811" refId="ref14146" refString="LAPPARENT DE BROIN F., BOUR R., PARHAM J. F. & PERALA J. 2006 a. - Eurotestudo, a new genus for the species Testudo hermanni Gmelin, 1789 (Chelonii, Testudinidae). Comptes Rendus Palevol 5 (6): 803 - 811. https: // doi. org / 10.1016 / j. crpv. 2006.03.002" type="journal article" year="2006">
|
||
Lapparent de Broin
|
||
<emphasis id="E8E2EABCFF9FDC55FB0DFCAAE2DFFCF3" box="[1194,1249,853,879]" italics="true" pageId="11" pageNumber="700">et al.</emphasis>
|
||
2006a
|
||
</bibRefCitation>
|
||
, b; Fritz & Bininda-Emonds 2007;
|
||
<bibRefCitation id="BE074B5FFF9FDC55FB9CFC89E333FC13" author="CORSINI J. A. & BOHME M. & JOYCE W. G." box="[1083,1293,885,911]" pageId="11" pageNumber="700" pagination="948 - 966" refId="ref12531" refString="CORSINI J. A., BOHME M. & JOYCE W. G. 2014. - Reappraisal of Testudo antiqua (Testudines, Testudinidae) from the Miocene of Hohenhowen, Germany. Journal of Paleontology 88 (5): 948 - 966. https: // doi. org / 10.1017 / S 0022336000057590" type="journal article" year="2014">
|
||
Corsini
|
||
<emphasis id="E8E2EABCFF9FDC55FB32FC8AE2F2FC13" box="[1173,1228,885,911]" italics="true" pageId="11" pageNumber="700">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="BE074B5FFF9FDC55FABAFC89E559FC33" author="DELFINO M. & LUJAN A. H. & CARMONA R. & ALBA D. M." pageId="11" pageNumber="700" pagination="215 - 225" refId="ref12771" refString="DELFINO M., LUJAN A. H., CARMONA R. & ALBA D. M. 2012. - Revision of the extinct Pleistocene tortoise Testudo lunellensis Almera and Bofill, 1903 from Cova de Gracia (Barcelona, Spain). Amphibia-Reptilia 33 (2): 215 - 225. https: // doi. org / 10.1163 / 156853812 X 636466" type="journal article" year="2012">
|
||
Delfino
|
||
<emphasis id="E8E2EABCFF9FDC55FADDFC8AE38EFC13" box="[1402,1456,885,911]" italics="true" pageId="11" pageNumber="700">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="BE074B5FFF9FDC55FCD0FC69E204FC33" author="LUJAN A. H. & DELFINO M. & ROBLES J. M. & ALBA D. M." box="[887,1082,917,943]" pageId="11" pageNumber="700" pagination="312 - 342" refId="ref14601" refString="LUJAN A. H., DELFINO M., ROBLES J. M. & ALBA D. M. 2016. - A review of the Miocene tortoise Testudo catalaunica Bataller, 1926 and a revised phylogeny of extinct species of genus Testudo (Testudines: Testudinidae). Zoological Journal of the Linnean Society 178 (2): 312 - 342. https: // doi. org / 10.1111 / zoj. 12414" type="journal article" year="2016">
|
||
Luján
|
||
<emphasis id="E8E2EABCFF9FDC55FC67FC6AE5C9FC33" box="[960,1015,917,943]" italics="true" pageId="11" pageNumber="700">et al.</emphasis>
|
||
2016
|
||
</bibRefCitation>
|
||
). Despite that, in the past, three extant genus-group taxa were distinguished as genera (see e.g.,
|
||
<bibRefCitation id="BE074B5FFF9FDC55FCC4FC28E314FC73" author="TURTLE TAXONOMY WORKING GROUP & VAN DIJK P. P. & IVERSON J. B. & RHODIN A. G. J. & SCHAFFER H. B. & BOUR R." box="[867,1322,980,1007]" pageId="11" pageNumber="700" refId="ref15888" refString="TURTLE TAXONOMY WORKING GROUP [VAN DIJK P. P., IVERSON J. B., RHODIN A. G. J., SCHAFFER H. B. & BOUR R.]. 2014. - Turtles of the world, 7 th edition: Annotated checklist of taxonomy, synonymy, distribution with maps, and conservation status. Chelonian Research Monographs 5: 000.329 - 000.479. https: // doi. org / 10.3854 / crm. 5.000. checklist. v 7.2014" type="journal volume" year="2014">Turtle Taxonomy Working Group 2014</bibRefCitation>
|
||
), currently, the use of a single genus (
|
||
<taxonomicName id="1D964D2DFF9FDC55FBECFC08E2A4FB92" authorityName="Linnaeus" authorityYear="1758" box="[1099,1178,1012,1038]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="700" phylum="Chordata" rank="genus" sensu="lato">
|
||
<emphasis id="E8E2EABCFF9FDC55FBECFC08E2A4FB92" box="[1099,1178,1012,1038]" italics="true" pageId="11" pageNumber="700">Testudo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F3D157C7FF9FDC55FB05FC08E281FB93" box="[1186,1215,1012,1039]" pageId="11" pageNumber="700" sensu="lato">s.l.</taxonomicNameLabel>
|
||
) and three subgenera (
|
||
<taxonomicName id="1D964D2DFF9FDC55FC91FBE8E5BBFBB2" authorityName="Linnaeus" authorityYear="1758" box="[822,901,1044,1070]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="700" phylum="Chordata" rank="genus">
|
||
<emphasis id="E8E2EABCFF9FDC55FC91FBE8E5BBFBB2" box="[822,901,1044,1070]" italics="true" pageId="11" pageNumber="700">Testudo</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="1D964D2DFF9FDC55FC34FBE9E235FBB3" box="[915,1035,1045,1071]" class="Reptilia" family="Testudinidae" genus="Agrionemys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="700" phylum="Chordata" rank="genus">
|
||
<emphasis id="E8E2EABCFF9FDC55FC34FBE9E235FBB3" box="[915,1035,1045,1071]" italics="true" pageId="11" pageNumber="700">Agrionemys</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="1D964D2DFF9FDC55FBE5FBE8E2A3FBB2" authorityName="Merrem" authorityYear="1820" box="[1090,1181,1044,1070]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="703" phylum="Chordata" rank="subGenus" subGenus="Chersine">
|
||
<emphasis id="E8E2EABCFF9FDC55FBE5FBE8E2A3FBB2" box="[1090,1181,1044,1070]" italics="true" pageId="11" pageNumber="700">Chersine</emphasis>
|
||
</taxonomicName>
|
||
) is being better accepted (see
|
||
<bibRefCitation id="BE074B5FFF9FDC55FCFBFBC8E228FBD2" author="LUJAN A. H. & DELFINO M. & ROBLES J. M. & ALBA D. M." box="[860,1046,1076,1102]" pageId="11" pageNumber="700" pagination="312 - 342" refId="ref14601" refString="LUJAN A. H., DELFINO M., ROBLES J. M. & ALBA D. M. 2016. - A review of the Miocene tortoise Testudo catalaunica Bataller, 1926 and a revised phylogeny of extinct species of genus Testudo (Testudines: Testudinidae). Zoological Journal of the Linnean Society 178 (2): 312 - 342. https: // doi. org / 10.1111 / zoj. 12414" type="journal article" year="2016">
|
||
Luján
|
||
<emphasis id="E8E2EABCFF9FDC55FC04FBC9E5E9FBD2" box="[931,983,1076,1102]" italics="true" pageId="11" pageNumber="700">et al.</emphasis>
|
||
2016
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="BE074B5FFF9FDC55FB83FBC8E2DBFBD2" author="GRACIA E. V. A. & RAMIREZ M. & DELFINO M. & ANADON J. D. & GIMENEZ A. & FAHD S. & CORTI C. & JDEIDI T. B. & FRITZ U." box="[1060,1253,1076,1102]" pageId="11" pageNumber="700" pagination="641 - 654" refId="ref13115" refString="GRACIA E. V. A., VARGAS- RAMIREZ M., DELFINO M., ANADON J. D., GIMENEZ A., FAHD S., CORTI C., JDEIDI T. B. & FRITZ U. 2017. - Expansion after expansion: dissecting the phylogeography of the widely distributed spur-thighed tortoise, Testudo graeca (Testudines: Testudinidae). Biological Journal of the Linnean Society 121 (3): 641 - 654. https: // doi. org / 10.1093 / biolinnean / blx 007" type="journal article" year="2017">
|
||
Graciá
|
||
<emphasis id="E8E2EABCFF9FDC55FBD5FBC9E298FBD2" box="[1138,1190,1076,1102]" italics="true" pageId="11" pageNumber="700">et al.</emphasis>
|
||
2017
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="BE074B5FFF9FDC55FB55FBC8E225FBF2" author="TURTLE TAXONOMY WORKING GROUP & RHODIN A. G. J. & IVERSON J. B. & BOUR R. & FRITZ U. & GEORGES A. & SHAFFER H. B. & VAN DIJK P. P." pageId="11" pageNumber="700" pagination="1 - 292" refId="ref15978" refString="TURTLE TAXONOMY WORKING GROUP [RHODIN A. G. J., IVERSON J. B., BOUR R., FRITZ U., GEORGES A., SHAFFER H. B. & VAN DIJK P. P. 2017. - Turtles of the world: annotated checklist and atlas of taxonomy, synonymy, distribution, and conservation status, 8 th edition. Chelonian Research Monographs 7: 1 - 292. https: // doi. org / 10.3854 / crm. 7. checklist. atlas. v 8.2017" type="journal article" year="2017">Turtle Taxonomy Working Group 2017</bibRefCitation>
|
||
). Among extinct taxa, the taxonomy of
|
||
<taxonomicName id="1D964D2DFF9FDC55FCEBFB88E5A2FB12" authorityName="Linnaeus" authorityYear="1758" box="[844,924,1140,1166]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="700" phylum="Chordata" rank="genus" sensu="lato">
|
||
<emphasis id="E8E2EABCFF9FDC55FCEBFB88E5A2FB12" box="[844,924,1140,1166]" italics="true" pageId="11" pageNumber="700">Testudo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F3D157C7FF9FDC55FC02FB88E5FBFB12" box="[933,965,1140,1166]" pageId="11" pageNumber="700" sensu="lato">s.l.</taxonomicNameLabel>
|
||
also included the distinction of the extinct genus
|
||
<taxonomicName id="1D964D2DFF9FDC55FCD0FB68E309FB32" authority="Lapparent de Broin, 2000" authorityName="Lapparent de Broin" authorityYear="2000" box="[887,1335,1172,1198]" class="Reptilia" family="Testudinidae" genus="Paleotestudo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="700" phylum="Chordata" rank="genus">
|
||
<emphasis id="E8E2EABCFF9FDC55FCD0FB68E5C2FB32" box="[887,1020,1172,1198]" italics="true" pageId="11" pageNumber="700">Paleotestudo</emphasis>
|
||
Lapparent de Broin, 2000
|
||
</taxonomicName>
|
||
. Although the latter has not been the object of an exhaustive review, results presented by
|
||
<bibRefCitation id="BE074B5FFF9FDC55FBACFB28E2E7FB72" author="LUJAN A. H. & DELFINO M. & ROBLES J. M. & ALBA D. M." box="[1035,1241,1236,1262]" pageId="11" pageNumber="700" pagination="312 - 342" refId="ref14601" refString="LUJAN A. H., DELFINO M., ROBLES J. M. & ALBA D. M. 2016. - A review of the Miocene tortoise Testudo catalaunica Bataller, 1926 and a revised phylogeny of extinct species of genus Testudo (Testudines: Testudinidae). Zoological Journal of the Linnean Society 178 (2): 312 - 342. https: // doi. org / 10.1111 / zoj. 12414" type="journal article" year="2016">
|
||
Luján
|
||
<emphasis id="E8E2EABCFF9FDC55FBF5FB28E2B9FB72" box="[1106,1159,1236,1262]" italics="true" pageId="11" pageNumber="700">et al.</emphasis>
|
||
(2016)
|
||
</bibRefCitation>
|
||
clearly pointed out that
|
||
<taxonomicName id="1D964D2DFF9FDC55FCF9FB0FE5E0FA91" authorityName="Lapparent de Broin" authorityYear="2000" box="[862,990,1267,1293]" class="Reptilia" family="Testudinidae" genus="Paleotestudo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="700" phylum="Chordata" rank="genus">
|
||
<emphasis id="E8E2EABCFF9FDC55FCF9FB0FE5E0FA91" box="[862,990,1267,1293]" italics="true" pageId="11" pageNumber="700">Paleotestudo</emphasis>
|
||
</taxonomicName>
|
||
is a junior subjective synonym of subgenus
|
||
<taxonomicName id="1D964D2DFF9FDC55FC8CFAEFE5B8FAB1" authorityName="Merrem" authorityYear="1820" box="[811,902,1299,1325]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="700" phylum="Chordata" rank="subGenus" subGenus="Chersine">
|
||
<emphasis id="E8E2EABCFF9FDC55FC8CFAEFE5B8FAB1" box="[811,902,1299,1325]" italics="true" pageId="11" pageNumber="700">Chersine</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="DA2936AEFF9FDC55FCE4FACFE31CF8D0" blockId="11.[811,1457,758,1868]" pageId="11" pageNumber="700">
|
||
Regarding the species
|
||
<taxonomicName id="1D964D2DFF9FDC55FB8EFACFE3AAFAD1" authority="Toula, 1896" authorityName="Toula" authorityYear="1896" box="[1065,1428,1331,1357]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="700" phylum="Chordata" rank="species" species="kalksburgensis">
|
||
<emphasis id="E8E2EABCFF9FDC55FB8EFACFE335FAD1" box="[1065,1291,1331,1357]" italics="true" pageId="11" pageNumber="700">Testudo kalksburgensis</emphasis>
|
||
<bibRefCitation id="BE074B5FFF9FDC55FAB7FACFE3AAFAD1" author="TOULA F." box="[1296,1428,1331,1357]" pageId="11" pageNumber="700" pagination="915 - 920" refId="ref15838" refString="TOULA F. 1896. - Ueber neue Wirbelthierreste aus dem Tertiar Oesterreichs und Rumeliens. I. Neue Landschildkrotenreste (Testudo kalksburgensis n. sp.) aus den mediterranen Strandbildungen von Kalksburg bei Wien. Zeitschrift der DeutschenGeologischen Gesellschaft 48 (4): 915 - 920." type="journal article" year="1896">Toula, 1896</bibRefCitation>
|
||
</taxonomicName>
|
||
, it was originally described by
|
||
<bibRefCitation id="BE074B5FFF9FDC55FBEBFAAFE2E2FAF2" author="TOULA F." box="[1100,1244,1363,1390]" pageId="11" pageNumber="700" pagination="915 - 920" refId="ref15838" refString="TOULA F. 1896. - Ueber neue Wirbelthierreste aus dem Tertiar Oesterreichs und Rumeliens. I. Neue Landschildkrotenreste (Testudo kalksburgensis n. sp.) aus den mediterranen Strandbildungen von Kalksburg bei Wien. Zeitschrift der DeutschenGeologischen Gesellschaft 48 (4): 915 - 920." type="journal article" year="1896">Toula (1896)</bibRefCitation>
|
||
based on
|
||
<specimenCount id="CC90FD27FF9FDC55FAE3FAA8E563FA12" pageId="11" pageNumber="700" type="generic">one specimen</specimenCount>
|
||
from Kalksburg, Vienna: a partial shell currently housed at the IGUW. At the beginning of the twentieth century, new fragmentary material, coming from the late Miocene locality of Au am Leithaberge, was referred to
|
||
<taxonomicName id="1D964D2DFF9FDC55FB5BFA28E391FA71" authorityName="Toula" authorityYear="1896" box="[1276,1455,1491,1517]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="700" phylum="Chordata" rank="species" species="kalksburgensis">
|
||
<emphasis id="E8E2EABCFF9FDC55FB5BFA28E391FA71" box="[1276,1455,1491,1517]" italics="true" pageId="11" pageNumber="700">T. kalksburgensis</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="BE074B5FFF9FDC55FCF7FA0FE21CF991" author="SIEBENROCK F." box="[848,1058,1523,1549]" pageId="11" pageNumber="700" pagination="357 - 362" refId="ref15633" refString="SIEBENROCK F. 1914. - Testudo kalksburgensis Toula aus dem Leithagebirge. Jahrbuch der Kaiserlich-Koniglichen Geologischen Reichsanstalt 64 (1 - 2): 357 - 362." type="journal article" year="1914">Siebenrock (1914)</bibRefCitation>
|
||
. Shortly after,
|
||
<bibRefCitation id="BE074B5FFF9FDC55FB6BFA0FE342F991" author="STAESCHE K." box="[1228,1404,1523,1549]" pageId="11" pageNumber="700" pagination="1 - 17" refId="ref15662" refString="STAESCHE K. 1931. - Die Schildkroten des Steinheimer Beckens A. Testudinidae. Palaeontographica Supplement 8: 1 - 17." type="journal article" year="1931">Staesche (1931)</bibRefCitation>
|
||
also erected
|
||
<taxonomicName id="1D964D2DFF9FDC55FC22F9EEE37AF9B0" box="[901,1348,1554,1581]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="700" phylum="Chordata" rank="variety" species="kalksburgensis" variety="steinheimensis">
|
||
<emphasis id="E8E2EABCFF9FDC55FC22F9EEE24AF9B0" box="[901,1140,1554,1580]" italics="true" pageId="11" pageNumber="700">Testudo kalksburgensis</emphasis>
|
||
var.
|
||
<emphasis id="E8E2EABCFF9FDC55FB08F9EEE37AF9B0" box="[1199,1348,1554,1580]" italics="true" pageId="11" pageNumber="700">steinheimensis</emphasis>
|
||
</taxonomicName>
|
||
based on various specimens from the middle Miocene (MN7) locality of Steinheim (
|
||
<collectingCountry id="A281763EFF9FDC55FC40F9AEE26BF9F0" box="[999,1109,1618,1644]" name="Germany" pageId="11" pageNumber="700">Germany</collectingCountry>
|
||
). Although
|
||
<taxonomicName id="1D964D2DFF9FDC55FB76F9AFE3BEF9F0" authorityName="Toula" authorityYear="1896" box="[1233,1408,1618,1644]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="700" phylum="Chordata" rank="species" species="kalksburgensis">
|
||
<emphasis id="E8E2EABCFF9FDC55FB76F9AFE3BEF9F0" box="[1233,1408,1618,1644]" italics="true" pageId="11" pageNumber="700">T. kalksburgensis</emphasis>
|
||
</taxonomicName>
|
||
was considered a junior subjective synonym of
|
||
<taxonomicName id="1D964D2DFF9FDC55FB48F98FE559F930" authority="Bronn, 1831" authorityName="Bronn" authorityYear="1831" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="700" phylum="Chordata" rank="species" species="antiqua">
|
||
<emphasis id="E8E2EABCFF9FDC55FB48F98FE362F910" box="[1263,1372,1651,1676]" italics="true" pageId="11" pageNumber="700">T. antiqua</emphasis>
|
||
Bronn, 1831
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="BE074B5FFF9FDC55FC2CF96EE275F931" author="GLAESSNER M. F." box="[907,1099,1682,1709]" pageId="11" pageNumber="700" pagination="353 - 387" refId="ref13212" refString="GLAESSNER M. F. 1933. - Die Tertiar schildkroten Niederosterreichs. Neues Jahrbuch fur Mineralogie, Geologie, und Palaontologie, Abteilung B 69: 353 - 387." type="journal article" year="1933">Glaessner (1933)</bibRefCitation>
|
||
, subsequently
|
||
<bibRefCitation id="BE074B5FFF9FDC55FB4DF96EE391F931" author="MLYNARSKI M." box="[1258,1455,1682,1709]" pageId="11" pageNumber="700" refId="ref14954" refString="MLYNARSKI M. 1976. - Encyclopedia of Paleoherpetology. Part 7. Testudines. Gustav Fischer Verlag, Stuttgart, 130 p." type="book" year="1976">Młynarski (1976)</bibRefCitation>
|
||
resurrected the species once again. Indeed, the validity of
|
||
<taxonomicName id="1D964D2DFF9FDC55FC89F92FE5DDF970" authorityName="Toula" authorityYear="1896" box="[814,995,1746,1772]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="700" phylum="Chordata" rank="species" species="kalksburgensis">
|
||
<emphasis id="E8E2EABCFF9FDC55FC89F92FE5DDF970" box="[814,995,1746,1772]" italics="true" pageId="11" pageNumber="700">T. kalksburgensis</emphasis>
|
||
</taxonomicName>
|
||
was also confirmed by latter studies (see
|
||
<bibRefCitation id="BE074B5FFF9FDC55FC89F90EE5DEF890" author="MLYNARSKI M." box="[814,992,1778,1804]" pageId="11" pageNumber="700" pagination="1 - 35" refId="ref14978" refString="MLYNARSKI M. 1980. - Die Schildkroten des Steinheimer Beckens B. Chelydridae mit einem Nachtrag zu den Testudinoidea. Palaeontographica Supplement B 8 (2): 1 - 35." type="journal article" year="1980">Młynarski 1980</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="BE074B5FFF9FDC55FC49F90EE37AF890" author="BACHMAYER M. F. & MLYNARSKI M." box="[1006,1348,1778,1804]" pageId="11" pageNumber="700" pagination="111 - 119" refId="ref11639" refString="BACHMAYER M. F. & MLYNARSKI M. 1981. - Testudo kalksburgensis Toula, 1896, eine valide Schildkrotenart aus den miozanen Strandbildungen von Kalksburg bei Wien. Sitzungsberichte Osterreichische Akademie der Wissenschaften, Mathematisch. Naturwissenschaftliche Klasse 190: 111 - 119." type="journal article" year="1981">Bachmayer & Młynarski 1981</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="BE074B5FFF9FDC55FAF5F90EE558F8B0" author="SCHLEICH H. - H." pageId="11" pageNumber="700" pagination="1 - 372" refId="ref15544" refString="SCHLEICH H. - H. 1981. - Jungtertiare Schildkroten Suddeutschlands unter besonderer Berucksichtigung der Fundstelle Sandelzhausen. Courier Forschunginstitut Senckenberg 48: 1 - 372." type="journal article" year="1981">Schleich 1981</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="BE074B5FFF9FDC55FCD5F8EEE241F8B0" author="GEMEL R. & RAUSCHER K." box="[882,1151,1810,1836]" pageId="11" pageNumber="700" pagination="63 - 86" refId="ref13089" refString="GEMEL R. & RAUSCHER K. 2000. - Fossile Schildkroten aus Osterreich (Reptilia, Testudiens. Stapfia 69: 63 - 86." type="journal article" year="2000">Gemel & Rauscher 2000</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="BE074B5FFF9FDC55FB2BF8EEE32DF8B0" author="GEMEL R." box="[1164,1299,1810,1836]" pageId="11" pageNumber="700" pagination="373 - 393" refId="ref13060" refString="GEMEL R. 2002. - Weitere Schildkrotenreste aus dem Karpatium des Korneuburger Beckens (Untermiozan; Niederosterreich). Beitrage zur Palaeontologie 27: 373 - 393." type="journal article" year="2002">Gemel 2002</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="BE074B5FFF9FDC55FAB8F8EEE559F8D0" author="DANILOV I. G. & CERNANSKY A. & SYROMYATNIKOVA E. V. & JONIAK P." pageId="11" pageNumber="700" pagination="423 - 442" refId="ref12710" refString="DANILOV I. G., CERNANSKY A., SYROMYATNIKOVA E. V. & JONIAK P. 2012. - Fossil turtles of Slovakia: new material and a review of the previous record. Amphibia-Reptilia 33 (3 - 4): 423 - 442. https: // doi. org / 10.1163 / 15685381 - 00002846" type="journal article" year="2012">
|
||
Danilov
|
||
<emphasis id="E8E2EABCFF9FDC55FADAF8EEE38EF8B0" box="[1405,1456,1810,1836]" italics="true" pageId="11" pageNumber="700">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="BE074B5FFF9FDC55FCD2F8CDE20DF8D7" author="LUJAN A. H. & DELFINO M. & ROBLES J. M. & ALBA D. M." box="[885,1075,1841,1867]" pageId="11" pageNumber="700" pagination="312 - 342" refId="ref14601" refString="LUJAN A. H., DELFINO M., ROBLES J. M. & ALBA D. M. 2016. - A review of the Miocene tortoise Testudo catalaunica Bataller, 1926 and a revised phylogeny of extinct species of genus Testudo (Testudines: Testudinidae). Zoological Journal of the Linnean Society 178 (2): 312 - 342. https: // doi. org / 10.1111 / zoj. 12414" type="journal article" year="2016">
|
||
Luján
|
||
<emphasis id="E8E2EABCFF9FDC55FC1AF8CEE5CCF8D7" box="[957,1010,1841,1867]" italics="true" pageId="11" pageNumber="700">et al.</emphasis>
|
||
2016
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="BE074B5FFF9FDC55FBE6F8CDE32CF8D0" author="BREZINA J. & ALBA D. M. & IVANOV M. & HANACEK M. & LUJAN A. H." box="[1089,1298,1841,1868]" pageId="11" pageNumber="700" pagination="110473" refId="ref12051" refString="BREZINA J., ALBA D. M., IVANOV M., HANACEK M. & LUJAN A. H. 2021. - A middle Miocene vertebrate assemblage from the Czech part of the Vienna Basin: Implications for the paleoenvironments of the Central Paratethys. Palaeogeography, Palaeoclimatology, Palaeoecology 575: 110473. https: // doi. org / 10.1016 / j. palaeo. 2021.110473" type="journal article" year="2021">
|
||
Březina
|
||
<emphasis id="E8E2EABCFF9FDC55FB3BF8CEE2F7F8D7" box="[1180,1225,1841,1867]" italics="true" pageId="11" pageNumber="700">et al</emphasis>
|
||
. 2021
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |