627 lines
98 KiB
XML
627 lines
98 KiB
XML
<document id="678FFDABC3A2F6F9DD5B97010A02543A" ID-CLB-Dataset="3587" ID-DOI="10.5252/geodiversitas2021v43a20" ID-GBIF-Dataset="8e41f2cf-1620-45db-847c-54fc788f51ad" ID-ISSN="1638-9395" ID-Zenodo-Dep="5679113" ID-ZooBank="urn:lsid:zoobank.org:pub:04F82471-9F26-4935-97CE-1A286E08C958" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="felipe" checkinTime="1636557091778" checkinUser="felipe" docAuthor="Luján, Àngel H., Čerňanský, Andrej, Bonilla-Salomón, Isaac, Březina, Jakub & Ivanov, Martin" docDate="2021" docId="523F87B8FF91DC57FC75FDCAE539F911" docLanguage="en" docName="Geodiversitas.43.20.691-707.pdf" docOrigin="Geodiversitas 43 (20)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Ptychogaster (Ptychogaster)" docType="treatment" docVersion="10" lastPageNumber="698" masterDocId="AE06FFC0FF94DC5EFFA7FFFCE63EFF9C" masterDocTitle="Fossil turtles from the early Miocene localities of Mokrá-Quarry (Burdigalian, MN 4), South Moravian Region, Czech Republic" masterLastPageNumber="707" masterPageNumber="691" pageNumber="694" updateTime="1728566026018" updateUser="guilherme" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="A70B88D15DAE03D97A567F185FE7DD6F">Fossil turtles from the early Miocene localities of Mokrá-Quarry (Burdigalian, MN 4), South Moravian Region, Czech Republic</mods:title>
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<mods:name id="21687658121044ADA317587191942AF7" type="personal">
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<mods:namePart id="D56BC90745BA247550B8756F292F2965">Luján, Àngel H.</mods:namePart>
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<mods:affiliation id="B9B76D1153098669BB6C80DDECA41903">Masaryk University, Faculty of Science, Department of Geological Sciences, 611 37 Brno (Czechia) and Comenius University in Bratislava, Faculty of Natural Sciences, Department of Ecology, 842 15 Bratislava (Slovakia) and Institut Català de Paleontologia Miquel Crusafont, Universitat Autònoma de Barcelona, Edifici ICTA, 08193 Cerdanyola del Vallès (Spain)</mods:affiliation>
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<mods:nameIdentifier id="FAACA098924C0EB19740D6C6DAE982B2" type="email">angel.lujan@icp.cat</mods:nameIdentifier>
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<mods:roleTerm id="B79A5FCEAE97F3BF54FC2ADE6A549B7D">Author</mods:roleTerm>
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<mods:namePart id="BB5AF0E8BF45F8D6D921C1D9F7E59E37">Čerňanský, Andrej</mods:namePart>
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<mods:affiliation id="C48CDE55B35353D643C7BFFF9F397EFA">Comenius University in Bratislava, Faculty of Natural Sciences, Department of Ecology, 842 15 Bratislava (Slovakia)</mods:affiliation>
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<mods:nameIdentifier id="03FE1F3981C1866FA64FC50CAE5C9FB0" type="email">cernansky.paleontology@gmail.com</mods:nameIdentifier>
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<mods:roleTerm id="A00837202D89A74F948FEAD673146A12">Author</mods:roleTerm>
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<mods:namePart id="AC12A02F3B5524FB99CC746769BE5BBD">Bonilla-Salomón, Isaac</mods:namePart>
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<mods:affiliation id="6BB1A327EAE027D2AE09E7DA57A550D6">Department of Geology and Paleontology, Faculty of Natural Sciences, Comenius University, Mlynská dolina, Ilkovičova, 842 15 Bratislava (Slovakia)</mods:affiliation>
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<mods:nameIdentifier id="8B11D786EBC94FFE8D484E045E008A8D" type="email">salomon1@uniba.sk</mods:nameIdentifier>
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<mods:namePart id="2570BDA52AFC576E926279BBA2077FA2">Březina, Jakub</mods:namePart>
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<mods:affiliation id="3BD745D68CA971E8D866E3D0E336BF03">Moravian Museum, Department of Geology and Paleontology, 659 37 Brno (Czechia)</mods:affiliation>
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<mods:nameIdentifier id="8BA398D8F27D821379E7644F5222440C" type="email">jbrezina@mzm.cz</mods:nameIdentifier>
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<mods:namePart id="322DE29CA06C8B1AE2D457702FA25608">Ivanov, Martin</mods:namePart>
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<mods:affiliation id="D62E127E8F546208FECE527AE819E895">Masaryk University, Faculty of Science, Department of Geological Sciences, 611 37 Brno (Czechia)</mods:affiliation>
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<mods:nameIdentifier id="76ECE7A129AC884B996482C0BF8C8A9C" type="email">mivanov@sci.muni.cz</mods:nameIdentifier>
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<mods:title id="FC3DA9387C56680B9DCE7F0EEAC554B4">Geodiversitas</mods:title>
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<mods:part id="A221999E37B83B2F5F76168789930B86">
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<mods:date id="88E366A3A41FC4FBA6BFAA2464AA3449">2021</mods:date>
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<mods:number id="BD56951A5C6E37A17AA5B09F2BE6F4B7">2021-10-26</mods:number>
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<mods:number id="B08B45779A33C46678FDCCA9EDC85ECD">43</mods:number>
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<mods:number id="E6760ED440093F1DC181B849FC9D08EB">20</mods:number>
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<mods:identifier id="E2A238FD090202681F95CD155D4848E2" type="CLB-Dataset">3587</mods:identifier>
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<mods:identifier id="FDF6BE1CCEEDB116471387E8422801FC" type="DOI">10.5252/geodiversitas2021v43a20</mods:identifier>
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<mods:identifier id="7B007732E2989002147DC7382E02546F" type="ISSN">1638-9395</mods:identifier>
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<treatment id="523F87B8FF91DC57FC75FDCAE539F911" ID-DOI="http://doi.org/10.5281/zenodo.5699189" ID-GBIF-Taxon="190303802" ID-Zenodo-Dep="5699189" LSID="urn:lsid:plazi:treatment:523F87B8FF91DC57FC75FDCAE539F911" httpUri="http://treatment.plazi.org/id/523F87B8FF91DC57FC75FDCAE539F911" lastPageId="9" lastPageNumber="698" pageId="5" pageNumber="694">
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<subSubSection id="928C6525FF91DC5BFC75FDCAE334FDCD" box="[978,1290,566,593]" pageId="5" pageNumber="694" type="nomenclature">
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<paragraph id="DA2936AEFF91DC5BFC75FDCAE334FDCD" blockId="5.[978,1290,566,624]" box="[978,1290,566,593]" pageId="5" pageNumber="694">
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<heading id="816181C2FF91DC5BFC75FDCAE334FDCD" box="[978,1290,566,593]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="694" reason="2">
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<taxonomicName id="1D964D2DFF91DC5BFC75FDCAE2DBFDCD" authority="(Ptychogaster)" box="[978,1253,566,593]" class="Reptilia" family="Geoemydidae" genus="Ptychogaster" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="694" phylum="Chordata" rank="subGenus" subGenus="Ptychogaster">
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<emphasis id="E8E2EABCFF91DC5BFC75FDCAE271FDCC" box="[978,1103,566,592]" italics="true" pageId="5" pageNumber="694">Ptychogaster</emphasis>
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(
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<emphasis id="E8E2EABCFF91DC5BFBF8FDCAE2E2FDCC" box="[1119,1244,566,592]" italics="true" pageId="5" pageNumber="694">Ptychogaster</emphasis>
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)
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</taxonomicName>
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sp.
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="928C6525FF91DC56FB90FDAAE2C6FA12" lastPageId="8" lastPageNumber="697" pageId="5" pageNumber="694" type="description">
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<paragraph id="DA2936AEFF91DC5BFB90FDAAE29BFDEC" blockId="5.[978,1290,566,624]" box="[1079,1189,598,624]" pageId="5" pageNumber="694">
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(
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<figureCitation id="42AD2A2BFF91DC5BFBE7FDAAE2BEFDEC" box="[1088,1152,598,624]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="5" pageNumber="694">Figs 2</figureCitation>
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;
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<figureCitation id="42AD2A2BFF91DC5BFB29FDAAE2A2FDEC" box="[1166,1180,598,624]" captionStart="FIG" captionStartId="7.[132,143,1894,1911]" captionTargetBox="[139,1445,213,1863]" captionTargetId="figure-16@7.[133,1455,207,1869]" captionTargetPageId="7" captionText="FIG. 3. — Plastral remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-F, Pal. 1315 (hypoplastron); A, B, visceral; C, D, ventral; E, F, left lateral; G-L, Pal. 1316 (hypoplastron); G, H, visceral; I, J, ventral; K, L, right lateral; M-R, Pal. 1317 (hypoplastron); M, N, visceral; O, P, ventral; Q, R, right lateral; S-V, Pal. 1318 (xiphiplastron); S, T, visceral; U, V, ventral views; W, reconstruction of plastron. Thick lines indicate to scute sulci, dotted lines denote plate sutures, dashed lines denote completely ligamentous kinesis (between hypoplastra and peripherals and the hinge between hyoplastron and hypoplastron), and oblique lines indicate missing plate portions. Abbreviations: Ab, abdominal; An, anal; ent, entoplastron; epi, epiplastron; hyo, hyoplastron; hyp, hypoplastron; Fe, femoral; Gu, gular; Hu, humeral; Pe, pectoral; xi, xiphiplastron. Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5679125" httpUri="https://zenodo.org/record/5679125/files/figure.png" pageId="5" pageNumber="694">3</figureCitation>
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)
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</paragraph>
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<paragraph id="DA2936AEFF91DC5BFC8AFD73E248FD5D" blockId="5.[813,1455,655,705]" pageId="5" pageNumber="694">
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LOCALITIES. —
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<materialsCitation id="6AFE3CF3FF91DC5BFC6CFD73E248FD5D" ID-GBIF-Occurrence="3400720303" pageId="5" pageNumber="694">MWQ1/2001, MWQ2/2003, MWQTC/2001, MWQ4/2018 and MCQ3/2005.</materialsCitation>
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</paragraph>
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<paragraph id="DA2936AEFF91DC5BFC8AFD25E2EDFC43" blockId="5.[811,1457,729,991]" pageId="5" pageNumber="694">
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STUDIED MATERIAL. —
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<materialsCitation id="6AFE3CF3FF91DC5BFBBCFD26E2EDFC43" ID-GBIF-Occurrence="3400720307" pageId="5" pageNumber="694">
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<emphasis id="E8E2EABCFF91DC5BFBBCFD26E286FD6E" bold="true" box="[1051,1208,730,754]" pageId="5" pageNumber="694">
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<collectingCountry id="A281763EFF91DC5BFBBCFD26E286FD6E" box="[1051,1208,730,754]" name="Czech Republic" pageId="5" pageNumber="694">Czech Republic</collectingCountry>
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</emphasis>
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.
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<collectingRegion id="1852F84CFF91DC5BFB60FD25E392FD6D" box="[1223,1452,729,753]" country="Czech Republic" name="Jihomoravsky" pageId="5" pageNumber="694">South Moravia Region</collectingRegion>
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, Mokrá-Quarry, carapace remains (
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<figureCitation id="42AD2A2BFF91DC5BFB25FD0FE282FC97" box="[1154,1212,755,779]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="5" pageNumber="694">Fig. 2</figureCitation>
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A-R’): Pal. 1300, nuchal and left peripherals 1-3; Pal. 1301, nuchal; Pal. 1302, neural 4; Pal. 1303, neural 5; Pal. 1304, neural 7; Pal. 1305, suprapygal 1; Pal. 1306, suprapygal 2; Pal. 1307, left costal 1; Pal. 1308, right costal 6; Pal. 1309, right costal 6; Pal. 1310, left costal 8; Pal. 1311, right costal 8; Pal. 1312, right peripheral 1; Pal. 1313, left peripheral 7; Pal. 1314, left peripheral 8. Plastral remains (
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<figureCitation id="42AD2A2BFF91DC5BFB7BFC6EE32CFC36" box="[1244,1298,914,938]" captionStart="FIG" captionStartId="7.[132,143,1894,1911]" captionTargetBox="[139,1445,213,1863]" captionTargetId="figure-16@7.[133,1455,207,1869]" captionTargetPageId="7" captionText="FIG. 3. — Plastral remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-F, Pal. 1315 (hypoplastron); A, B, visceral; C, D, ventral; E, F, left lateral; G-L, Pal. 1316 (hypoplastron); G, H, visceral; I, J, ventral; K, L, right lateral; M-R, Pal. 1317 (hypoplastron); M, N, visceral; O, P, ventral; Q, R, right lateral; S-V, Pal. 1318 (xiphiplastron); S, T, visceral; U, V, ventral views; W, reconstruction of plastron. Thick lines indicate to scute sulci, dotted lines denote plate sutures, dashed lines denote completely ligamentous kinesis (between hypoplastra and peripherals and the hinge between hyoplastron and hypoplastron), and oblique lines indicate missing plate portions. Abbreviations: Ab, abdominal; An, anal; ent, entoplastron; epi, epiplastron; hyo, hyoplastron; hyp, hypoplastron; Fe, femoral; Gu, gular; Hu, humeral; Pe, pectoral; xi, xiphiplastron. Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5679125" httpUri="https://zenodo.org/record/5679125/files/figure.png" pageId="5" pageNumber="694">Fig. 3</figureCitation>
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A-R): Pal. 1315, left hypoplastron; Pal. 1316, right hypoplastron; Pal. 1317, right hypoplastron; Pal. 1318, left xiphiplastron.
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</materialsCitation>
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</paragraph>
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<paragraph id="DA2936AEFF91DC5BFC8AFBE9E583FBB1" blockId="5.[810,1457,1045,1900]" box="[813,957,1045,1071]" pageId="5" pageNumber="694">DESCRIPTION</paragraph>
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<paragraph id="DA2936AEFF91DC5BFC8AFBC8E294F9D0" blockId="5.[810,1457,1045,1900]" pageId="5" pageNumber="694">
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The ptychogasterid material (Pal. 1300-1318) from Mokrá- Quarry consists of numerous disarticulated plates, which present preserved parts of the almost the entire shell. However, some plates are missing from the carapace (i.e., pygal, costals 2-4 and 7, neurals 1-3 and both medial and posterior peripherals). The plastron is represented only by the hypoplastron and the xiphiplastron. The following description is based on all available material, however, not all preserved plates are depicted in
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<figureCitation id="42AD2A2BFF91DC5BFC0CFACFE233FAD1" box="[939,1037,1331,1357]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="5" pageNumber="694">Figures 2</figureCitation>
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and
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<figureCitation id="42AD2A2BFF91DC5BFBE5FACFE26EFAD1" box="[1090,1104,1331,1357]" captionStart="FIG" captionStartId="7.[132,143,1894,1911]" captionTargetBox="[139,1445,213,1863]" captionTargetId="figure-16@7.[133,1455,207,1869]" captionTargetPageId="7" captionText="FIG. 3. — Plastral remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-F, Pal. 1315 (hypoplastron); A, B, visceral; C, D, ventral; E, F, left lateral; G-L, Pal. 1316 (hypoplastron); G, H, visceral; I, J, ventral; K, L, right lateral; M-R, Pal. 1317 (hypoplastron); M, N, visceral; O, P, ventral; Q, R, right lateral; S-V, Pal. 1318 (xiphiplastron); S, T, visceral; U, V, ventral views; W, reconstruction of plastron. Thick lines indicate to scute sulci, dotted lines denote plate sutures, dashed lines denote completely ligamentous kinesis (between hypoplastra and peripherals and the hinge between hyoplastron and hypoplastron), and oblique lines indicate missing plate portions. Abbreviations: Ab, abdominal; An, anal; ent, entoplastron; epi, epiplastron; hyo, hyoplastron; hyp, hypoplastron; Fe, femoral; Gu, gular; Hu, humeral; Pe, pectoral; xi, xiphiplastron. Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5679125" httpUri="https://zenodo.org/record/5679125/files/figure.png" pageId="5" pageNumber="694">3</figureCitation>
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. According to the dimensions of the nuchal plate (Pal. 1300-1301), costals (Pal. 1307-1311) and peripherals (Pal. 1312-1314), the shell would have been relatively large (>
|
||
<quantity id="1D6E9B4BFF91DC5BFC4CFA6FE20DFA32" box="[1003,1075,1427,1454]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="694" unit="cm" value="20.0">20 cm</quantity>
|
||
:
|
||
<figureCitation id="42AD2A2BFF91DC5BFB9AFA6FE243FA31" box="[1085,1149,1427,1453]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="5" pageNumber="694">Figs 2</figureCitation>
|
||
;
|
||
<figureCitation id="42AD2A2BFF91DC5BFB2CFA6FE2A7FA31" box="[1163,1177,1427,1453]" captionStart="FIG" captionStartId="7.[132,143,1894,1911]" captionTargetBox="[139,1445,213,1863]" captionTargetId="figure-16@7.[133,1455,207,1869]" captionTargetPageId="7" captionText="FIG. 3. — Plastral remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-F, Pal. 1315 (hypoplastron); A, B, visceral; C, D, ventral; E, F, left lateral; G-L, Pal. 1316 (hypoplastron); G, H, visceral; I, J, ventral; K, L, right lateral; M-R, Pal. 1317 (hypoplastron); M, N, visceral; O, P, ventral; Q, R, right lateral; S-V, Pal. 1318 (xiphiplastron); S, T, visceral; U, V, ventral views; W, reconstruction of plastron. Thick lines indicate to scute sulci, dotted lines denote plate sutures, dashed lines denote completely ligamentous kinesis (between hypoplastra and peripherals and the hinge between hyoplastron and hypoplastron), and oblique lines indicate missing plate portions. Abbreviations: Ab, abdominal; An, anal; ent, entoplastron; epi, epiplastron; hyo, hyoplastron; hyp, hypoplastron; Fe, femoral; Gu, gular; Hu, humeral; Pe, pectoral; xi, xiphiplastron. Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5679125" httpUri="https://zenodo.org/record/5679125/files/figure.png" pageId="5" pageNumber="694">3</figureCitation>
|
||
). Ornamentation of both carapace and plastron external surfaces is absent. As it is typical for
|
||
<taxonomicName id="1D964D2DFF91DC5BFCD5FA2FE5D1FA71" authorityName="Pomel" authorityYear="1847" box="[882,1007,1491,1517]" class="Reptilia" family="Geoemydidae" genus="Ptychogaster" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="694" phylum="Chordata" rank="genus">
|
||
<emphasis id="E8E2EABCFF91DC5BFCD5FA2FE5D1FA71" box="[882,1007,1491,1517]" italics="true" pageId="5" pageNumber="694">Ptychogaster</emphasis>
|
||
</taxonomicName>
|
||
plates, dermal grooves are well developed. Unlike
|
||
<taxonomicName id="1D964D2DFF91DC5BFCD9FA0FE5F2F991" authorityName="Linnaeus" authorityYear="1758" box="[894,972,1523,1549]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="694" phylum="Chordata" rank="genus">
|
||
<emphasis id="E8E2EABCFF91DC5BFCD9FA0FE5F2F991" box="[894,972,1523,1549]" italics="true" pageId="5" pageNumber="694">Testudo</emphasis>
|
||
</taxonomicName>
|
||
, neither growth lines nor sutures (i.e., completely fused to each other: see e.g.,
|
||
<figureCitation id="42AD2A2BFF91DC5BFB05F9EEE2E0F9B1" box="[1186,1246,1554,1581]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="5" pageNumber="694">Fig. 2</figureCitation>
|
||
A-D) are preserved, so it is impossible to evaluate them.
|
||
</paragraph>
|
||
<paragraph id="DA2936AEFF91DC5BFCE4F9AEE391F8F0" blockId="5.[810,1457,1045,1900]" pageId="5" pageNumber="694">
|
||
The nuchal plate is hexagonal in outline, wider than long (Pal. 1301:
|
||
<figureCitation id="42AD2A2BFF91DC5BFC09F98EE5D3F910" box="[942,1005,1650,1676]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="5" pageNumber="694">Fig. 2</figureCitation>
|
||
E-H). It contacts the first pair of peripherals, the first pair of costals and the neural 1. The anterior border possesses a shallow nuchal notch, affecting the nuchal border and the first peripherals (Pal. 1300:
|
||
<figureCitation id="42AD2A2BFF91DC5BFAA7F92EE34CF970" box="[1280,1394,1746,1772]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="5" pageNumber="694">Fig. 2A, B</figureCitation>
|
||
). The anteroposterolateral sides of the nuchal are rather equal in length, whereas the posterior border is narrow and slightly convex anteriorly (
|
||
<figureCitation id="42AD2A2BFF91DC5BFC5FF8CDE256F8D0" box="[1016,1128,1841,1868]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="5" pageNumber="694">Fig. 2E, F</figureCitation>
|
||
). In lateral view, the nuchal is vaguely curved (
|
||
<figureCitation id="42AD2A2BFF91DC5BFC47F8ADE278F8F0" box="[992,1094,1873,1900]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="5" pageNumber="694">Fig. 2I, J</figureCitation>
|
||
). Two transversal thickenings are
|
||
</paragraph>
|
||
<caption id="8EE96626FF91DC5BFF22F856E391F875" ID-DOI="http://doi.org/10.5281/zenodo.5679119" ID-Zenodo-Dep="5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="5" pageNumber="694" startId="5.[133,143,1962,1979]" targetBox="[142,1449,216,1871]" targetPageId="6">
|
||
<paragraph id="DA2936AEFF91DC5BFF22F856E391F875" blockId="5.[132,1456,1962,2025]" pageId="5" pageNumber="694">
|
||
FIG. 2. — Carapace remains of
|
||
<taxonomicName id="1D964D2DFF91DC5BFE23F856E450F827" authority="(Ptychogaster)" box="[388,622,1962,1979]" class="Reptilia" family="Geoemydidae" genus="Ptychogaster" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="694" phylum="Chordata" rank="subGenus" subGenus="Ptychogaster">
|
||
<emphasis id="E8E2EABCFF91DC5BFE23F856E7CCF827" box="[388,498,1962,1979]" italics="true" pageId="5" pageNumber="694">Ptychogaster</emphasis>
|
||
(
|
||
<emphasis id="E8E2EABCFF91DC5BFE5CF856E457F827" box="[507,617,1962,1979]" italics="true" pageId="5" pageNumber="694">Ptychogaster</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
sp. from Mokrá-Quarry localities:
|
||
<emphasis id="E8E2EABCFF91DC5BFC2FF856E594F827" bold="true" box="[904,938,1962,1979]" pageId="5" pageNumber="694">A -D</emphasis>
|
||
, Pal. 1300 (nuchal and peripherals 1-3);
|
||
<emphasis id="E8E2EABCFF91DC5BFB50F856E33AF827" bold="true" box="[1271,1284,1962,1979]" pageId="5" pageNumber="694">A</emphasis>
|
||
,
|
||
<emphasis id="E8E2EABCFF91DC5BFAAAF856E324F827" bold="true" box="[1293,1306,1962,1979]" pageId="5" pageNumber="694">B</emphasis>
|
||
, dorsal;
|
||
<emphasis id="E8E2EABCFF91DC5BFAC5F856E34EF827" bold="true" box="[1378,1392,1962,1979]" pageId="5" pageNumber="694">C</emphasis>
|
||
,
|
||
<emphasis id="E8E2EABCFF91DC5BFADEF856E3B9F827" bold="true" box="[1401,1415,1962,1979]" pageId="5" pageNumber="694">D</emphasis>
|
||
, visceral;
|
||
<emphasis id="E8E2EABCFF91DC5BFF11F83DE6EBF84E" bold="true" box="[182,213,1985,2002]" pageId="5" pageNumber="694">E -L</emphasis>
|
||
, Pal.1301 (nuchal);
|
||
<emphasis id="E8E2EABCFF91DC5BFEDCF83DE7B9F84E" bold="true" box="[379,391,1985,2002]" pageId="5" pageNumber="694">E</emphasis>
|
||
,
|
||
<emphasis id="E8E2EABCFF91DC5BFE36F83DE7A2F84E" bold="true" box="[401,412,1985,2002]" pageId="5" pageNumber="694">F</emphasis>
|
||
, dorsal;
|
||
<emphasis id="E8E2EABCFF91DC5BFE44F83DE7CFF84E" bold="true" box="[483,497,1985,2002]" pageId="5" pageNumber="694">G</emphasis>
|
||
,
|
||
<emphasis id="E8E2EABCFF91DC5BFE5DF83DE436F84E" bold="true" box="[506,520,1985,2002]" pageId="5" pageNumber="694">H</emphasis>
|
||
, visceral;
|
||
<emphasis id="E8E2EABCFF91DC5BFDFCF83DE45FF84E" bold="true" box="[603,609,1985,2002]" pageId="5" pageNumber="694">I</emphasis>
|
||
,
|
||
<emphasis id="E8E2EABCFF91DC5BFDCDF83DE44AF84E" bold="true" box="[618,628,1985,2002]" pageId="5" pageNumber="694">J</emphasis>
|
||
, left lateral;
|
||
<emphasis id="E8E2EABCFF91DC5BFD7FF83DE4DBF84E" bold="true" box="[728,741,1985,2002]" pageId="5" pageNumber="694">K</emphasis>
|
||
,
|
||
<emphasis id="E8E2EABCFF91DC5BFD48F83DE4C4F84E" bold="true" box="[751,762,1985,2002]" pageId="5" pageNumber="694">L</emphasis>
|
||
, anterior;
|
||
<emphasis id="E8E2EABCFF91DC5BFCEAF83DE560F84E" bold="true" box="[845,862,1985,2002]" pageId="5" pageNumber="694">M</emphasis>
|
||
,
|
||
<emphasis id="E8E2EABCFF91DC5BFCCFF83DE548F84E" bold="true" box="[872,886,1985,2002]" pageId="5" pageNumber="694">N</emphasis>
|
||
, Pal. 1302 (neural 4);
|
||
<emphasis id="E8E2EABCFF91DC5BFB81F83DE209F84E" bold="true" box="[1062,1079,1985,2002]" pageId="5" pageNumber="694">M</emphasis>
|
||
, dorsal;
|
||
<emphasis id="E8E2EABCFF91DC5BFBD9F83DE2B2F84E" bold="true" box="[1150,1164,1985,2002]" pageId="5" pageNumber="694">N</emphasis>
|
||
, visceral;
|
||
<emphasis id="E8E2EABCFF91DC5BFB79F83DE33DF84E" bold="true" box="[1246,1283,1985,2002]" pageId="5" pageNumber="694">O -Q</emphasis>
|
||
, Pal. 1303 (neural 5);
|
||
<emphasis id="E8E2EABCFF91DC5BFF23F824E698F875" bold="true" box="[132,166,2008,2025]" pageId="5" pageNumber="694">O -P</emphasis>
|
||
, dorsal;
|
||
<emphasis id="E8E2EABCFF91DC5BFF4AF824E6C2F875" bold="true" box="[237,252,2008,2025]" pageId="5" pageNumber="694">Q</emphasis>
|
||
, visceral;
|
||
<emphasis id="E8E2EABCFF91DC5BFEE9F824E765F875" bold="true" box="[334,347,2008,2025]" pageId="5" pageNumber="694">R</emphasis>
|
||
,
|
||
<emphasis id="E8E2EABCFF91DC5BFEC2F824E74FF875" bold="true" box="[357,369,2008,2025]" pageId="5" pageNumber="694">S</emphasis>
|
||
, Pal. 1304 (neural 7);
|
||
<emphasis id="E8E2EABCFF91DC5BFD85F824E411F875" bold="true" box="[546,559,2008,2025]" pageId="5" pageNumber="694">R</emphasis>
|
||
, dorsal;
|
||
<emphasis id="E8E2EABCFF91DC5BFDD0F824E4BDF875" bold="true" box="[631,643,2008,2025]" pageId="5" pageNumber="694">S</emphasis>
|
||
, visceral.
|
||
<emphasis id="E8E2EABCFF91DC5BFD72F824E4DEF875" bold="true" box="[725,736,2008,2025]" pageId="5" pageNumber="694">T</emphasis>
|
||
,
|
||
<emphasis id="E8E2EABCFF91DC5BFD4DF824E4C6F875" bold="true" box="[746,760,2008,2025]" pageId="5" pageNumber="694">U</emphasis>
|
||
, Pal. 1305 (suprapygals 1-2);
|
||
<emphasis id="E8E2EABCFF91DC5BFC4AF824E5C6F875" bold="true" box="[1005,1016,2008,2025]" pageId="5" pageNumber="694">T</emphasis>
|
||
, dorsal;
|
||
<emphasis id="E8E2EABCFF91DC5BFB98F824E273F875" bold="true" box="[1087,1101,2008,2025]" pageId="5" pageNumber="694">U</emphasis>
|
||
, visceral;
|
||
<emphasis id="E8E2EABCFF91DC5BFB07F824E292F875" bold="true" box="[1184,1196,2008,2025]" pageId="5" pageNumber="694">V</emphasis>
|
||
,
|
||
<emphasis id="E8E2EABCFF91DC5BFB12F824E2F9F875" bold="true" box="[1205,1223,2008,2025]" pageId="5" pageNumber="694">W</emphasis>
|
||
, Pal. 1307 (costal 1);
|
||
<emphasis id="E8E2EABCFF91DC5BFAD1F824E3BCF875" bold="true" box="[1398,1410,2008,2025]" pageId="5" pageNumber="694">V</emphasis>
|
||
, dor-
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="8EE96626FF92DC58FF23F881E729F875" ID-DOI="http://doi.org/10.5281/zenodo.5679123" ID-Zenodo-Dep="5679123" httpUri="https://zenodo.org/record/5679123/files/figure.png" pageId="6" pageNumber="695" startId="6.[132,158,1917,1934]" subCaptionStartIDs="6.[257,375,1985,2002]" subCaptionStarts="Abbr" targetBox="[142,1449,216,1871]" targetPageId="6">
|
||
<paragraph id="DA2936AEFF92DC58FF23F881E729F875" blockId="6.[132,1457,1917,2025]" pageId="6" pageNumber="695">
|
||
sal;
|
||
<emphasis id="E8E2EABCFF92DC58FF01F881E686F812" bold="true" box="[166,184,1917,1934]" pageId="6" pageNumber="695">W</emphasis>
|
||
, visceral;
|
||
<emphasis id="E8E2EABCFF92DC58FEACF881E715F812" bold="true" box="[267,299,1917,1934]" pageId="6" pageNumber="695">X -Z</emphasis>
|
||
, Pal. 1308 (costal 5);
|
||
<emphasis id="E8E2EABCFF92DC58FE7AF881E7D7F812" bold="true" box="[477,489,1917,1934]" pageId="6" pageNumber="695">X</emphasis>
|
||
,
|
||
<emphasis id="E8E2EABCFF92DC58FE54F881E7C1F812" bold="true" box="[499,511,1917,1934]" pageId="6" pageNumber="695">Y</emphasis>
|
||
, dorsal;
|
||
<emphasis id="E8E2EABCFF92DC58FDE0F881E46DF812" bold="true" box="[583,595,1917,1934]" pageId="6" pageNumber="695">Z</emphasis>
|
||
, visceral;
|
||
<emphasis id="E8E2EABCFF92DC58FD00F881E4EDF812" bold="true" box="[679,723,1917,1934]" pageId="6" pageNumber="695">A’-C’</emphasis>
|
||
, Pal. 1309 (costal 6);
|
||
<emphasis id="E8E2EABCFF92DC58FC22F881E5A9F812" bold="true" box="[901,919,1917,1934]" pageId="6" pageNumber="695">A’</emphasis>
|
||
,
|
||
<emphasis id="E8E2EABCFF92DC58FC06F881E58DF812" bold="true" box="[929,947,1917,1934]" pageId="6" pageNumber="695">B’</emphasis>
|
||
, dorsal;
|
||
<emphasis id="E8E2EABCFF92DC58FC5CF881E230F812" bold="true" box="[1019,1038,1917,1934]" pageId="6" pageNumber="695">C’</emphasis>
|
||
, visceral;
|
||
<emphasis id="E8E2EABCFF92DC58FBC6F881E2B2F812" bold="true" box="[1121,1164,1917,1934]" pageId="6" pageNumber="695">D’-F’</emphasis>
|
||
, Pal. 1310 (costal 8);
|
||
<emphasis id="E8E2EABCFF92DC58FA99F881E36FF812" bold="true" box="[1342,1361,1917,1934]" pageId="6" pageNumber="695">D’</emphasis>
|
||
,
|
||
<emphasis id="E8E2EABCFF92DC58FAFCF881E352F812" bold="true" box="[1371,1388,1917,1934]" pageId="6" pageNumber="695">E’</emphasis>
|
||
, dorsal;
|
||
<emphasis id="E8E2EABCFF92DC58FF23F868E6AAF839" bold="true" box="[132,148,1940,1957]" pageId="6" pageNumber="695">F’</emphasis>
|
||
, visceral;
|
||
<emphasis id="E8E2EABCFF92DC58FF4FF868E72CF839" bold="true" box="[232,274,1940,1957]" pageId="6" pageNumber="695">G’-J’</emphasis>
|
||
, Pal. 1312 (peripheral 1);
|
||
<emphasis id="E8E2EABCFF92DC58FE43F868E7C9F839" bold="true" box="[484,503,1940,1957]" pageId="6" pageNumber="695">G’</emphasis>
|
||
,
|
||
<emphasis id="E8E2EABCFF92DC58FDA5F868E42BF839" bold="true" box="[514,533,1940,1957]" pageId="6" pageNumber="695">H’</emphasis>
|
||
, dorsal;
|
||
<emphasis id="E8E2EABCFF92DC58FDFBF868E459F839" bold="true" box="[604,615,1940,1957]" pageId="6" pageNumber="695">I’</emphasis>
|
||
,
|
||
<emphasis id="E8E2EABCFF92DC58FDD6F868E4BFF839" bold="true" box="[625,641,1940,1957]" pageId="6" pageNumber="695">J’</emphasis>
|
||
, visceral;
|
||
<emphasis id="E8E2EABCFF92DC58FD74F868E53EF839" bold="true" box="[723,768,1940,1957]" pageId="6" pageNumber="695">K’-N’</emphasis>
|
||
, Pal. 1313 (peripheral 7);
|
||
<emphasis id="E8E2EABCFF92DC58FC74F868E5D8F839" bold="true" box="[979,998,1940,1957]" pageId="6" pageNumber="695">K’</emphasis>
|
||
,
|
||
<emphasis id="E8E2EABCFF92DC58FC48F868E5C1F839" bold="true" box="[1007,1023,1940,1957]" pageId="6" pageNumber="695">L’</emphasis>
|
||
, dorsal;
|
||
<emphasis id="E8E2EABCFF92DC58FBE0F868E263F839" bold="true" box="[1095,1117,1940,1957]" pageId="6" pageNumber="695">M’</emphasis>
|
||
,
|
||
<emphasis id="E8E2EABCFF92DC58FBC0F868E244F839" bold="true" box="[1127,1146,1940,1957]" pageId="6" pageNumber="695">N’</emphasis>
|
||
, visceral;
|
||
<emphasis id="E8E2EABCFF92DC58FB6AF868E2C5F839" bold="true" box="[1229,1275,1940,1957]" pageId="6" pageNumber="695">O’-R’</emphasis>
|
||
, Pal. 1314 (peripheral 8);
|
||
<emphasis id="E8E2EABCFF92DC58FF39F856E68CF827" bold="true" box="[158,178,1962,1979]" pageId="6" pageNumber="695">O’</emphasis>
|
||
,
|
||
<emphasis id="E8E2EABCFF92DC58FF1BF856E6F0F827" bold="true" box="[188,206,1962,1979]" pageId="6" pageNumber="695">P’</emphasis>
|
||
, dorsal;
|
||
<emphasis id="E8E2EABCFF92DC58FEB1F856E714F827" bold="true" box="[278,298,1962,1979]" pageId="6" pageNumber="695">Q’</emphasis>
|
||
,
|
||
<emphasis id="E8E2EABCFF92DC58FE93F856E779F827" bold="true" box="[308,327,1962,1979]" pageId="6" pageNumber="695">R’</emphasis>
|
||
, visceral views;
|
||
<emphasis id="E8E2EABCFF92DC58FE68F856E7DEF827" bold="true" box="[463,480,1962,1979]" pageId="6" pageNumber="695">S’</emphasis>
|
||
, reconstruction of carapace. Thick lines indicate to scute sulci, dotted lines sutures and oblique lines denote missing plate portions. Abbreviations:
|
||
<emphasis id="E8E2EABCFF92DC58FED9F83DE7A9F84E" bold="true" box="[382,407,1985,2002]" pageId="6" pageNumber="695">Ce</emphasis>
|
||
, cervical;
|
||
<emphasis id="E8E2EABCFF92DC58FE4CF83DE43FF84E" bold="true" box="[491,513,1985,2002]" pageId="6" pageNumber="695">co</emphasis>
|
||
, costal;
|
||
<emphasis id="E8E2EABCFF92DC58FDEFF83DE45AF84E" bold="true" box="[584,612,1985,2002]" pageId="6" pageNumber="695">Ma</emphasis>
|
||
, marginal;
|
||
<emphasis id="E8E2EABCFF92DC58FD67F83DE4E8F84E" bold="true" box="[704,726,1985,2002]" pageId="6" pageNumber="695">ne</emphasis>
|
||
, neural;
|
||
<emphasis id="E8E2EABCFF92DC58FCBAF83DE50DF84E" bold="true" box="[797,819,1985,2002]" pageId="6" pageNumber="695">nu</emphasis>
|
||
, nuchal;
|
||
<emphasis id="E8E2EABCFF92DC58FC27F83DE5A3F84E" bold="true" box="[896,925,1985,2002]" pageId="6" pageNumber="695">per</emphasis>
|
||
, peripheral;
|
||
<emphasis id="E8E2EABCFF92DC58FBA2F83DE228F84E" bold="true" box="[1029,1046,1985,2002]" pageId="6" pageNumber="695">Pl</emphasis>
|
||
, pleural;
|
||
<emphasis id="E8E2EABCFF92DC58FBC4F83DE246F84E" bold="true" box="[1123,1144,1985,2002]" pageId="6" pageNumber="695">py</emphasis>
|
||
, pygal;
|
||
<emphasis id="E8E2EABCFF92DC58FB1DF83DE2F1F84E" bold="true" box="[1210,1231,1985,2002]" pageId="6" pageNumber="695">sp</emphasis>
|
||
, suprapygal;
|
||
<emphasis id="E8E2EABCFF92DC58FAE7F83DE368F84E" bold="true" box="[1344,1366,1985,2002]" pageId="6" pageNumber="695">Ve</emphasis>
|
||
, vertebral. Scale bars:
|
||
<quantity id="1D6E9B4BFF92DC58FF40F824E72DF875" box="[231,275,2008,2025]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="695" unit="cm" value="1.0">1 cm</quantity>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="8EE96626FF93DC59FF23F89AE579F875" ID-DOI="http://doi.org/10.5281/zenodo.5679125" ID-Zenodo-Dep="5679125" httpUri="https://zenodo.org/record/5679125/files/figure.png" pageId="7" pageNumber="696" startId="7.[132,143,1894,1911]" targetBox="[139,1445,213,1863]" targetPageId="7">
|
||
<paragraph id="DA2936AEFF93DC59FF23F89AE579F875" blockId="7.[132,1457,1894,2025]" pageId="7" pageNumber="696">
|
||
FIG. 3. — Plastral remains of
|
||
<taxonomicName id="1D964D2DFF93DC59FED2F89AE45EF8EB" authority="(Ptychogaster)" box="[373,608,1894,1911]" class="Reptilia" family="Geoemydidae" genus="Ptychogaster" kingdom="Animalia" order="Testudines" pageId="7" pageNumber="696" phylum="Chordata" rank="subGenus" subGenus="Ptychogaster">
|
||
<emphasis id="E8E2EABCFF93DC59FED2F89AE7DDF8EB" box="[373,483,1894,1911]" italics="true" pageId="7" pageNumber="696">Ptychogaster</emphasis>
|
||
(
|
||
<emphasis id="E8E2EABCFF93DC59FE4AF89AE465F8EB" box="[493,603,1894,1911]" italics="true" pageId="7" pageNumber="696">Ptychogaster</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
sp. from Mokrá-Quarry localities:
|
||
<emphasis id="E8E2EABCFF93DC59FCDAF89AE5A2F8EB" bold="true" box="[893,924,1894,1911]" pageId="7" pageNumber="696">A -F</emphasis>
|
||
, Pal. 1315 (hypoplastron);
|
||
<emphasis id="E8E2EABCFF93DC59FBDBF89AE2B7F8EB" bold="true" box="[1148,1161,1894,1911]" pageId="7" pageNumber="696">A</emphasis>
|
||
,
|
||
<emphasis id="E8E2EABCFF93DC59FB33F89AE29FF8EB" bold="true" box="[1172,1185,1894,1911]" pageId="7" pageNumber="696">B</emphasis>
|
||
, visceral;
|
||
<emphasis id="E8E2EABCFF93DC59FB52F89AE33DF8EB" bold="true" box="[1269,1283,1894,1911]" pageId="7" pageNumber="696">C</emphasis>
|
||
,
|
||
<emphasis id="E8E2EABCFF93DC59FAA9F89AE322F8EB" bold="true" box="[1294,1308,1894,1911]" pageId="7" pageNumber="696">D</emphasis>
|
||
, ventral;
|
||
<emphasis id="E8E2EABCFF93DC59FACEF89AE34BF8EB" bold="true" box="[1385,1397,1894,1911]" pageId="7" pageNumber="696">E</emphasis>
|
||
,
|
||
<emphasis id="E8E2EABCFF93DC59FA27F89AE3B5F8EB" bold="true" box="[1408,1419,1894,1911]" pageId="7" pageNumber="696">F</emphasis>
|
||
, left lateral;
|
||
<emphasis id="E8E2EABCFF93DC59FF65F881E6DCF812" bold="true" box="[194,226,1917,1934]" pageId="7" pageNumber="696">G -L</emphasis>
|
||
, Pal. 1316 (hypoplastron);
|
||
<emphasis id="E8E2EABCFF93DC59FE64F881E7EFF812" bold="true" box="[451,465,1917,1934]" pageId="7" pageNumber="696">G</emphasis>
|
||
,
|
||
<emphasis id="E8E2EABCFF93DC59FE7BF881E7D4F812" bold="true" box="[476,490,1917,1934]" pageId="7" pageNumber="696">H</emphasis>
|
||
, visceral;
|
||
<emphasis id="E8E2EABCFF93DC59FD98F881E47BF812" bold="true" box="[575,581,1917,1934]" pageId="7" pageNumber="696">I</emphasis>
|
||
,
|
||
<emphasis id="E8E2EABCFF93DC59FDE8F881E467F812" bold="true" box="[591,601,1917,1934]" pageId="7" pageNumber="696">J</emphasis>
|
||
, ventral;
|
||
<emphasis id="E8E2EABCFF93DC59FD00F881E48AF812" bold="true" box="[679,692,1917,1934]" pageId="7" pageNumber="696">K</emphasis>
|
||
,
|
||
<emphasis id="E8E2EABCFF93DC59FD67F881E4F5F812" bold="true" box="[704,715,1917,1934]" pageId="7" pageNumber="696">L</emphasis>
|
||
, right lateral;
|
||
<emphasis id="E8E2EABCFF93DC59FC99F881E55DF812" bold="true" box="[830,867,1917,1934]" pageId="7" pageNumber="696">M -R</emphasis>
|
||
, Pal. 1317 (hypoplastron);
|
||
<emphasis id="E8E2EABCFF93DC59FBE2F881E268F812" bold="true" box="[1093,1110,1917,1934]" pageId="7" pageNumber="696">M</emphasis>
|
||
,
|
||
<emphasis id="E8E2EABCFF93DC59FBC6F881E251F812" bold="true" box="[1121,1135,1917,1934]" pageId="7" pageNumber="696">N</emphasis>
|
||
, visceral;
|
||
<emphasis id="E8E2EABCFF93DC59FB64F881E2ECF812" bold="true" box="[1219,1234,1917,1934]" pageId="7" pageNumber="696">O</emphasis>
|
||
,
|
||
<emphasis id="E8E2EABCFF93DC59FB7AF881E2D7F812" bold="true" box="[1245,1257,1917,1934]" pageId="7" pageNumber="696">P</emphasis>
|
||
, ventral;
|
||
<emphasis id="E8E2EABCFF93DC59FA90F881E378F812" bold="true" box="[1335,1350,1917,1934]" pageId="7" pageNumber="696">Q</emphasis>
|
||
,
|
||
<emphasis id="E8E2EABCFF93DC59FAF7F881E363F812" bold="true" box="[1360,1373,1917,1934]" pageId="7" pageNumber="696">R</emphasis>
|
||
, right lateral;
|
||
<emphasis id="E8E2EABCFF93DC59FF0AF868E6F3F839" bold="true" box="[173,205,1940,1957]" pageId="7" pageNumber="696">S -V</emphasis>
|
||
, Pal. 1318 (xiphiplastron);
|
||
<emphasis id="E8E2EABCFF93DC59FE0EF868E78BF839" bold="true" box="[425,437,1940,1957]" pageId="7" pageNumber="696">S</emphasis>
|
||
,
|
||
<emphasis id="E8E2EABCFF93DC59FE67F868E7F5F839" bold="true" box="[448,459,1940,1957]" pageId="7" pageNumber="696">T</emphasis>
|
||
, visceral;
|
||
<emphasis id="E8E2EABCFF93DC59FDB8F868E413F839" bold="true" box="[543,557,1940,1957]" pageId="7" pageNumber="696">U</emphasis>
|
||
,
|
||
<emphasis id="E8E2EABCFF93DC59FD9FF868E47AF839" bold="true" box="[568,580,1940,1957]" pageId="7" pageNumber="696">V</emphasis>
|
||
, ventral views;
|
||
<emphasis id="E8E2EABCFF93DC59FD62F868E4E9F839" bold="true" box="[709,727,1940,1957]" pageId="7" pageNumber="696">W</emphasis>
|
||
, reconstruction of plastron. Thick lines indicate to scute sulci, dotted lines denote plate sutures, dashed lines denote completely ligamentous kinesis (between hypoplastra and peripherals and the hinge between hyoplastron and hypoplastron), and oblique lines indicate missing plate portions. Abbreviations:
|
||
<emphasis id="E8E2EABCFF93DC59FDDBF83DE4AAF84E" bold="true" box="[636,660,1985,2002]" pageId="7" pageNumber="696">Ab</emphasis>
|
||
, abdominal;
|
||
<emphasis id="E8E2EABCFF93DC59FCA6F83DE527F84E" bold="true" box="[769,793,1985,2002]" pageId="7" pageNumber="696">An</emphasis>
|
||
, anal;
|
||
<emphasis id="E8E2EABCFF93DC59FCF6F83DE553F84E" bold="true" box="[849,877,1985,2002]" pageId="7" pageNumber="696">ent</emphasis>
|
||
, entoplastron;
|
||
<emphasis id="E8E2EABCFF93DC59FC4BF83DE239F84E" bold="true" box="[1004,1031,1985,2002]" pageId="7" pageNumber="696">epi</emphasis>
|
||
, epiplastron;
|
||
<emphasis id="E8E2EABCFF93DC59FBDDF83DE2A4F84E" bold="true" box="[1146,1178,1985,2002]" pageId="7" pageNumber="696">hyo</emphasis>
|
||
, hyoplastron;
|
||
<emphasis id="E8E2EABCFF93DC59FAB5F83DE30CF84E" bold="true" box="[1298,1330,1985,2002]" pageId="7" pageNumber="696">hyp</emphasis>
|
||
, hypoplastron;
|
||
<emphasis id="E8E2EABCFF93DC59FF23F824E6A4F875" bold="true" box="[132,154,2008,2025]" pageId="7" pageNumber="696">Fe</emphasis>
|
||
, femoral;
|
||
<emphasis id="E8E2EABCFF93DC59FF4AF824E738F875" bold="true" box="[237,262,2008,2025]" pageId="7" pageNumber="696">Gu</emphasis>
|
||
, gular;
|
||
<emphasis id="E8E2EABCFF93DC59FEE2F824E760F875" bold="true" box="[325,350,2008,2025]" pageId="7" pageNumber="696">Hu</emphasis>
|
||
, humeral;
|
||
<emphasis id="E8E2EABCFF93DC59FE11F824E7F3F875" bold="true" box="[438,461,2008,2025]" pageId="7" pageNumber="696">Pe</emphasis>
|
||
, pectoral;
|
||
<emphasis id="E8E2EABCFF93DC59FD81F824E40BF875" bold="true" box="[550,565,2008,2025]" pageId="7" pageNumber="696">xi</emphasis>
|
||
, xiphiplastron. Scale bars: 1 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="DA2936AEFF9CDC56FF23FF2BE449FE2D" blockId="8.[130,777,215,2028]" pageId="8" pageNumber="697">
|
||
developed on the internal surface of this bone (
|
||
<figureCitation id="42AD2A2BFF9CDC56FD2EFF2BE6ADFE8D" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="8" pageNumber="697">Fig. 2C, D, 2</figureCitation>
|
||
G-L). The cervical scute is present anteriorly, both dorsally and viscerally. It is a relatively large and trapezoidal element that is longer than wide (
|
||
<figureCitation id="42AD2A2BFF9CDC56FE36FECBE7C0FECD" box="[401,510,311,337]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="8" pageNumber="697">Fig. 2E, F</figureCitation>
|
||
). The lateral edges of the cervical are slightly curved medially both in dorsal and visceral sides (
|
||
<figureCitation id="42AD2A2BFF9CDC56FF58FE8BE750FE0D" box="[255,366,375,401]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="8" pageNumber="697">Fig. 2A, B</figureCitation>
|
||
, E-H). The overlap of this scute is less developed on the ventral surface (
|
||
<figureCitation id="42AD2A2BFF9CDC56FE57FE6AE457FE2D" box="[496,617,406,433]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="8" pageNumber="697">Fig. 2G, H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="DA2936AEFF9CDC56FF3CFE4AE53CFD2C" blockId="8.[130,777,215,2028]" pageId="8" pageNumber="697">
|
||
According to the preserved neural plates, an alternating between octagonal and hexagonal plates forms the neural series: neural 4 octagonal (Pal. 1302:
|
||
<figureCitation id="42AD2A2BFF9CDC56FDA6FE0AE446FD8D" box="[513,632,502,529]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="8" pageNumber="697">Fig. 2M, N</figureCitation>
|
||
) and neural 5 and 7 hexagonal with short sides behind (Pal. 1303-1304:
|
||
<figureCitation id="42AD2A2BFF9CDC56FF21FDCAE6FBFDCC" box="[134,197,566,592]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="8" pageNumber="697">Fig. 2</figureCitation>
|
||
O-S). A weak medial keel develops on the neural 7 (Pal. 1304:
|
||
<figureCitation id="42AD2A2BFF9CDC56FF50FDAAE765FDEC" box="[247,347,598,624]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="8" pageNumber="697">Fig. 2R, S</figureCitation>
|
||
) and the suprapygals 1-2 (Pal.1305-1306:
|
||
<figureCitation id="42AD2A2BFF9CDC56FF25FD8AE6CDFD0C" box="[130,243,630,656]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="8" pageNumber="697">Fig. 2T, U</figureCitation>
|
||
). In visceral view, remains of the thoracic vertebrae attachments are present in all neural plates (
|
||
<figureCitation id="42AD2A2BFF9CDC56FDF9FD6AE4CAFD2C" box="[606,756,662,688]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="8" pageNumber="697">Fig. 2N, Q, S</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="DA2936AEFF9CDC56FF3CFD49E41BFB93" blockId="8.[130,777,215,2028]" pageId="8" pageNumber="697">
|
||
The suprapygal plate 1 is trapezoidal with a wider posterior part (Pal. 1305:
|
||
<figureCitation id="42AD2A2BFF9CDC56FEE7FD29E7ABFD6C" box="[320,405,725,752]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="8" pageNumber="697">Fig. 2T</figureCitation>
|
||
). The anterior border is concave, whereas the posterior one is slightly convex. It contacts the neural 8 anteriorly, costals 8 laterally and suprapygal 2 posteriorly. Suprapygal 2 is hexagonal, wider than long, and much wider than suprapygal 1 (
|
||
<figureCitation id="42AD2A2BFF9CDC56FE45FCA9E468FCEC" box="[482,598,853,880]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="8" pageNumber="697">Fig. 2T, U</figureCitation>
|
||
). It contacts the suprapygal 1 and the posteromedial sides of costals 8 anteriorly, peripherals 11 laterally and pygal posteriorly. The anterolateral sides of the suprapygal 2 are slightly longer anteroposteriorly, compared to the posterolateral ones. The posterior side is vaguely convex anteriorly (
|
||
<figureCitation id="42AD2A2BFF9CDC56FE02FC08E426FB93" box="[421,536,1012,1039]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="8" pageNumber="697">Fig. 2T, U</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="DA2936AEFF9CDC56FF3CFBE8E4BEF991" blockId="8.[130,777,215,2028]" pageId="8" pageNumber="697">
|
||
The vertebral scute series is partially preserved, which is quadrangular and slightly narrower than the costal series. Vertebral 1 contacts the cervical and marginals 1-2 anteriorly (
|
||
<figureCitation id="42AD2A2BFF9CDC56FF2AFB88E6C4FB13" box="[141,250,1140,1167]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="8" pageNumber="697">Fig. 2B, F</figureCitation>
|
||
). It seems to be lyre-shaped and covers the lateral corners of the nuchal and costals 1 (Pal. 1300-1301:
|
||
<figureCitation id="42AD2A2BFF9CDC56FD16FB68E6EBFB52" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="8" pageNumber="697">Fig. 2A, B, E, F</figureCitation>
|
||
). According to preserved portion of the vertebral 3, it expanded at least on costals 5 and neural 5 (
|
||
<figureCitation id="42AD2A2BFF9CDC56FD21FB28E4C5FB72" box="[646,763,1236,1262]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="8" pageNumber="697">Fig. 2X, Y</figureCitation>
|
||
). The sulcus between the vertebrals 3-4 is wavy in its medial part, and more specifically in the part that is crossing the neural 5 (
|
||
<figureCitation id="42AD2A2BFF9CDC56FF62FACFE707FAD2" box="[197,313,1331,1358]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="8" pageNumber="697">Fig. 2O, P</figureCitation>
|
||
). Vertebral 4 likely contacts the vertebral 3 anteriorly, pleurals 3-4 laterally and vertebral 5 posteriorly (
|
||
<figureCitation id="42AD2A2BFF9CDC56FF2AFA8FE6E2FA12" box="[141,220,1395,1422]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="8" pageNumber="697">Fig. 2P</figureCitation>
|
||
, A’, B’, D’, E’). Vertebral 5 is the widest vertebral scute, contacting with the vertebral 4 anteriorly, pleural 4 anterolaterally and marginals 11-12 posteriorly. It expands on costals 8, neural 8, peripheral 12 and pygal, and therefore covers the entire surface of the suprapygals 1-2.
|
||
</paragraph>
|
||
<paragraph id="DA2936AEFF9CDC56FF3CF9EEE376FE8D" blockId="8.[130,777,215,2028]" lastBlockId="8.[811,1458,215,1422]" pageId="8" pageNumber="697">
|
||
Although not fully preserved costals, the costal plate 1 is much longer than the rest of costal (Pal. 1307:
|
||
<figureCitation id="42AD2A2BFF9CDC56FD26F9CEE4C5F9D1" box="[641,763,1586,1613]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="8" pageNumber="697">Fig. 2V, W</figureCitation>
|
||
). It is trapezoidal and always contacts the peripheral plates 1-3 anterolaterally, nuchal anteromedially and neurals 1-2 medially. The anterior border is sinuous to articulate with the corresponding peripherals. Costal 6 is similar in regards of its shape, being much wider than long (Pal. 1308-9:
|
||
<figureCitation id="42AD2A2BFF9CDC56FD2FF92EE4FDF970" box="[648,707,1746,1772]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="8" pageNumber="697">Fig. 2</figureCitation>
|
||
X-B’). The medial side of Pal. 1308 shows a short anteromedial and long posteromedial sides (
|
||
<figureCitation id="42AD2A2BFF9CDC56FE2AF8EEE7F9F8B0" box="[397,455,1810,1836]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="8" pageNumber="697">Fig. 2</figureCitation>
|
||
X-Z). However, the medial side of Pal. 1308 is most likely rounded (
|
||
<figureCitation id="42AD2A2BFF9CDC56FDADF8CDE476F8D0" box="[522,584,1841,1868]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="8" pageNumber="697">Fig. 2</figureCitation>
|
||
A’-C’). Costal 8 is narrow, slightly wider than long, contacting the costal 7 anteriorly, neural 8 and suprapygal 1 medially, and suprapygal 2 and peripherals 11-12 posteriorly (Pal. 1310-11:
|
||
<figureCitation id="42AD2A2BFF9CDC56FD25F86DE480F837" box="[642,702,1937,1963]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="8" pageNumber="697">Fig. 2</figureCitation>
|
||
D’-F’). The pleural scutes are not preserved with the exception of the first one. The marginopleural sulcus is entirely situated on peripheral plates, at least in both anterior and posterior part of the carapace (
|
||
<figureCitation id="42AD2A2BFF9CDC56FC7BFF0BE29DFE8E" box="[988,1187,247,274]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="8" pageNumber="697">Fig. 2A, B, E, F, V</figureCitation>
|
||
, A’, B’, D’, E’).
|
||
</paragraph>
|
||
<paragraph id="DA2936AEFF9CDC56FCE4FEEBE261FC13" blockId="8.[811,1458,215,1422]" pageId="8" pageNumber="697">
|
||
The peripheral plates 1-3 are longer than wide and slightly trapezoidal (
|
||
<figureCitation id="42AD2A2BFF9CDC56FC12FECBE5C9FECD" box="[949,1015,311,337]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="8" pageNumber="697">Fig. 2</figureCitation>
|
||
A-D, G’-J’). They are completely fused together (i.e., sutures are not visible) and crossed not only by the intermarginal sulcus, but also by the pleuro-marginal sulcus, unlike in
|
||
<taxonomicName id="1D964D2DFF9CDC56FC51FE6AE278FE2C" authorityName="Linnaeus" authorityYear="1758" box="[1014,1094,406,432]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="8" pageNumber="697" phylum="Chordata" rank="genus">
|
||
<emphasis id="E8E2EABCFF9CDC56FC51FE6AE278FE2C" box="[1014,1094,406,432]" italics="true" pageId="8" pageNumber="697">Testudo</emphasis>
|
||
</taxonomicName>
|
||
. Peripherals 1-2 are prominent in anterior direction in Pal. 1300. Peripheral 7, which is partially preserved, is rectangular (Pal. 1313:
|
||
<figureCitation id="42AD2A2BFF9CDC56FA86FE2AE361FE6C" box="[1313,1375,470,496]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="8" pageNumber="697">Fig. 2</figureCitation>
|
||
K’-N’). In internal view, it displays a rough elongated area for the cartilaginous union of the inguinal process (
|
||
<figureCitation id="42AD2A2BFF9CDC56FAB4FDEAE36DFDAC" box="[1299,1363,534,560]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="8" pageNumber="697">Fig. 2</figureCitation>
|
||
M’, N’). Pal. 1313 is the last peripheral involved in the shell bridge and also displays a weak lateral ridge on its external side. Peripheral 8 is rectangular and hosts both the pleuromarginal sulcus and intermarginal sulci (i.e., between marginals 8-9), which are situated far from the costoperipheral suture (Pal. 1314:
|
||
<figureCitation id="42AD2A2BFF9CDC56FC00FD29E5DAFD6C" box="[935,996,725,752]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="8" pageNumber="697">Fig. 2</figureCitation>
|
||
O’-R’). The marginal scute 1 is rectangular, slightly wider than long, whereas marginal 2 is trapezoidal. Marginal 3 is approximately as wide as long. Marginal 8, the only complete scute from the bridge area, is rectangular and higher than wide. The ventral overlap of all preserved marginals is well developed.
|
||
</paragraph>
|
||
<paragraph id="DA2936AEFF9CDC56FCE4FC69E2C6FA12" blockId="8.[811,1458,215,1422]" pageId="8" pageNumber="697">
|
||
The preserved portion of the hyo-hypoplastral hinge is straight and approximately transversal (Pal. 1377:
|
||
<figureCitation id="42AD2A2BFF9CDC56FA95FC49E353FC53" box="[1330,1389,949,975]" captionStart="FIG" captionStartId="7.[132,143,1894,1911]" captionTargetBox="[139,1445,213,1863]" captionTargetId="figure-16@7.[133,1455,207,1869]" captionTargetPageId="7" captionText="FIG. 3. — Plastral remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-F, Pal. 1315 (hypoplastron); A, B, visceral; C, D, ventral; E, F, left lateral; G-L, Pal. 1316 (hypoplastron); G, H, visceral; I, J, ventral; K, L, right lateral; M-R, Pal. 1317 (hypoplastron); M, N, visceral; O, P, ventral; Q, R, right lateral; S-V, Pal. 1318 (xiphiplastron); S, T, visceral; U, V, ventral views; W, reconstruction of plastron. Thick lines indicate to scute sulci, dotted lines denote plate sutures, dashed lines denote completely ligamentous kinesis (between hypoplastra and peripherals and the hinge between hyoplastron and hypoplastron), and oblique lines indicate missing plate portions. Abbreviations: Ab, abdominal; An, anal; ent, entoplastron; epi, epiplastron; hyo, hyoplastron; hyp, hypoplastron; Fe, femoral; Gu, gular; Hu, humeral; Pe, pectoral; xi, xiphiplastron. Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5679125" httpUri="https://zenodo.org/record/5679125/files/figure.png" pageId="8" pageNumber="697">Fig. 3</figureCitation>
|
||
M-P), which is located at the mid of the peripheral 6. The hypoplastron contact anteriorly with the hypoplastron through a hinge and laterally with the peripherals 6-7 through a completely ligamentous union in the inguinal process (Pal. 1315- 17:
|
||
<figureCitation id="42AD2A2BFF9CDC56FCF0FBA8E5ABFBF2" box="[855,917,1108,1134]" captionStart="FIG" captionStartId="7.[132,143,1894,1911]" captionTargetBox="[139,1445,213,1863]" captionTargetId="figure-16@7.[133,1455,207,1869]" captionTargetPageId="7" captionText="FIG. 3. — Plastral remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-F, Pal. 1315 (hypoplastron); A, B, visceral; C, D, ventral; E, F, left lateral; G-L, Pal. 1316 (hypoplastron); G, H, visceral; I, J, ventral; K, L, right lateral; M-R, Pal. 1317 (hypoplastron); M, N, visceral; O, P, ventral; Q, R, right lateral; S-V, Pal. 1318 (xiphiplastron); S, T, visceral; U, V, ventral views; W, reconstruction of plastron. Thick lines indicate to scute sulci, dotted lines denote plate sutures, dashed lines denote completely ligamentous kinesis (between hypoplastra and peripherals and the hinge between hyoplastron and hypoplastron), and oblique lines indicate missing plate portions. Abbreviations: Ab, abdominal; An, anal; ent, entoplastron; epi, epiplastron; hyo, hyoplastron; hyp, hypoplastron; Fe, femoral; Gu, gular; Hu, humeral; Pe, pectoral; xi, xiphiplastron. Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5679125" httpUri="https://zenodo.org/record/5679125/files/figure.png" pageId="8" pageNumber="697">Fig. 3</figureCitation>
|
||
A-R). The visceral overlap of both abdominal and femoral scutes on hypoplastra is well developed (
|
||
<figureCitation id="42AD2A2BFF9CDC56FA8EFB88E39CFB13" box="[1321,1442,1140,1167]" captionStart="FIG" captionStartId="7.[132,143,1894,1911]" captionTargetBox="[139,1445,213,1863]" captionTargetId="figure-16@7.[133,1455,207,1869]" captionTargetPageId="7" captionText="FIG. 3. — Plastral remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-F, Pal. 1315 (hypoplastron); A, B, visceral; C, D, ventral; E, F, left lateral; G-L, Pal. 1316 (hypoplastron); G, H, visceral; I, J, ventral; K, L, right lateral; M-R, Pal. 1317 (hypoplastron); M, N, visceral; O, P, ventral; Q, R, right lateral; S-V, Pal. 1318 (xiphiplastron); S, T, visceral; U, V, ventral views; W, reconstruction of plastron. Thick lines indicate to scute sulci, dotted lines denote plate sutures, dashed lines denote completely ligamentous kinesis (between hypoplastra and peripherals and the hinge between hyoplastron and hypoplastron), and oblique lines indicate missing plate portions. Abbreviations: Ab, abdominal; An, anal; ent, entoplastron; epi, epiplastron; hyo, hyoplastron; hyp, hypoplastron; Fe, femoral; Gu, gular; Hu, humeral; Pe, pectoral; xi, xiphiplastron. Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5679125" httpUri="https://zenodo.org/record/5679125/files/figure.png" pageId="8" pageNumber="697">Fig. 3M, N</figureCitation>
|
||
). The xiphiplastron is trapezoidal and its posterior tip is rather rounded. The abdominofemoral sulcus, developed on the ventral surface of each hypoplastron, is concave (
|
||
<figureCitation id="42AD2A2BFF9CDC56FA8CFB28E39FFB72" box="[1323,1441,1236,1262]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="8" pageNumber="697">Fig. 2N, O</figureCitation>
|
||
). Moreover, the latter does not reach the inguinal process laterally, but is located slightly below it. The femoroanal sulcus is oblique, whereas the anal scute is triangular with rounded lateral borders (
|
||
<figureCitation id="42AD2A2BFF9CDC56FC7AFAAFE223FAF1" box="[989,1053,1363,1389]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="8" pageNumber="697">Fig. 2</figureCitation>
|
||
S-V). The dorsal overlap of the anal scute is moderately developed (
|
||
<figureCitation id="42AD2A2BFF9CDC56FBDAFA8FE2D4FA12" box="[1149,1258,1395,1422]" captionStart="FIG" captionStartId="5.[133,143,1962,1979]" captionTargetId="figure-19@6.[134,1454,213,1873]" captionTargetPageId="6" captionText="FIG. 2. — Carapace remains of Ptychogaster (Ptychogaster) sp. from Mokrá-Quarry localities: A-D, Pal. 1300 (nuchal and peripherals 1-3); A, B, dorsal; C, D, visceral;E-L, Pal.1301 (nuchal); E, F, dorsal; G, H, visceral; I, J, left lateral; K, L, anterior; M, N, Pal. 1302 (neural 4); M, dorsal; N, visceral;O-Q, Pal. 1303 (neural 5); O-P, dorsal; Q, visceral;R, S, Pal. 1304 (neural 7); R, dorsal; S, visceral.T, U, Pal. 1305 (suprapygals 1-2); T, dorsal; U, visceral; V, W, Pal. 1307 (costal 1); V, dor-" figureDoi="http://doi.org/10.5281/zenodo.5679119" httpUri="https://zenodo.org/record/5679119/files/figure.png" pageId="8" pageNumber="697">Fig. 2S, T</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="928C6525FF9CDC56FC8AFA4FE5AFFA57" box="[813,913,1459,1485]" pageId="8" pageNumber="697" type="discussion">
|
||
<paragraph id="DA2936AEFF9CDC56FC8AFA4FE5AFFA57" blockId="8.[811,1457,1459,2027]" box="[813,913,1459,1485]" pageId="8" pageNumber="697">REMARKS</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="928C6525FF9CDC57FC8AFA2FE539F911" lastPageId="9" lastPageNumber="698" pageId="8" pageNumber="697" type="reference_group">
|
||
<paragraph id="DA2936AEFF9CDC56FC8AFA2FE591F877" blockId="8.[811,1457,1459,2027]" pageId="8" pageNumber="697">
|
||
<taxonomicName id="1D964D2DFF9CDC56FC8AFA2FE5E1FA71" baseAuthorityName="De Stefano" baseAuthorityYear="1903" box="[813,991,1491,1517]" class="Reptilia" family="Ptychogasteridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="8" pageNumber="697" phylum="Chordata" rank="family">Ptychogasteridae</taxonomicName>
|
||
is a geoemydid family which includes turtles of medium body size. This clade originated in Europe during the Eocene and its members are characterized mainly by a plastral kinesis (
|
||
<bibRefCitation id="BE074B5FFF9CDC56FC49F9CFE33DF9D1" author="LAPPARENT DE BROIN F." box="[1006,1283,1586,1613]" pageId="8" pageNumber="697" pagination="155 - 217" refId="ref14120" refString="LAPPARENT DE BROIN F. 2001. - The European turtle fauna from the Triassic to the Present. Dumerilia 4: 155 - 217." type="journal article" year="2001">Lapparent de Broin 2001</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="BE074B5FFF9CDC56FAB5F9CEE3A1F9D0" author="CLAUDE J." box="[1298,1439,1586,1612]" pageId="8" pageNumber="697" pagination="34 - 55" refId="ref12459" refString="CLAUDE J. 2006. - Convergence induced by plastral kinesis and phylogenetic constraints in Testudinoidea: a geometric morphometric assessment. Russia Journal of Herpetology 13 (Suppl.): 34 - 55." type="journal article" year="2006">Claude 2006</bibRefCitation>
|
||
). Thus, the anterior part of the plastron is firmly connected to the carapace, whereas the posterior one is movable thanks to a hinge situated between hyo- hypoplastra and peripherals 6. Some studies claimed that ptychogasterids constitute a monophyletic clade diagnosable by several synapomorphies (
|
||
<bibRefCitation id="BE074B5FFF9CDC56FC91F90EE5ECF890" author="HERVET S." box="[822,978,1778,1804]" pageId="8" pageNumber="697" pagination="13 - 78" refId="ref13243" refString="HERVET S. 2004 a. - Systematique du groupe " Palaeochelys sensu lato - Mauremys " (Chelonii, Testudinoidea) duTertiaire d'Europe occidentale: principaux resultats. Annales de Paleontologie 90 (1): 13 - 78. https: // doi. org / 10.1016 / j. annPal. 2003.12.002" type="journal article" year="2004">Hervet 2004a</bibRefCitation>
|
||
, b, 2006). However, the inclusion within ptychogasterids of non-kinetic extinct geoemydids genera, such as
|
||
<taxonomicName id="1D964D2DFF9CDC56FCDDF8CEE5D5F8D7" authorityName="Hervet" authorityYear="2006" box="[890,1003,1842,1867]" class="Reptilia" family="Geoemydidae" genus="Merovemys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="8" pageNumber="697" phylum="Chordata" rank="genus">
|
||
<emphasis id="E8E2EABCFF9CDC56FCDDF8CEE5D5F8D7" box="[890,1003,1842,1867]" italics="true" pageId="8" pageNumber="697">Merovemys</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="1D964D2DFF9CDC56FC5DF8CDE2B5F8D7" authorityName="Boda" authorityYear="1927" box="[1018,1163,1841,1867]" class="Reptilia" family="Emydidae" genus="Clemmydopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="8" pageNumber="697" phylum="Chordata" rank="genus">
|
||
<emphasis id="E8E2EABCFF9CDC56FC5DF8CDE2B5F8D7" box="[1018,1163,1841,1867]" italics="true" pageId="8" pageNumber="697">Clemmydopsis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="1D964D2DFF9CDC56FB1AF8CDE376F8D7" authorityName="Hervet" authorityYear="2004" box="[1213,1352,1841,1867]" class="Reptilia" family="Emydidae" genus="Hummelemys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="8" pageNumber="697" phylum="Chordata" rank="genus">
|
||
<emphasis id="E8E2EABCFF9CDC56FB1AF8CDE376F8D7" box="[1213,1352,1841,1867]" italics="true" pageId="8" pageNumber="697">Hummelemys</emphasis>
|
||
</taxonomicName>
|
||
, has been questioned (see
|
||
<bibRefCitation id="BE074B5FFF9CDC56FC7FF8ADE281F8F0" author="CLAUDE J. & TONG H." box="[984,1215,1873,1900]" pageId="8" pageNumber="697" pagination="3 - 45" refId="ref12495" refString="CLAUDE J. & TONG H. 2004. - Early Eocene testudinoid turtles from Saint-Papoul, France, with comments on the early evolution of modern Testudinoidea. Oryctos 5: 3 - 45." type="journal article" year="2004">Claude & Tong 2004</bibRefCitation>
|
||
). A more comprehensive phylogenetic analysis would be required in the future to know what genera belong to subfamily
|
||
<taxonomicName id="1D964D2DFF9CDC56FB70F86DE3B1F837" authorityName="De Stefano" authorityYear="1903" box="[1239,1423,1937,1963]" class="Reptilia" family="Geoemydidae" kingdom="Animalia" order="Testudines" pageId="8" pageNumber="697" phylum="Chordata" rank="subFamily" subFamily="Ptychogasterinae">Ptychogasterinae</taxonomicName>
|
||
, as well as to further clarify which are the closest extinct relatives of the latter.
|
||
</paragraph>
|
||
<caption id="8EE96626FF9DDC57FF23FB26E2D2FA84" ID-DOI="http://doi.org/10.5281/zenodo.5679129" ID-Zenodo-Dep="5679129" httpUri="https://zenodo.org/record/5679129/files/figure.png" pageId="9" pageNumber="698" startId="9.[132,143,1242,1259]" targetBox="[138,1448,214,1205]" targetPageId="9">
|
||
<paragraph id="DA2936AEFF9DDC57FF23FB26E2D2FA84" blockId="9.[132,1456,1242,1304]" pageId="9" pageNumber="698">
|
||
FIG. 4. — Shell and postcranial remains of
|
||
<taxonomicName id="1D964D2DFF9DDC57FE4AFB26E466FB77" box="[493,600,1242,1259]" class="Reptilia" family="Testudinidae" kingdom="Animalia" order="Testudines" pageId="9" pageNumber="698" phylum="Chordata" rank="family">Testudinidae</taxonomicName>
|
||
indet. from MWQ4/2018:
|
||
<emphasis id="E8E2EABCFF9DDC57FC91FB26E566FB77" bold="true" box="[822,856,1242,1259]" pageId="9" pageNumber="698">A -D</emphasis>
|
||
, Pal. 1363 (shell fragment);
|
||
<emphasis id="E8E2EABCFF9DDC57FBE2FB26E26CFB77" bold="true" box="[1093,1106,1242,1259]" pageId="9" pageNumber="698">A</emphasis>
|
||
,
|
||
<emphasis id="E8E2EABCFF9DDC57FBF9FB26E255FB77" bold="true" box="[1118,1131,1242,1259]" pageId="9" pageNumber="698">B</emphasis>
|
||
, dorsal;
|
||
<emphasis id="E8E2EABCFF9DDC57FB12FB26E2FDFB77" bold="true" box="[1205,1219,1242,1259]" pageId="9" pageNumber="698">C</emphasis>
|
||
,
|
||
<emphasis id="E8E2EABCFF9DDC57FB68FB26E2E3FB77" bold="true" box="[1231,1245,1242,1259]" pageId="9" pageNumber="698">D</emphasis>
|
||
, visceral;
|
||
<emphasis id="E8E2EABCFF9DDC57FA94FB26E36AFB77" bold="true" box="[1331,1364,1242,1259]" pageId="9" pageNumber="698">E -H</emphasis>
|
||
, Pal. 1364 (scapula);
|
||
<emphasis id="E8E2EABCFF9DDC57FF7DFB0CE6D8FA9D" bold="true" box="[218,230,1264,1281]" pageId="9" pageNumber="698">E</emphasis>
|
||
, lateral;
|
||
<emphasis id="E8E2EABCFF9DDC57FE89FB0CE707FA9D" bold="true" box="[302,313,1264,1281]" pageId="9" pageNumber="698">F</emphasis>
|
||
, medial;
|
||
<emphasis id="E8E2EABCFF9DDC57FE21FB0CE7AAFA9D" bold="true" box="[390,404,1264,1281]" pageId="9" pageNumber="698">G</emphasis>
|
||
, anterior;
|
||
<emphasis id="E8E2EABCFF9DDC57FE4DFB0CE7C6FA9D" bold="true" box="[490,504,1264,1281]" pageId="9" pageNumber="698">H</emphasis>
|
||
, dorsal;
|
||
<emphasis id="E8E2EABCFF9DDC57FDE6FB0CE461FA9D" bold="true" box="[577,607,1264,1281]" pageId="9" pageNumber="698">I -M</emphasis>
|
||
, Pal. 1365 (fibula);
|
||
<emphasis id="E8E2EABCFF9DDC57FCA7FB0CE538FA9D" bold="true" box="[768,774,1264,1281]" pageId="9" pageNumber="698">I</emphasis>
|
||
, dorsal;
|
||
<emphasis id="E8E2EABCFF9DDC57FCE8FB0CE567FA9D" bold="true" box="[847,857,1264,1281]" pageId="9" pageNumber="698">J</emphasis>
|
||
, ventral;
|
||
<emphasis id="E8E2EABCFF9DDC57FC00FB0CE58AFA9D" bold="true" box="[935,948,1264,1281]" pageId="9" pageNumber="698">K</emphasis>
|
||
, posterior;
|
||
<emphasis id="E8E2EABCFF9DDC57FBB2FB0CE21EFA9D" bold="true" box="[1045,1056,1264,1281]" pageId="9" pageNumber="698">L</emphasis>
|
||
, anterior;
|
||
<emphasis id="E8E2EABCFF9DDC57FBD2FB0CE2B8FA9D" bold="true" box="[1141,1158,1264,1281]" pageId="9" pageNumber="698">M</emphasis>
|
||
, lateral views. Dotted lines indicate plate sutures and oblique lines denote missing plate portions. Abbreviations:
|
||
<emphasis id="E8E2EABCFF9DDC57FCA1FAFBE52FFA84" bold="true" box="[774,785,1287,1304]" pageId="9" pageNumber="698">fi</emphasis>
|
||
, fibula;
|
||
<emphasis id="E8E2EABCFF9DDC57FCF3FAFBE557FA84" bold="true" box="[852,873,1287,1304]" pageId="9" pageNumber="698">sc</emphasis>
|
||
, scapula;
|
||
<emphasis id="E8E2EABCFF9DDC57FC19FAFBE5F0FA84" bold="true" box="[958,974,1287,1304]" pageId="9" pageNumber="698">sf</emphasis>
|
||
, shell fragment. Scale bars: 1 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="DA2936AEFF9DDC57FF3CFAAFE539F911" blockId="9.[130,775,1363,1677]" pageId="9" pageNumber="698">
|
||
As for the genus
|
||
<emphasis id="E8E2EABCFF9DDC57FEFFFAAFE7E5FAF1" box="[344,475,1363,1389]" italics="true" pageId="9" pageNumber="698">
|
||
<taxonomicName id="1D964D2DFF9DDC57FEFFFAAFE7E9FAF1" ID-CoL="63QTH" authorityName="Pomel" authorityYear="1847" box="[344,471,1363,1389]" class="Reptilia" family="Geoemydidae" genus="Ptychogaster" kingdom="Animalia" order="Testudines" pageId="9" pageNumber="698" phylum="Chordata" rank="genus">Ptychogaster</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
it was originally erected by
|
||
<bibRefCitation id="BE074B5FFF9DDC57FF21FA8FE72FFA12" author="POMEL A." box="[134,273,1395,1422]" pageId="9" pageNumber="698" pagination="378 - 385" refId="ref15164" refString="POMEL A. 1847. - Note sur les animaux fossiles decouverts dans le departement de l'Allier. Bulletin de la Societe geologique du France 4: 378 - 385. https: // www. biodiversitylibrary. org / page / 54331323" type="journal article" year="1847">Pomel (1847)</bibRefCitation>
|
||
from the early Miocene (MN2) of Saint-Gerandle-Puy,
|
||
<collectingCountry id="A281763EFF9DDC57FF7FFA68E71BFA32" box="[216,293,1428,1454]" name="France" pageId="9" pageNumber="698">France</collectingCountry>
|
||
. Two subgenera are currently distinguished within
|
||
<taxonomicName id="1D964D2DFF9DDC57FF68FA4FE775FA51" ID-CoL="63QTH" authorityName="Pomel" authorityYear="1847" box="[207,331,1459,1485]" class="Reptilia" family="Geoemydidae" genus="Ptychogaster" kingdom="Animalia" order="Testudines" pageId="9" pageNumber="698" phylum="Chordata" rank="genus">
|
||
<emphasis id="E8E2EABCFF9DDC57FF68FA4FE775FA51" box="[207,331,1459,1485]" italics="true" pageId="9" pageNumber="698">Ptychogaster</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<emphasis id="E8E2EABCFF9DDC57FEF0FA48E758FA51" box="[343,358,1460,1485]" italics="true" pageId="9" pageNumber="698">P</emphasis>
|
||
. (
|
||
<taxonomicName id="1D964D2DFF9DDC57FEDBFA4FE7C6FA51" ID-CoL="63QTH" authorityName="Pomel" authorityYear="1847" box="[380,504,1459,1485]" class="Reptilia" family="Geoemydidae" genus="Ptychogaster" kingdom="Animalia" order="Testudines" pageId="9" pageNumber="698" phylum="Chordata" rank="genus">
|
||
<emphasis id="E8E2EABCFF9DDC57FEDBFA4FE7C6FA51" box="[380,504,1459,1485]" italics="true" pageId="9" pageNumber="698">Ptychogaster</emphasis>
|
||
</taxonomicName>
|
||
); and
|
||
<emphasis id="E8E2EABCFF9DDC57FD9FFA48E479FA51" box="[568,583,1460,1485]" italics="true" pageId="9" pageNumber="698">P</emphasis>
|
||
. (
|
||
<taxonomicName id="1D964D2DFF9DDC57FDF9FA4FE4C9FA51" authorityName="Bergounioux" authorityYear="1957" box="[606,759,1459,1485]" class="Reptilia" family="Emydidae" genus="Temnoclemmys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="9" pageNumber="698" phylum="Chordata" rank="genus">
|
||
<emphasis id="E8E2EABCFF9DDC57FDF9FA4FE4C9FA51" box="[606,759,1459,1485]" italics="true" pageId="9" pageNumber="698">Temnoclemmys</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="1D964D2DFF9DDC57FF23FA2FE4A7FA71" authority="Pomel, 1847" authorityName="Pomel" authorityYear="1847" box="[132,665,1491,1517]" class="Reptilia" family="Geoemydidae" genus="Ptychogaster" kingdom="Animalia" order="Testudines" pageId="9" pageNumber="698" phylum="Chordata" rank="species" species="emydoides" subGenus="Ptychogaster">
|
||
<emphasis id="E8E2EABCFF9DDC57FF23FA2FE73FFA71" box="[132,257,1491,1517]" italics="true" pageId="9" pageNumber="698">Ptychogaster</emphasis>
|
||
(
|
||
<emphasis id="E8E2EABCFF9DDC57FEB5FA2FE7B1FA71" box="[274,399,1491,1517]" italics="true" pageId="9" pageNumber="698">Ptychogaster</emphasis>
|
||
)
|
||
<emphasis id="E8E2EABCFF9DDC57FE07FA2FE439FA71" box="[416,519,1491,1517]" italics="true" pageId="9" pageNumber="698">emydoides</emphasis>
|
||
<bibRefCitation id="BE074B5FFF9DDC57FDA9FA2FE4A7FA71" author="POMEL A." box="[526,665,1491,1517]" pageId="9" pageNumber="698" pagination="378 - 385" refId="ref15164" refString="POMEL A. 1847. - Note sur les animaux fossiles decouverts dans le departement de l'Allier. Bulletin de la Societe geologique du France 4: 378 - 385. https: // www. biodiversitylibrary. org / page / 54331323" type="journal article" year="1847">Pomel, 1847</bibRefCitation>
|
||
</taxonomicName>
|
||
(i.e.,
|
||
<typeStatus id="052D880CFF9DDC57FD7EFA2FE539FA71" box="[729,775,1491,1517]" pageId="9" pageNumber="698">type</typeStatus>
|
||
species of subgenus
|
||
<taxonomicName id="1D964D2DFF9DDC57FEFEFA0FE7E8F991" ID-CoL="63QTH" authorityName="Pomel" authorityYear="1847" box="[345,470,1523,1549]" class="Reptilia" family="Geoemydidae" genus="Ptychogaster" kingdom="Animalia" order="Testudines" pageId="9" pageNumber="698" phylum="Chordata" rank="genus">
|
||
<emphasis id="E8E2EABCFF9DDC57FEFEFA0FE7E8F991" box="[345,470,1523,1549]" italics="true" pageId="9" pageNumber="698">Ptychogaster</emphasis>
|
||
</taxonomicName>
|
||
) has a complex taxonomical and nomenclatural history, with many junior subjective synonyms (
|
||
<bibRefCitation id="BE074B5FFF9DDC57FF6CF9CEE76AF9D1" author="SCHAFER D." box="[203,340,1586,1613]" pageId="9" pageNumber="698" refId="ref15504" refString="SCHAFER D. 2013. - Die Schildkroten der Gattung Ptychogaster Pomel, 1847 (Reptilia, Testudines). PhD dissertation, Ludwig- Maximilians-Universitat Munchen, 387 p. (in German with English summary; unpublished)." type="book" year="2013">Schäfer 2013</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="BE074B5FFF9DDC57FEF8F9CEE433F9D1" author="LUJAN A. H. & DELFINO M. & VILAR I. & ALBA D. M." box="[351,525,1586,1613]" pageId="9" pageNumber="698" pagination="277 - 295" refId="ref14529" refString="LUJAN A. H., DELFINO M., CASANOVAS- VILAR I. & ALBA D. M. 2014. - Taxonomy of subgenus Temnoclemmys Bergounioux, 1958 (Testudines: Geoemydidae: Ptychogasterinae) based on new material from the Valles-Penedes Basin. Comptes Rendus Paleovol 13 (4): 277 - 295. https: // doi. org / 10.1016 / j. crpv. 2013.11.004" type="journal article" year="2014">
|
||
Luján
|
||
<emphasis id="E8E2EABCFF9DDC57FE06F9CFE7ECF9D0" box="[417,466,1586,1612]" italics="true" pageId="9" pageNumber="698">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
). Although one revision of the genus
|
||
<taxonomicName id="1D964D2DFF9DDC57FEB0F9AEE7ABF9F0" ID-CoL="63QTH" authorityName="Pomel" authorityYear="1847" box="[279,405,1618,1644]" class="Reptilia" family="Geoemydidae" genus="Ptychogaster" kingdom="Animalia" order="Testudines" pageId="9" pageNumber="698" phylum="Chordata" rank="genus">
|
||
<emphasis id="E8E2EABCFF9DDC57FEB0F9AEE7ABF9F0" box="[279,405,1618,1644]" italics="true" pageId="9" pageNumber="698">Ptychogaster</emphasis>
|
||
</taxonomicName>
|
||
has recently been performed, the results of this PhD thesis are still unpublished (
|
||
<bibRefCitation id="BE074B5FFF9DDC57FDCBF98EE4C9F910" author="SCHAFER D." box="[620,759,1650,1676]" pageId="9" pageNumber="698" refId="ref15504" refString="SCHAFER D. 2013. - Die Schildkroten der Gattung Ptychogaster Pomel, 1847 (Reptilia, Testudines). PhD dissertation, Ludwig- Maximilians-Universitat Munchen, 387 p. (in German with English summary; unpublished)." type="book" year="2013">Schäfer 2013</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |