270 lines
23 KiB
XML
270 lines
23 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1030.62544" ID-GBIF-Dataset="91fec855-7fa9-4ad6-84d7-9b9d35e23e8b" ID-PMC="PMC8060247" ID-Pensoft-Pub="1313-2970-1030-1" ID-Pensoft-UUID="875199397EEE5F7898EA1DB25DA62D25" ID-PubMed="33958904" ModsDocID="1313-2970-1030-1" checkinTime="1618371561543" checkinUser="pensoft" docAuthor="Means, Jackson C., Hennen, Derek A. & Marek, Paul E." docDate="2021" docId="18A65F983CB756BC8C5E1E693A77AEAE" docLanguage="en" docName="ZooKeys 1030: 1-180" docOrigin="ZooKeys 1030" docPubDate="2021-04-13" docSource="http://dx.doi.org/10.3897/zookeys.1030.62544" docTitle="Nannaria equalis Chamberlin 1949" docType="treatment" docVersion="5" id="875199397EEE5F7898EA1DB25DA62D25" lastPageNumber="1" masterDocId="875199397EEE5F7898EA1DB25DA62D25" masterDocTitle="A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae)" masterLastPageNumber="180" masterPageNumber="1" pageNumber="1" updateTime="1673961214978" updateUser="julia">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Means, Jackson C.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7377-0696</mods:nameIdentifier>
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<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
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<mods:nameIdentifier type="email">mjacks4@vt.edu</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hennen, Derek A.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7005-1151</mods:nameIdentifier>
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<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Marek, Paul E.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7048-2514</mods:nameIdentifier>
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<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-04-13</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1030</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>180</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1030.62544</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1030.62544</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1030-1</mods:identifier>
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<mods:identifier type="Pensoft-UUID">875199397EEE5F7898EA1DB25DA62D25</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="181004758" LSID="urn:lsid:plazi:treatment:18A65F983CB756BC8C5E1E693A77AEAE" httpUri="http://treatment.plazi.org/id/18A65F983CB756BC8C5E1E693A77AEAE" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName ID-CoL="45M4M" LSID="18A65F98-3CB7-56BC-8C5E-1E693A77AEAE" authority="Chamberlin, 1949" authorityName="Chamberlin" authorityYear="1949" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Nannaria equalis" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="equalis">Nannaria equalis Chamberlin, 1949</taxonomicName>
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<figureCitation captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Nannaria equalis ♂ (VMNH, NAN 0335) left gonopod A anterior view; red arrow indicates large, straight prefemoral process B posterior view; red triangle indicates sharp prefemoral spine. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure54" httpUri="https://binary.pensoft.net/fig/531157" pageId="0" pageNumber="1">Fig. 54 Vernacular name: "The Knoxville Twisted-Claw Millipede"</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Chamberlin" authorityYear="1949" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Nannaria equalis" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="equalis">Nannaria equalis</taxonomicName>
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Chamberlin, 1949: 4, fig. 4.
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<bibRefCitation DOI="https://doi.org/10.5479/si.03629236.212" author="Chamberlin, RV" journalOrPublisher="United States National Museum Bulletin" pageId="0" pageNumber="1" pagination="1 - 236" refId="B22" refString="Chamberlin, RV, Hoffman, RL, 1958. Checklist of the millipeds of North America. United States National Museum Bulletin 212: 1 - 236, DOI: https://doi.org/10.5479/si.03629236.212" title="Checklist of the millipeds of North America." url="https://doi.org/10.5479/si.03629236.212" volume="212" year="1958">Chamberlin and Hoffman 1958</bibRefCitation>
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: 40.
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<bibRefCitation author="Hoffman, RL" journalOrPublisher="Virginia Museum of Natural History Special Publication Number" pageId="0" pageNumber="1" pagination="1 - 584" refId="B33" refString="Hoffman, RL, 1999. Checklist of the millipeds of North and Middle America. Virginia Museum of Natural History Special Publication Number 8: 1 - 584" title="Checklist of the millipeds of North and Middle America." volume="8" year="1999">Hoffman 1999</bibRefCitation>
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: 366.
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<bibRefCitation author="Marek, PE" journalOrPublisher="Virginia Museum of Natural History Publications" pageId="0" pageNumber="1" pagination="1 - 117" refId="B46" refString="Marek, PE, Tanabe, T, Sierwald, P, 2014. A species catalog of the millipede family Xystodesmidae (Diplopoda: Polydesmida). Virginia Museum of Natural History Publications 17: 1 - 117" title="A species catalog of the millipede family Xystodesmidae (Diplopoda: Polydesmida)." volume="17" year="2014">Marek et al. 2014</bibRefCitation>
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: 37.
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<bibRefCitation DOI="https://doi.org/10.1093/isd/ixab003" author="Means, JC" journalOrPublisher="Insect Systematics and Diversity" pageId="0" pageNumber="1" pagination="1 - 26" refId="B52" refString="Means, JC, Hennen, DA, Tanabe, T, Marek, PE, 2021. Phylogenetic systematics of the millipede family Xystodesmidae. Insect Systematics and Diversity 5: 1 - 26, DOI: https://doi.org/10.1093/isd/ixab003" title="Phylogenetic systematics of the millipede family Xystodesmidae." url="https://doi.org/10.1093/isd/ixab003" volume="5" year="2021">Means et al. 2021</bibRefCitation>
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: S69-S70.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<materialsCitation ID-GBIF-Occurrence="3071355452" collectingDate="2005-05-10" collectorName="J. Sevier, For, Suppl." country="United States" county="Knox County" location="Knox County" longitude="17.233902" specimenCount="1" specimenCount-male="1" stateProvince="Tennessee">
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<collectingCountry name="United States of America">United States</collectingCountry>
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-
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<emphasis bold="true" pageId="0" pageNumber="1">
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<collectingRegion country="United States of America" name="Tennessee">Tennessee</collectingRegion>
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</emphasis>
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•
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<specimenCount type="male">1 ♂</specimenCount>
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;
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<collectingCounty>Knox County</collectingCounty>
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, site 2 mesic cove hardwoods; from label:
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<geoCoordinate degrees="17.233902" direction="east" orientation="longitude" precision="1" value="17.233902">17.233902E</geoCoordinate>
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3993447N;
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<collectingDate value="2005-05-10">10 May 2005</collectingDate>
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;
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<collectorName>J. Sevier</collectorName>
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leg.; VMNH NAN0335.
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<collectorName>For</collectorName>
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detailed collection data see
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<collectorName>Suppl.</collectorName>
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material 7
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Adult males of
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. equalis" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="equalis">
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<emphasis italics="true" pageId="0" pageNumber="1">N. equalis</emphasis>
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</taxonomicName>
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are distinct from other
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<taxonomicName authorityName="Chamberlin" authorityYear="1918" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Nannaria</emphasis>
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</taxonomicName>
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and the nearby
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. monsdomia" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="monsdomia">
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<emphasis italics="true" pageId="0" pageNumber="1">N. monsdomia</emphasis>
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</taxonomicName>
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sp. nov. and
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. scutellaria" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="scutellaria">
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<emphasis italics="true" pageId="0" pageNumber="1">N. scutellaria</emphasis>
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</taxonomicName>
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based on the following combination of characters:
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Gonopods</emphasis>
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.
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</emphasis>
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Gonopodal acropodite straight before apex, not slightly curving before apex as in
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. monsdomia" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="monsdomia">
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<emphasis italics="true" pageId="0" pageNumber="1">N. monsdomia</emphasis>
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</taxonomicName>
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sp. nov. Distal zone much reduced, rounded, simple, not with small, lobed lateral flange as in
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. monsdomia" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="monsdomia">
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<emphasis italics="true" pageId="0" pageNumber="1">N. monsdomia</emphasis>
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</taxonomicName>
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sp. nov., or bent at 90° with pronounced medial flange as in
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. scutellaria" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="scutellaria">
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<emphasis italics="true" pageId="0" pageNumber="1">N. scutellaria</emphasis>
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</taxonomicName>
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. Acropodite with medial swelling, lacking in
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. scutellaria" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="scutellaria">
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<emphasis italics="true" pageId="0" pageNumber="1">N. scutellaria</emphasis>
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</taxonomicName>
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. Telopodite basal zone height ca. 1/2 length of acropodite, not> 2/3 length as in
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. monsdomia" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="monsdomia">
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<emphasis italics="true" pageId="0" pageNumber="1">N. monsdomia</emphasis>
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</taxonomicName>
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sp. nov. or <1/3 length as in
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. scutellaria" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="scutellaria">
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<emphasis italics="true" pageId="0" pageNumber="1">N. scutellaria</emphasis>
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</taxonomicName>
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. Prefemur with large, straight prefemoral process (Fig.
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<figureCitation captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Nannaria equalis ♂ (VMNH, NAN 0335) left gonopod A anterior view; red arrow indicates large, straight prefemoral process B posterior view; red triangle indicates sharp prefemoral spine. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure54" httpUri="https://binary.pensoft.net/fig/531157" pageId="0" pageNumber="1">54A</figureCitation>
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, red arrow), not curving medially, crossing acropodite, and expanding before tip, as in
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. monsdomia" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="monsdomia">
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<emphasis italics="true" pageId="0" pageNumber="1">N. monsdomia</emphasis>
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</taxonomicName>
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sp. nov., or curving laterally as in
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. scutellaria" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="scutellaria">
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<emphasis italics="true" pageId="0" pageNumber="1">N. scutellaria</emphasis>
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</taxonomicName>
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. Prefemoral spine sharp, curving laterally (Fig.
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<figureCitation captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. Nannaria equalis ♂ (VMNH, NAN 0335) left gonopod A anterior view; red arrow indicates large, straight prefemoral process B posterior view; red triangle indicates sharp prefemoral spine. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure54" httpUri="https://binary.pensoft.net/fig/531157" pageId="0" pageNumber="1">54B</figureCitation>
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, red triangle), not widely separated from prefemoral process and rounded, as in
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. monsdomia" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="monsdomia">
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<emphasis italics="true" pageId="0" pageNumber="1">N. monsdomia</emphasis>
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</taxonomicName>
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sp. nov., or lacking as in
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. scutellaria" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="scutellaria">
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<emphasis italics="true" pageId="0" pageNumber="1">N. scutellaria</emphasis>
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</taxonomicName>
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. Acropodite appearing as straight line in medial view, therefore the medial view was not illustrated.
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Color</emphasis>
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.
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</emphasis>
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Chamberlin reports that the paranota of
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<taxonomicName authorityName="Chamberlin" authorityYear="1949" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria equalis" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="equalis">
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<emphasis italics="true" pageId="0" pageNumber="1">Nannaria equalis</emphasis>
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</taxonomicName>
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are yellow in color, though this would be unique in the
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<taxonomicName authorityName="Chamberlin" authorityYear="1918" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Nannaria</emphasis>
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</taxonomicName>
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and has not been confirmed by the authors. It is likely Chamberlin was referring to a preserved specimen whose color had faded in alcohol.
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</paragraph>
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<caption doi="10.3897/zookeys.1030.62544.figure54" httpUri="https://binary.pensoft.net/fig/531157" pageId="0" pageNumber="1" start="Figure 54" startId="F54">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 54.</emphasis>
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<taxonomicName authorityName="Chamberlin" authorityYear="1949" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria equalis" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="equalis">
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<emphasis italics="true" pageId="0" pageNumber="1">Nannaria equalis</emphasis>
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</taxonomicName>
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♂ (VMNH, NAN0335) left gonopod
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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anterior view; red arrow indicates large, straight prefemoral process
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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posterior view; red triangle indicates sharp prefemoral spine. Scale bar: 0.5 mm.
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</paragraph>
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</caption>
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<caption doi="10.3897/zookeys.1030.62544.figure55" httpUri="https://binary.pensoft.net/fig/531158" pageId="0" pageNumber="1" start="Figure 55" startId="F55">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 55.</emphasis>
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<taxonomicName authorityName="Means & Hennen & Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria fritzae" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="fritzae">
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<emphasis italics="true" pageId="0" pageNumber="1">Nannaria fritzae</emphasis>
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</taxonomicName>
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sp. nov. ♂ holotype (VTEC, MPE02359) left gonopod
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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anterior view; red arrow indicates tooth-like medial flange
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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medial view; red triangle indicates small, thorn-like prefemoral spine
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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posterior view. Scale bar: 0.5 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Measurements.</paragraph>
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<paragraph pageId="0" pageNumber="1">♂ (VMNH, NAN0335): BL = 27.2, CW = 4.0, IW = 1.7, ISW = 0.9, B11W = 4.7, B11H = 2.8.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="variation">
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<paragraph pageId="0" pageNumber="1">Variation.</paragraph>
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<paragraph pageId="0" pageNumber="1">No known variation.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Known only from the type locality (Tennessee: Knox County; Suppl. material 7; Fig.
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<figureCitation captionStart="Figure 126" captionStartId="F126" captionText="Figure 126. Distribution map of Nannaria minor species group collection localities in the central Appalachians." figureDoi="10.3897/zookeys.1030.62544.figure126" httpUri="https://binary.pensoft.net/fig/531229" pageId="0" pageNumber="1">126</figureCitation>
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). Distribution area: N/A; status: MRE.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="ecology">
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<paragraph pageId="0" pageNumber="1">Ecology.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<bibRefCitation author="Chamberlin, RV" journalOrPublisher="Proceedings of the Biological Society of Washington" pageId="0" pageNumber="1" pagination="3 - 6" publicationUrl="https://www.biodiversitylibrary.org/page/35878747" refId="B20" refString="Chamberlin, RV, 1949. A new genus and four new species in the diplopod family Xystodesmidae. Proceedings of the Biological Society of Washington 62: 3 - 6, https://www.biodiversitylibrary.org/page/35878747" title="A new genus and four new species in the diplopod family Xystodesmidae." url="https://www.biodiversitylibrary.org/page/35878747" volume="62" year="1949">Chamberlin (1949)</bibRefCitation>
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gives no information on the ecology of
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<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. equalis" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="equalis">
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<emphasis italics="true" pageId="0" pageNumber="1">N. equalis</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<bibRefCitation author="Chamberlin, RV" journalOrPublisher="Proceedings of the Biological Society of Washington" pageId="0" pageNumber="1" pagination="3 - 6" publicationUrl="https://www.biodiversitylibrary.org/page/35878747" refId="B20" refString="Chamberlin, RV, 1949. A new genus and four new species in the diplopod family Xystodesmidae. Proceedings of the Biological Society of Washington 62: 3 - 6, https://www.biodiversitylibrary.org/page/35878747" title="A new genus and four new species in the diplopod family Xystodesmidae." url="https://www.biodiversitylibrary.org/page/35878747" volume="62" year="1949">Chamberlin (1949)</bibRefCitation>
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provided no etymology for the name
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<emphasis italics="true" pageId="0" pageNumber="1">equalis</emphasis>
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, though it is reasonable to assume that it is in reference to the prefemoral process nearly equaling in length the acropodite, as Chamberlin mentions in his description.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type locality">
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<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="1">United States, Tennessee, Knox County, Knoxville.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="notes">
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<paragraph pageId="0" pageNumber="1">Notes.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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In the original publication
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<bibRefCitation author="Chamberlin, RV" journalOrPublisher="Proceedings of the Biological Society of Washington" pageId="0" pageNumber="1" pagination="3 - 6" publicationUrl="https://www.biodiversitylibrary.org/page/35878747" refId="B20" refString="Chamberlin, RV, 1949. A new genus and four new species in the diplopod family Xystodesmidae. Proceedings of the Biological Society of Washington 62: 3 - 6, https://www.biodiversitylibrary.org/page/35878747" title="A new genus and four new species in the diplopod family Xystodesmidae." url="https://www.biodiversitylibrary.org/page/35878747" volume="62" year="1949">Chamberlin (1949</bibRefCitation>
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: 4) designated a male holotype and female paratype (originally as an
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<normalizedToken originalValue="“allotype”">"allotype"</normalizedToken>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |