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<document id="6E6F02F2BC6868BEB57FC45BF03E81CA" ID-DOI="10.5281/zenodo.8074943" ID-ISSN="2754-9844" ID-Zenodo-Dep="8074943" ID-ZooBank="BCAD06E8-0AFE-46ED-B7FA-930983CD44C4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1699388683108" checkinUser="felipe" docAuthor="Noyes, John Stuart" docDate="2023" docId="03BA87A7FF77FF04FDCFBCFAA739FC9E" docLanguage="en" docName="TaxMonogNeotropHymenopt.2.1-921.pdf.imf" docOrigin="Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) 2 (11)" docSource="http://dx.doi.org/10.5281/zenodo.8074943" docStyle="DocumentStyle:DAA2ADC4E839520FEEF1E0A10B782A41.2:TaxMonogNeotropHymenopt.2022-.journal_article" docStyleId="DAA2ADC4E839520FEEF1E0A10B782A41" docStyleName="TaxMonogNeotropHymenopt.2022-.journal_article" docStyleVersion="2" docTitle="Tico sinabelus Noyes 2023, sp.nov." docType="treatment" docUuid="3A846B79-6005-42B7-B117-56FF554F4E1A" docUuidSource="ZooBank" docVersion="3" lastPageNumber="162" masterDocId="FF83FFDFFFD5FFA7FFE2BC24A610FFC5" masterDocTitle="ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)" masterLastPageNumber="5239" masterPageNumber="5228" pageNumber="161" updateTime="1699893637884" updateUser="felipe" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="28AE6A313C5D62C79C5CCA88D8564ADA">ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)</mods:title>
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<mods:namePart id="82D70400856DECCC5779213278BDBCA9">Noyes, John Stuart</mods:namePart>
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<mods:title id="855BE305EDC8B0B97A7CB98F60AEE3A6">Taxonomic Monographs on Neotropical Hymenoptera</mods:title>
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<mods:date id="152BD42631E5F585EE72427320A5EC68">2023</mods:date>
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<mods:number id="BE2B98188C955BA334B7E014B60F0D68">2023-06-30</mods:number>
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<subSubSection id="C309653AFF77FF05FDCFBCFAA505FF33" box="[557,789,222,246]" pageId="162" pageNumber="161" type="nomenclature">
<paragraph id="8BAC36B1FF77FF05FDCFBCFAA505FF33" blockId="162.[537,808,222,273]" box="[557,789,222,246]" pageId="162" pageNumber="161">
<heading id="D0E481DDFF77FF05FDCFBCFAA505FF33" bold="true" box="[557,789,222,246]" centered="true" fontSize="10" level="2" pageId="162" pageNumber="161" reason="2">
<emphasis id="B967EAA3FF77FF05FDCFBCFAA505FF33" bold="true" box="[557,789,222,246]" pageId="162" pageNumber="161">
<taxonomicName id="4C134D32FF77FF05FDCFBCFAA4D4FF33" authority="Noyes, 2023" authorityName="Noyes" authorityYear="2023" box="[557,708,222,246]" class="Insecta" family="Derbidae" genus="Tico" kingdom="Animalia" order="Hemiptera" pageId="162" pageNumber="161" phylum="Arthropoda" rank="species" species="sinabelus" status="sp. nov.">
<emphasis id="B967EAA3FF77FF05FDCFBCFAA4D4FF33" bold="true" box="[557,708,222,246]" italics="true" pageId="162" pageNumber="161">Tico sinabelus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25457D8FF77FF05FD28BCFAA505FF33" box="[714,789,222,246]" pageId="162" pageNumber="161" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C309653AFF77FF05FDFBBCDDA538FED4" box="[537,808,249,273]" pageId="162" pageNumber="161" type="description">
<paragraph id="8BAC36B1FF77FF05FDFBBCDDA538FED4" blockId="162.[537,808,222,273]" box="[537,808,249,273]" pageId="162" pageNumber="161">
(
<figureCitation id="13282A34FF77FF05FDC2BCDDA4BCFED4" box="[544,684,249,273]" captionStart="Figs 396-411" captionStartId="765.[181,226,1526,1550]" captionTargetBox="[185,1163,231,1508]" captionTargetId="figure-10@765.[181,1163,223,1508]" captionTargetPageId="765" captionText="Figs 396-411. Mahencyrtus nitidus (396) antenna; (397) apex of fore wing venation; (398) ovipositor; (399) head, facial view, G; (400) antenna, G; (401) fore wing, G; (402) genitalia, G. Tico sinabelus (403) antenna; (404) apex of fore wing venation; (405) ovipositor; (406) fore wing; (407) hypopygium; (408) head, facial view; (409) mesosoma, dorsal view. Tico ryaspis (410) antenna; (411) head, facial view." pageId="162" pageNumber="161">Figs 403-409</figureCitation>
; Hab. E 66)
</paragraph>
</subSubSection>
<subSubSection id="C309653AFF77FF04FF57BD0AA771FDC4" lastPageId="163" lastPageNumber="162" pageId="162" pageNumber="161" type="diagnosis">
<paragraph id="8BAC36B1FF77FF05FF57BD0AA25FFE53" blockId="162.[181,1164,302,407]" pageId="162" pageNumber="161">
DIAGNOSIS.
<emphasis id="B967EAA3FF77FF05FEB2BD0AA78CFE83" box="[336,412,302,326]" italics="true" pageId="162" pageNumber="161">Female</emphasis>
(length about
<quantity id="4CEB9B54FF77FF05FDA3BD0AA49DFE83" box="[577,653,302,326]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="162" pageNumber="161" unit="mm" value="0.9">0.9mm</quantity>
): hind tibia completely pale yellow; tangent to frontovertex forming an angle of about 70° with that of face; scape not longer than minimum width of frontovertex; funicle segments (
<figureCitation id="13282A34FF77FF05FDFFBD47A465FEBE" box="[541,629,355,379]" captionStart="Figs 396-411" captionStartId="765.[181,226,1526,1550]" captionTargetBox="[185,1163,231,1508]" captionTargetId="figure-10@765.[181,1163,223,1508]" captionTargetPageId="765" captionText="Figs 396-411. Mahencyrtus nitidus (396) antenna; (397) apex of fore wing venation; (398) ovipositor; (399) head, facial view, G; (400) antenna, G; (401) fore wing, G; (402) genitalia, G. Tico sinabelus (403) antenna; (404) apex of fore wing venation; (405) ovipositor; (406) fore wing; (407) hypopygium; (408) head, facial view; (409) mesosoma, dorsal view. Tico ryaspis (410) antenna; (411) head, facial view." pageId="162" pageNumber="161">Fig. 403</figureCitation>
) not longer than broad; mesopleuron not extending past posterior margin of propodeum and not quite touching base of gaster.
<emphasis id="B967EAA3FF77FF05FC4ABD5AA5CCFE53" box="[936,988,382,406]" italics="true" pageId="162" pageNumber="161">Male</emphasis>
: unknown.
</paragraph>
<paragraph id="8BAC36B1FF77FF05FF57BD97A293FE0E" blockId="162.[181,1165,435,1687]" box="[181,1155,435,459]" pageId="162" pageNumber="161">
Female
<emphasis id="B967EAA3FF77FF05FEE4BD97A71EFE0E" bold="true" box="[262,270,435,459]" pageId="162" pageNumber="161">(</emphasis>
<typeStatus id="54A88813FF77FF05FEEDBD97A778FE0E" box="[271,360,435,459]" pageId="162" pageNumber="161" type="holotype">holotype</typeStatus>
<emphasis id="B967EAA3FF77FF05FE85BD97A77FFE0E" bold="true" box="[359,367,435,459]" pageId="162" pageNumber="161">)</emphasis>
: length, including ovipositor,
<quantity id="4CEB9B54FF77FF05FD40BD97A4E8FE0E" box="[674,760,435,459]" metricMagnitude="-4" metricUnit="m" metricValue="9.2" pageId="162" pageNumber="161" unit="mm" value="0.92">0.92mm</quantity>
; excluding ovipositor,
<quantity id="4CEB9B54FF77FF05FC3DBD97A223FE0E" box="[991,1075,435,459]" metricMagnitude="-4" metricUnit="m" metricValue="8.9" pageId="162" pageNumber="161" unit="mm" value="0.89">0.89mm</quantity>
(CPD).
</paragraph>
<paragraph id="8BAC36B1FF77FF05FF39BDEAA7F8FC6E" blockId="162.[181,1165,435,1687]" pageId="162" pageNumber="161">Head dark brown to black with a metallic sheen; occiput with a weak coppery purple sheen; frontovertex generally with a weak, mixed brassy, dark green and coppery-purple sheen, mostly coppery purple above scrobes; scrobal area, temple and gena with a weak coppery purple sheen, scrobes weakly metallic dark green; frontovertex, gena and interantennal prominence with a few scattered, inconspicuous, pale brown setae; maxillary palpus white; radicle dark brown; scape dark brown in proximal 0.5X, apex pale yellow; pedicel and flagellum mainly pale yellow, but pedicel brown at base and distal segments of clava slightly dusky; thorax dark brown to black with a weak metallic lustre; pronotum with a weak coppery purple sheen; mesoscutum and axilla with a weak brassy sheen and weak green and coppery purple reflections; tegula dark purple brown with a weak coppery purple sheen; scutellum dorsally slightly shiny with a weak coppery purple sheen with some dark green reflections, more dark green towards apex and side, extreme apex and side more strongly shining coppery and metallic green; mesopleuron with a moderate purple sheen, posteriorly more strongly coppery; fore coxa white; fore femur white to very pale yellow, tibia and tarsus very pale yellow; mid coxa orange-brown, ventrally mostly brown; mid leg otherwise pale yellow; hind coxa brown; hind femur dark brown, pale orange at apex, tibia and tarsus very pale orange; wings hyaline; propodeum dark brown with a weak purple sheen, side with a weak metallic green and coppery sheen; gaster dark brown with coppery purple, brassy, green and blue reflections; gonostylus brown.</paragraph>
<paragraph id="8BAC36B1FF77FF05FF39BF8AA6E4FA49" blockId="162.[181,1165,435,1687]" pageId="162" pageNumber="161">
Head (
<figureCitation id="13282A34FF77FF05FEC3BF8AA769FC03" box="[289,377,942,966]" captionStart="Figs 396-411" captionStartId="765.[181,226,1526,1550]" captionTargetBox="[185,1163,231,1508]" captionTargetId="figure-10@765.[181,1163,223,1508]" captionTargetPageId="765" captionText="Figs 396-411. Mahencyrtus nitidus (396) antenna; (397) apex of fore wing venation; (398) ovipositor; (399) head, facial view, G; (400) antenna, G; (401) fore wing, G; (402) genitalia, G. Tico sinabelus (403) antenna; (404) apex of fore wing venation; (405) ovipositor; (406) fore wing; (407) hypopygium; (408) head, facial view; (409) mesosoma, dorsal view. Tico ryaspis (410) antenna; (411) head, facial view." pageId="162" pageNumber="161">Fig. 408</figureCitation>
) about 2.5X as wide as frontovertex, in profile about 1.3X as high as deep, tangent to frontovertex forming an angle of about 70° with that of face; occipital margin sharp, carinate; an indistinct, shallow, shiny-bottomed, oval depression present near occipital margin adjacent to eye; ocelli forming an angle of about 85°; frontovertex narrowest about level with anterior ocellus, with distinct, coarse, polygonally reticulate sculpture of mesh size generally subequal to eye facet, not shallower above top of scrobes; temple and gena with much shallower, fine, elongate, polygonally reticulate sculpture; top of scrobes with shallow, regular, polygonally reticulate sculpture, scrobes smooth; interantennal prominence and mouth margin with slightly shallower, polygonally reticulate sculpture; scrobes shallow, indistinctly margined, narrowly ∩-shaped; interantennal prominence dorsally acute; antenna as in
<figureCitation id="13282A34FF77FF05FC9EB8BAA5C9FB73" box="[892,985,1182,1206]" captionStart="Figs 396-411" captionStartId="765.[181,226,1526,1550]" captionTargetBox="[185,1163,231,1508]" captionTargetId="figure-10@765.[181,1163,223,1508]" captionTargetPageId="765" captionText="Figs 396-411. Mahencyrtus nitidus (396) antenna; (397) apex of fore wing venation; (398) ovipositor; (399) head, facial view, G; (400) antenna, G; (401) fore wing, G; (402) genitalia, G. Tico sinabelus (403) antenna; (404) apex of fore wing venation; (405) ovipositor; (406) fore wing; (407) hypopygium; (408) head, facial view; (409) mesosoma, dorsal view. Tico ryaspis (410) antenna; (411) head, facial view." pageId="162" pageNumber="161">Fig. 408</figureCitation>
; scape relatively short and stout, about 3.6X as long as broad; funicle with segments subquadrate, but distinctly increasing in size and width distad so that F6 is largest; clava about 3X as long as broad and about 1.2X as wide as F6, not curved downwards towards apex, about as long as F3-F6 combined, sutures parallel, sensory area small, at apex only giving it a rounded appearance; eye reaching occipital margin, clothed with sparse, very inconspicuous setae, each clearly shorter than diameter of facet; clypeal margin clearly concave; mandible with 3 acute teeth. Relative measurements: HW 48,
<collectingRegion id="49D7F853FF77FF05FEE9B97EA720FAB4" box="[267,304,1370,1393]" country="Swaziland" name="Hhohho" pageId="162" pageNumber="161">HH</collectingRegion>
38, FV 19.5, POL 9.5, OOL 2, OCL 2.5, AOL 7.5, EL 28, EW 24.5, MS 15.5, SL 18, SW 5.
</paragraph>
<paragraph id="8BAC36B1FF77FF04FF39B9AAA734FED4" blockId="162.[181,1165,435,1687]" lastBlockId="163.[181,1164,222,460]" lastPageId="163" lastPageNumber="162" pageId="162" pageNumber="161">
Thorax (
<figureCitation id="13282A34FF77FF05FED6B9AAA79DFA63" box="[308,397,1422,1446]" captionStart="Figs 396-411" captionStartId="765.[181,226,1526,1550]" captionTargetBox="[185,1163,231,1508]" captionTargetId="figure-10@765.[181,1163,223,1508]" captionTargetPageId="765" captionText="Figs 396-411. Mahencyrtus nitidus (396) antenna; (397) apex of fore wing venation; (398) ovipositor; (399) head, facial view, G; (400) antenna, G; (401) fore wing, G; (402) genitalia, G. Tico sinabelus (403) antenna; (404) apex of fore wing venation; (405) ovipositor; (406) fore wing; (407) hypopygium; (408) head, facial view; (409) mesosoma, dorsal view. Tico ryaspis (410) antenna; (411) head, facial view." pageId="162" pageNumber="161">Fig. 409</figureCitation>
) with pronotum hardly exposed behind head, its posterior margin broadly concave, not angular medially; pronotum and mesoscutum with sculpture generally similar to that in ocellar area but clearly finer and shallower; axilla clearly raised, dorsally with shallow, polygonally reticulate sculpture, posterior face virtually completely smooth and shiny; scutellum mainly with fine, striate-reticulate sculpture that is clearly deeper than sculpture on mesoscutum and more or less arranged in whorls, extreme side and apex completely smooth and shiny; visible part of mesoscutum about 1.8X as broad as long; scutellum about 1.2X as broad as long, dorsally slightly convex, apex with a pair of slender, elongate setae; hind femur about 3.8X as long as broad; fore wing with venation and setation as in
<figureCitation id="13282A34FF77FF05FD4DBA40A52DF9B9" box="[687,829,1636,1660]" captionStart="Figs 396-411" captionStartId="765.[181,226,1526,1550]" captionTargetBox="[185,1163,231,1508]" captionTargetId="figure-10@765.[181,1163,223,1508]" captionTargetPageId="765" captionText="Figs 396-411. Mahencyrtus nitidus (396) antenna; (397) apex of fore wing venation; (398) ovipositor; (399) head, facial view, G; (400) antenna, G; (401) fore wing, G; (402) genitalia, G. Tico sinabelus (403) antenna; (404) apex of fore wing venation; (405) ovipositor; (406) fore wing; (407) hypopygium; (408) head, facial view; (409) mesosoma, dorsal view. Tico ryaspis (410) antenna; (411) head, facial view." pageId="162" pageNumber="161">Figs 404, 406</figureCitation>
; mesopleuron not extending past posterior margin of propodeum and not quite touching base of gaster; propodeum (
<figureCitation id="13282A34FF77FF05FBC8BA5AA295F952" box="[1066,1157,1662,1687]" captionStart="Figs 396-411" captionStartId="765.[181,226,1526,1550]" captionTargetBox="[185,1163,231,1508]" captionTargetId="figure-10@765.[181,1163,223,1508]" captionTargetPageId="765" captionText="Figs 396-411. Mahencyrtus nitidus (396) antenna; (397) apex of fore wing venation; (398) ovipositor; (399) head, facial view, G; (400) antenna, G; (401) fore wing, G; (402) genitalia, G. Tico sinabelus (403) antenna; (404) apex of fore wing venation; (405) ovipositor; (406) fore wing; (407) hypopygium; (408) head, facial view; (409) mesosoma, dorsal view. Tico ryaspis (410) antenna; (411) head, facial view." pageId="162" pageNumber="161">Fig. 409</figureCitation>
) medially about 0.1X as long as scutellum. Relative measurements: FWL 48, FWW 20; HWL 30, HWW 7.5.
</paragraph>
<paragraph id="8BAC36B1FF76FF04FF39BD30A4EDFEA4" blockId="163.[181,1164,222,460]" pageId="163" pageNumber="162">Gaster with hypopygium reaching about 0.8X towards apex; last tergite about 0.8X as long as mid tibia, with apex broadly rounded; ovipositor slightly exserted, the exserted part about 0.3X as long as mid tibial spur or about 0.1X as long as mid tibia.</paragraph>
<paragraph id="8BAC36B1FF76FF04FF39BD40A51DFE53" blockId="163.[181,1164,222,460]" pageId="163" pageNumber="162">
<typeStatus id="54A88813FF76FF04FF39BD40A727FEB9" box="[219,311,356,380]" pageId="163" pageNumber="162" type="paratype">Paratype</typeStatus>
. Funicle with linear sensilla present only on F4-F6; hypopygium
<figureCitation id="13282A34FF76FF04FC27BD40A20CFEB9" box="[965,1052,356,380]" captionStart="Figs 396-411" captionStartId="765.[181,226,1526,1550]" captionTargetBox="[185,1163,231,1508]" captionTargetId="figure-10@765.[181,1163,223,1508]" captionTargetPageId="765" captionText="Figs 396-411. Mahencyrtus nitidus (396) antenna; (397) apex of fore wing venation; (398) ovipositor; (399) head, facial view, G; (400) antenna, G; (401) fore wing, G; (402) genitalia, G. Tico sinabelus (403) antenna; (404) apex of fore wing venation; (405) ovipositor; (406) fore wing; (407) hypopygium; (408) head, facial view; (409) mesosoma, dorsal view. Tico ryaspis (410) antenna; (411) head, facial view." pageId="163" pageNumber="162">Fig. 407</figureCitation>
; ovipositor
<figureCitation id="13282A34FF76FF04FF57BD5AA71DFE53" box="[181,269,382,406]" captionStart="Figs 396-411" captionStartId="765.[181,226,1526,1550]" captionTargetBox="[185,1163,231,1508]" captionTargetId="figure-10@765.[181,1163,223,1508]" captionTargetPageId="765" captionText="Figs 396-411. Mahencyrtus nitidus (396) antenna; (397) apex of fore wing venation; (398) ovipositor; (399) head, facial view, G; (400) antenna, G; (401) fore wing, G; (402) genitalia, G. Tico sinabelus (403) antenna; (404) apex of fore wing venation; (405) ovipositor; (406) fore wing; (407) hypopygium; (408) head, facial view; (409) mesosoma, dorsal view. Tico ryaspis (410) antenna; (411) head, facial view." pageId="163" pageNumber="162">Fig. 405</figureCitation>
. Relative measurements: OL 36, GL 7.5 [MT 62].
</paragraph>
<paragraph id="8BAC36B1FF76FF04FF39BDBDA783FE09" blockId="163.[181,1164,222,460]" pageId="163" pageNumber="162">
Variation. Females vary in overall length from
<quantity id="4CEB9B54FF76FF04FD5FBDBDA559FE74" box="[701,841,409,433]" metricMagnitude="-4" metricUnit="m" metricValue="8.95" metricValueMax="9.399999999999999" metricValueMin="8.5" pageId="163" pageNumber="162" unit="mm" value="0.895" valueMax="0.94" valueMin="0.85">0.85-0.94mm</quantity>
, otherwise little variation noted in material examined.
</paragraph>
<paragraph id="8BAC36B1FF76FF04FF57BDCDA771FDC4" blockId="163.[181,353,489,513]" box="[181,353,489,513]" pageId="163" pageNumber="162">Male. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C309653AFF76FF04FF57BE3BA7F9FDF3" box="[181,489,542,567]" pageId="163" pageNumber="162" type="distribution">
<paragraph id="8BAC36B1FF76FF04FF57BE3BA7F9FDF3" blockId="163.[181,489,542,567]" box="[181,489,542,567]" pageId="163" pageNumber="162">
DISTRIBUTION.
<collectingCountry id="F3047621FF76FF04FE91BE3BA7F5FDF3" box="[371,485,542,567]" name="Costa Rica" pageId="163" pageNumber="162">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C309653AFF76FF04FF57BE70A76EFDA9" box="[181,382,596,620]" pageId="163" pageNumber="162" type="biology_ecology">
<paragraph id="8BAC36B1FF76FF04FF57BE70A76EFDA9" blockId="163.[181,382,596,620]" box="[181,382,596,620]" pageId="163" pageNumber="162">HOSTS. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C309653AFF76FF04FF57BEAFA739FC9E" pageId="163" pageNumber="162" type="materials_examined">
<paragraph id="8BAC36B1FF76FF04FF57BEAFA7BDFD65" blockId="163.[181,1165,651,859]" box="[181,429,651,672]" pageId="163" pageNumber="162">MATERIAL EXAMINED.</paragraph>
<paragraph id="8BAC36B1FF76FF04FF39BE82A739FC9E" blockId="163.[181,1165,651,859]" pageId="163" pageNumber="162">
Type material.
<materialsCitation id="3B7B3CECFF76FF04FE86BE82A4BDFD13" collectingDate="2000-01-04" collectionCode="INBio, OET" country="Costa Rica" elevation="100" latitude="10.4" location="Est. Biol. La Selva" longLatPrecision="1298" longitude="-84.01667" pageId="163" pageNumber="162" specimenCount="1" stateProvince="Heredia" typeStatus="holotype">
<typeStatus id="54A88813FF76FF04FE86BE82A7A8FD79" box="[356,440,678,700]" pageId="163" pageNumber="162" type="holotype">Holotype</typeStatus>
E:
<emphasis id="B967EAA3FF76FF04FE3EBE81A477FD7E" bold="true" box="[476,615,677,699]" pageId="163" pageNumber="162">
<collectingCountry id="F3047621FF76FF04FE3EBE81A477FD7E" box="[476,615,677,699]" name="Costa Rica" pageId="163" pageNumber="162">COSTA RICA</collectingCountry>
</emphasis>
,
<emphasis id="B967EAA3FF76FF04FD91BE81A4D1FD7E" bold="true" box="[627,705,677,699]" pageId="163" pageNumber="162">
<collectingRegion id="49D7F853FF76FF04FD91BE81A4D1FD7E" box="[627,705,677,699]" country="Costa Rica" name="Heredia" pageId="163" pageNumber="162">Heredia</collectingRegion>
</emphasis>
,
<location id="8ECC606AFF76FF04FD2EBE82A56AFD79" LSID="urn:lsid:plazi:treatment:03BA87A7FF77FF04FDCFBCFAA739FC9E:8ECC606AFF76FF04FD2EBE82A56AFD79" box="[716,890,678,700]" country="Costa Rica" latitude="10.4" longLatPrecision="1298" longitude="-84.01667" name="Est. Biol. La Selva" pageId="163" pageNumber="162" stateProvince="Heredia">Est. Biol. La Selva</location>
,
<geoCoordinate id="EE275076FF76FF04FC61BE82A5C2FD7E" box="[899,978,678,699]" degrees="10" direction="north" minutes="24" orientation="latitude" pageId="163" pageNumber="162" precision="925" value="10.4">10°24N</geoCoordinate>
<geoCoordinate id="EE275076FF76FF04FC3ABE82A23BFD7E" box="[984,1067,678,699]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="163" pageNumber="162" precision="925" value="-84.01667">84°01W</geoCoordinate>
,
<quantity id="4CEB9B54FF76FF04FBD7BE81A298FD7E" box="[1077,1160,677,699]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="163" pageNumber="162" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">
<elevation id="003ED182FF76FF04FBD7BE81A298FD7E" box="[1077,1160,677,699]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="163" pageNumber="162" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">50-150m</elevation>
</quantity>
, FOT/45/01-40,
<taxonomicName id="4C134D32FF76FF04FEA6BEE5A7C4FD13" box="[324,468,705,726]" class="Liliopsida" family="Poaceae" genus="Pouruma" kingdom="Plantae" order="Poales" pageId="163" pageNumber="162" phylum="Tracheophyta" rank="species" species="minor">
<emphasis id="B967EAA3FF76FF04FEA6BEE5A7C4FD13" box="[324,468,705,726]" italics="true" pageId="163" pageNumber="162">Pouruma minor</emphasis>
</taxonomicName>
,
<date id="FFAD1071FF76FF04FE02BEE4A43AFD13" box="[480,554,704,726]" pageId="163" pageNumber="162" value="2000-01-04">
<collectingDate id="EFE9E999FF76FF04FE02BEE4A43AFD13" box="[480,554,704,726]" pageId="163" pageNumber="162" value="2000-01-04">4.i.2000</collectingDate>
</date>
(
<collectionCode id="ED02AE74FF76FF04FDD4BEE4A461FD13" box="[566,625,704,726]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="163" pageNumber="162">INBio</collectionCode>
/
<collectionCode id="ED02AE74FF76FF04FD9BBEE4A4B7FD13" box="[633,679,704,726]" pageId="163" pageNumber="162">OET</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7B3CECFF76FF04FD54BEE5A482FD35" collectingDate="1996-03-24" collectingDateMax="1996-03-26" collectingDateMin="1996-03-24" collectorName="L. Masner" country="Costa Rica" elevation="700" location="Alajuela" pageId="163" pageNumber="162" specimenCount="1" stateProvince="Alajuela" typeStatus="paratype">
<typeStatus id="54A88813FF76FF04FD54BEE5A502FD13" box="[694,786,705,726]" pageId="163" pageNumber="162" type="paratype">Paratypes</typeStatus>
:
<emphasis id="B967EAA3FF76FF04FCF8BEE4A5B5FD13" bold="true" box="[794,933,704,726]" pageId="163" pageNumber="162">
<collectingCountry id="F3047621FF76FF04FCF8BEE4A5B5FD13" box="[794,933,704,726]" name="Costa Rica" pageId="163" pageNumber="162">COSTA RICA</collectingCountry>
</emphasis>
, 1E,
<emphasis id="B967EAA3FF76FF04FC3BBEE4A23BFD13" bold="true" box="[985,1067,704,726]" pageId="163" pageNumber="162">
<collectingRegion id="49D7F853FF76FF04FC3BBEE4A23BFD13" box="[985,1067,704,726]" country="Costa Rica" name="Alajuela" pageId="163" pageNumber="162">Alajuela</collectingRegion>
</emphasis>
, Est. Biol. San Ramon,
<quantity id="4CEB9B54FF76FF04FECCBEFFA774FD35" box="[302,356,731,752]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="163" pageNumber="162" unit="m" value="700.0">
<elevation id="003ED182FF76FF04FECCBEFFA774FD35" box="[302,356,731,752]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="163" pageNumber="162" unit="m" value="700.0">700m</elevation>
</quantity>
,
<date id="FFAD1071FF76FF04FE92BEFFA7E0FD34" box="[368,496,731,753]" pageId="163" pageNumber="162" value="1996-03-24" valueMax="1996-03-26" valueMin="1996-03-24">
<collectingDate id="EFE9E999FF76FF04FE92BEFFA7E0FD34" box="[368,496,731,753]" pageId="163" pageNumber="162" value="1996-03-24" valueMax="1996-03-26" valueMin="1996-03-24">24-26.iii.1996</collectingDate>
</date>
(
<collectorName id="26E65367FF76FF04FE1DBEFFA473FD35" box="[511,611,731,752]" pageId="163" pageNumber="162">L. Masner</collectorName>
); 1E
</materialsCitation>
,
<materialsCitation id="3B7B3CECFF76FF04FD7DBEFFA56DFCE0" collectingDate="1993-03-05" collectingDateMax="1999-12-29" collectingDateMin="1993-03-05" collectionCode="INBio, OET" country="Costa Rica" elevation="100" latitude="10.4" location="Est. Biol. La Selva" longLatPrecision="1298" longitude="-84.01667" municipality="Est. Biol. La Selva" pageId="163" pageNumber="162" specimenCount="1" stateProvince="Heredia" typeStatus="paratype">
<emphasis id="B967EAA3FF76FF04FD7DBEFFA4FDFD34" bold="true" box="[671,749,731,753]" pageId="163" pageNumber="162">
<collectingRegion id="49D7F853FF76FF04FD7DBEFFA4FDFD34" box="[671,749,731,753]" country="Costa Rica" name="Heredia" pageId="163" pageNumber="162">Heredia</collectingRegion>
</emphasis>
,
<collectingMunicipality id="6BC8ACCBFF76FF04FD18BEFFA5BEFD34" box="[762,942,731,753]" pageId="163" pageNumber="162">Est. Biol. La Selva</collectingMunicipality>
,
<geoCoordinate id="EE275076FF76FF04FC5BBEFFA218FD35" box="[953,1032,731,752]" degrees="10" direction="north" minutes="24" orientation="latitude" pageId="163" pageNumber="162" precision="925" value="10.4">10°24N</geoCoordinate>
<geoCoordinate id="EE275076FF76FF04FBEDBEFFA273FD35" box="[1039,1123,731,752]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="163" pageNumber="162" precision="925" value="-84.01667">84°01W</geoCoordinate>
,
<quantity id="4CEB9B54FF76FF04FB8CBEFFA6FBFCCE" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="163" pageNumber="162" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">
<elevation id="003ED182FF76FF04FB8CBEFFA6FBFCCE" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="163" pageNumber="162" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">50- 150m</elevation>
</quantity>
, fog
<taxonomicName id="4C134D32FF76FF04FEFBBED2A7EDFCCE" authority=", FOT" authorityName="FOT" box="[281,509,758,779]" class="Magnoliopsida" family="Meliaceae" genus="Carapa" kingdom="Plantae" order="Sapindales" pageId="163" pageNumber="162" phylum="Tracheophyta" rank="species" species="guianensis">
<emphasis id="B967EAA3FF76FF04FEFBBED2A7D6FCCE" box="[281,454,758,779]" italics="true" pageId="163" pageNumber="162">Carapa guianensis</emphasis>
, FOT
</taxonomicName>
/02,
<date id="FFAD1071FF76FF04FDC7BED1A46CFCCE" box="[549,636,757,779]" pageId="163" pageNumber="162" value="1993-03-05">
<collectingDate id="EFE9E999FF76FF04FDC7BED1A46CFCCE" box="[549,636,757,779]" pageId="163" pageNumber="162" value="1993-03-05">5.iii.1993</collectingDate>
</date>
(INBio-OET); 1E,
<collectingRegion id="49D7F853FF76FF04FCD0BED1A56CFCCE" box="[818,892,757,779]" country="Costa Rica" name="Heredia" pageId="163" pageNumber="162">Heredia</collectingRegion>
,
<location id="8ECC606AFF76FF04FC64BED2A225FCCE" LSID="urn:lsid:plazi:treatment:03BA87A7FF77FF04FDCFBCFAA739FC9E:8ECC606AFF76FF04FC64BED2A225FCCE" box="[902,1077,757,779]" country="Costa Rica" latitude="10.4" longLatPrecision="1298" longitude="-84.01667" municipality="Est. Biol. La Selva" name="Est. Biol. La Selva" pageId="163" pageNumber="162" stateProvince="Heredia">Est. Biol. La Selva</location>
,
<geoCoordinate id="EE275076FF76FF04FBDCBED2A29DFCCE" box="[1086,1165,758,779]" degrees="10" direction="north" minutes="24" orientation="latitude" pageId="163" pageNumber="162" precision="925" value="10.4">10°24N</geoCoordinate>
<geoCoordinate id="EE275076FF76FF04FF57BF34A719FCE0" box="[181,265,784,805]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="163" pageNumber="162" precision="925" value="-84.01667">84°01W</geoCoordinate>
,
<quantity id="4CEB9B54FF76FF04FEF0BF34A775FCE3" box="[274,357,784,806]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="163" pageNumber="162" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">
<elevation id="003ED182FF76FF04FEF0BF34A775FCE3" box="[274,357,784,806]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="163" pageNumber="162" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">50-150m</elevation>
</quantity>
, canopy fog, FOT/42/01-40,
<date id="FFAD1071FF76FF04FD8FBF34A4C4FCE3" box="[621,724,784,806]" pageId="163" pageNumber="162" value="1999-12-29">
<collectingDate id="EFE9E999FF76FF04FD8FBF34A4C4FCE3" box="[621,724,784,806]" pageId="163" pageNumber="162" value="1999-12-29">29.xii.1999</collectingDate>
</date>
(
<collectionCode id="ED02AE74FF76FF04FD02BF34A50BFCE3" box="[736,795,784,806]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="163" pageNumber="162">INBio</collectionCode>
/
<collectionCode id="ED02AE74FF76FF04FCC0BF34A540FCE3" box="[802,848,784,806]" pageId="163" pageNumber="162">OET</collectionCode>
); 1E
</materialsCitation>
,
<materialsCitation id="3B7B3CECFF76FF04FC65BF34A573FC84" collectingDate="2002-02-20" collectorName="PN Corcovado &amp; Sirena, LS &amp; J. A. Azofeifa" country="Costa Rica" elevation="70" location="Puntarenas" pageId="163" pageNumber="162" specimenCount="1" stateProvince="Puntarenas" typeStatus="paratype">
<emphasis id="B967EAA3FF76FF04FC65BF34A5E5FCE0" bold="true" box="[903,1013,784,805]" pageId="163" pageNumber="162">
<collectingRegion id="49D7F853FF76FF04FC65BF34A5E5FCE0" box="[903,1013,784,805]" country="Costa Rica" name="Puntarenas" pageId="163" pageNumber="162">Puntarenas</collectingRegion>
</emphasis>
,
<collectorName id="26E65367FF76FF04FBE3BF35A299FCE3" box="[1025,1161,784,806]" pageId="163" pageNumber="162">PN Corcovado</collectorName>
, Send. Los Patos/
<collectorName id="26E65367FF76FF04FEAFBF0FA7BAFC85" box="[333,426,811,833]" pageId="163" pageNumber="162">Sirena, LS</collectorName>
509200 275700,
<quantity id="4CEB9B54FF76FF04FDA4BF0FA461FC85" box="[582,625,811,832]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="163" pageNumber="162" unit="m" value="70.0">
<elevation id="003ED182FF76FF04FDA4BF0FA461FC85" box="[582,625,811,832]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="163" pageNumber="162" unit="m" value="70.0">70m</elevation>
</quantity>
,
<date id="FFAD1071FF76FF04FD9BBF0FA4C5FC84" box="[633,725,811,833]" pageId="163" pageNumber="162" value="2002-02-20">
<collectingDate id="EFE9E999FF76FF04FD9BBF0FA4C5FC84" box="[633,725,811,833]" pageId="163" pageNumber="162" value="2002-02-20">20.ii.2002</collectingDate>
</date>
(
<collectorName id="26E65367FF76FF04FD03BF0FA54DFC84" box="[737,861,811,833]" pageId="163" pageNumber="162">J.A.Azofeifa</collectorName>
)
</materialsCitation>
.
<materialsCitation id="3B7B3CECFF76FF04FC89BF0FA736FC9E" collectionCode="MZUCR, NHMUK" country="Costa Rica" pageId="163" pageNumber="162" specimenCount="2" stateProvince="Puntarenas" typeStatus="holotype">
<typeStatus id="54A88813FF76FF04FC89BF0FA5AFFC84" box="[875,959,811,833]" pageId="163" pageNumber="162" type="holotype">Holotype</typeStatus>
in
<collectionCode id="ED02AE74FF76FF04FC38BF0FA23DFC85" box="[986,1069,811,832]" pageId="163" pageNumber="162">MZUCR</collectionCode>
,
<typeStatus id="54A88813FF76FF04FBD7BF0FA29CFC85" box="[1077,1164,811,832]" pageId="163" pageNumber="162" type="paratype">paratypes</typeStatus>
in
<collectionCode id="ED02AE74FF76FF04FF2FBF62A736FC9E" box="[205,294,838,859]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="163" pageNumber="162" type="Museum">NHMUK</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>