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<document id="E3EB3E072AFC761EC2BE8F4273E5AAAD" ID-DOI="10.5281/zenodo.8074943" ID-ISSN="2754-9844" ID-Zenodo-Dep="8074943" ID-ZooBank="BCAD06E8-0AFE-46ED-B7FA-930983CD44C4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1699388683108" checkinUser="felipe" docAuthor="Noyes, John Stuart" docDate="2023" docId="03BA87A7FF73FF0FFDCAB98DA42EFDF3" docLanguage="en" docName="TaxMonogNeotropHymenopt.2.1-921.pdf.imf" docOrigin="Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) 2 (11)" docSource="http://dx.doi.org/10.5281/zenodo.8074943" docStyle="DocumentStyle:DAA2ADC4E839520FEEF1E0A10B782A41.2:TaxMonogNeotropHymenopt.2022-.journal_article" docStyleId="DAA2ADC4E839520FEEF1E0A10B782A41" docStyleName="TaxMonogNeotropHymenopt.2022-.journal_article" docStyleVersion="2" docTitle="Georynus aitne Noyes 2023, sp.nov." docType="treatment" docUuid="2A1EC43C-70A3-40F0-B48B-DDBF645658E2" docUuidSource="ZooBank" docVersion="3" lastPageNumber="167" masterDocId="FF83FFDFFFD5FFA7FFE2BC24A610FFC5" masterDocTitle="ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)" masterLastPageNumber="5239" masterPageNumber="5228" pageNumber="165" updateTime="1699893637884" updateUser="felipe" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="C73B92B27AFA6888AE934BD0D5668478">ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)</mods:title>
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<mods:namePart id="D455C566856C206C6033F39F74B868E4">Noyes, John Stuart</mods:namePart>
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<paragraph id="8BAC36B1FF73FF01FDCAB98DA50AFA07" blockId="166.[537,808,1449,1500]" box="[552,794,1449,1474]" pageId="166" pageNumber="165">
<heading id="D0E481DDFF73FF01FDCAB98DA50AFA07" bold="true" box="[552,794,1449,1474]" centered="true" fontSize="10" level="2" pageId="166" pageNumber="165" reason="2">
<emphasis id="B967EAA3FF73FF01FDCAB98DA50AFA07" bold="true" box="[552,794,1449,1474]" pageId="166" pageNumber="165">
<taxonomicName id="4C134D32FF73FF01FDCAB98DA4D8FA04" authority="Noyes, 2023" authorityName="Noyes" authorityYear="2023" box="[552,712,1449,1473]" class="Liliopsida" family="Poaceae" genus="Georynus" kingdom="Plantae" order="Poales" pageId="166" pageNumber="165" phylum="Tracheophyta" rank="species" species="aitne" status="sp. nov.">
<emphasis id="B967EAA3FF73FF01FDCAB98DA4D8FA04" bold="true" box="[552,712,1449,1473]" italics="true" pageId="166" pageNumber="165">Georynus aitne</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25457D8FF73FF01FD2CB98EA50AFA07" box="[718,794,1450,1474]" pageId="166" pageNumber="165" rank="species">sp.nov.</taxonomicNameLabel>
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<subSubSection id="C309653AFF73FF01FDFBB9E0A538FA19" box="[537,808,1476,1500]" pageId="166" pageNumber="165" type="description">
<paragraph id="8BAC36B1FF73FF01FDFBB9E0A538FA19" blockId="166.[537,808,1449,1500]" box="[537,808,1476,1500]" pageId="166" pageNumber="165">
(
<figureCitation id="13282A34FF73FF01FDC2B9E0A4BCFA19" box="[544,684,1476,1500]" captionStart="Figs 412-425" captionStartId="766.[181,226,1526,1550]" captionTargetBox="[182,1163,227,1508]" captionTargetId="figure-9@766.[181,1163,223,1508]" captionTargetPageId="766" captionText="Figs 412-425. Tico ryaspis (412) fore wing; (413) hypopygium; (414) apex of fore wing venation; (415) ovipositor; (416) mesosoma, dorsal view. Georynus aitne (417) antenna; (418) apex of fore wing venation; (419) fore wing; (420) mesosoma, dorsal view; (421) hypopygium; (422) ovipositor. Georynus dionysus, HT (423) apex of fore wing venation; (424) ovipositor; (425) antenna (a, outer aspect of clava; b, inner aspect of clava)." pageId="166" pageNumber="165">Figs 417-422</figureCitation>
; Hab. E 68)
</paragraph>
</subSubSection>
<subSubSection id="C309653AFF73FF00FF57B9DEA771F991" lastPageId="167" lastPageNumber="166" pageId="166" pageNumber="165" type="diagnosis">
<paragraph id="8BAC36B1FF73FF00FF57B9DEA70BFED4" blockId="166.[181,1165,1530,1689]" lastBlockId="167.[181,1164,222,273]" lastPageId="167" lastPageNumber="166" pageId="166" pageNumber="165">
DIAGNOSIS.
<emphasis id="B967EAA3FF73FF01FEAEB9DEA788F9D7" box="[332,408,1530,1554]" italics="true" pageId="166" pageNumber="165">Female</emphasis>
(length about
<quantity id="4CEB9B54FF73FF01FDCCB9DEA4B1F9D7" box="[558,673,1530,1554]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.8" metricValueMin="1.3" pageId="166" pageNumber="165" unit="mm" value="1.55" valueMax="1.8" valueMin="1.3">1.3-1.8mm</quantity>
): fore wing (
<figureCitation id="13282A34FF73FF01FCC7B9DEA56DF9D7" box="[805,893,1530,1554]" captionStart="Figs 412-425" captionStartId="766.[181,226,1526,1550]" captionTargetBox="[182,1163,227,1508]" captionTargetId="figure-9@766.[181,1163,223,1508]" captionTargetPageId="766" captionText="Figs 412-425. Tico ryaspis (412) fore wing; (413) hypopygium; (414) apex of fore wing venation; (415) ovipositor; (416) mesosoma, dorsal view. Georynus aitne (417) antenna; (418) apex of fore wing venation; (419) fore wing; (420) mesosoma, dorsal view; (421) hypopygium; (422) ovipositor. Georynus dionysus, HT (423) apex of fore wing venation; (424) ovipositor; (425) antenna (a, outer aspect of clava; b, inner aspect of clava)." pageId="166" pageNumber="165">Fig. 419</figureCitation>
) completely hyaline; head with 4 or 5 setae medially in front of anterior ocellus; area immediately below eye completely smooth and shiny; antenna (
<figureCitation id="13282A34FF73FF01FE39BA14A423F98D" box="[475,563,1584,1608]" captionStart="Figs 412-425" captionStartId="766.[181,226,1526,1550]" captionTargetBox="[182,1163,227,1508]" captionTargetId="figure-9@766.[181,1163,223,1508]" captionTargetPageId="766" captionText="Figs 412-425. Tico ryaspis (412) fore wing; (413) hypopygium; (414) apex of fore wing venation; (415) ovipositor; (416) mesosoma, dorsal view. Georynus aitne (417) antenna; (418) apex of fore wing venation; (419) fore wing; (420) mesosoma, dorsal view; (421) hypopygium; (422) ovipositor. Georynus dionysus, HT (423) apex of fore wing venation; (424) ovipositor; (425) antenna (a, outer aspect of clava; b, inner aspect of clava)." pageId="166" pageNumber="165">Fig. 417</figureCitation>
) with scape about 4.0X as long as broad; funicle with all segments slightly longer than broad; clava with sensory area extending about 0.3X as long ventral surface; fore wing (
<figureCitation id="13282A34FF73FF01FE62BA41A7C8F9BB" box="[384,472,1637,1662]" captionStart="Figs 412-425" captionStartId="766.[181,226,1526,1550]" captionTargetBox="[182,1163,227,1508]" captionTargetId="figure-9@766.[181,1163,223,1508]" captionTargetPageId="766" captionText="Figs 412-425. Tico ryaspis (412) fore wing; (413) hypopygium; (414) apex of fore wing venation; (415) ovipositor; (416) mesosoma, dorsal view. Georynus aitne (417) antenna; (418) apex of fore wing venation; (419) fore wing; (420) mesosoma, dorsal view; (421) hypopygium; (422) ovipositor. Georynus dionysus, HT (423) apex of fore wing venation; (424) ovipositor; (425) antenna (a, outer aspect of clava; b, inner aspect of clava)." pageId="166" pageNumber="165">Fig. 419</figureCitation>
) with linea calva open near posterior wing margin; costal cell with two lines of setae on ventral surface in apical half; syntergum about 1.3X as long as mid tibia; ovipositor about 2X as long as mid tibia; gonostylus about 2.5X as long as mid tibial spur.
<emphasis id="B967EAA3FF72FF00FBB3BCFAA295FF33" box="[1105,1157,222,246]" italics="true" pageId="167" pageNumber="166">Male</emphasis>
: unknown.
</paragraph>
<paragraph id="8BAC36B1FF72FF00FF57BD0BA293FE82" blockId="167.[181,1165,303,1566]" box="[181,1155,303,327]" pageId="167" pageNumber="166">
Female (
<typeStatus id="54A88813FF72FF00FEEFBD0BA77CFE82" box="[269,364,303,327]" pageId="167" pageNumber="166" type="holotype">holotype</typeStatus>
): length, including ovipositor,
<quantity id="4CEB9B54FF72FF00FD40BD0BA4E9FE82" box="[674,761,303,327]" metricMagnitude="-3" metricUnit="m" metricValue="1.76" pageId="167" pageNumber="166" unit="mm" value="1.76">1.76mm</quantity>
; excluding ovipositor,
<quantity id="4CEB9B54FF72FF00FC3DBD0BA223FE82" box="[991,1075,303,327]" metricMagnitude="-3" metricUnit="m" metricValue="1.61" pageId="167" pageNumber="166" unit="mm" value="1.61">1.61mm</quantity>
(CPD).
</paragraph>
<paragraph id="8BAC36B1FF72FF00FF39BD6EA7A4FCA7" blockId="167.[181,1165,303,1566]" pageId="167" pageNumber="166">
Head mainly black; frontovertex generally with a weak brassy and coppery purple sheen with some blue reflections; scrobes with a weak dark green sheen mixed coppery purple, interantennal prominence with a weak coppery purple sheen; temple and gena with a weak dark metallic green sheen, position of malar sulcus broadly marked metallic coppery purple; frontovertex, gena and interantennal prominence with short, moderately conspicuous, dark brown setae; clava brown; pronotum black with a coppery purple and brassy sheen; propleuron and prosternum dark brown; mesoscutum black with a slight metallic dark blue and purple sheen; tegula dark brown with a slight purple and brassy sheen; axilla dark brown with a purple sheen; scutellum largely metallic dark green, purplish anteriorly between axillae, extreme apex shining metallic green, side largely coppery mixed metallic green; dorsum of thorax clothed with dark brown setae; metanotum dark orange-brown with a weak brassy sheen; mesopleuron dark purple-brown with a weak brassy sheen; mesosternum dark brown; fore femur proximally white, very pale orange distally, tibia and tarsus very pale orange; mid femur mostly white, apex pale orange, tibia largely white, but very pale orange at apices, spur very pale orange, tarsus pale orange; hind femur white, apical one-third pale orange, tibia white proximally, apex very pale orange, tarsus very pale orange; fore wing (
<figureCitation id="13282A34FF72FF00FF5FBEFAA706FD33" box="[189,278,734,758]" captionStart="Figs 412-425" captionStartId="766.[181,226,1526,1550]" captionTargetBox="[182,1163,227,1508]" captionTargetId="figure-9@766.[181,1163,223,1508]" captionTargetPageId="766" captionText="Figs 412-425. Tico ryaspis (412) fore wing; (413) hypopygium; (414) apex of fore wing venation; (415) ovipositor; (416) mesosoma, dorsal view. Georynus aitne (417) antenna; (418) apex of fore wing venation; (419) fore wing; (420) mesosoma, dorsal view; (421) hypopygium; (422) ovipositor. Georynus dionysus, HT (423) apex of fore wing venation; (424) ovipositor; (425) antenna (a, outer aspect of clava; b, inner aspect of clava)." pageId="167" pageNumber="166">Fig. 419</figureCitation>
) hyaline; propodeum dark orange-brown with a weak purple and brassy sheen, setae anterior to and outside spiracle white, translucent and moderately conspicuous; gaster dark brown dorsally mainly with a coppery and purple sheen, but mixed brassy and blue, side and venter mainly with a mixed green, blue, brassy, purple and coppery sheen; gonostylus dark brown with apex widely pale orange.
</paragraph>
<paragraph id="8BAC36B1FF72FF00FF39BF41A551FB05" blockId="167.[181,1165,303,1566]" pageId="167" pageNumber="166">
Head about 8.3X as wide as frontovertex, with 4 or 5 setae medially in front of anterior ocellus, a line of sparse setae along inner eye margin continued along lower eye margin; ocelli forming an angle of about 35°; temple and gena slightly more shiny than frontovertex, with slightly shallower, longitudinally elongate, strigose to striate-reticulate sculpture, area immediately below eye completely smooth and very shiny; eye separated from scrobal margin by less than 0.5X diameter of ocellus, area between eye and scrobe with shallow, transversely elongate, imbricate-reticulate sculpture; sculpture on interantennal prominence polygonally reticulate but a little shallower that on frontovertex; antenna as in
<figureCitation id="13282A34FF72FF00FD73B805A4FBFBFC" box="[657,747,1057,1081]" captionStart="Figs 412-425" captionStartId="766.[181,226,1526,1550]" captionTargetBox="[182,1163,227,1508]" captionTargetId="figure-9@766.[181,1163,223,1508]" captionTargetPageId="766" captionText="Figs 412-425. Tico ryaspis (412) fore wing; (413) hypopygium; (414) apex of fore wing venation; (415) ovipositor; (416) mesosoma, dorsal view. Georynus aitne (417) antenna; (418) apex of fore wing venation; (419) fore wing; (420) mesosoma, dorsal view; (421) hypopygium; (422) ovipositor. Georynus dionysus, HT (423) apex of fore wing venation; (424) ovipositor; (425) antenna (a, outer aspect of clava; b, inner aspect of clava)." pageId="167" pageNumber="166">Fig. 417</figureCitation>
; scape hardly broadened, about 4.1X as long as broad; funicle with all segments only slightly longer than broad, becoming wider distad; clava slightly broadened, about 1.2X as wide as F6, nearly as long as F2-F6 combined, outer suture slightly oblique, sensory area enlarged, extending ventrally about 0.3X length of clava, apex conspicuously obliquely truncate. Relative measurements: HW 70.5,
<collectingRegion id="49D7F853FF72FF00FC46B8A9A5D9FB61" box="[932,969,1165,1188]" country="Swaziland" name="Hhohho" pageId="167" pageNumber="166">HH</collectingRegion>
69, FV 8.5, POL 4, OOL 0, OCL 8.5, AOL 10, EL 52, EW 44, MS 28, SL 37, SW 9.
</paragraph>
<paragraph id="8BAC36B1FF72FF00FF39B8E7A5E2FA82" blockId="167.[181,1165,303,1566]" pageId="167" pageNumber="166">
Thorax (
<figureCitation id="13282A34FF72FF00FED7B8E7A79EFB1E" box="[309,398,1219,1243]" captionStart="Figs 412-425" captionStartId="766.[181,226,1526,1550]" captionTargetBox="[182,1163,227,1508]" captionTargetId="figure-9@766.[181,1163,223,1508]" captionTargetPageId="766" captionText="Figs 412-425. Tico ryaspis (412) fore wing; (413) hypopygium; (414) apex of fore wing venation; (415) ovipositor; (416) mesosoma, dorsal view. Georynus aitne (417) antenna; (418) apex of fore wing venation; (419) fore wing; (420) mesosoma, dorsal view; (421) hypopygium; (422) ovipositor. Georynus dionysus, HT (423) apex of fore wing venation; (424) ovipositor; (425) antenna (a, outer aspect of clava; b, inner aspect of clava)." pageId="167" pageNumber="166">Fig. 420</figureCitation>
) with visible part of mesoscutum about 1.9X as broad as long; scutellum about as long as broad; fore wing with venation and distribution of setae as in
<figureCitation id="13282A34FF72FF00FC17B8FAA298FB33" box="[1013,1160,1246,1270]" captionStart="Figs 412-425" captionStartId="766.[181,226,1526,1550]" captionTargetBox="[182,1163,227,1508]" captionTargetId="figure-9@766.[181,1163,223,1508]" captionTargetPageId="766" captionText="Figs 412-425. Tico ryaspis (412) fore wing; (413) hypopygium; (414) apex of fore wing venation; (415) ovipositor; (416) mesosoma, dorsal view. Georynus aitne (417) antenna; (418) apex of fore wing venation; (419) fore wing; (420) mesosoma, dorsal view; (421) hypopygium; (422) ovipositor. Georynus dionysus, HT (423) apex of fore wing venation; (424) ovipositor; (425) antenna (a, outer aspect of clava; b, inner aspect of clava)." pageId="167" pageNumber="166">Figs 418, 419</figureCitation>
; mid tibial spur shorter than basitarsus; propodeum (
<figureCitation id="13282A34FF72FF00FD33B8DDA53BFAD4" box="[721,811,1273,1297]" captionStart="Figs 412-425" captionStartId="766.[181,226,1526,1550]" captionTargetBox="[182,1163,227,1508]" captionTargetId="figure-9@766.[181,1163,223,1508]" captionTargetPageId="766" captionText="Figs 412-425. Tico ryaspis (412) fore wing; (413) hypopygium; (414) apex of fore wing venation; (415) ovipositor; (416) mesosoma, dorsal view. Georynus aitne (417) antenna; (418) apex of fore wing venation; (419) fore wing; (420) mesosoma, dorsal view; (421) hypopygium; (422) ovipositor. Georynus dionysus, HT (423) apex of fore wing venation; (424) ovipositor; (425) antenna (a, outer aspect of clava; b, inner aspect of clava)." pageId="167" pageNumber="166">Fig. 420</figureCitation>
) medially about 0.16X as long as scutellum, with some shallow, irregular sculpture medially; about 7 or 8 setae anterior to and outside spiracle. Relative measurements: FWL 72, FWW 29; HWL 52, HWW 12.
</paragraph>
<paragraph id="8BAC36B1FF72FF00FF39B96EA224FA52" blockId="167.[181,1165,303,1566]" pageId="167" pageNumber="166">Gaster with hypopygium reaching about 0.5X towards apex (but slightly retracted towards anterior); syntergum about 1.3X as long as mid tibia, with apex very acute; ovipositor slightly exserted, the exserted part about as long as mid tibial spur or 0.25X as long as mid tibia.</paragraph>
<paragraph id="8BAC36B1FF72FF00FF39B9BFA510FA08" blockId="167.[181,1165,303,1566]" pageId="167" pageNumber="166">
<typeStatus id="54A88813FF72FF00FF39B9BFA727FA76" box="[219,311,1435,1459]" pageId="167" pageNumber="166" type="paratype">Paratype</typeStatus>
. Funicle with linear sensilla present only on F4-F6; hypopygium Fig, 421; ovipositor
<figureCitation id="13282A34FF72FF00FF57B991A71DFA08" box="[181,269,1461,1485]" captionStart="Figs 412-425" captionStartId="766.[181,226,1526,1550]" captionTargetBox="[182,1163,227,1508]" captionTargetId="figure-9@766.[181,1163,223,1508]" captionTargetPageId="766" captionText="Figs 412-425. Tico ryaspis (412) fore wing; (413) hypopygium; (414) apex of fore wing venation; (415) ovipositor; (416) mesosoma, dorsal view. Georynus aitne (417) antenna; (418) apex of fore wing venation; (419) fore wing; (420) mesosoma, dorsal view; (421) hypopygium; (422) ovipositor. Georynus dionysus, HT (423) apex of fore wing venation; (424) ovipositor; (425) antenna (a, outer aspect of clava; b, inner aspect of clava)." pageId="167" pageNumber="166">Fig. 422</figureCitation>
. Relative measurements: OL 72, GS 24 [MT 35].
</paragraph>
<paragraph id="8BAC36B1FF72FF00FF39B9F4A7FCF9DB" blockId="167.[181,1165,303,1566]" pageId="167" pageNumber="166">
Variation. Females vary in length from
<quantity id="4CEB9B54FF72FF00FD81B9F4A4FEFA2D" box="[611,750,1488,1512]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.81" metricValueMin="1.29" pageId="167" pageNumber="166" unit="mm" value="1.55" valueMax="1.81" valueMin="1.29">1.29-1.81mm</quantity>
, some funicle segments may be quadrate, the purple area in the position of the malar sulcus may be absent, and there may be only 4 setae outside the propodeal spiracle.
</paragraph>
<paragraph id="8BAC36B1FF72FF00FF57BA18A771F991" blockId="167.[181,353,1596,1620]" box="[181,353,1596,1620]" pageId="167" pageNumber="166">Male. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C309653AFF72FF00FF57BA56A7F9F94F" box="[181,489,1650,1674]" pageId="167" pageNumber="166" type="distribution">
<paragraph id="8BAC36B1FF72FF00FF57BA56A7F9F94F" blockId="167.[181,489,1650,1674]" box="[181,489,1650,1674]" pageId="167" pageNumber="166">
DISTRIBUTION.
<collectingCountry id="F3047621FF72FF00FE91BA56A7F5F94F" box="[371,485,1650,1674]" name="Costa Rica" pageId="167" pageNumber="166">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C309653AFF7DFF0FFF57BCFBA76EFF33" box="[181,382,222,247]" pageId="168" pageNumber="167" type="biology_ecology">
<paragraph id="8BAC36B1FF7DFF0FFF57BCFBA76EFF33" blockId="168.[181,382,222,247]" box="[181,382,222,247]" pageId="168" pageNumber="167">HOSTS. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C309653AFF7DFF0FFF57BD32A42EFDF3" pageId="168" pageNumber="167" type="materials_examined">
<paragraph id="8BAC36B1FF7DFF0FFF57BD32A7BDFEEE" blockId="168.[181,1165,278,566]" box="[181,429,278,299]" pageId="168" pageNumber="167">MATERIAL EXAMINED.</paragraph>
<paragraph id="8BAC36B1FF7DFF0FFF39BD15A42EFDF3" blockId="168.[181,1165,278,566]" pageId="168" pageNumber="167">
Type material.
<materialsCitation id="3B7B3CECFF7DFF0FFEBDBD14A71EFEA4" collectingDate="1985-10-05" collectingDateMax="1985-10-26" collectingDateMin="1985-10-05" collectorName="Janzen &amp; Gauld" country="Costa Rica" location="Sta" pageId="168" pageNumber="167" specimenCount="1" stateProvince="Guanacaste" typeStatus="holotype">
<typeStatus id="54A88813FF7DFF0FFEBDBD14A7A3FE83" box="[351,435,304,326]" pageId="168" pageNumber="167" type="holotype">Holotype</typeStatus>
E:
<emphasis id="B967EAA3FF7DFF0FFE30BD14A44AFE83" bold="true" box="[466,602,304,326]" pageId="168" pageNumber="167">
<collectingCountry id="F3047621FF7DFF0FFE30BD14A44AFE83" box="[466,602,304,326]" name="Costa Rica" pageId="168" pageNumber="167">COSTA RICA</collectingCountry>
</emphasis>
,
<emphasis id="B967EAA3FF7DFF0FFD81BD14A4C4FE83" bold="true" box="[611,724,304,326]" pageId="168" pageNumber="167">
<collectingRegion id="49D7F853FF7DFF0FFD81BD14A4C4FE83" box="[611,724,304,326]" country="Costa Rica" name="Guanacaste" pageId="168" pageNumber="167">Guanacaste</collectingRegion>
</emphasis>
,
<location id="8ECC606AFF7DFF0FFD3CBD14A4EBFE80" LSID="urn:lsid:plazi:treatment:03BA87A7FF73FF0FFDCAB98DA42EFDF3:8ECC606AFF7DFF0FFD3CBD14A4EBFE80" box="[734,763,304,325]" country="Costa Rica" name="Sta" pageId="168" pageNumber="167" stateProvince="Guanacaste">Sta</location>
<taxonomicName id="4C134D32FF7DFF0FFD1CBD15A5B8FE83" authority="NP, Hacienda" authorityName="NP, Hacienda" box="[766,936,304,326]" class="Magnoliopsida" family="Rosaceae" genus="Rosa" kingdom="Plantae" order="Rosales" pageId="168" pageNumber="167" phylum="Tracheophyta" rank="genus">Rosa NP,Hacienda</taxonomicName>
4-C,
<date id="FFAD1071FF7DFF0FFC34BD14A254FE83" box="[982,1092,304,326]" pageId="168" pageNumber="167" value="1985-10-05" valueMax="1985-10-26" valueMin="1985-10-05">
<collectingDate id="EFE9E999FF7DFF0FFC34BD14A254FE83" box="[982,1092,304,326]" pageId="168" pageNumber="167" value="1985-10-05" valueMax="1985-10-26" valueMin="1985-10-05">5-26.x.1985</collectingDate>
</date>
(
<collectorName id="26E65367FF7DFF0FFBAFBD14A29CFE80" box="[1101,1164,304,325]" pageId="168" pageNumber="167">Janzen</collectorName>
&amp;
<collectorName id="26E65367FF7DFF0FFF2FBD6FA718FEA4" box="[205,264,331,353]" pageId="168" pageNumber="167">Gauld</collectorName>
)
</materialsCitation>
.
<materialsCitation id="3B7B3CECFF7DFF0FFEF5BD6FA75DFDDF" collectingDate="1985-09-14" collectingDateMax="2000-11-12" collectingDateMin="1985-09-14" collectionCode="LN" collectorName="Janzen &amp; Gauld &amp; PN Palo Verde &amp; I. Jimenez" country="Costa Rica" county="Santa" elevation="385020140" location="Sector Palo Verde" municipality="Cerro Guayacan" pageId="168" pageNumber="167" specimenCode="LN 260952, LN 259350, LN 259750, LN 259450, LN 260770" specimenCount="1" stateProvince="Guanacaste" typeStatus="paratype">
<typeStatus id="54A88813FF7DFF0FFEF5BD6FA763FEA5" box="[279,371,331,352]" pageId="168" pageNumber="167" type="paratype">Paratypes</typeStatus>
:
<emphasis id="B967EAA3FF7DFF0FFE99BD6FA416FEA4" bold="true" box="[379,518,331,353]" pageId="168" pageNumber="167">
<collectingCountry id="F3047621FF7DFF0FFE99BD6FA416FEA4" box="[379,518,331,353]" name="Costa Rica" pageId="168" pageNumber="167">COSTA RICA</collectingCountry>
</emphasis>
, 6E,
<emphasis id="B967EAA3FF7DFF0FFDDBBD6FA4BAFEA4" bold="true" box="[569,682,331,353]" pageId="168" pageNumber="167">
<collectingRegion id="49D7F853FF7DFF0FFDDBBD6FA4BAFEA4" box="[569,682,331,353]" country="Costa Rica" name="Guanacaste" pageId="168" pageNumber="167">Guanacaste</collectingRegion>
</emphasis>
,
<collectingCounty id="62CD4E3DFF7DFF0FFD54BD6FA4F8FEA5" box="[694,744,331,352]" pageId="168" pageNumber="167">Santa</collectingCounty>
<taxonomicName id="4C134D32FF7DFF0FFD0CBD6FA58EFEA4" authority="NP, Hacienda" authorityName="NP, Hacienda" box="[750,926,331,353]" class="Magnoliopsida" family="Rosaceae" genus="Rosa" kingdom="Plantae" order="Rosales" pageId="168" pageNumber="167" phylum="Tracheophyta" rank="genus">Rosa NP, Hacienda</taxonomicName>
, various dates
<date id="FFAD1071FF7DFF0FFBC6BD6FA295FEA4" box="[1060,1157,331,353]" pageId="168" pageNumber="167" value="1985-09-14">
<collectingDate id="EFE9E999FF7DFF0FFBC6BD6FA295FEA4" box="[1060,1157,331,353]" pageId="168" pageNumber="167" value="1985-09-14">14.ix.1985</collectingDate>
</date>
-
<date id="FFAD1071FF7DFF0FFF57BD41A70DFEBE" box="[181,285,357,379]" pageId="168" pageNumber="167" value="1986-12-20">
<collectingDate id="EFE9E999FF7DFF0FFF57BD41A70DFEBE" box="[181,285,357,379]" pageId="168" pageNumber="167" value="1986-12-20">20.xii.1986</collectingDate>
</date>
(
<collectorName id="26E65367FF7DFF0FFECABD42A777FEBE" box="[296,359,358,379]" pageId="168" pageNumber="167">Janzen</collectorName>
&amp;
<collectorName id="26E65367FF7DFF0FFE61BD41A7AEFEBE" box="[387,446,357,379]" pageId="168" pageNumber="167">Gauld</collectorName>
); 1E,
<collectingRegion id="49D7F853FF7DFF0FFE14BD42A471FEBE" box="[502,609,358,379]" country="Costa Rica" name="Guanacaste" pageId="168" pageNumber="167">Guanacaste</collectingRegion>
,
<collectorName id="26E65367FF7DFF0FFD8BBD42A4E0FEBE" box="[617,752,357,379]" pageId="168" pageNumber="167">PN Palo Verde</collectorName>
,
<collectingMunicipality id="6BC8ACCBFF7DFF0FFD1BBD42A59EFEB9" box="[761,910,358,380]" pageId="168" pageNumber="167">Cerro Guayacán</collectingMunicipality>
,
<specimenCode id="DBB59ECAFF7DFF0FFC75BD42A5EFFEBE" box="[919,1023,357,379]" collectionCode="LN" pageId="168" pageNumber="167">LN 260952</specimenCode>
<quantity id="4CEB9B54FF7DFF0FFBE6BD41A298FEBE" box="[1028,1160,357,379]" metricMagnitude="8" metricUnit="m" metricValue="3.8502014" pageId="168" pageNumber="167" unit="m" value="3.8502014E8">
<elevation id="003ED182FF7DFF0FFBE6BD41A298FEBE" box="[1028,1160,357,379]" metricMagnitude="8" metricUnit="m" metricValue="3.8502014" pageId="168" pageNumber="167" unit="m" value="3.8502014E8">385020, 140m</elevation>
</quantity>
,
<date id="FFAD1071FF7DFF0FFF57BDA4A756FE53" box="[181,326,384,406]" pageId="168" pageNumber="167" value="1999-05-19" valueMax="1999-06-15" valueMin="1999-05-19">
<collectingDate id="EFE9E999FF7DFF0FFF57BDA4A756FE53" box="[181,326,384,406]" pageId="168" pageNumber="167" value="1999-05-19" valueMax="1999-06-15" valueMin="1999-05-19">19.v-15.vi.1999</collectingDate>
</date>
(
<collectorName id="26E65367FF7DFF0FFEB5BDA4A7ABFE53" box="[343,443,384,406]" pageId="168" pageNumber="167">I. Jiménez</collectorName>
); 3E,
<collectingRegion id="49D7F853FF7DFF0FFE19BDA4A476FE50" box="[507,614,384,405]" country="Costa Rica" name="Guanacaste" pageId="168" pageNumber="167">Guanacaste</collectingRegion>
,
<collectorName id="26E65367FF7DFF0FFD91BDA5A511FE53" box="[627,769,384,406]" pageId="168" pageNumber="167">PN Palo Verde</collectorName>
,
<location id="8ECC606AFF7DFF0FFCECBDA4A5AAFE53" LSID="urn:lsid:plazi:treatment:03BA87A7FF73FF0FFDCAB98DA42EFDF3:8ECC606AFF7DFF0FFCECBDA4A5AAFE53" box="[782,954,384,406]" country="Costa Rica" county="Santa" municipality="Cerro Guayacan" name="Sector Palo Verde" pageId="168" pageNumber="167" stateProvince="Guanacaste">Sector Palo Verde</location>
,
<location id="8ECC606AFF7DFF0FFC25BDA4A271FE53" LSID="urn:lsid:plazi:treatment:03BA87A7FF73FF0FFDCAB98DA42EFDF3:8ECC606AFF7DFF0FFC25BDA4A271FE53" box="[967,1121,384,406]" country="Costa Rica" county="Santa" municipality="Cerro Guayacan" name="Cerro Guayacan" pageId="168" pageNumber="167" stateProvince="Guanacaste">Cerro Guayacán</location>
,
<specimenCode id="DBB59ECAFF7DFF0FFB8CBDA5A6E9FE74" collectionCode="LN" pageId="168" pageNumber="167">LN 259350</specimenCode>
<quantity id="4CEB9B54FF7DFF0FFF1DBDBFA794FE75" box="[255,388,411,433]" metricMagnitude="8" metricUnit="m" metricValue="3.8960014" pageId="168" pageNumber="167" unit="m" value="3.8960014E8">
<elevation id="003ED182FF7DFF0FFF1DBDBFA794FE75" box="[255,388,411,433]" metricMagnitude="8" metricUnit="m" metricValue="3.8960014" pageId="168" pageNumber="167" unit="m" value="3.8960014E8">389600, 140m</elevation>
</quantity>
, vii-viii.1999 (
<collectorName id="26E65367FF7DFF0FFDEFBDBFA47DFE74" box="[525,621,411,433]" pageId="168" pageNumber="167">I. Jiménez</collectorName>
); 1E,
<collectingRegion id="49D7F853FF7DFF0FFD44BDBFA501FE75" box="[678,785,411,432]" country="Costa Rica" name="Guanacaste" pageId="168" pageNumber="167">Guanacaste</collectingRegion>
,
<collectorName id="26E65367FF7DFF0FFCF8BDBFA5B1FE74" box="[794,929,411,433]" pageId="168" pageNumber="167">PN Palo Verde</collectorName>
,
<location id="8ECC606AFF7DFF0FFC49BDBFA25FFE74" LSID="urn:lsid:plazi:treatment:03BA87A7FF73FF0FFDCAB98DA42EFDF3:8ECC606AFF7DFF0FFC49BDBFA25FFE74" box="[939,1103,411,433]" country="Costa Rica" county="Santa" municipality="Cerro Guayacan" name="Sector Palo Verde" pageId="168" pageNumber="167" stateProvince="Guanacaste">Sector Palo Verde</location>
,
<location id="8ECC606AFF7DFF0FFBBABDBFA702FE09" LSID="urn:lsid:plazi:treatment:03BA87A7FF73FF0FFDCAB98DA42EFDF3:8ECC606AFF7DFF0FFBBABDBFA702FE09" country="Costa Rica" county="Santa" municipality="Cerro Guayacan" name="Cerro Guayacan" pageId="168" pageNumber="167" stateProvince="Guanacaste">Cerro Guayacán</location>
,
<specimenCode id="DBB59ECAFF7DFF0FFEFFBD92A797FE0E" box="[285,391,437,459]" collectionCode="LN" pageId="168" pageNumber="167">LN 259750</specimenCode>
<quantity id="4CEB9B54FF7DFF0FFE6DBD91A406FE0E" box="[399,534,437,459]" metricMagnitude="8" metricUnit="m" metricValue="3.885000999999999" pageId="168" pageNumber="167" unit="m" value="3.885001E8">
<elevation id="003ED182FF7DFF0FFE6DBD91A406FE0E" box="[399,534,437,459]" metricMagnitude="8" metricUnit="m" metricValue="3.885000999999999" pageId="168" pageNumber="167" unit="m" value="3.885001E8">388500, 100m</elevation>
</quantity>
, #57963,
<date id="FFAD1071FF7DFF0FFD91BD91A50BFE0E" box="[627,795,437,459]" pageId="168" pageNumber="167" value="2000-07-11" valueMax="2000-08-10" valueMin="2000-07-11">
<collectingDate id="EFE9E999FF7DFF0FFD91BD91A50BFE0E" box="[627,795,437,459]" pageId="168" pageNumber="167" value="2000-07-11" valueMax="2000-08-10" valueMin="2000-07-11">11.vii-10.viii.2000</collectingDate>
</date>
(
<collectorName id="26E65367FF7DFF0FFCC9BD92A59DFE0E" box="[811,909,437,459]" pageId="168" pageNumber="167">I. Jiménez</collectorName>
); 1E,
<collectingRegion id="49D7F853FF7DFF0FFC2BBD92A224FE0E" box="[969,1076,438,459]" country="Costa Rica" name="Guanacaste" pageId="168" pageNumber="167">Guanacaste</collectingRegion>
,
<collectorName id="26E65367FF7DFF0FFBDDBD92A6FCFE23" pageId="168" pageNumber="167">PN Palo Verde</collectorName>
,
<location id="8ECC606AFF7DFF0FFF17BDF4A789FE23" LSID="urn:lsid:plazi:treatment:03BA87A7FF73FF0FFDCAB98DA42EFDF3:8ECC606AFF7DFF0FFF17BDF4A789FE23" box="[245,409,464,486]" country="Costa Rica" county="Santa" municipality="Cerro Guayacan" name="Sector Palo Verde" pageId="168" pageNumber="167" stateProvince="Guanacaste">Sector Palo Verde</location>
,
<location id="8ECC606AFF7DFF0FFE40BDF4A428FE23" LSID="urn:lsid:plazi:treatment:03BA87A7FF73FF0FFDCAB98DA42EFDF3:8ECC606AFF7DFF0FFE40BDF4A428FE23" box="[418,568,464,486]" country="Costa Rica" county="Santa" municipality="Cerro Guayacan" name="Cerro Guayacan" pageId="168" pageNumber="167" stateProvince="Guanacaste">Cerro Guayacán</location>
,
<specimenCode id="DBB59ECAFF7DFF0FFDA3BDF5A4B9FE23" box="[577,681,464,486]" collectionCode="LN" pageId="168" pageNumber="167">LN 259450</specimenCode>
<quantity id="4CEB9B54FF7DFF0FFD4CBDF4A523FE23" box="[686,819,464,486]" metricMagnitude="8" metricUnit="m" metricValue="3.8875015" pageId="168" pageNumber="167" unit="m" value="3.8875015E8">
<elevation id="003ED182FF7DFF0FFD4CBDF4A523FE23" box="[686,819,464,486]" metricMagnitude="8" metricUnit="m" metricValue="3.8875015" pageId="168" pageNumber="167" unit="m" value="3.8875015E8">388750, 150m</elevation>
</quantity>
, #53297,
<date id="FFAD1071FF7DFF0FFC69BDF4A5FDFE23" box="[907,1005,464,486]" pageId="168" pageNumber="167" value="1999-06-15">
<collectingDate id="EFE9E999FF7DFF0FFC69BDF4A5FDFE23" box="[907,1005,464,486]" pageId="168" pageNumber="167" value="1999-06-15">15.vi.1999</collectingDate>
</date>
(
<collectorName id="26E65367FF7DFF0FFC18BDF4A24AFE23" box="[1018,1114,464,486]" pageId="168" pageNumber="167">I. Jiménez</collectorName>
); 2E,
<collectingRegion id="49D7F853FF7DFF0FFF57BDCFA730FDC5" box="[181,288,491,512]" country="Costa Rica" name="Guanacaste" pageId="168" pageNumber="167">Guanacaste</collectingRegion>
,
<collectorName id="26E65367FF7DFF0FFEC9BDCFA7A6FDC4" box="[299,438,491,513]" pageId="168" pageNumber="167">PN Palo Verde</collectorName>
,
<location id="8ECC606AFF7DFF0FFE23BDCFA461FDC4" LSID="urn:lsid:plazi:treatment:03BA87A7FF73FF0FFDCAB98DA42EFDF3:8ECC606AFF7DFF0FFE23BDCFA461FDC4" box="[449,625,491,513]" country="Costa Rica" county="Santa" municipality="Cerro Guayacan" name="Camino Alto Viejo" pageId="168" pageNumber="167" stateProvince="Guanacaste">Camino Alto Viejo</location>
,
<specimenCode id="DBB59ECAFF7DFF0FFD9FBDCFA4F6FDC4" box="[637,742,491,513]" collectionCode="LN" pageId="168" pageNumber="167">LN 260770</specimenCode>
386300,
<quantity id="4CEB9B54FF7DFF0FFCDDBDCFA57AFDC5" box="[831,874,491,512]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="168" pageNumber="167" unit="m" value="40.0">
<elevation id="003ED182FF7DFF0FFCDDBDCFA57AFDC5" box="[831,874,491,512]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="168" pageNumber="167" unit="m" value="40.0">40m</elevation>
</quantity>
, ix-x.2000 and
<date id="FFAD1071FF7DFF0FFC1EBDCFA29CFDC4" box="[1020,1164,491,513]" pageId="168" pageNumber="167" value="2000-10-12" valueMax="2000-11-12" valueMin="2000-10-12">
<collectingDate id="EFE9E999FF7DFF0FFC1EBDCFA29CFDC4" box="[1020,1164,491,513]" pageId="168" pageNumber="167" value="2000-10-12" valueMax="2000-11-12" valueMin="2000-10-12">12.x-12.xi.2000</collectingDate>
</date>
(
<collectorName id="26E65367FF7DFF0FFF5FBE22A70FFDDE" box="[189,287,517,539]" pageId="168" pageNumber="167">I. Jiménez</collectorName>
); 1E
</materialsCitation>
,
<materialsCitation id="3B7B3CECFF7DFF0FFEBBBE21A206FDDE" box="[345,1046,517,539]" collectingDate="1994-09-09" collectorName="M. Cerda &amp; P. Hanson" country="Costa Rica" elevation="1100" location="Santo Domingo" pageId="168" pageNumber="167" specimenCount="1" stateProvince="Heredia" typeStatus="paratype">
<emphasis id="B967EAA3FF7DFF0FFEBBBE21A7B7FDDE" bold="true" box="[345,423,517,539]" pageId="168" pageNumber="167">
<collectingRegion id="49D7F853FF7DFF0FFEBBBE21A7B7FDDE" box="[345,423,517,539]" country="Costa Rica" name="Heredia" pageId="168" pageNumber="167">Heredia</collectingRegion>
</emphasis>
,
<location id="8ECC606AFF7DFF0FFE51BE22A455FDDE" LSID="urn:lsid:plazi:treatment:03BA87A7FF73FF0FFDCAB98DA42EFDF3:8ECC606AFF7DFF0FFE51BE22A455FDDE" box="[435,581,517,539]" country="Costa Rica" name="Santo Domingo" pageId="168" pageNumber="167" stateProvince="Heredia">Santo Domingo</location>
,
<quantity id="4CEB9B54FF7DFF0FFDADBE22A480FDDE" box="[591,656,518,539]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="168" pageNumber="167" unit="m" value="1100.0">
<elevation id="003ED182FF7DFF0FFDADBE22A480FDDE" box="[591,656,518,539]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="168" pageNumber="167" unit="m" value="1100.0">1100m</elevation>
</quantity>
, cafetal,
<date id="FFAD1071FF7DFF0FFD0BBE21A52FFDDE" box="[745,831,517,539]" pageId="168" pageNumber="167" value="1994-09-09">
<collectingDate id="EFE9E999FF7DFF0FFD0BBE21A52FFDDE" box="[745,831,517,539]" pageId="168" pageNumber="167" value="1994-09-09">9.ix.1994</collectingDate>
</date>
(
<collectorName id="26E65367FF7DFF0FFCAEBE22A5B5FDDE" box="[844,933,517,539]" pageId="168" pageNumber="167">M. Cerda</collectorName>
,
<collectorName id="26E65367FF7DFF0FFC52BE22A21FFDDE" box="[944,1039,518,539]" pageId="168" pageNumber="167">P. Hanson</collectorName>
)
</materialsCitation>
.
<materialsCitation id="3B7B3CECFF7DFF0FFBFDBE21A42AFDF3" collectionCode="MZUCR, NHMUK" country="Costa Rica" pageId="168" pageNumber="167" specimenCount="2" stateProvince="Heredia" typeStatus="holotype">
<typeStatus id="54A88813FF7DFF0FFBFDBE21A263FDDE" box="[1055,1139,517,539]" pageId="168" pageNumber="167" type="holotype">Holotype</typeStatus>
in
<collectionCode id="ED02AE74FF7DFF0FFF57BE04A718FDF0" box="[181,264,544,565]" pageId="168" pageNumber="167">MZUCR</collectionCode>
,
<typeStatus id="54A88813FF7DFF0FFEF3BE05A778FDF3" box="[273,360,545,566]" pageId="168" pageNumber="167" type="paratype">paratypes</typeStatus>
in
<collectionCode id="ED02AE74FF7DFF0FFE67BE04A7C8FDF0" box="[389,472,544,565]" pageId="168" pageNumber="167">MZUCR</collectionCode>
,
<collectionCode id="ED02AE74FF7DFF0FFE03BE05A42AFDF3" box="[481,570,545,566]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="168" pageNumber="167" type="Museum">NHMUK</collectionCode>
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.
</paragraph>
</subSubSection>
</treatment>
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