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<document id="0489A1A752306C9C8F1B932F38E8AE81" ID-DOI="10.5281/zenodo.8074943" ID-ISSN="2754-9844" ID-Zenodo-Dep="8074943" ID-ZooBank="BCAD06E8-0AFE-46ED-B7FA-930983CD44C4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1699388683108" checkinUser="felipe" docAuthor="Noyes, John Stuart" docDate="2023" docId="03BA87A7FF3BFF57FDE8BE86A7FCFE9A" docLanguage="en" docName="TaxMonogNeotropHymenopt.2.1-921.pdf.imf" docOrigin="Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) 2 (11)" docSource="http://dx.doi.org/10.5281/zenodo.8074943" docStyle="DocumentStyle:DAA2ADC4E839520FEEF1E0A10B782A41.2:TaxMonogNeotropHymenopt.2022-.journal_article" docStyleId="DAA2ADC4E839520FEEF1E0A10B782A41" docStyleName="TaxMonogNeotropHymenopt.2022-.journal_article" docStyleVersion="2" docTitle="Cheiloneurus catreta Noyes 2023, sp.nov." docType="treatment" docUuid="D7E511DE-725F-48E7-86EE-E73FB2AB2C0C" docUuidSource="ZooBank" docVersion="3" lastPageNumber="239" masterDocId="FF83FFDFFFD5FFA7FFE2BC24A610FFC5" masterDocTitle="ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)" masterLastPageNumber="5239" masterPageNumber="5228" pageNumber="237" updateTime="1699893637884" updateUser="felipe" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="E50813B9CD5436ACE6DA10DB8BD9EE87">ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)</mods:title>
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<mods:namePart id="5739A0750B71C1518194C1F7E221FB64">Noyes, John Stuart</mods:namePart>
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<mods:title id="83FF1AF78537C88DA5FABDDB8A812DE5">Taxonomic Monographs on Neotropical Hymenoptera</mods:title>
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<mods:date id="B2C8DAB05FEBB851A5AEEBA9D934F21D">2023</mods:date>
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<mods:number id="E120FBAC60A6DA056B5D042700DF4E7A">2023-06-30</mods:number>
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<subSubSection id="C309653AFF3BFF49FDE8BE86A528FD7F" box="[522,824,674,698]" pageId="238" pageNumber="237" type="nomenclature">
<paragraph id="8BAC36B1FF3BFF49FDE8BE86A528FD7F" blockId="238.[522,824,674,725]" box="[522,824,674,698]" pageId="238" pageNumber="237">
<heading id="D0E481DDFF3BFF49FDE8BE86A528FD7F" bold="true" box="[522,824,674,698]" centered="true" fontSize="10" level="2" pageId="238" pageNumber="237" reason="2">
<emphasis id="B967EAA3FF3BFF49FDE8BE86A528FD7F" bold="true" box="[522,824,674,698]" pageId="238" pageNumber="237">
<taxonomicName id="4C134D32FF3BFF49FDE8BE86A4F7FD7C" authority="Noyes, 2023" authorityName="Noyes" authorityYear="2023" box="[522,743,674,698]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="238" pageNumber="237" phylum="Arthropoda" rank="species" species="catreta" status="sp. nov.">
<emphasis id="B967EAA3FF3BFF49FDE8BE86A4F7FD7C" bold="true" box="[522,743,674,698]" italics="true" pageId="238" pageNumber="237">Cheiloneurus catreta</emphasis>
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<subSubSection id="C309653AFF3BFF49FDFBBE99A538FD10" box="[537,808,700,725]" pageId="238" pageNumber="237" type="description">
<paragraph id="8BAC36B1FF3BFF49FDFBBE99A538FD10" blockId="238.[522,824,674,725]" box="[537,808,700,725]" pageId="238" pageNumber="237">
(
<figureCitation id="13282A34FF3BFF49FDC2BE99A4BBFD11" box="[544,683,700,725]" captionStart-0="Figs 583-595" captionStart-1="Figs 596-607" captionStartId-0="778.[181,226,1526,1550]" captionStartId-1="779.[181,226,1526,1550]" captionTargetBox-0="[181,1163,227,1504]" captionTargetBox-1="[181,1163,229,1508]" captionTargetId-0="figure-9@778.[181,1163,223,1508]" captionTargetId-1="figure-10@779.[181,1163,223,1508]" captionTargetPageId-0="778" captionTargetPageId-1="779" captionText-0="Figs 583-595. Cheiloneurus parvus (583) antenna (a, outer aspect of clava; b, inner aspect of clava); (584) ovipositor; (585) hypopygium; (586) fore wing; (587) apex of fore wing venation. Cheiloneurus semis (588) fore wing; (589) hypopygium; (590) apex of fore wing venation; (591) antenna (a, outer aspect of clava; b, inner aspect of clava); (592) mesosoma, dorsal view. Cheiloneurus catreta (593) antenna (a, outer aspect of clava; b, inner aspect of clava); (594) apex of fore wing venation; (595) fore wing." captionText-1="Figs 596-607. Cheiloneurus catreta (596) mesosoma, dorsal view. Cheiloneurus herndonia, HT (597) mesosoma, dorsal view; (598) antenna (a, outer aspect of clava; b, inner aspect of clava); (599) apex of fore wing venation; (600) fore wing. Cheiloneurus exeres (601) fore wing; (602) antenna (a, outer aspect of clava; b, inner aspect of clava); (603) apex of fore wing venation. Cheiloneurus fidenas (604) fore wing; (605) antenna (a, outer aspect of clava; b, inner aspect of clava); (606) apex of fore wing venation; (607) mesosoma, dorsal view." pageId="238" pageNumber="237">Figs 593-596</figureCitation>
; Hab. E 90)
</paragraph>
</subSubSection>
<subSubSection id="C309653AFF3BFF48FF57BED6A771FA19" lastPageId="239" lastPageNumber="238" pageId="238" pageNumber="237" type="diagnosis">
<paragraph id="8BAC36B1FF3BFF49FF57BED6A783FA90" blockId="238.[181,1165,754,1365]" pageId="238" pageNumber="237">
DIAGNOSIS.
<emphasis id="B967EAA3FF3BFF49FEB3BED6A78DFCCF" box="[337,413,754,778]" italics="true" pageId="238" pageNumber="237">Female</emphasis>
(length about
<quantity id="4CEB9B54FF3BFF49FDA3BED6A4A6FCCF" box="[577,694,754,778]" metricMagnitude="-3" metricUnit="m" metricValue="1.2999999999999998" metricValueMax="1.4" metricValueMin="1.2" pageId="238" pageNumber="237" unit="mm" value="1.2999999999999998" valueMax="1.4" valueMin="1.2">1.2-1.4mm</quantity>
): body generally dark brown to black with a metallic sheen; antenna (
<figureCitation id="13282A34FF3BFF49FE2CBF29A438FCE1" box="[462,552,780,805]" captionStart="Figs 583-595" captionStartId="778.[181,226,1526,1550]" captionTargetBox="[181,1163,227,1504]" captionTargetId="figure-9@778.[181,1163,223,1508]" captionTargetPageId="778" captionText="Figs 583-595. Cheiloneurus parvus (583) antenna (a, outer aspect of clava; b, inner aspect of clava); (584) ovipositor; (585) hypopygium; (586) fore wing; (587) apex of fore wing venation. Cheiloneurus semis (588) fore wing; (589) hypopygium; (590) apex of fore wing venation; (591) antenna (a, outer aspect of clava; b, inner aspect of clava); (592) mesosoma, dorsal view. Cheiloneurus catreta (593) antenna (a, outer aspect of clava; b, inner aspect of clava); (594) apex of fore wing venation; (595) fore wing." pageId="238" pageNumber="237">Fig. 593</figureCitation>
) with scape dark brown with apex white; pedicel dorsally dark brown, orange-brown ventrally, apex white; funicle white to pale orange-brown; clava pale orange, proximally brownish; posterior part of mesoscutum with silvery setae; all coxae white; hind tibia dark brown, legs otherwise white to pale orange mixed dark brown; fore wing (
<figureCitation id="13282A34FF3BFF49FB81BF79A6CCFC4A" captionStart="Figs 583-595" captionStartId="778.[181,226,1526,1550]" captionTargetBox="[181,1163,227,1504]" captionTargetId="figure-9@778.[181,1163,223,1508]" captionTargetPageId="778" captionText="Figs 583-595. Cheiloneurus parvus (583) antenna (a, outer aspect of clava; b, inner aspect of clava); (584) ovipositor; (585) hypopygium; (586) fore wing; (587) apex of fore wing venation. Cheiloneurus semis (588) fore wing; (589) hypopygium; (590) apex of fore wing venation; (591) antenna (a, outer aspect of clava; b, inner aspect of clava); (592) mesosoma, dorsal view. Cheiloneurus catreta (593) antenna (a, outer aspect of clava; b, inner aspect of clava); (594) apex of fore wing venation; (595) fore wing." pageId="238" pageNumber="237">Fig. 595</figureCitation>
) hyaline; head about 5.2X as wide as frontovertex, in facial view very slightly longer than broad and subcircular, gena weakly and fairly evenly curved; frontovertex naked medially below anterior ocellus; area between eye and scrobe about 1.5X as wide as anterior ocellus, with distinct sculpture, scrobal margin rounded; interantennal prominence dorsally rounded with only a few setae; mandible with two long, acute, lower teeth and a slightly shorter upper tooth; scape about 2X as long as broad; funicle with F1 and F2 quadrate, other segments transverse; flagellum very slightly longer than head width; linear sensilla on F5-F6; clava 3-segmented, slightly longer than funicle, sensory area extending slightly less than 0.5X along ventral surface; mesoscutum (
<figureCitation id="13282A34FF3BFF49FB81B816A6CCFBA0" captionStart="Figs 596-607" captionStartId="779.[181,226,1526,1550]" captionTargetBox="[181,1163,229,1508]" captionTargetId="figure-10@779.[181,1163,223,1508]" captionTargetPageId="779" captionText="Figs 596-607. Cheiloneurus catreta (596) mesosoma, dorsal view. Cheiloneurus herndonia, HT (597) mesosoma, dorsal view; (598) antenna (a, outer aspect of clava; b, inner aspect of clava); (599) apex of fore wing venation; (600) fore wing. Cheiloneurus exeres (601) fore wing; (602) antenna (a, outer aspect of clava; b, inner aspect of clava); (603) apex of fore wing venation. Cheiloneurus fidenas (604) fore wing; (605) antenna (a, outer aspect of clava; b, inner aspect of clava); (606) apex of fore wing venation; (607) mesosoma, dorsal view." pageId="238" pageNumber="237">Fig. 596</figureCitation>
) mainly with polygonally reticulate sculpture; scutellum with distinct, apical tuft of setae; wings fully developed; fore wing (
<figureCitation id="13282A34FF3BFF49FDC6B843A46EFBBA" box="[548,638,1127,1151]" captionStart="Figs 583-595" captionStartId="778.[181,226,1526,1550]" captionTargetBox="[181,1163,227,1504]" captionTargetId="figure-9@778.[181,1163,223,1508]" captionTargetPageId="778" captionText="Figs 583-595. Cheiloneurus parvus (583) antenna (a, outer aspect of clava; b, inner aspect of clava); (584) ovipositor; (585) hypopygium; (586) fore wing; (587) apex of fore wing venation. Cheiloneurus semis (588) fore wing; (589) hypopygium; (590) apex of fore wing venation; (591) antenna (a, outer aspect of clava; b, inner aspect of clava); (592) mesosoma, dorsal view. Cheiloneurus catreta (593) antenna (a, outer aspect of clava; b, inner aspect of clava); (594) apex of fore wing venation; (595) fore wing." pageId="238" pageNumber="237">Fig. 595</figureCitation>
) about 2.8-3.0X as long as broad; parastigma not curved downwards; costal cell dorsally with two or three setae at apex and a single complete line of setae ventrally; area below proximal part of parastigma with a group of about 10 pale setae; apices of postmarginal and stigmal veins (
<figureCitation id="13282A34FF3BFF49FDBCB893A4A4FB0A" box="[606,692,1207,1231]" captionStart="Figs 583-595" captionStartId="778.[181,226,1526,1550]" captionTargetBox="[181,1163,227,1504]" captionTargetId="figure-9@778.[181,1163,223,1508]" captionTargetPageId="778" captionText="Figs 583-595. Cheiloneurus parvus (583) antenna (a, outer aspect of clava; b, inner aspect of clava); (584) ovipositor; (585) hypopygium; (586) fore wing; (587) apex of fore wing venation. Cheiloneurus semis (588) fore wing; (589) hypopygium; (590) apex of fore wing venation; (591) antenna (a, outer aspect of clava; b, inner aspect of clava); (592) mesosoma, dorsal view. Cheiloneurus catreta (593) antenna (a, outer aspect of clava; b, inner aspect of clava); (594) apex of fore wing venation; (595) fore wing." pageId="238" pageNumber="237">Fig. 594</figureCitation>
) not connected by a naked hyaline area; apical bristle of postmarginal vein about 0.5X as long as marginal vein; mid tibial spur clearly longer than basitarsus; propodeum (
<figureCitation id="13282A34FF3BFF49FE3DB8C9A427FAC0" box="[479,567,1261,1285]" captionStart="Figs 596-607" captionStartId="779.[181,226,1526,1550]" captionTargetBox="[181,1163,229,1508]" captionTargetId="figure-10@779.[181,1163,223,1508]" captionTargetPageId="779" captionText="Figs 596-607. Cheiloneurus catreta (596) mesosoma, dorsal view. Cheiloneurus herndonia, HT (597) mesosoma, dorsal view; (598) antenna (a, outer aspect of clava; b, inner aspect of clava); (599) apex of fore wing venation; (600) fore wing. Cheiloneurus exeres (601) fore wing; (602) antenna (a, outer aspect of clava; b, inner aspect of clava); (603) apex of fore wing venation. Cheiloneurus fidenas (604) fore wing; (605) antenna (a, outer aspect of clava; b, inner aspect of clava); (606) apex of fore wing venation; (607) mesosoma, dorsal view." pageId="238" pageNumber="237">Fig. 596</figureCitation>
) with about 15-18 setae around each spiracle, none down side; syntergum about as long as mid tibia; ovipositor about 3.6X gonostylus or 1.3X mid tibia; gonostylus about as long as mid tibial spur; exserted part of ovipositor about 0.25X mid tibial spur.
<emphasis id="B967EAA3FF3BFF49FF0EB919A730FA90" box="[236,288,1341,1365]" italics="true" pageId="238" pageNumber="237">Male</emphasis>
: unknown.
</paragraph>
<paragraph id="8BAC36B1FF3BFF49FF57B956A293FA4F" blockId="238.[181,1164,1394,1685]" box="[181,1155,1394,1418]" pageId="238" pageNumber="237">
Female (
<typeStatus id="54A88813FF3BFF49FEECB956A77DFA4F" box="[270,365,1394,1418]" pageId="238" pageNumber="237" type="holotype">holotype</typeStatus>
): length, including ovipositor,
<quantity id="4CEB9B54FF3BFF49FD40B956A4E9FA4F" box="[674,761,1394,1418]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" pageId="238" pageNumber="237" unit="mm" value="1.35">1.35mm</quantity>
; excluding ovipositor,
<quantity id="4CEB9B54FF3BFF49FC3DB956A223FA4F" box="[991,1075,1394,1418]" metricMagnitude="-3" metricUnit="m" metricValue="1.31" pageId="238" pageNumber="237" unit="mm" value="1.31">1.31mm</quantity>
(CPD).
</paragraph>
<paragraph id="8BAC36B1FF3BFF48FF39B9A9A6EDFDF3" blockId="238.[181,1164,1394,1685]" lastBlockId="239.[181,1165,222,1447]" lastPageId="239" lastPageNumber="238" pageId="238" pageNumber="237">
Head black with a very dull, dark metallic green sheen on frontovertex with slight purplish reflections, entire scrobal area with a distinct, deep, purple sheen; frontovertex with inconspicuous dark brown setae; temple and posterior part of gena below eye metallic green mixed brassy; remainder of malar space coppery; antenna (
<figureCitation id="13282A34FF3BFF49FD95B9F9A4DDFA30" box="[631,717,1501,1525]" captionStart="Figs 583-595" captionStartId="778.[181,226,1526,1550]" captionTargetBox="[181,1163,227,1504]" captionTargetId="figure-9@778.[181,1163,223,1508]" captionTargetPageId="778" captionText="Figs 583-595. Cheiloneurus parvus (583) antenna (a, outer aspect of clava; b, inner aspect of clava); (584) ovipositor; (585) hypopygium; (586) fore wing; (587) apex of fore wing venation. Cheiloneurus semis (588) fore wing; (589) hypopygium; (590) apex of fore wing venation; (591) antenna (a, outer aspect of clava; b, inner aspect of clava); (592) mesosoma, dorsal view. Cheiloneurus catreta (593) antenna (a, outer aspect of clava; b, inner aspect of clava); (594) apex of fore wing venation; (595) fore wing." pageId="238" pageNumber="237">Fig. 593</figureCitation>
) with radicle dark brown; scape dark brown with apex white; pedicel dorsally dark brown in proximal two-thirds or so, apex white, orange-brown ventrally; funicle with F1-F2 white, F2 dusky distally; F3-F5 dusky yellow to pale orange-brown, F6 pale orange; clava pale orange with basal segment brownish proximally; thorax mostly dark brown to black; pronotum with a dull coppery-purple sheen; anterior one-third or so of mesoscutum relatively dull and clothed in dark brown setae, posterior two-thirds with slight metallic blue and purple reflections and clothed in moderately dense, slightly silvery, translucent setae, posterior margin dark purple; tegula dark brown; scutellum clothed with dark brown setae, mostly matt, posteriorly with a coppery purple sheen, sides and apex shining metallic green with some coppery reflections; metanotum with slight purple sheen; mesopleuron dark purple-brown; fore coxa white; fore femur white with a broad, subapical brown band; tibia with a broad subbasal brown band, narrowly pale orange proximally, distal half pale orange; mid coxa white, brown towards base; mid femur white with a narrow, dark brown, subapical band; mid tibia white with a brown subbasal band; hind coxa white; hind femur white with apical half dark brown; hind tibia dark brown, narrowly pale yellow proximally, apical one-third or so white; all tarsi very pale orange; pretarsi brownish; fore wing (
<figureCitation id="13282A34FF3AFF48FDD4BD90A49CFE09" box="[566,652,436,460]" captionStart="Figs 583-595" captionStartId="778.[181,226,1526,1550]" captionTargetBox="[181,1163,227,1504]" captionTargetId="figure-9@778.[181,1163,223,1508]" captionTargetPageId="778" captionText="Figs 583-595. Cheiloneurus parvus (583) antenna (a, outer aspect of clava; b, inner aspect of clava); (584) ovipositor; (585) hypopygium; (586) fore wing; (587) apex of fore wing venation. Cheiloneurus semis (588) fore wing; (589) hypopygium; (590) apex of fore wing venation; (591) antenna (a, outer aspect of clava; b, inner aspect of clava); (592) mesosoma, dorsal view. Cheiloneurus catreta (593) antenna (a, outer aspect of clava; b, inner aspect of clava); (594) apex of fore wing venation; (595) fore wing." pageId="239" pageNumber="238">Fig. 595</figureCitation>
) completely hyaline, venation pale yellow-brown; propodeum dark purple-brown; sides weakly metallic green with conspicuous silvery setae in spiracular area; gaster dark purple-brown, Gt1 distinctly metallic blue; remainder of gaster mixed brassy, sides of apical tergites weakly metallic green with purple reflections; gonostylus dark brown.
</paragraph>
<paragraph id="8BAC36B1FF3AFF48FF39BE1DA5ACFC54" blockId="239.[181,1165,222,1447]" pageId="239" pageNumber="238">
Head about 5.2X as wide as frontovertex with occipital margin sharp and without a median groove to foramen; ocelli forming an angle of about 45°; frontovertex slightly shiny, with conspicuous reticulate sculpture of mesh size generally slightly smaller than diameter of eye facet; scrobes, interantennal prominence and mouth margin with shallower sculpture; temple and gena with distinct irregular, reticulate sculpture, of larger mesh than the frontovertex and distinctly elongate posteriorly; scrobes moderately deep, not meeting, dorsal and lateral margins weakly rounded; antenna as in
<figureCitation id="13282A34FF3AFF48FE7BBEFDA7FFFD34" box="[409,495,729,753]" captionStart="Figs 583-595" captionStartId="778.[181,226,1526,1550]" captionTargetBox="[181,1163,227,1504]" captionTargetId="figure-9@778.[181,1163,223,1508]" captionTargetPageId="778" captionText="Figs 583-595. Cheiloneurus parvus (583) antenna (a, outer aspect of clava; b, inner aspect of clava); (584) ovipositor; (585) hypopygium; (586) fore wing; (587) apex of fore wing venation. Cheiloneurus semis (588) fore wing; (589) hypopygium; (590) apex of fore wing venation; (591) antenna (a, outer aspect of clava; b, inner aspect of clava); (592) mesosoma, dorsal view. Cheiloneurus catreta (593) antenna (a, outer aspect of clava; b, inner aspect of clava); (594) apex of fore wing venation; (595) fore wing." pageId="239" pageNumber="238">Fig. 593</figureCitation>
; scape strongly broadened and flattened, about 2X as long as broad; funicle with F1 and F2 more or less quadrate; remaining segments transverse; clava about long as funicle, both sutures slightly oblique, sensory area forming an oblique truncation that extends about half-way along clava; eye reaching occipital margin and clothed in fairly conspicuous setae that are each slightly longer than diameter of facet; malar sulcus absent; mandible with 2 long, acute lower teeth and short upper, third tooth. Relative measurements: HW 52,
<collectingRegion id="49D7F853FF3AFF48FC2CBF7BA5E3FCB3" box="[974,1011,863,886]" country="Swaziland" name="Hhohho" pageId="239" pageNumber="238">HH</collectingRegion>
53, FV 10, OD 4, POL 5, OOL 0.5, OCL 6, AOL 9.5, EL 38, EW 29, MS 27, SL 31, SW 16.
</paragraph>
<paragraph id="8BAC36B1FF3AFF48FF39BFB0A29CFB17" blockId="239.[181,1165,222,1447]" pageId="239" pageNumber="238">
Thorax (
<figureCitation id="13282A34FF3AFF48FEDABFB0A783FC69" box="[312,403,916,940]" captionStart="Figs 596-607" captionStartId="779.[181,226,1526,1550]" captionTargetBox="[181,1163,229,1508]" captionTargetId="figure-10@779.[181,1163,223,1508]" captionTargetPageId="779" captionText="Figs 596-607. Cheiloneurus catreta (596) mesosoma, dorsal view. Cheiloneurus herndonia, HT (597) mesosoma, dorsal view; (598) antenna (a, outer aspect of clava; b, inner aspect of clava); (599) apex of fore wing venation; (600) fore wing. Cheiloneurus exeres (601) fore wing; (602) antenna (a, outer aspect of clava; b, inner aspect of clava); (603) apex of fore wing venation. Cheiloneurus fidenas (604) fore wing; (605) antenna (a, outer aspect of clava; b, inner aspect of clava); (606) apex of fore wing venation; (607) mesosoma, dorsal view." pageId="239" pageNumber="238">Fig. 596</figureCitation>
) with shallow, irregular, polygonally reticulate sculpture on pronotum, generally of mesh size a little larger than diameter of facet; sculpture on anterior part of mesoscutum coarser and finer than that on pronotum, but posterior part with shallower sculpture similar to that on pronotum; sculpture on axilla slightly coarser and reticulate; scutellum mostly with coarse, slightly longitudinally elongate, polygonally reticulate sculpture, extreme apex and sides completely smooth; visible part of mesoscutum about 2X as broad as long; scutellum about as long as broad, subapical tuft distinct, setae about half as long as scutellum; hind femur about 3.5X as long as broad; fore wing with venation and setation as in
<figureCitation id="13282A34FF3AFF48FCB5B86BA5F6FBA2" box="[855,998,1103,1127]" captionStart="Figs 583-595" captionStartId="778.[181,226,1526,1550]" captionTargetBox="[181,1163,227,1504]" captionTargetId="figure-9@778.[181,1163,223,1508]" captionTargetPageId="778" captionText="Figs 583-595. Cheiloneurus parvus (583) antenna (a, outer aspect of clava; b, inner aspect of clava); (584) ovipositor; (585) hypopygium; (586) fore wing; (587) apex of fore wing venation. Cheiloneurus semis (588) fore wing; (589) hypopygium; (590) apex of fore wing venation; (591) antenna (a, outer aspect of clava; b, inner aspect of clava); (592) mesosoma, dorsal view. Cheiloneurus catreta (593) antenna (a, outer aspect of clava; b, inner aspect of clava); (594) apex of fore wing venation; (595) fore wing." pageId="239" pageNumber="238">Figs 594, 595</figureCitation>
; costal cell with 2 or 3 setae dorsally at apex; no naked streak present between apices of postmarginal and stigmal veins (
<figureCitation id="13282A34FF3AFF48FF1CB8A0A74CFB59" box="[254,348,1156,1180]" captionStart="Figs 583-595" captionStartId="778.[181,226,1526,1550]" captionTargetBox="[181,1163,227,1504]" captionTargetId="figure-9@778.[181,1163,223,1508]" captionTargetPageId="778" captionText="Figs 583-595. Cheiloneurus parvus (583) antenna (a, outer aspect of clava; b, inner aspect of clava); (584) ovipositor; (585) hypopygium; (586) fore wing; (587) apex of fore wing venation. Cheiloneurus semis (588) fore wing; (589) hypopygium; (590) apex of fore wing venation; (591) antenna (a, outer aspect of clava; b, inner aspect of clava); (592) mesosoma, dorsal view. Cheiloneurus catreta (593) antenna (a, outer aspect of clava; b, inner aspect of clava); (594) apex of fore wing venation; (595) fore wing." pageId="239" pageNumber="238">Fig. 594</figureCitation>
); propodeum (
<figureCitation id="13282A34FF3AFF48FE1BB8A0A444FB59" box="[505,596,1156,1180]" captionStart="Figs 596-607" captionStartId="779.[181,226,1526,1550]" captionTargetBox="[181,1163,229,1508]" captionTargetId="figure-10@779.[181,1163,223,1508]" captionTargetPageId="779" captionText="Figs 596-607. Cheiloneurus catreta (596) mesosoma, dorsal view. Cheiloneurus herndonia, HT (597) mesosoma, dorsal view; (598) antenna (a, outer aspect of clava; b, inner aspect of clava); (599) apex of fore wing venation; (600) fore wing. Cheiloneurus exeres (601) fore wing; (602) antenna (a, outer aspect of clava; b, inner aspect of clava); (603) apex of fore wing venation. Cheiloneurus fidenas (604) fore wing; (605) antenna (a, outer aspect of clava; b, inner aspect of clava); (606) apex of fore wing venation; (607) mesosoma, dorsal view." pageId="239" pageNumber="238">Fig. 596</figureCitation>
) with numerous setae in spiracular area, side naked, medially about 0.1X as long as scutellum and medially with a small area of fine, very shallow, polygonally reticulate sculpture. Relative measurements: FWL 59, FWW 20; HWL 47, HWW 10.
</paragraph>
<paragraph id="8BAC36B1FF3AFF48FF39B8F0A26FFAC2" blockId="239.[181,1165,222,1447]" pageId="239" pageNumber="238">Gaster with hypopygium reaching about half-way to apex; ovipositor hardly exserted, the exserted part about one-quarter as long as mid tibial spur; syntergum about as long as mid tibia.</paragraph>
<paragraph id="8BAC36B1FF3AFF48FF39B92EA781FA92" blockId="239.[181,1165,222,1447]" pageId="239" pageNumber="238">
<typeStatus id="54A88813FF3AFF48FF39B92EA727FAE7" box="[219,311,1290,1314]" pageId="239" pageNumber="238" type="paratype">Paratype</typeStatus>
. Funicle with linear sensilla present only on F5-F6; gaster without “gland-like” structures on Gt1 or Gt5; gonostylus about as long as mid tibial spur. Relative measurements: OL 41, GL 11.5 [MT 31].
</paragraph>
<paragraph id="8BAC36B1FF3AFF48FF39B97DA708FA62" blockId="239.[181,1165,222,1447]" pageId="239" pageNumber="238">
Variation. Females vary in overall length from
<quantity id="4CEB9B54FF3AFF48FD5FB97EA557FAB7" box="[701,839,1370,1394]" metricMagnitude="-3" metricUnit="m" metricValue="1.325" metricValueMax="1.44" metricValueMin="1.21" pageId="239" pageNumber="238" unit="mm" value="1.325" valueMax="1.44" valueMin="1.21">1.21-1.44mm</quantity>
, the clava may be hardly dusky basally and the fore wing may be 2.8-3.0X as long as broad, otherwise very little in material available.
</paragraph>
<paragraph id="8BAC36B1FF3AFF48FF57B9E0A771FA19" blockId="239.[181,353,1476,1500]" box="[181,353,1476,1500]" pageId="239" pageNumber="238">Male. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C309653AFF3AFF48FF57B9DEA7F9F9D4" box="[181,489,1529,1554]" pageId="239" pageNumber="238" type="distribution">
<paragraph id="8BAC36B1FF3AFF48FF57B9DEA7F9F9D4" blockId="239.[181,489,1529,1554]" box="[181,489,1529,1554]" pageId="239" pageNumber="238">
DISTRIBUTION.
<collectingCountry id="F3047621FF3AFF48FE91B9DEA7F5F9D4" box="[371,485,1529,1554]" name="Costa Rica" pageId="239" pageNumber="238">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C309653AFF3AFF48FF57BA0BA76EF982" box="[181,382,1583,1607]" pageId="239" pageNumber="238" type="biology_ecology">
<paragraph id="8BAC36B1FF3AFF48FF57BA0BA76EF982" blockId="239.[181,382,1583,1607]" box="[181,382,1583,1607]" pageId="239" pageNumber="238">HOSTS. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C309653AFF3AFF57FF57BA42A260FECB" lastPageId="240" lastPageNumber="239" pageId="239" pageNumber="238" type="materials_examined">
<paragraph id="8BAC36B1FF3AFF48FF57BA42A29DF953" blockId="239.[181,1165,1638,1686]" pageId="239" pageNumber="238">
<materialsCitation id="3B7B3CECFF3AFF48FF57BA42A29DF953" country="Costa Rica" elevation="100" latitude="10.4" location="Est. Biol. La Selva" longLatPrecision="1298" longitude="-84.01667" pageId="239" pageNumber="238" specimenCount="1" stateProvince="Heredia" typeStatus="holotype">
MATERIAL EXAMINED. Type material.
<typeStatus id="54A88813FF3AFF48FE86BAA4A7A8F953" box="[356,440,1664,1686]" pageId="239" pageNumber="238" type="holotype">Holotype</typeStatus>
E:
<emphasis id="B967EAA3FF3AFF48FE3EBAA4A477F953" bold="true" box="[476,615,1664,1686]" pageId="239" pageNumber="238">
<collectingCountry id="F3047621FF3AFF48FE3EBAA4A477F953" box="[476,615,1664,1686]" name="Costa Rica" pageId="239" pageNumber="238">COSTA RICA</collectingCountry>
</emphasis>
,
<emphasis id="B967EAA3FF3AFF48FD91BAA4A4D1F953" bold="true" box="[627,705,1664,1686]" pageId="239" pageNumber="238">
<collectingRegion id="49D7F853FF3AFF48FD91BAA4A4D1F953" box="[627,705,1664,1686]" country="Costa Rica" name="Heredia" pageId="239" pageNumber="238">Heredia</collectingRegion>
</emphasis>
,
<location id="8ECC606AFF3AFF48FD2EBAA5A56AF953" LSID="urn:lsid:plazi:treatment:03BA87A7FF3BFF57FDE8BE86A7FCFE9A:8ECC606AFF3AFF48FD2EBAA5A56AF953" box="[716,890,1664,1686]" country="Costa Rica" latitude="10.4" longLatPrecision="1298" longitude="-84.01667" name="Est. Biol. La Selva" pageId="239" pageNumber="238" stateProvince="Heredia">Est. Biol. La Selva</location>
,
<geoCoordinate id="EE275076FF3AFF48FC61BAA4A5C2F950" box="[899,978,1664,1685]" degrees="10" direction="north" minutes="24" orientation="latitude" pageId="239" pageNumber="238" precision="925" value="10.4">10°24N</geoCoordinate>
<geoCoordinate id="EE275076FF3AFF48FC3ABAA4A23BF950" box="[984,1067,1664,1685]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="239" pageNumber="238" precision="925" value="-84.01667">84°01W</geoCoordinate>
,
<quantity id="4CEB9B54FF3AFF48FBD7BAA4A298F953" box="[1077,1160,1664,1686]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="239" pageNumber="238" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">
<elevation id="003ED182FF3AFF48FBD7BAA4A298F953" box="[1077,1160,1664,1686]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="239" pageNumber="238" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">50-150m</elevation>
</quantity>
,
</materialsCitation>
</paragraph>
<paragraph id="8BAC36B1FF25FF57FF57BCFBA260FECB" blockId="240.[181,1164,222,271]" pageId="240" pageNumber="239">
<materialsCitation id="3B7B3CECFF25FF57FF57BCFBA556FF31" box="[181,838,222,244]" collectingDate="2000-01-05" collectionCode="INBio, OET" pageId="240" pageNumber="239" specimenCount="1" typeStatus="paratype">
canopy fog
<taxonomicName id="4C134D32FF25FF57FEFDBCFBA79BFF31" box="[287,395,223,244]" class="Insecta" family="Tachinidae" genus="Eugenia" kingdom="Animalia" order="Diptera" pageId="240" pageNumber="239" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B967EAA3FF25FF57FEFDBCFBA77BFF31" box="[287,363,223,244]" italics="true" pageId="240" pageNumber="239">Eugenia</emphasis>
sp.
</taxonomicName>
, FOT/76/1-40,
<date id="FFAD1071FF25FF57FDF4BCFAA470FF31" box="[534,608,222,244]" pageId="240" pageNumber="239" value="2000-01-05">
<collectingDate id="EFE9E999FF25FF57FDF4BCFAA470FF31" box="[534,608,222,244]" pageId="240" pageNumber="239" value="2000-01-05">5.i.2000</collectingDate>
</date>
(
<collectionCode id="ED02AE74FF25FF57FD89BCFAA4B6FF31" box="[619,678,222,244]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="240" pageNumber="239">INBio</collectionCode>
/
<collectionCode id="ED02AE74FF25FF57FD4CBCFAA4CCFF31" box="[686,732,222,244]" pageId="240" pageNumber="239">OET</collectionCode>
).
<typeStatus id="54A88813FF25FF57FD08BCFBA556FF31" box="[746,838,223,244]" pageId="240" pageNumber="239" type="paratype">Paratypes</typeStatus>
</materialsCitation>
:
<materialsCitation id="3B7B3CECFF25FF57FCAFBCFAA536FECA" collectingDate="xi. 1993 and ix. 1995" collectionCode="ALAS" collectorName="La Selva" country="Costa Rica" latitude="10.433333" location="Heredia" longLatPrecision="1298" longitude="-84.01667" pageId="240" pageNumber="239" specimenCount="1" stateProvince="Heredia" typeStatus="holotype">
<emphasis id="B967EAA3FF25FF57FCAFBCFAA5C7FF31" bold="true" box="[845,983,222,244]" pageId="240" pageNumber="239">
<collectingCountry id="F3047621FF25FF57FCAFBCFAA5C7FF31" box="[845,983,222,244]" name="Costa Rica" pageId="240" pageNumber="239">COSTA RICA</collectingCountry>
</emphasis>
, 2E,
<emphasis id="B967EAA3FF25FF57FBEBBCFAA247FF31" bold="true" box="[1033,1111,222,244]" pageId="240" pageNumber="239">
<collectingRegion id="49D7F853FF25FF57FBEBBCFAA247FF31" box="[1033,1111,222,244]" country="Costa Rica" name="Heredia" pageId="240" pageNumber="239">Heredia</collectingRegion>
</emphasis>
, OTS
<collectorName id="26E65367FF25FF57FF57BCDDA718FECA" box="[181,264,249,271]" pageId="240" pageNumber="239">La Selva</collectorName>
,
<geoCoordinate id="EE275076FF25FF57FEF3BCDDA74FFECA" box="[273,351,249,271]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="240" pageNumber="239" precision="925" value="10.433333">10°26N</geoCoordinate>
<geoCoordinate id="EE275076FF25FF57FE87BCDDA7A8FECB" box="[357,440,249,270]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="240" pageNumber="239" precision="925" value="-84.01667">84°01W</geoCoordinate>
,
<collectingDate id="EFE9E999FF25FF57FE23BCDDA467FECA" box="[449,631,249,271]" isEnumeration="true" pageId="240" pageNumber="239">
<date id="FFAD1071FF25FF57FE23BCDDA416FECA" box="[449,518,249,271]" pageId="240" pageNumber="239" value="1993-11">xi.1993</date>
and
<date id="FFAD1071FF25FF57FDD0BCDDA467FECA" box="[562,631,249,271]" pageId="240" pageNumber="239" value="1995-09">ix.1995</date>
</collectingDate>
(
<collectionCode id="ED02AE74FF25FF57FD61BCDDA4D3FECA" box="[643,707,249,271]" pageId="240" pageNumber="239">ALAS</collectionCode>
).
<typeStatus id="54A88813FF25FF57FD30BCDDA536FECA" box="[722,806,249,271]" pageId="240" pageNumber="239" type="holotype">Holotype</typeStatus>
</materialsCitation>
<materialsCitation id="3B7B3CECFF25FF57FCCEBCDDA5E6FECB" box="[812,1014,249,271]" collectionCode="MZUCR" country="Costa Rica" pageId="240" pageNumber="239" specimenCount="1" stateProvince="Heredia" typeStatus="paratype">
in
<collectionCode id="ED02AE74FF25FF57FCA1BCDDA586FECB" box="[835,918,249,270]" pageId="240" pageNumber="239">MZUCR</collectionCode>
,
<typeStatus id="54A88813FF25FF57FC7DBCDDA5E6FECB" box="[927,1014,249,270]" pageId="240" pageNumber="239" type="paratype">paratypes</typeStatus>
</materialsCitation>
in
<collectionCode id="ED02AE74FF25FF57FBF1BCDDA27CFECB" box="[1043,1132,249,270]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="240" pageNumber="239" type="Museum">NHMUK</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C309653AFF25FF57FF57BD09A7FCFE9A" pageId="240" pageNumber="239" type="discussion">
<paragraph id="8BAC36B1FF25FF57FF57BD09A7FCFE9A" blockId="240.[181,1164,301,351]" pageId="240" pageNumber="239">
COMMENTS. The
<typeStatus id="54A88813FF25FF57FE60BD09A7CBFE80" box="[386,475,301,325]" pageId="240" pageNumber="239" type="holotype">holotype</typeStatus>
appears to have F1 and F2 partially fused on both antennae, but is otherwise in perfect condition.
</paragraph>
</subSubSection>
</treatment>
</document>