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<document id="BC21972078CAC2F78C69970436684214" ID-DOI="10.5281/zenodo.8074943" ID-ISSN="2754-9844" ID-Zenodo-Dep="8074943" ID-ZooBank="BCAD06E8-0AFE-46ED-B7FA-930983CD44C4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="admin" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="admin" checkinTime="1699388683108" checkinUser="felipe" docAuthor="Noyes, John Stuart" docDate="2023" docId="03BA87A7FF33FF40FDE8BCFAA4BFFB21" docLanguage="en" docName="TaxMonogNeotropHymenopt.2.1-921.pdf" docOrigin="Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) 2 (1)" docSource="http://dx.doi.org/10.5281/zenodo.8074943" docStyle="DocumentStyle:DAA2ADC4E839520FEEF1E0A10B782A41.1:TaxMonogNeotropHymenopt.2022-.journal_article" docStyleId="DAA2ADC4E839520FEEF1E0A10B782A41" docStyleName="TaxMonogNeotropHymenopt.2022-.journal_article" docStyleVersion="1" docTitle="Cheiloneurus roellus Noyes 2023, sp.nov." docType="treatment" docUuid="DA846BDD-F65A-472F-B470-6E073E52E014" docUuidSource="ZooBank" docVersion="2" lastPageNumber="230" masterDocId="FF83FFDFFFD5FFA7FFE2BC24A610FFC5" masterDocTitle="ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)" masterLastPageNumber="921" masterPageNumber="1" pageNumber="229" updateTime="1699637169064" updateUser="admin" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="96D9E861565E91E79BFCE59298C874CD">ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)</mods:title>
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<mods:namePart id="5EB0F2BA8BFE07428E714C777AE696A2">Noyes, John Stuart</mods:namePart>
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<mods:title id="56D1ACD9BF948B9DFE8276A3228FB8BF">Taxonomic Monographs on Neotropical Hymenoptera</mods:title>
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<mods:date id="D1EE16340B60A5DE5844A5CD38C6A179">2023</mods:date>
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<subSubSection id="C309653AFF33FF41FDE8BCFAA528FF33" box="[522,824,222,246]" pageId="230" pageNumber="229" type="nomenclature">
<paragraph id="8BAC36B1FF33FF41FDE8BCFAA528FF33" blockId="230.[522,824,222,273]" box="[522,824,222,246]" pageId="230" pageNumber="229">
<heading id="D0E481DDFF33FF41FDE8BCFAA528FF33" bold="true" box="[522,824,222,246]" centered="true" fontSize="10" level="2" pageId="230" pageNumber="229" reason="2">
<emphasis id="B967EAA3FF33FF41FDE8BCFAA528FF33" bold="true" box="[522,824,222,246]" pageId="230" pageNumber="229">
<taxonomicName id="4C134D32FF33FF41FDE8BCFAA4F7FF33" authority="Noyes, 2023" authorityName="Noyes" authorityYear="2023" box="[522,743,222,246]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="230" pageNumber="229" phylum="Arthropoda" rank="species" species="roellus" status="sp.nov.">
<emphasis id="B967EAA3FF33FF41FDE8BCFAA4F7FF33" bold="true" box="[522,743,222,246]" italics="true" pageId="230" pageNumber="229">Cheiloneurus roellus</emphasis>
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<subSubSection id="C309653AFF33FF41FDFBBCDDA539FED4" box="[537,809,248,273]" pageId="230" pageNumber="229" type="description">
<paragraph id="8BAC36B1FF33FF41FDFBBCDDA539FED4" blockId="230.[522,824,222,273]" box="[537,809,248,273]" pageId="230" pageNumber="229">
(
<figureCitation id="13282A34FF33FF41FDC2BCDDA4BBFED5" box="[544,683,248,273]" captionStart-0="Figs 553-568" captionStart-1="Figs 569-582" captionStartId-0="776.[181,226,1526,1550]" captionStartId-1="777.[181,226,1526,1550]" captionTargetBox-0="[185,1163,227,1508]" captionTargetBox-1="[185,1163,231,1508]" captionTargetId-0="figure-9@776.[181,1163,223,1508]" captionTargetId-1="figure-10@777.[181,1163,223,1508]" captionTargetPageId-0="776" captionTargetPageId-1="777" captionText-0="Figs 553-568. Cheiloneurus danessa, HT (553) fore wing. Cheiloneurus idris (554) ovipositor; (555) apex of fore wing venation; (556) head, facial view; (557) fore wing; (558) antenna; (559) hypopygium; (560) mesosoma, dorsal view. Cheiloneurus socrates, HT (561) antenna; (562) fore wing; (563) apex of fore wing venation; (564) mesosoma, dorsal view; (565) ovipositor; (566) gastral tergites showing “glands” on Gt1 and Gt5. Cheiloneurus roellus, HT (567) antenna; (568) mesosoma, dorsal view." captionText-1="Figs 569-582. Cheiloneurus roellus, HT (569) ovipositor; (570) apex of fore wing venation; (571) fore wing. Cheiloneurus dalycera (572) mesosoma, dorsal view; (573) antenna; (574) apex of fore wing venation; (575) fore wing. Cheiloneurus ection, HT (576) antenna (clava slightly collapsed); (577) ovipositor; (578) fore wing; (579) apex of fore wing venation; (580) mesosoma, dorsal view. Cheiloneurus parvus (581) head, facial view (reconstructed); (582) mesosoma, dorsal view." pageId="230" pageNumber="229">Figs 567-571</figureCitation>
; Hab. E 87)
</paragraph>
</subSubSection>
<subSubSection id="C309653AFF33FF40FF57BD0AA771FC35" lastPageId="231" lastPageNumber="230" pageId="230" pageNumber="229" type="diagnosis">
<paragraph id="8BAC36B1FF33FF41FF57BD0AA476FC54" blockId="230.[181,1165,302,913]" pageId="230" pageNumber="229">
DIAGNOSIS.
<emphasis id="B967EAA3FF33FF41FEADBD0AA78BFE83" box="[335,411,302,326]" italics="true" pageId="230" pageNumber="229">Female</emphasis>
(length about
<quantity id="4CEB9B54FF33FF41FDD8BD0AA4BEFE83" box="[570,686,302,326]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="230" pageNumber="229" unit="mm" value="1.25" valueMax="1.3" valueMin="1.2">1.2-1.3mm</quantity>
): body generally dark brown to black with a weak to moderate metallic sheen; scape (
<figureCitation id="13282A34FF33FF41FDB6BD6DA4BAFEA5" box="[596,682,328,353]" captionStart="Figs 553-568" captionStartId="776.[181,226,1526,1550]" captionTargetBox="[185,1163,227,1508]" captionTargetId="figure-9@776.[181,1163,223,1508]" captionTargetPageId="776" captionText="Figs 553-568. Cheiloneurus danessa, HT (553) fore wing. Cheiloneurus idris (554) ovipositor; (555) apex of fore wing venation; (556) head, facial view; (557) fore wing; (558) antenna; (559) hypopygium; (560) mesosoma, dorsal view. Cheiloneurus socrates, HT (561) antenna; (562) fore wing; (563) apex of fore wing venation; (564) mesosoma, dorsal view; (565) ovipositor; (566) gastral tergites showing “glands” on Gt1 and Gt5. Cheiloneurus roellus, HT (567) antenna; (568) mesosoma, dorsal view." pageId="230" pageNumber="229">Fig. 567</figureCitation>
) dark brown, apex yellow; pedicel brown, apex yellow; funicle and clava pale testaceous yellow; mesoscutum with pale, silvery setae; coxae white, mid coxa brown proximally; legs otherwise white to pale orange with brown areas; fore wing hyaline; head about 3.7X as wide as frontovertex, in facial view about as broad as broad as long, subcircular, gena weakly and evenly curved; frontovertex naked medially below anterior ocellus; area between eye and scrobe about 2X as wide as diameter of anterior ocellus, with distinct sculpture, scrobal margin rounded; interantennal prominence dorsally rounded with only a few long setae; mandible with three sharp teeth, upper tooth shortest; antenna (
<figureCitation id="13282A34FF33FF41FC1ABE27A240FDDE" box="[1016,1104,515,539]" captionStart="Figs 553-568" captionStartId="776.[181,226,1526,1550]" captionTargetBox="[185,1163,227,1508]" captionTargetId="figure-9@776.[181,1163,223,1508]" captionTargetPageId="776" captionText="Figs 553-568. Cheiloneurus danessa, HT (553) fore wing. Cheiloneurus idris (554) ovipositor; (555) apex of fore wing venation; (556) head, facial view; (557) fore wing; (558) antenna; (559) hypopygium; (560) mesosoma, dorsal view. Cheiloneurus socrates, HT (561) antenna; (562) fore wing; (563) apex of fore wing venation; (564) mesosoma, dorsal view; (565) ovipositor; (566) gastral tergites showing “glands” on Gt1 and Gt5. Cheiloneurus roellus, HT (567) antenna; (568) mesosoma, dorsal view." pageId="230" pageNumber="229">Fig. 567</figureCitation>
) with scape about 4X as long as broad; funicle with all segments subquadrate, becoming larger distad; head about as wide as length of flagellum; linear sensilla on F5-F6; clava 3-segmented, about as long as F2-F6 combined, sensory area small, extending less than 0.1X along ventral margin; mesoscutum(
<figureCitation id="13282A34FF33FF41FEDDBE4AA783FD43" box="[319,403,622,646]" captionStart="Figs 553-568" captionStartId="776.[181,226,1526,1550]" captionTargetBox="[185,1163,227,1508]" captionTargetId="figure-9@776.[181,1163,223,1508]" captionTargetPageId="776" captionText="Figs 553-568. Cheiloneurus danessa, HT (553) fore wing. Cheiloneurus idris (554) ovipositor; (555) apex of fore wing venation; (556) head, facial view; (557) fore wing; (558) antenna; (559) hypopygium; (560) mesosoma, dorsal view. Cheiloneurus socrates, HT (561) antenna; (562) fore wing; (563) apex of fore wing venation; (564) mesosoma, dorsal view; (565) ovipositor; (566) gastral tergites showing “glands” on Gt1 and Gt5. Cheiloneurus roellus, HT (567) antenna; (568) mesosoma, dorsal view." pageId="230" pageNumber="229">Fig. 568</figureCitation>
) with polygonally reticulate sculpture; scutellum with a distinct apical tuft of setae; wings fully developed; fore wing (
<figureCitation id="13282A34FF33FF41FD9DBEADA4C7FD64" box="[639,727,649,673]" captionStart="Figs 569-582" captionStartId="777.[181,226,1526,1550]" captionTargetBox="[185,1163,231,1508]" captionTargetId="figure-10@777.[181,1163,223,1508]" captionTargetPageId="777" captionText="Figs 569-582. Cheiloneurus roellus, HT (569) ovipositor; (570) apex of fore wing venation; (571) fore wing. Cheiloneurus dalycera (572) mesosoma, dorsal view; (573) antenna; (574) apex of fore wing venation; (575) fore wing. Cheiloneurus ection, HT (576) antenna (clava slightly collapsed); (577) ovipositor; (578) fore wing; (579) apex of fore wing venation; (580) mesosoma, dorsal view. Cheiloneurus parvus (581) head, facial view (reconstructed); (582) mesosoma, dorsal view." pageId="230" pageNumber="229">Fig. 571</figureCitation>
) about 2.7X as long as broad; parastigma slightly swollen, not curved downwards; apex of stigmal vein with 4 campaniform sensilla; costal cell dorsally with only 1 or 2 setae at apex and a single, complete line of setae ventrally; area below proximal part of parastigma with a group of about 15 pale setae; apices of postmarginal and stigmal veins not connected by a naked hyaline area (
<figureCitation id="13282A34FF33FF41FD2BBED7A50EFCCE" box="[713,798,755,779]" captionStart="Figs 569-582" captionStartId="777.[181,226,1526,1550]" captionTargetBox="[185,1163,231,1508]" captionTargetId="figure-10@777.[181,1163,223,1508]" captionTargetPageId="777" captionText="Figs 569-582. Cheiloneurus roellus, HT (569) ovipositor; (570) apex of fore wing venation; (571) fore wing. Cheiloneurus dalycera (572) mesosoma, dorsal view; (573) antenna; (574) apex of fore wing venation; (575) fore wing. Cheiloneurus ection, HT (576) antenna (clava slightly collapsed); (577) ovipositor; (578) fore wing; (579) apex of fore wing venation; (580) mesosoma, dorsal view. Cheiloneurus parvus (581) head, facial view (reconstructed); (582) mesosoma, dorsal view." pageId="230" pageNumber="229">Fig. 570</figureCitation>
); apical bristle of postmarginal vein about 0.5X as long as marginal vein; mid tibial spur slightly longer than basitarsus; propodeum with about 15 setae around each spiracle, not extending down side; gaster without “glands” on Gt1 and Gt5; syntergum about 1.2X as long as mid tibia; ovipositor about 2.7X gonostylus or 1.7X mid tibia; gonostylus about 1.8X as long as mid tibial spur; exserted part of ovipositor about 0.4X as long as mid tibial spur.
<emphasis id="B967EAA3FF33FF41FE5DBF5DA7E3FC54" box="[447,499,889,913]" italics="true" pageId="230" pageNumber="229">Male</emphasis>
: unknown.
</paragraph>
<paragraph id="8BAC36B1FF33FF41FF57BF8AA293FC03" blockId="230.[181,1165,942,1686]" box="[181,1155,942,966]" pageId="230" pageNumber="229">
Female (
<typeStatus id="54A88813FF33FF41FEECBF8AA77DFC03" box="[270,365,942,966]" pageId="230" pageNumber="229">holotype</typeStatus>
): length, including ovipositor,
<quantity id="4CEB9B54FF33FF41FD40BF8AA4E9FC03" box="[674,761,942,966]" metricMagnitude="-3" metricUnit="m" metricValue="1.21" pageId="230" pageNumber="229" unit="mm" value="1.21">1.21mm</quantity>
; excluding ovipositor,
<quantity id="4CEB9B54FF33FF41FC3DBF8AA223FC03" box="[991,1075,942,966]" metricMagnitude="-3" metricUnit="m" metricValue="1.18" pageId="230" pageNumber="229" unit="mm" value="1.18">1.18mm</quantity>
(CPD).
</paragraph>
<paragraph id="8BAC36B1FF33FF41FF39BFEDA475FA33" blockId="230.[181,1165,942,1686]" pageId="230" pageNumber="229">
Head black with weak, blue and coppery purple reflections on frontovertex; scrobal area mostly with a purple sheen merging to coppery laterally on gena; mouth margin with a slight metallic green sheen; temple metallic green; frontovertex with inconspicuous dark brown setae; antenna (
<figureCitation id="13282A34FF33FF41FEF3B83DA777FBF4" box="[273,359,1049,1073]" captionStart="Figs 553-568" captionStartId="776.[181,226,1526,1550]" captionTargetBox="[185,1163,227,1508]" captionTargetId="figure-9@776.[181,1163,223,1508]" captionTargetPageId="776" captionText="Figs 553-568. Cheiloneurus danessa, HT (553) fore wing. Cheiloneurus idris (554) ovipositor; (555) apex of fore wing venation; (556) head, facial view; (557) fore wing; (558) antenna; (559) hypopygium; (560) mesosoma, dorsal view. Cheiloneurus socrates, HT (561) antenna; (562) fore wing; (563) apex of fore wing venation; (564) mesosoma, dorsal view; (565) ovipositor; (566) gastral tergites showing “glands” on Gt1 and Gt5. Cheiloneurus roellus, HT (567) antenna; (568) mesosoma, dorsal view." pageId="230" pageNumber="229">Fig. 567</figureCitation>
) with radicle dark brown; scape dark brown, apical one-third yellowish; pedicel brown, apical one-third yellow; flagellum pale testaceous-yellow; thorax mostly dark brown to black; pronotum with a slight purple sheen; mesoscutum clothed in pale, silvery setae and with a weak, blue sheen mixed purple and posterior margin dark purple; tegula dark brown; axilla with a weak dark blue and purple sheen; scutellum matt, clothed with dark brown setae, sides and apex shining green with a distinct dark blue and purple sheen; metanotum with slight purple sheen; mesopleuron in posterior half with a coppery-purple sheen, anterior half with a weak violet sheen; fore coxa white; fore femur white proximally, apical half dark brown, tibia narrowly white proximally, remainder of basal half dark brown, apex very pale orange; mid coxa brown proximally, otherwise white; mid femur white in proximal one-third, and extreme apex, otherwise dark brown, tibia narrowly white proximally, a dark brown subbasal band extending a little less than half-way, remainder very pale orange; hind coxa white; hind femur white in proximal one-third, otherwise dark brown, tibia narrowly white proximally, a dark brown subbasal band extending a little more than half-way, remainder white; pretarsus slightly dusky; fore wing completely hyaline (
<figureCitation id="13282A34FF33FF41FBCBB950A290FA49" box="[1065,1152,1396,1420]" captionStart="Figs 569-582" captionStartId="777.[181,226,1526,1550]" captionTargetBox="[185,1163,231,1508]" captionTargetId="figure-10@777.[181,1163,223,1508]" captionTargetPageId="777" captionText="Figs 569-582. Cheiloneurus roellus, HT (569) ovipositor; (570) apex of fore wing venation; (571) fore wing. Cheiloneurus dalycera (572) mesosoma, dorsal view; (573) antenna; (574) apex of fore wing venation; (575) fore wing. Cheiloneurus ection, HT (576) antenna (clava slightly collapsed); (577) ovipositor; (578) fore wing; (579) apex of fore wing venation; (580) mesosoma, dorsal view. Cheiloneurus parvus (581) head, facial view (reconstructed); (582) mesosoma, dorsal view." pageId="230" pageNumber="229">Fig. 571</figureCitation>
), venation mostly pale yellow-brown, submarginal vein partially hyaline; propodeum dark purple-brown; sides metallic green with conspicuous silvery setae in spiracular area; gaster dark purple-brown, Gt1 largely dark metallic blue with some violet reflections; remainder of gaster mixed purple and brassy; gonostylus dark brown.
</paragraph>
<paragraph id="8BAC36B1FF33FF41FF39B9DDA29DF953" blockId="230.[181,1165,942,1686]" pageId="230" pageNumber="229">Head about 3.7X as wide as frontovertex, not subtriangular in profile, occipital margin sharp, occiput without a shallow, median groove; ocelli forming an angle of about 65°; frontovertex slightly shiny, with conspicuous reticulate sculpture of mesh size generally very slightly smaller than diameter of eye facet; scrobes and mouth margin with shallower sculpture; gena with elongate sculpture; temple almost smooth, with shallow irregular, reticulate sculpture, of larger mesh than frontovertex and distinctly elongate posteriorly; scrobes shallow, meeting dorsally, dorsal and</paragraph>
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<pageTitle id="CB8CEED6FF32FF40FF57BCA6A6F3FF5A" box="[181,227,130,159]" pageId="231" pageNumber="230">230</pageTitle>
</pageNumber>
</paragraph>
<paragraph id="8BAC36B1FF32FF40FEB5BCA4A5FBFF5D" blockId="231.[343,1003,128,152]" box="[343,1003,128,152]" pageId="231" pageNumber="230">
<pageTitle id="CB8CEED6FF32FF40FEB5BCA4A5FBFF5D" box="[343,1003,128,152]" pageId="231" pageNumber="230">
Taxonomic Monographs on Neotropical
<taxonomicName id="4C134D32FF32FF40FD11BCA4A5C7FF5D" authority=", Volume" authorityName="Volume" box="[755,983,128,152]" class="Insecta" kingdom="Animalia" order="Hymenoptera" pageId="231" pageNumber="230" phylum="Arthropoda" rank="order">Hymenoptera, Volume</taxonomicName>
2
</pageTitle>
</paragraph>
<paragraph id="8BAC36B1FF32FF40FF57BCFAA597FE71" blockId="231.[181,1165,222,954]" pageId="231" pageNumber="230">
lateral margins weakly rounded; antenna as in
<figureCitation id="13282A34FF32FF40FD7ABCFAA4E1FF33" box="[664,753,222,246]" captionStart="Figs 553-568" captionStartId="776.[181,226,1526,1550]" captionTargetBox="[185,1163,227,1508]" captionTargetId="figure-9@776.[181,1163,223,1508]" captionTargetPageId="776" captionText="Figs 553-568. Cheiloneurus danessa, HT (553) fore wing. Cheiloneurus idris (554) ovipositor; (555) apex of fore wing venation; (556) head, facial view; (557) fore wing; (558) antenna; (559) hypopygium; (560) mesosoma, dorsal view. Cheiloneurus socrates, HT (561) antenna; (562) fore wing; (563) apex of fore wing venation; (564) mesosoma, dorsal view; (565) ovipositor; (566) gastral tergites showing “glands” on Gt1 and Gt5. Cheiloneurus roellus, HT (567) antenna; (568) mesosoma, dorsal view." pageId="231" pageNumber="230">Fig. 567</figureCitation>
; scape slightly broadened and flattened, about 4X as long as broad; all funicle segments subquadrate, broader and larger distad; clava slender, about as long as F2-F6 combined, sutures not oblique, sensory area small, at apex only and giving it a rounded appearance; eye very nearly reaching occipital margin, separated by less than diameter of a facet, clothed with conspicuous setae that are each at least as long as diameter of facet; inner eye margins not converging anteriorly; malar sulcus absent; mandible with three sharp teeth, upper tooth shortest. Relative measurements: HW 48,
<collectingRegion id="49D7F853FF32FF40FC83BDA6A596FE5C" box="[865,902,386,409]" country="Swaziland" name="Hhohho" pageId="231" pageNumber="230">HH</collectingRegion>
49, FV 13, OD 3.5, POL 6.5, OOL 1, OCL 5, AOL 7.5, EL 33, EW 26.5, MS 21, SL 27.5, SW 7.
</paragraph>
<paragraph id="8BAC36B1FF32FF40FF39BD9CA5D3FD25" blockId="231.[181,1165,222,954]" pageId="231" pageNumber="230">
Thorax (
<figureCitation id="13282A34FF32FF40FEDABD9CA783FE15" box="[312,403,440,464]" captionStart="Figs 553-568" captionStartId="776.[181,226,1526,1550]" captionTargetBox="[185,1163,227,1508]" captionTargetId="figure-9@776.[181,1163,223,1508]" captionTargetPageId="776" captionText="Figs 553-568. Cheiloneurus danessa, HT (553) fore wing. Cheiloneurus idris (554) ovipositor; (555) apex of fore wing venation; (556) head, facial view; (557) fore wing; (558) antenna; (559) hypopygium; (560) mesosoma, dorsal view. Cheiloneurus socrates, HT (561) antenna; (562) fore wing; (563) apex of fore wing venation; (564) mesosoma, dorsal view; (565) ovipositor; (566) gastral tergites showing “glands” on Gt1 and Gt5. Cheiloneurus roellus, HT (567) antenna; (568) mesosoma, dorsal view." pageId="231" pageNumber="230">Fig. 568</figureCitation>
) with shallow, irregular, polygonally reticulate sculpture on pronotum, generally of mesh size subequal to diameter of facet; sculpture on mesoscutum generally of similar depth to that on pronotum, but more regular and of smaller mesh; sculpture on axilla similar to that on mesoscutum, but slightly deeper; scutellum with conspicuously deeper, polygonally reticulate sculpture; apex and sides narrowly completely smooth; visible part of mesoscutum about 2X as broad as long; scutellum very slightly broader than long, with a distinct subapical tuft, setae about half as long as scutellum; hind femur about 3.5X as long as broad; fore wing with venation and setation as in
<figureCitation id="13282A34FF32FF40FEA3BE52A7DFFD4B" box="[321,463,630,654]" captionStart="Figs 569-582" captionStartId="777.[181,226,1526,1550]" captionTargetBox="[185,1163,231,1508]" captionTargetId="figure-10@777.[181,1163,223,1508]" captionTargetPageId="777" captionText="Figs 569-582. Cheiloneurus roellus, HT (569) ovipositor; (570) apex of fore wing venation; (571) fore wing. Cheiloneurus dalycera (572) mesosoma, dorsal view; (573) antenna; (574) apex of fore wing venation; (575) fore wing. Cheiloneurus ection, HT (576) antenna (clava slightly collapsed); (577) ovipositor; (578) fore wing; (579) apex of fore wing venation; (580) mesosoma, dorsal view. Cheiloneurus parvus (581) head, facial view (reconstructed); (582) mesosoma, dorsal view." pageId="231" pageNumber="230">Figs 570, 571</figureCitation>
; costal cell with only 1 seta dorsally at apex; no naked streak present between apices of postmarginal and stigmal veins; propodeum (
<figureCitation id="13282A34FF32FF40FCACBEB6A5B7FD6F" box="[846,935,658,682]" captionStart="Figs 553-568" captionStartId="776.[181,226,1526,1550]" captionTargetBox="[185,1163,227,1508]" captionTargetId="figure-9@776.[181,1163,223,1508]" captionTargetPageId="776" captionText="Figs 553-568. Cheiloneurus danessa, HT (553) fore wing. Cheiloneurus idris (554) ovipositor; (555) apex of fore wing venation; (556) head, facial view; (557) fore wing; (558) antenna; (559) hypopygium; (560) mesosoma, dorsal view. Cheiloneurus socrates, HT (561) antenna; (562) fore wing; (563) apex of fore wing venation; (564) mesosoma, dorsal view; (565) ovipositor; (566) gastral tergites showing “glands” on Gt1 and Gt5. Cheiloneurus roellus, HT (567) antenna; (568) mesosoma, dorsal view." pageId="231" pageNumber="230">Fig. 568</figureCitation>
) with numerous setae in spiracular area, side naked, medially about 0.1X as long as scutellum and without distinct sculpture. Relative measurements: FWL 55, FWW 20.5; HWL 44, HWW 10.
</paragraph>
<paragraph id="8BAC36B1FF32FF40FF39BEC7A731FCF4" blockId="231.[181,1165,222,954]" pageId="231" pageNumber="230">Gaster with hypopygium reaching about half-way to apex; syntergum slightly longer than mid tibia and with apex acute; ovipositor slightly exserted, the exserted part about 0.4X as long as mid tibial spur.</paragraph>
<paragraph id="8BAC36B1FF32FF40FF39BF11A448FC41" blockId="231.[181,1165,222,954]" pageId="231" pageNumber="230">
<typeStatus id="54A88813FF32FF40FF39BF11A727FC88" box="[219,311,821,845]" pageId="231" pageNumber="230">Paratype</typeStatus>
. Funicle with linear sensilla present only on 5-F6; gaster without “glands” on Gt1 and Gt5; ovipositor as in Fig, 569, gonostylus about 2X as long as mid tibial spur. Relative measurements: OL 45.5, GL 17 [MT 27].
</paragraph>
<paragraph id="8BAC36B1FF32FF40FF39BFA3A70FFC7F" blockId="231.[181,1165,222,954]" pageId="231" pageNumber="230">
Variation. Females vary in overall length from
<quantity id="4CEB9B54FF32FF40FD3BBFA3A574FC5A" box="[729,868,903,927]" metricMagnitude="-3" metricUnit="m" metricValue="1.24" metricValueMax="1.27" metricValueMin="1.21" pageId="231" pageNumber="230" unit="mm" value="1.24" valueMax="1.27" valueMin="1.21">1.21-1.27mm</quantity>
, otherwise little in material examined.
</paragraph>
<paragraph id="8BAC36B1FF32FF40FF57BFFCA771FC35" blockId="231.[181,353,984,1008]" box="[181,353,984,1008]" pageId="231" pageNumber="230">Male. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C309653AFF32FF40FF57B82BA7F9FBE2" box="[181,489,1039,1063]" pageId="231" pageNumber="230" type="distribution">
<paragraph id="8BAC36B1FF32FF40FF57B82BA7F9FBE2" blockId="231.[181,489,1039,1063]" box="[181,489,1039,1063]" pageId="231" pageNumber="230">
DISTRIBUTION.
<collectingCountry id="F3047621FF32FF40FE91B82BA7F5FBE2" box="[371,485,1039,1063]" name="Costa Rica" pageId="231" pageNumber="230">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C309653AFF32FF40FF57B861A76EFB98" box="[181,382,1093,1117]" pageId="231" pageNumber="230" type="biology_ecology">
<paragraph id="8BAC36B1FF32FF40FF57B861A76EFB98" blockId="231.[181,382,1093,1117]" box="[181,382,1093,1117]" pageId="231" pageNumber="230">HOSTS. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C309653AFF32FF40FF57B859A4BFFB21" pageId="231" pageNumber="230" type="materials_examined">
<paragraph id="8BAC36B1FF32FF40FF57B859A7BDFB57" blockId="231.[181,1164,1149,1253]" box="[181,429,1149,1170]" pageId="231" pageNumber="230">MATERIAL EXAMINED.</paragraph>
<paragraph id="8BAC36B1FF32FF40FF39B8BDA4BFFB21" blockId="231.[181,1164,1149,1253]" pageId="231" pageNumber="230">
Type material.
<materialsCitation id="3B7B3CECFF32FF40FE80B8BCA737FB0C" collectingDate="1996-01-24" collectingDateMax="1996-02-27" collectingDateMin="1996-01-24" collectionCode="ACG, MT, YPT" collectorName="J. S. Noyes" country="Costa Rica" elevation="700" location="Pitilla" pageId="231" pageNumber="230" specimenCount="1" stateProvince="Guanacaste" typeStatus="holotype">
<typeStatus id="54A88813FF32FF40FE80B8BCA7A6FB6B" box="[354,438,1176,1198]" pageId="231" pageNumber="230">Holotype</typeStatus>
E:
<emphasis id="B967EAA3FF32FF40FE3AB8BCA472FB6B" bold="true" box="[472,610,1176,1198]" pageId="231" pageNumber="230">
<collectingCountry id="F3047621FF32FF40FE3AB8BCA472FB6B" box="[472,610,1176,1198]" name="Costa Rica" pageId="231" pageNumber="230">COSTA RICA</collectingCountry>
</emphasis>
,
<emphasis id="B967EAA3FF32FF40FD8EB8BCA4CDFB6B" bold="true" box="[620,733,1176,1198]" pageId="231" pageNumber="230">
<collectingRegion id="49D7F853FF32FF40FD8EB8BCA4CDFB6B" box="[620,733,1176,1198]" country="Costa Rica" name="Guanacaste" pageId="231" pageNumber="230">Guanacaste</collectingRegion>
</emphasis>
,
<location id="8ECC606AFF32FF40FD0AB8BCA50EFB6B" LSID="urn:lsid:plazi:treatment:03BA87A7FF33FF40FDE8BCFAA4BFFB21:8ECC606AFF32FF40FD0AB8BCA50EFB6B" box="[744,798,1176,1198]" country="Costa Rica" name="Pitilla" pageId="231" pageNumber="230" stateProvince="Guanacaste">Pitilla</location>
(
<collectionCode id="ED02AE74FF32FF40FCCBB8BCA54DFB6B" box="[809,861,1176,1198]" pageId="231" pageNumber="230">ACG</collectionCode>
),
<quantity id="4CEB9B54FF32FF40FC8EB8BCA5B1FB68" box="[876,929,1176,1197]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="231" pageNumber="230" unit="m" value="700.0">
<elevation id="003ED182FF32FF40FC8EB8BCA5B1FB68" box="[876,929,1176,1197]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="231" pageNumber="230" unit="m" value="700.0">700m</elevation>
</quantity>
,
<collectionCode id="ED02AE74FF32FF40FC48B8BCA5DCFB68" box="[938,972,1176,1197]" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="231" pageNumber="230">MT</collectionCode>
/
<collectionCode id="ED02AE74FF32FF40FC31B8BCA5EEFB68" box="[979,1022,1176,1197]" pageId="231" pageNumber="230">YPT</collectionCode>
,
<date id="FFAD1071FF32FF40FBE4B8BCA29CFB6B" box="[1030,1164,1176,1198]" pageId="231" pageNumber="230" value="1996-01-24" valueMax="1996-02-27" valueMin="1996-01-24">
<collectingDate id="EFE9E999FF32FF40FBE4B8BCA29CFB6B" box="[1030,1164,1176,1198]" pageId="231" pageNumber="230" value="1996-01-24" valueMax="1996-02-27" valueMin="1996-01-24">24.i-27.ii.1996</collectingDate>
</date>
(
<collectorName id="26E65367FF32FF40FF5FB890A731FB0C" box="[189,289,1204,1225]" pageId="231" pageNumber="230">J.S. Noyes</collectorName>
)
</materialsCitation>
.
<materialsCitation id="3B7B3CECFF32FF40FECDB890A701FB20" collectingDate="1989-04" collectorName="I. Gauld" country="Costa Rica" elevation="700" location="Santa Cecilia" pageId="231" pageNumber="230" specimenCount="1" stateProvince="Guanacaste" typeStatus="paratype">
<typeStatus id="54A88813FF32FF40FECDB890A79BFB0C" box="[303,395,1204,1225]" pageId="231" pageNumber="230">Paratypes</typeStatus>
:
<emphasis id="B967EAA3FF32FF40FE71B897A40DFB0C" bold="true" box="[403,541,1203,1225]" pageId="231" pageNumber="230">
<collectingCountry id="F3047621FF32FF40FE71B897A40DFB0C" box="[403,541,1203,1225]" name="Costa Rica" pageId="231" pageNumber="230">COSTA RICA</collectingCountry>
</emphasis>
, 1E,
<emphasis id="B967EAA3FF32FF40FDB2B897A4D1FB0C" bold="true" box="[592,705,1203,1225]" pageId="231" pageNumber="230">
<collectingRegion id="49D7F853FF32FF40FDB2B897A4D1FB0C" box="[592,705,1203,1225]" country="Costa Rica" name="Guanacaste" pageId="231" pageNumber="230">Guanacaste</collectingRegion>
</emphasis>
, Estac. Pitilla,
<quantity id="4CEB9B54FF32FF40FCABB897A561FB0C" box="[841,881,1203,1225]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="231" pageNumber="230" unit="km" value="9.0">9km</quantity>
S
<location id="8ECC606AFF32FF40FC6AB890A212FB0C" LSID="urn:lsid:plazi:treatment:03BA87A7FF33FF40FDE8BCFAA4BFFB21:8ECC606AFF32FF40FC6AB890A212FB0C" box="[904,1026,1203,1225]" country="Costa Rica" name="Santa Cecilia" pageId="231" pageNumber="230" stateProvince="Guanacaste">Santa Cecilia</location>
,
<quantity id="4CEB9B54FF32FF40FBE8B890A250FB0C" box="[1034,1088,1204,1225]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="231" pageNumber="230" unit="m" value="700.0">
<elevation id="003ED182FF32FF40FBE8B890A250FB0C" box="[1034,1088,1204,1225]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="231" pageNumber="230" unit="m" value="700.0">700m</elevation>
</quantity>
,
<date id="FFAD1071FF32FF40FBABB897A29CFB0C" box="[1097,1164,1203,1225]" pageId="231" pageNumber="230" value="1989-04">
<collectingDate id="EFE9E999FF32FF40FBABB897A29CFB0C" box="[1097,1164,1203,1225]" pageId="231" pageNumber="230" value="1989-04">iv.1989</collectingDate>
</date>
(
<collectorName id="26E65367FF32FF40FF5FB8EBA71BFB20" box="[189,267,1231,1253]" pageId="231" pageNumber="230">I. Gauld</collectorName>
)
</materialsCitation>
.
<materialsCitation id="3B7B3CECFF32FF40FEF8B8EBA4BBFB21" box="[282,683,1231,1253]" collectionCode="MZUCR, NHMUK" country="Costa Rica" pageId="231" pageNumber="230" specimenCount="2" stateProvince="Guanacaste" typeStatus="holotype">
<typeStatus id="54A88813FF32FF40FEF8B8EBA77EFB20" box="[282,366,1231,1253]" pageId="231" pageNumber="230">Holotype</typeStatus>
in
<collectionCode id="ED02AE74FF32FF40FE69B8EBA7CEFB21" box="[395,478,1231,1252]" pageId="231" pageNumber="230">MZUCR</collectionCode>
,
<typeStatus id="54A88813FF32FF40FE05B8EBA425FB21" box="[487,565,1231,1252]" pageId="231" pageNumber="230">paratype</typeStatus>
in
<collectionCode id="ED02AE74FF32FF40FDB0B8EBA4BBFB21" box="[594,683,1231,1252]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="231" pageNumber="230" type="Museum">NHMUK</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>