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<document id="C0AF566BA8B92630A647F3D07EFAB6A5" ID-DOI="10.11646/zootaxa.3949.3.3" ID-GBIF-Dataset="25ece91c-6566-4122-a406-236f147b58b1" ID-ISSN="1175-5326" ID-Zenodo-Dep="238900" ID-ZooBank="CBEF804D-5FBF-48D2-8CCD-036B70FE6ECD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461046281391" checkinUser="plazi" docAuthor="Arias, Andrés &amp; Paxton, Hannelore" docDate="2015" docId="03BA8797FFA1FFBAFF2BF92FFD09C4BC" docLanguage="en" docName="zt03949p369.pdf" docOrigin="Zootaxa 3949 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Aponuphis willsiei Cantone &amp; Bellan 1996" docType="treatment" docVersion="15" lastPageNumber="364" masterDocId="FF83FFEFFFB3FFA9FFBCFFE0FFA7C254" masterDocTitle="Onuphis and Aponuphis (Annelida: Onuphidae) from southwestern Europe, with the description of a new species" masterLastPageNumber="369" masterPageNumber="345" pageNumber="363" updateTime="1698577207772" updateUser="plazi">
<mods:mods id="AC3993740E61809A17C08FD65DF9871F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="018C001045BE7FB2620FFF3E23506C30">Onuphis and Aponuphis (Annelida: Onuphidae) from southwestern Europe, with the description of a new species</mods:title>
</mods:titleInfo>
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<mods:namePart id="53D87DB76171624A1629659CDA92FB9C">Arias, Andrés</mods:namePart>
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<mods:namePart id="8DDD776FCB727DBB055B35FD356C200F">Paxton, Hannelore</mods:namePart>
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<mods:title id="C9531811C9BCBD1265AA63E10BC4B5D9">Zootaxa</mods:title>
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<mods:date id="375C8FAC0747DEBD195B4E9AF9498995">2015</mods:date>
<mods:detail id="E3694EB73E5FBDEEC95B1882D243F8D0" type="volume">
<mods:number id="0B2E4699F2D11A5F23DFCAFD48FF9BF4">3949</mods:number>
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<mods:number id="C2FE136306C12C4FEE95E2648B31C870">3</mods:number>
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<mods:classification id="FCBECCD56CE1295C0CC8CE26D1EE91DC">journal article</mods:classification>
<mods:identifier id="626B36B01E0B1D967259B8F344ADDFA6" type="DOI">10.11646/zootaxa.3949.3.3</mods:identifier>
<mods:identifier id="1284A6CB20D33DF00F210E6E07C04CA5" type="GBIF-Dataset">25ece91c-6566-4122-a406-236f147b58b1</mods:identifier>
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<treatment id="03BA8797FFA1FFBAFF2BF92FFD09C4BC" ID-DOI="http://doi.org/10.5281/zenodo.5217760" ID-GBIF-Taxon="119610581" ID-Zenodo-Dep="5217760" LSID="urn:lsid:plazi:treatment:03BA8797FFA1FFBAFF2BF92FFD09C4BC" httpUri="http://treatment.plazi.org/id/03BA8797FFA1FFBAFF2BF92FFD09C4BC" lastPageId="19" lastPageNumber="364" pageId="18" pageNumber="363">
<subSubSection id="C309650AFFA1FFBBFF2BF92FFEA4C55F" pageId="18" pageNumber="363" type="nomenclature">
<paragraph id="8BAC3681FFA1FFBBFF2BF92FFD0EC4BD" blockId="18.[151,681,1743,1803]" box="[151,681,1743,1769]" pageId="18" pageNumber="363">
<heading id="D0E481EDFFA1FFBBFF2BF92FFD0EC4BD" bold="true" box="[151,681,1743,1769]" fontSize="11" level="1" pageId="18" pageNumber="363" reason="1">
<taxonomicName id="4C134D02FFA1FFBBFF2BF92FFD0EC4BD" ID-CoL="67P42" authority="Cantone &amp; Bellan, 1996" authorityName="Cantone &amp; Bellan" authorityYear="1996" box="[151,681,1743,1769]" class="Polychaeta" family="Onuphidae" genus="Aponuphis" kingdom="Animalia" order="Eunicida" pageId="18" pageNumber="363" phylum="Annelida" rank="species" species="willsiei">
<emphasis id="B967EA93FFA1FFBBFF2BF92FFD0EC4BD" bold="true" box="[151,681,1743,1769]" pageId="18" pageNumber="363">
<emphasis id="B967EA93FFA1FFBBFF2BF92FFED0C4BD" bold="true" box="[151,375,1743,1769]" italics="true" pageId="18" pageNumber="363">Aponuphis willsiei</emphasis>
<bibRefCitation id="EF824B70FFA1FFBBFEC2F92FFD0EC4BD" author="Cantone" box="[382,681,1743,1769]" pageId="18" pageNumber="363" refString="Cantone, G. &amp; Bellan, G. (1996) Aponuphis willsiei, una nuova specie di Onuphidae (Annelida, Polychaeta) delle coste marsigliesi. Animalia, 21 (1 / 3), 27 - 30." type="journal article" year="1996">Cantone &amp; Bellan, 1996</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BAC3681FFA1FFBBFF2BF912FEA4C55F" blockId="18.[151,681,1743,1803]" box="[151,259,1778,1803]" pageId="18" pageNumber="363">
<figureCitation id="13282A04FFA1FFBBFF2BF912FEA4C55F" box="[151,259,1778,1803]" captionStart="FIGURE 10" captionStartId="19.[151,250,1106,1128]" captionTargetBox="[151,1436,517,1084]" captionTargetId="figure@19.[151,1436,517,1085]" captionTargetPageId="19" captionText="FIGURE 10. Aponuphis willsiei. Scanning electron micrographs. A, bidentate compound hooks of chaetiger 3; detailed view of bidentate pseudocompound hook of chaetiger 2; C, pectinate chaeta from chaetiger 15." httpUri="https://zenodo.org/record/238909/files/figure.png" pageId="18" pageNumber="363">Figure 10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C309650AFFA1FFBBFF2BF8DAFDDAC504" box="[151,637,1850,1872]" pageId="18" pageNumber="363" type="reference_group">
<paragraph id="8BAC3681FFA1FFBBFF2BF8DAFDDAC504" blockId="18.[151,637,1850,1872]" box="[151,637,1850,1872]" pageId="18" pageNumber="363">
<treatmentCitationGroup id="AB0311AFFFA1FFBBFF2BF8DAFDDAC504" box="[151,637,1850,1872]" pageId="18" pageNumber="363">
<taxonomicName id="4C134D02FFA1FFBBFF2BF8DAFDDEC504" ID-CoL="67P42" authority="Cantone &amp; Bellan, 1996: 27" authorityName="Cantone &amp; Bellan" authorityPageNumber="27" authorityYear="1996" box="[151,633,1850,1872]" class="Polychaeta" family="Onuphidae" genus="Aponuphis" kingdom="Animalia" order="Eunicida" pageId="18" pageNumber="363" phylum="Annelida" rank="species" species="willsiei">
<emphasis id="B967EA93FFA1FFBBFF2BF8DAFEF5C504" box="[151,338,1850,1872]" italics="true" pageId="18" pageNumber="363">Aponuphis willsiei</emphasis>
<treatmentCitation id="0AB21090FFA1FFBBFEE5F8DAFDDEC504" author="Cantone" box="[345,633,1850,1872]" page="27" pageId="18" pageNumber="363" year="1996">
<bibRefCitation id="EF824B70FFA1FFBBFEE5F8DAFDF4C504" author="Cantone" box="[345,595,1850,1872]" pageId="18" pageNumber="363" refString="Cantone, G. &amp; Bellan, G. (1996) Aponuphis willsiei, una nuova specie di Onuphidae (Annelida, Polychaeta) delle coste marsigliesi. Animalia, 21 (1 / 3), 27 - 30." type="journal article" year="1996">Cantone &amp; Bellan, 1996</bibRefCitation>
: 27
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C309650AFFA1FFBAFF2BF89CFCE5C349" lastPageId="19" lastPageNumber="364" pageId="18" pageNumber="363" type="materials_examined">
<paragraph id="8BAC3681FFA1FFBBFF2BF89CFA3AC58A" blockId="18.[151,1437,1916,2014]" pageId="18" pageNumber="363">
<emphasis id="B967EA93FFA1FFBBFF2BF89CFE26C5C1" bold="true" box="[151,385,1916,1941]" pageId="18" pageNumber="363">Material examined.</emphasis>
Non-type material. One specimen (
<collectionCode id="ED02AE44FFA1FFBBFCABF89DFCCAC5C1" LSID="urn:lsid:biocol.org:col:35275" box="[791,877,1917,1941]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="18" pageNumber="363">MNCN</collectionCode>
16.01/15244) Fauna IV off shore Alborán Island Lighthouse (35º 55.68’–
<geoCoordinate id="EE275046FFA1FFBBFE0EF840FDE0C5EC" box="[434,583,1952,1977]" direction="north" orientation="latitude" pageId="18" pageNumber="363" precision="9" value="35.927166">35º 55.63’ N</geoCoordinate>
, 0 3 º03.25’– 03º
<geoCoordinate id="EE275046FFA1FFBBFCB0F840FCD5C5ED" box="[780,882,1952,1977]" direction="west" orientation="longitude" pageId="18" pageNumber="363" precision="9" value="-0.0445">0 2.67 W</geoCoordinate>
), E.
<collectingCountry id="F3047611FFA1FFBBFC1BF840FC4DC5ED" box="[935,1002,1952,1977]" name="Spain" pageId="18" pageNumber="363">Spain</collectingCountry>
, Alborán Sea, western Mediterranean,
<quantity id="4CEB9B64FFA1FFBBFF2BF825FF5AC589" box="[151,253,1989,2013]" metricMagnitude="1" metricUnit="m" metricValue="4.1" metricValueMax="4.9" metricValueMin="3.3" pageId="18" pageNumber="363" unit="m" value="41.0" valueMax="49.0" valueMin="33.0">33–49 m</quantity>
,
<date id="FFAD1041FFA1FFBBFEB6F825FE35C58A" box="[266,402,1989,2014]" pageId="18" pageNumber="363" value="1996-07-23">23 Jul 1996</date>
;
<specimenCount id="9D15FD08FFA1FFBBFE23F825FD8DC58A" box="[415,554,1989,2014]" pageId="18" pageNumber="363" type="generic">4 specimens</specimenCount>
(AM W.47515) Mazarrón Port, Murcia,
<collectingCountry id="F3047611FFA1FFBBFC46F825FB9AC58A" box="[1018,1085,1989,2014]" name="Spain" pageId="18" pageNumber="363">Spain</collectingCountry>
, dredged from
<quantity id="4CEB9B64FFA1FFBBFB56F825FAF9C589" box="[1258,1374,1989,2014]" metricMagnitude="1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="18" pageNumber="363" unit="m" value="75.0" valueMax="100.0" valueMin="50.0">50–100 m</quantity>
, coll.
</paragraph>
<paragraph id="8BAC3681FFA0FFBAFF2BFF78FE1BC2AC" blockId="19.[151,1437,151,464]" pageId="19" pageNumber="364">
<date id="FFAD1041FFA0FFBAFF2BFF78FE8BC2E4" box="[151,300,152,176]" pageId="19" pageNumber="364" value="2012-03-20">20 Mar 2012</date>
;
<specimenCount id="9D15FD08FFA0FFBAFE8BFF77FE65C2E4" box="[311,450,151,176]" pageId="19" pageNumber="364" type="generic">5 specimens</specimenCount>
(
<collectionCode id="ED02AE44FFA0FFBAFE6CFF78FD81C2E4" LSID="urn:lsid:biocol.org:col:35275" box="[464,550,152,176]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="19" pageNumber="364">MNCN</collectionCode>
16.01/16173) Mazarrón Port, Murcia,
<collectingCountry id="F3047611FFA0FFBAFC67FF77FBB9C2E4" box="[987,1054,151,176]" name="Spain" pageId="19" pageNumber="364">Spain</collectingCountry>
, dredged from
<quantity id="4CEB9B64FFA0FFBAFB74FF77FA9BC2FB" box="[1224,1340,151,176]" metricMagnitude="1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="19" pageNumber="364" unit="m" value="75.0" valueMax="100.0" valueMin="50.0">50–100 m</quantity>
, coll.
<date id="FFAD1041FFA0FFBAFA3CFF78FEAFC280" pageId="19" pageNumber="364" value="2012-03-20">20 Mar 2012</date>
;
<specimenCount id="9D15FD08FFA0FFBAFEA9FF5CFE07C281" box="[277,416,188,213]" pageId="19" pageNumber="364" type="generic">4 specimens</specimenCount>
coated with gold (
<collectionCode id="ED02AE44FFA0FFBAFDC8FF5CFD6DC280" LSID="urn:lsid:biocol.org:col:35275" box="[628,714,188,212]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="19" pageNumber="364">MNCN</collectionCode>
16.01/16174) Mazarrón Port, Murcia,
<collectingCountry id="F3047611FFA0FFBAFB3FFF5CFB61C281" box="[1155,1222,188,213]" name="Spain" pageId="19" pageNumber="364">Spain</collectingCountry>
, dredged from
<quantity id="4CEB9B64FFA0FFBAFACEFF5CFF47C2A3" metricMagnitude="1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="19" pageNumber="364" unit="m" value="75.0" valueMax="100.0" valueMin="50.0">50– 100 m</quantity>
, coll.
<date id="FFAD1041FFA0FFBAFE98FF00FE1FC2AC" box="[292,440,224,248]" pageId="19" pageNumber="364" value="2012-03-20">20 Mar 2012</date>
.
</paragraph>
<paragraph id="8BAC3681FFA0FFBAFF7BFEE4FCE5C349" blockId="19.[151,1437,151,464]" box="[199,834,260,285]" pageId="19" pageNumber="364">
<emphasis id="B967EA93FFA0FFBAFF7BFEE4FEC2C349" bold="true" box="[199,357,260,285]" pageId="19" pageNumber="364">
<typeStatus id="54A88823FFA0FFBAFF7BFEE4FEA5C349" box="[199,258,260,285]" pageId="19" pageNumber="364">Type</typeStatus>
locality.
</emphasis>
Gulf of Marseille,
<collectingCountry id="F3047611FFA0FFBAFD80FEE5FD2CC348" box="[572,651,261,284]" name="France" pageId="19" pageNumber="364">France</collectingCountry>
, Mediterranean.
</paragraph>
</subSubSection>
<subSubSection id="C309650AFFA0FFBAFF7BFEC7FECEC39B" pageId="19" pageNumber="364" type="diagnosis">
<paragraph id="8BAC3681FFA0FFBAFF7BFEC7FECEC39B" blockId="19.[151,1437,151,464]" pageId="19" pageNumber="364">
<emphasis id="B967EA93FFA0FFBAFF7BFEC7FEE6C314" bold="true" box="[199,321,295,320]" pageId="19" pageNumber="364">Diagnosis.</emphasis>
Antennae up to chaetiger 10; three to six ceratophoral rings on antennae. Anterior four chaetigers with four to five bidentate pseudocompound to compound hooks (
<figureCitation id="13282A04FFA0FFBAFCC1FEACFC77C330" box="[893,976,332,357]" captionStart="FIGURE 10" captionStartId="19.[151,250,1106,1128]" captionTargetBox="[151,1436,517,1084]" captionTargetId="figure@19.[151,1436,517,1085]" captionTargetPageId="19" captionText="FIGURE 10. Aponuphis willsiei. Scanning electron micrographs. A, bidentate compound hooks of chaetiger 3; detailed view of bidentate pseudocompound hook of chaetiger 2; C, pectinate chaeta from chaetiger 15." httpUri="https://zenodo.org/record/238909/files/figure.png" pageId="19" pageNumber="364">Fig. 10</figureCitation>
), chaetiger 5 with two to three hooks; all hooks of almost equal thickness, slender long-appendaged hooks absent. Ventral cirri subulate on first four chaetigers. Subacicular hooks from chaetiger 9–10. Branchiae absent. No colour pattern. Maximum width without parapodia 1.0 mm.
</paragraph>
</subSubSection>
<caption id="DF6C6609FFA0FFBAFF2BFBB2FB87C6D3" httpUri="https://zenodo.org/record/238909/files/figure.png" pageId="19" pageNumber="364" targetBox="[151,1436,517,1084]" targetPageId="19">
<paragraph id="8BAC3681FFA0FFBAFF2BFBB2FB87C6D3" blockId="19.[151,1435,1106,1159]" pageId="19" pageNumber="364">
<emphasis id="B967EA93FFA0FFBAFF2BFBB2FE87C633" bold="true" box="[151,288,1106,1128]" pageId="19" pageNumber="364">FIGURE 10.</emphasis>
<taxonomicName id="4C134D02FFA0FFBAFE94FBB2FE43C63C" box="[296,484,1106,1128]" class="Polychaeta" family="Onuphidae" genus="Aponuphis" kingdom="Animalia" order="Eunicida" pageId="19" pageNumber="364" phylum="Annelida" rank="species" species="willsiei">
<emphasis id="B967EA93FFA0FFBAFE94FBB2FE43C63C" box="[296,484,1106,1128]" italics="true" pageId="19" pageNumber="364">Aponuphis willsiei</emphasis>
</taxonomicName>
. Scanning electron micrographs. A, bidentate compound hooks of chaetiger 3; detailed view of bidentate pseudocompound hook of chaetiger 2; C, pectinate chaeta from chaetiger 15.
</paragraph>
</caption>
<subSubSection id="C309650AFFA0FFBAFF7BFB53FEC9C7F0" pageId="19" pageNumber="364" type="discussion">
<paragraph id="8BAC3681FFA0FFBAFF7BFB53FEC9C7F0" blockId="19.[151,1437,1203,1768]" pageId="19" pageNumber="364">
<emphasis id="B967EA93FFA0FFBAFF7BFB53FE9CC698" bold="true" box="[199,315,1203,1228]" pageId="19" pageNumber="364">Remarks.</emphasis>
This is the first confirmed record of this species from Iberian Peninsula waters.
<bibRefCitation id="EF824B70FFA0FFBAFB6DFB54FA3BC698" author="Amoureux" box="[1233,1436,1204,1228]" pageId="19" pageNumber="364" refString="Amoureux, L. (1972) Annelides polychetes recueillies sur les pentes du talus continental, au large de la Galice (Espagne). Campagnes 1967 et 1968 de la &quot; Thalassa &quot;. Cahiers de Biologie Marine, 13, 63 - 89." type="journal article" year="1972">Amoureux (1972)</bibRefCitation>
reported one incomplete
<quantity id="4CEB9B64FFA0FFBAFE0CFB38FE4FC6BB" box="[432,488,1240,1264]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="364" unit="cm" value="1.0">1 cm</quantity>
long specimen without branchiae among the
<taxonomicName id="4C134D02FFA0FFBAFC57FB38FBC8C6BB" box="[1003,1135,1239,1263]" class="Polychaeta" family="Onuphidae" genus="Hyalinoecia" kingdom="Animalia" order="Eunicida" pageId="19" pageNumber="364" phylum="Annelida" rank="species" species="bilineata">
<emphasis id="B967EA93FFA0FFBAFC57FB38FBC8C6BB" box="[1003,1135,1239,1263]" italics="true" pageId="19" pageNumber="364">A. bilineata</emphasis>
</taxonomicName>
(as
<taxonomicName id="4C134D02FFA0FFBAFB22FB37FA35C6BB" box="[1182,1426,1239,1263]" class="Polychaeta" family="Onuphidae" genus="Hyalinoecia" kingdom="Animalia" order="Eunicida" pageId="19" pageNumber="364" phylum="Annelida" rank="species" species="bilineata">
<emphasis id="B967EA93FFA0FFBAFB22FB37FA35C6BB" box="[1182,1426,1239,1263]" italics="true" pageId="19" pageNumber="364">Hyalinoecia bilineata</emphasis>
</taxonomicName>
) specimens collected on the continental slope of Galicia (northwestern
<collectingCountry id="F3047611FFA0FFBAFC60FB1BFBB9C740" box="[988,1054,1275,1300]" name="Spain" pageId="19" pageNumber="364">Spain</collectingCountry>
) and he considered it to be the abranchiate Mediterranean form reported by Bellan. However, we did not find any specimens belonging to this species among the samples we examined from the area. Since the species has not been reported again from the Galician coasts, we consider this record as questionable, since it may represent merely a juvenile phase of another
<taxonomicName id="4C134D02FFA0FFBAFF2BFA6CFEA8C7F0" box="[151,271,1420,1444]" class="Polychaeta" family="Onuphidae" genus="Aponuphis" kingdom="Animalia" order="Eunicida" pageId="19" pageNumber="364" phylum="Annelida" rank="genus">
<emphasis id="B967EA93FFA0FFBAFF2BFA6CFEA8C7F0" box="[151,271,1420,1444]" italics="true" pageId="19" pageNumber="364">Aponuphis</emphasis>
</taxonomicName>
species.
</paragraph>
</subSubSection>
<subSubSection id="C309650AFFA0FFBAFF7BFA4FFD43C4F4" pageId="19" pageNumber="364" type="biology_ecology">
<paragraph id="8BAC3681FFA0FFBAFF7BFA4FFD43C4F4" blockId="19.[151,1437,1203,1768]" pageId="19" pageNumber="364">
<emphasis id="B967EA93FFA0FFBAFF7BFA4FFE80C79C" bold="true" box="[199,295,1455,1480]" pageId="19" pageNumber="364">Biology.</emphasis>
One specimen preserved in its tube was found with two juveniles closely appressed to its body. The juveniles were 2.3 and
<quantity id="4CEB9B64FFA0FFBAFE23FA33FE50C7BF" box="[415,503,1491,1516]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="19" pageNumber="364" unit="mm" value="2.5">2.5 mm</quantity>
long for 17 and 18 chaetigers and measured
<quantity id="4CEB9B64FFA0FFBAFC45FA34FBF6C7BF" box="[1017,1105,1492,1516]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="19" pageNumber="364" unit="mm" value="0.3">0.3 mm</quantity>
in width. The juveniles have well developed frontal lips; palps and antennae with two ceratophoral rings; parapodia are well developed, anterior ones with dorsal cirri; subulate ventral cirri on chaetigers 1 and 2. The chaetal distribution is very similar to that described for
<taxonomicName id="4C134D02FFA0FFBAFE87F9A0FE0DC403" box="[315,426,1600,1623]" class="Polychaeta" family="Onuphidae" genus="Aponuphis" kingdom="Animalia" order="Eunicida" pageId="19" pageNumber="364" phylum="Annelida" rank="species" species="ornata">
<emphasis id="B967EA93FFA0FFBAFE87F9A0FE0DC403" box="[315,426,1600,1623]" italics="true" pageId="19" pageNumber="364">A. ornata</emphasis>
</taxonomicName>
above with chaetigers 1 and 2 with bidentate compound hooks, limbate and anterior provisional SAHs from chaetiger 3–8, permanent SAHs from chaetiger 9–14, posterior provisional hooks in the last few chaetigers, and pectinate chaetae are absent.
</paragraph>
</subSubSection>
<subSubSection id="C309650AFFA0FFBAFF7BF94BFD09C4BC" pageId="19" pageNumber="364" type="distribution">
<paragraph id="8BAC3681FFA0FFBAFF7BF94BFD09C4BC" blockId="19.[151,1437,1203,1768]" pageId="19" pageNumber="364">
<emphasis id="B967EA93FFA0FFBAFF7BF94BFEC7C490" bold="true" box="[199,352,1707,1732]" pageId="19" pageNumber="364">Distribution.</emphasis>
Mediterranean, this species is only known from the Gulf of Marseille, the southwestern Iberian Peninsula (Murcia,
<collectingCountry id="F3047611FFA0FFBAFECFF92FFE13C4BC" box="[371,436,1743,1768]" name="Spain" pageId="19" pageNumber="364">Spain</collectingCountry>
) and the Alborán Sea.
</paragraph>
</subSubSection>
</treatment>
</document>
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