304 lines
28 KiB
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304 lines
28 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/contrib.entomol.74.e114543" ID-Pensoft-Pub="2511-6428-1-13" ID-Pensoft-UUID="F4C1C6A2DFD95D3BAF5AF1CE09226837" ID-ZooBank="589077DFF8BB45ABB30D7E0DE1C307FE" ModsDocID="2511-6428-74-1-13" checkinTime="1706792588976" checkinUser="pensoft" docAuthor="Irmler, Ulrich" docDate="2024" docId="5A76D65FBDC95AA9B12B33A3F1686DBD" docLanguage="en" docName="ContribEntomol 74(1): 13-34" docOrigin="Contributions to Entomology 74 (1)" docPubDate="2024-01-31" docSource="http://dx.doi.org/10.3897/contrib.entomol.74.e114543" docTitle="Somoleptus umicola Irmler 2024, sp. nov." docType="treatment" docUuid="CF53A865-0698-413F-B9F5-F47D51DFFCAC" docUuidSource="ZooBank" docVersion="1" id="F4C1C6A2DFD95D3BAF5AF1CE09226837" lastPageNumber="13" masterDocId="F4C1C6A2DFD95D3BAF5AF1CE09226837" masterDocTitle="Additions to Neotropical species of the genera Lithocharodes Sharp, 1881 and Somoleptus Sharp, 1881 (Coleoptera, Staphylinidae)" masterLastPageNumber="34" masterPageNumber="13" pageNumber="13" updateTime="1706792588976" updateUser="pensoft">
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<mods:mods id="3713E0C39AA0BDBDD87A954495E92C10" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="8C365242E87F9BB6C56BEBA724FBF9C9">
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<mods:title id="3B4F56E5370A516DC75166D684AF1185">Additions to Neotropical species of the genera Lithocharodes Sharp, 1881 and Somoleptus Sharp, 1881 (Coleoptera, Staphylinidae)</mods:title>
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<mods:name id="4A26F259A9834A094158BE488610D16E" type="personal">
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<mods:roleTerm id="6D98674E5792EBF3FBC6A453E4B9B1AA">Author</mods:roleTerm>
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<mods:namePart id="9A15DBCE7C7CB141D1D0059F29C30D36">Irmler, Ulrich</mods:namePart>
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<mods:affiliation id="FA5196DACD7D0783AFA84B5E862BED12">Institute for Ecosystem Research, University of Kiel, Olshausenstrasse 40, 24098 Kiel, Germany</mods:affiliation>
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<mods:nameIdentifier id="5806D806D0B0E4E80A24118616F6E1CC" type="email">uirmler@ecology.uni-kiel.de</mods:nameIdentifier>
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<mods:typeOfResource id="751EB802555EAE56832428F2467B3108">text</mods:typeOfResource>
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<mods:title id="75E0D5AB4B6D35CBC14A56AF72572AE9">Contributions to Entomology</mods:title>
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</mods:titleInfo>
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<mods:part id="ED2C0394EC108E584EA3C12C7EDC8C35">
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<mods:date id="6913C8B5243D812F656792176C397ADD">2024</mods:date>
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<mods:detail id="5B9EBD5F4B2B1A02BA361B5E635D38B5" type="pubDate">
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<mods:number id="7AA98D664E1D7D65E024A35858703CBC">2024-01-31</mods:number>
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<mods:detail id="1605A047100644B4CAF450F7BC148438" type="volume">
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<mods:number id="838506C2F52BF15EC0ED2CE665BA9E1E">74</mods:number>
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<mods:detail id="AD4798D7DC74F5AEFBA0A74730C69457" type="issue">
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<mods:number id="46D5E24A9A4CD62CF368B8BF7C91EFC8">1</mods:number>
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<mods:extent id="6DD85719E3325424558904E2FD7E1526" unit="page">
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<mods:start id="83894C9676546707F11D5B4E13AD2200">13</mods:start>
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<mods:end id="DC265F0542E707A37B5493F72A0A428F">34</mods:end>
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<mods:url id="69D25C6A517BAB1B34C6EB5FD418397F">http://dx.doi.org/10.3897/contrib.entomol.74.e114543</mods:url>
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</mods:location>
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<mods:classification id="7EE88A9CC92B2E17FC301DE3500C86CA">journal article</mods:classification>
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<mods:identifier id="1CA07F4AB8DA3F66C494D35B7CF82DE4" type="DOI">http://dx.doi.org/10.3897/contrib.entomol.74.e114543</mods:identifier>
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<mods:identifier id="74A607B85A19BC9C1609AD7586C576BC" type="Pensoft-Pub">2511-6428-1-13</mods:identifier>
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<mods:identifier id="2035E5A7861DAA7E9137FB0C8C046286" type="ZooBank">589077DFF8BB45ABB30D7E0DE1C307FE</mods:identifier>
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</mods:mods>
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<treatment id="5A76D65FBDC95AA9B12B33A3F1686DBD" LSID="urn:lsid:zoobank.org:act:CF53A865-0698-413F-B9F5-F47D51DFFCAC" httpUri="http://treatment.plazi.org/id/5A76D65FBDC95AA9B12B33A3F1686DBD" lastPageNumber="13" pageId="0" pageNumber="13">
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<subSubSection id="FD67955825B1B7AB51A53A19DD55D7D4" pageId="0" pageNumber="13" type="nomenclature">
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<paragraph id="F2428274D91BE81090F23F7F159DA116" pageId="0" pageNumber="13">
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<taxonomicName id="E87A8976E82BEE7B982D8357B21E4C25" LSID="https://zoobank.org/CF53A865-0698-413F-B9F5-F47D51DFFCAC" authority="Irmler, 2024" authorityName="Irmler" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Somoleptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Somoleptus umicola" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="umicola" status="sp. nov.">Somoleptus umicola</taxonomicName>
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<taxonomicNameLabel id="FB8000241AEB75FCC3B2DE5FA4CF12E7" pageId="0" pageNumber="13">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="4D54443EB6F37855A1A9692B2F9332AF" pageId="0" pageNumber="13" type="description">
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<paragraph id="C146471D9EF207D46038D889FCFB5A10" pageId="0" pageNumber="13">
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<figureCitation id="825BE53D31EC018BC116A9DFE7BBA1AE" captionStart="Figures 17, 18" captionStartId="F5" captionText="Figures 17, 18. 17. Somoleptus puntarenae; 18. S. umicola. a. Aedeagus; b. Paramere; c. Sternite VII; d. Tergite VII. Scale bars: 0.5 mm (a, c, d); 0.1 mm (b)." figureDoi="10.3897/contrib.entomol.74.e114543.figures17-18" httpUri="https://binary.pensoft.net/fig/975719" pageId="0" pageNumber="13">Figs 18a-d</figureCitation>
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<figureCitation id="A7C0734BD20E0E622652D3F8E5B51BF9" captionStart="Figure 21" captionStartId="F8" captionText="Figure 21. A. Fore-body of Somoleptus densiceps; B. S. guianensis; C. S. luteicornis; D. S. paramocola; E. S. puntarenae; F. S. umicola. Scale bars: 0.5 mm." figureDoi="10.3897/contrib.entomol.74.e114543.figure21" httpUri="https://binary.pensoft.net/fig/975722" pageId="0" pageNumber="13">, 21F</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="644A36FF2848A3071002B0D7FD1FA215" pageId="0" pageNumber="13" type="materials_examined">
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<paragraph id="E3D83FA0A600556C8A361AFAC5F5D168" pageId="0" pageNumber="13">Material examined.</paragraph>
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<paragraph id="64DDE01895DDE6E9FAEE0EE6565C31CE" pageId="0" pageNumber="13">
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<materialsCitation id="38564EB7DA028C4BD8AD41FFD5DEDA75" collectingDate="2004-06-07" collectorName="J. Ashe, Z. Falin, I. Hinojosa" country="COSTA RICA" elevation="1555" location="Puntarenas Prov." longitude="-83.01361" specimenCount="1" stateProvince="Puntarenas Prov." typeStatus="Holotype">
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<emphasis id="931751D9C9A90ABF054D2069CE1186A2" bold="true" italics="true" pageId="0" pageNumber="13">
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<typeStatus id="F71B111D6F1E3FBCD41BD6B631DB2DA1" type="holotype">Holotype</typeStatus>
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</emphasis>
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:
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<collectingCountry id="E1E0BD3DE928AA9C69D55B358625B596" name="Costa Rica">COSTA RICA</collectingCountry>
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male;
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<collectingRegion id="14F0599237E9463D0A34D9514ED7AFA1" country="Costa Rica" name="Puntarenas">Puntarenas Prov.</collectingRegion>
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; Altamira Biol. Stat.; 09°01'76"N,
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<geoCoordinate id="6EBAF2DE71BCB56A808CC19CCF122440" degrees="83" direction="west" minutes="00" orientation="longitude" precision="15" seconds="49" value="-83.01361">83°00'49"W</geoCoordinate>
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;
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<quantity id="D2C3C31BE136427D010FADCB6A11257D" metricMagnitude="3" metricUnit="m" metricValue="1.555" metricValueMax="1.6" metricValueMin="1.51" unit="m" value="1555.0" valueMax="1600.0" valueMin="1510.0">
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<elevation id="6B16C2D7F2E4D672E484CF1D63B21C4D" metricMagnitude="3" metricUnit="m" metricValue="1.555" metricValueMax="1.6" metricValueMin="1.51" unit="m" value="1555.0" valueMax="1600.0" valueMin="1510.0">1510-1600 m</elevation>
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</quantity>
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elevation; pyrethrum fogging moss-covered clay bank;
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<collectingDate id="33D915FB4A4EA0DC7AE5EA64A68470E1" value="2004-06-07">7 VI 2004</collectingDate>
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;
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<collectorName id="99707BFFF3A97707C744519940E35A8C">J. Ashe</collectorName>
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,
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<collectorName id="02161AC89100DB62C5D0D16A89058B35">Z. Falin</collectorName>
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,
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<collectorName id="12CC07A0A434307E402DF005DFBF11E9">I. Hinojosa</collectorName>
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leg.; KNHM CR1AFH04 151.
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</materialsCitation>
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</paragraph>
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<paragraph id="8CD13EE641F2396A91A1C9EB7ABF94C3" pageId="0" pageNumber="13">
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<materialsCitation id="95013A56AEEF12A6E31ECD1854B7610A" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" collectorName="J. Ashe & Z. Falin & I. Hinojosa" country="COSTA RICA" elevation="1555" location="COSTA RICA" longitude="-83.01361" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas" typeStatus="Paratype">
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<emphasis id="048DF53A5DFB8D230C15BAF68AF1B68B" bold="true" italics="true" pageId="0" pageNumber="13">
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<typeStatus id="4198EF72B2F8F7611795F96249F81B6D" type="paratype">Paratype</typeStatus>
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</emphasis>
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:
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<collectingCountry id="EA593B8866CE932D0DBEFDF97871E629" name="Costa Rica">COSTA RICA</collectingCountry>
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<specimenCount id="4B2BF25A26271E6EF0D50BA8FBDE7276" count="1" type="female">1 female</specimenCount>
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; same data as holotype KNHM
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="AD07C21ACE57650119C7327B8AD87B9D" pageId="0" pageNumber="13" type="diagnosis">
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<paragraph id="5440FFB1C074F4B420535061E8B33A47" pageId="0" pageNumber="13">Diagnosis.</paragraph>
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<paragraph id="5B2803B871D8F1B5FC05915AD4182277" pageId="0" pageNumber="13">
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The taxonomic position of the species is unclear. It has a large dorsal plate of the central lobe and a triangular prominence at posterior margin of sternite VII of male. These characters fit to the
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<taxonomicName id="579F6D60CF5E9D8AEAE79E21E7BCD37E" lsidName="S. longicollis" pageId="0" pageNumber="13" rank="species" species="longicollis">
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<emphasis id="D2EBD65462792FF084D7D21BF3B2C27A" italics="true" pageId="0" pageNumber="13">S. longicollis</emphasis>
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</taxonomicName>
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-subgroup. On the other hand, the endophallus is not sclerotised similar to the species of the
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<taxonomicName id="22BACE4D40CA933A5A96A5A06135636E" lsidName="S. admirabilis" pageId="0" pageNumber="13" rank="species" species="admirabilis">
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<emphasis id="22D59159BF0B2BE2C542F9BFB1D77D4F" italics="true" pageId="0" pageNumber="13">S. admirabilis</emphasis>
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</taxonomicName>
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-subgroup. The cones at the apex of the central lobe resemble those of several species of the
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<taxonomicName id="6430478518B10F246C77DB92776CCA99" lsidName="S. alticola" pageId="0" pageNumber="13" rank="species" species="alticola">
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<emphasis id="60E0339E89DFF719E11BBC2AA32229E5" italics="true" pageId="0" pageNumber="13">S. alticola</emphasis>
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</taxonomicName>
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-subgroup. According to the small size, shape of head and the proportion of the elytra,
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<taxonomicName id="CAC9AA6AEA234D22ABA6144B716E3607" authorityName="Irmler" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Somoleptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Somoleptus umicola" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="umicola">
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<emphasis id="E0FA1C5B1C0370243F4DEF8EFDBBA5A5" italics="true" pageId="0" pageNumber="13">Somoleptus umicola</emphasis>
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</taxonomicName>
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best resembles to
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<taxonomicName id="AD4AA6FB60F6603C27D9B94B841A61A4" authorityName="Sharp" authorityYear="1885" class="Insecta" family="Staphylinidae" genus="Somoleptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Somoleptus obsoletus" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="obsoletus">
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<emphasis id="F0742F7655C218EF9BBE84F917C7CE64" italics="true" pageId="0" pageNumber="13">Somoleptus obsoletus</emphasis>
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</taxonomicName>
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Sharp, 1885.
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<taxonomicName id="027130AA0E9D1410AB8461BD5A19E650" authorityName="Irmler" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Somoleptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Somoleptus umicola" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="umicola">
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<emphasis id="17BD211A2B371CEC6332B388EB21812E" italics="true" pageId="0" pageNumber="13">Somoleptus umicola</emphasis>
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</taxonomicName>
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is still smaller, such as
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<taxonomicName id="81F8583A07F7CB43AAF82FF27CCD54E1" lsidName="S. parvulus" pageId="0" pageNumber="13" rank="species" species="parvulus">
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<emphasis id="652B9652E1DCB43DA82948A95A07C60C" italics="true" pageId="0" pageNumber="13">S. parvulus</emphasis>
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</taxonomicName>
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Sharp, 1885. In contrast to
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<taxonomicName id="6E971A92D422DAE56C8773D51E439E33" lsidName="S. parvulus" pageId="0" pageNumber="13" rank="species" species="parvulus">
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<emphasis id="40B99AA37F1E63E030C49D51B4CEA8D2" italics="true" pageId="0" pageNumber="13">S. parvulus</emphasis>
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</taxonomicName>
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with light brown pronotum and sclerotised endophallus,
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<taxonomicName id="8907A4A12C4FE2DBE9A7126CC424482F" lsidName="S. umicola" pageId="0" pageNumber="13" rank="species" species="umicola">
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<emphasis id="48CAEAA7E7EF9AD51DA7B03300953EE7" italics="true" pageId="0" pageNumber="13">S. umicola</emphasis>
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</taxonomicName>
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is totally black and the endophallus is transparent.
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</paragraph>
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</subSubSection>
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<subSubSection id="B7EA93A531562C4BA9D8E5027EF35A10" pageId="0" pageNumber="13" type="description">
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<paragraph id="AC058C2F0C8FA34DE910324D53AAEFD7" pageId="0" pageNumber="13">Description.</paragraph>
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<paragraph id="D9A1227BDC6C5115C6C4C27ED744EC31" pageId="0" pageNumber="13">Length: 3.28 mm; Colouration: completely black; legs and antennae lighter brown.</paragraph>
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<paragraph id="CD44F9D30480BC5B5B35D10965F0B3DC" pageId="0" pageNumber="13">Head: 0.62 mm long, 0.49 mm wide; eyes slightly prominent; moderately large; PS:E ratio 3.1; sides nearly parallel; posterior angles sub-rectangular; posterior sides combined with posterior edge semi-circular; inter-antennal furrows distinct and long; reaching to mid-length of eyes; setiferous punctation deep and dense; on average, interstices between punctures shorter than diameter of punctures; on anterior head denser than on posterior head; surface without microsculpture; shiny; antennae with first antennomere elongate; half as long as head-length; second and third antennomere conical; longer than wide; combined half as long as first antennomere; following antennomeres wider than long; anteriad increasing in width. Pronotum: 0.71 mm long, 0.44 mm wide; widest at anterior third; conically narrowed to neck; posteriad, sides nearly parallel; posterior angles sub-rectangular; posterior margin slightly convex; setiferous punctation moderately deep and dense; on average, interstices as wide as diameter of punctures; small mid-line impunctate; adjacent to mid-line with irregular line of 22-24 punctures; surface without microsculpture; polished. Elytra: 0.69 mm long, 0.65 mm wide; humeral angles obtuse; sides nearly parallel; posterior margin convex; retreated to suture; setiferous punctation deep and dense; on average, interstices between punctures shorter than diameter of punctures; surface without microsculpture; polished. Abdomen with finer punctation than on elytra, but as dense; surface without microsculpture; polished; meso-tibia with three, meta-tibia with two ctenidia. Aedeagus egg-shaped; more acute at base and more oval at apex; dorsal plate nearly totally covering central lobe; endophallus transparent; apical orifice prominent; laterad to upper edge of orifice with two elongate slender cones; parameres bilobed; projecting apical cones; inner lobe broad at base; in middle, abruptly narrowed to acute apex; at outer edge, centrad with several setae; outer lobe slender; as long as inner lobe.</paragraph>
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<caption id="91507DBB95AACF0DD1E4AD1AFE90984F" doi="10.3897/contrib.entomol.74.e114543.figures17-18" httpUri="https://binary.pensoft.net/fig/975719" pageId="0" pageNumber="13" start="Figures 17, 18" startId="F5">
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<paragraph id="D391682CA20326C983A5E8A3F6B06838" pageId="0" pageNumber="13">
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<emphasis id="4663B610A7A12E2E24975A2106178B82" bold="true" pageId="0" pageNumber="13">Figures 17, 18.</emphasis>
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<emphasis id="1BE49ADCD4D9C6D1CAAD66C09F1D3D43" bold="true" pageId="0" pageNumber="13">17.</emphasis>
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<taxonomicName id="3FADDF563FEE5B01B7CAC5FD910246E5" authorityName="Irmler" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Somoleptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Somoleptus puntarenae" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="puntarenae">
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<emphasis id="E3E7C9AEE4E388593E843A6B3D6F003E" italics="true" pageId="0" pageNumber="13">Somoleptus puntarenae</emphasis>
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</taxonomicName>
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;
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<emphasis id="34814719ECAC4FB15DF5DA5A82659BB9" bold="true" pageId="0" pageNumber="13">18.</emphasis>
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<taxonomicName id="9FBEE9046A229854329FB856422EE231" lsidName="S. umicola" pageId="0" pageNumber="13" rank="species" species="umicola">
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<emphasis id="A1ECE87ADF03CA8F2018A51DA0956C1F" italics="true" pageId="0" pageNumber="13">S. umicola</emphasis>
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</taxonomicName>
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.
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<emphasis id="C6E6AEE7F6BBF905FFEC181B1C3399C2" bold="true" pageId="0" pageNumber="13">a.</emphasis>
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Aedeagus;
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<emphasis id="00986091BEFFC9427EE46ABA798E9ABE" bold="true" pageId="0" pageNumber="13">b.</emphasis>
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Paramere;
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<emphasis id="30D88DA9ED2702BC5FCBD14F7CBB625B" bold="true" pageId="0" pageNumber="13">c.</emphasis>
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Sternite VII;
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<emphasis id="19963F67A943FFE6F906A04F8ACBC6DA" bold="true" pageId="0" pageNumber="13">d.</emphasis>
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Tergite VII. Scale bars: 0.5 mm (
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<emphasis id="4A80A87AE0E5E0027356178FBD0F05DD" bold="true" pageId="0" pageNumber="13">a, c, d</emphasis>
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); 0.1 mm (
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<emphasis id="2F662EA899C34501CC271CE04E3AEB7F" bold="true" pageId="0" pageNumber="13">b</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="DB40F4FC58A19A06EBCB6FAC9ABBB574" doi="10.3897/contrib.entomol.74.e114543.figure19" httpUri="https://binary.pensoft.net/fig/975720" pageId="0" pageNumber="13" start="Figure 19" startId="F6">
|
|
<paragraph id="BFF911B940BE47F83B2AE0FB4D02F8F7" pageId="0" pageNumber="13">
|
|
<emphasis id="67AD220A4421DAE5AD74917AD8985AD4" bold="true" pageId="0" pageNumber="13">Figure 19.</emphasis>
|
|
<emphasis id="E6AFD4950700697CF7AE3251CA8ABF63" bold="true" pageId="0" pageNumber="13">A.</emphasis>
|
|
Fore-body of
|
|
<taxonomicName id="780F4C11602BA9C6762E3F4F1E461403" authorityName="Irmler" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Lithocharodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lithocharodes andersoni" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="andersoni">
|
|
<emphasis id="49234070C24B562E6194B5CFE2E5B4D1" italics="true" pageId="0" pageNumber="13">Lithocharodes andersoni</emphasis>
|
|
</taxonomicName>
|
|
;
|
|
<emphasis id="14559EE30E9D422D22B3FF88DF02344D" bold="true" pageId="0" pageNumber="13">B.</emphasis>
|
|
<taxonomicName id="80E67771903E61FDE56E19CA7543A9C2" lsidName="L. dubia" pageId="0" pageNumber="13" rank="species" species="dubia">
|
|
<emphasis id="93262106A226E9AF5A1FD6ECC7E3D39D" italics="true" pageId="0" pageNumber="13">L. dubia</emphasis>
|
|
</taxonomicName>
|
|
;
|
|
<emphasis id="94A5AE53230F1CF217E49D3B38C7D3D3" bold="true" pageId="0" pageNumber="13">C.</emphasis>
|
|
<taxonomicName id="C483AA011837EF48BFC36F6612100EDC" lsidName="L. esmeraldae" pageId="0" pageNumber="13" rank="species" species="esmeraldae">
|
|
<emphasis id="EE33B37AE1C9C2574C0DB3A55DECC47C" italics="true" pageId="0" pageNumber="13">L. esmeraldae</emphasis>
|
|
</taxonomicName>
|
|
;
|
|
<emphasis id="9A9AD098A25DD0B2BA2E56D8DB36C4AE" bold="true" pageId="0" pageNumber="13">D.</emphasis>
|
|
<taxonomicName id="63C3CA2436B118AC653756BAFE97762E" lsidName="L. falini" pageId="0" pageNumber="13" rank="species" species="falini">
|
|
<emphasis id="7B3F57233FDB798F40A0785736C5072A" italics="true" pageId="0" pageNumber="13">L. falini</emphasis>
|
|
</taxonomicName>
|
|
;
|
|
<emphasis id="788EADA412CF1A3BF89C6A1A04D1F4F1" bold="true" pageId="0" pageNumber="13">E.</emphasis>
|
|
<taxonomicName id="95791905F2806709048469A13AC3DC49" lsidName="L. hansoni" pageId="0" pageNumber="13" rank="species" species="hansoni">
|
|
<emphasis id="1894BD135BA7C690A5A530F6DED7772F" italics="true" pageId="0" pageNumber="13">L. hansoni</emphasis>
|
|
</taxonomicName>
|
|
;
|
|
<emphasis id="3C4822C9CC585CC81A4DFBBEC25AC510" bold="true" pageId="0" pageNumber="13">F.</emphasis>
|
|
<taxonomicName id="4D78CC55B178E0F3CB6E2D88340BD90D" lsidName="L. hibbsi" pageId="0" pageNumber="13" rank="species" species="hibbsi">
|
|
<emphasis id="9A08AB52CF5D2F7C01FC1E49FBD8E47D" italics="true" pageId="0" pageNumber="13">L. hibbsi</emphasis>
|
|
</taxonomicName>
|
|
. Scale bars: 0.5 mm.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="1EDD057D0F46C33C1F4378771E525711" doi="10.3897/contrib.entomol.74.e114543.figure20" httpUri="https://binary.pensoft.net/fig/975721" pageId="0" pageNumber="13" start="Figure 20" startId="F7">
|
|
<paragraph id="8AE6C4C737DBCDD7D69B1E7D759BA2B8" pageId="0" pageNumber="13">
|
|
<emphasis id="074CAF12C592E216A183DB44955D6091" bold="true" pageId="0" pageNumber="13">Figure 20.</emphasis>
|
|
<emphasis id="55165470B224356EC0F95CBFF8BDC615" bold="true" pageId="0" pageNumber="13">A.</emphasis>
|
|
Fore-body of
|
|
<taxonomicName id="743A1161F675C43290BE2BEF5A93A7A0" authorityName="Irmler" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Lithocharodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lithocharodes lituratus" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="lituratus">
|
|
<emphasis id="6403C50EFEC63FADA337A3D8B84648E9" italics="true" pageId="0" pageNumber="13">Lithocharodes lituratus</emphasis>
|
|
</taxonomicName>
|
|
;
|
|
<emphasis id="E19A7E20381CF160E929E88F13AFD9FC" bold="true" pageId="0" pageNumber="13">B.</emphasis>
|
|
<taxonomicName id="036D6A479B27F543C04E1450B210576F" lsidName="L. montanus" pageId="0" pageNumber="13" rank="species" species="montanus">
|
|
<emphasis id="597F3761B0D978554CF418744F895E8C" italics="true" pageId="0" pageNumber="13">L. montanus</emphasis>
|
|
</taxonomicName>
|
|
;
|
|
<emphasis id="D1C3DB37D219F5739B80813507934E68" bold="true" pageId="0" pageNumber="13">C.</emphasis>
|
|
<taxonomicName id="090B0C69E19847266A78B7643D0EE758" lsidName="L. parallelus" pageId="0" pageNumber="13" rank="species" species="parallelus">
|
|
<emphasis id="5F551AE40DF4EF02C48DBF8CC80B7A86" italics="true" pageId="0" pageNumber="13">L. parallelus</emphasis>
|
|
</taxonomicName>
|
|
;
|
|
<emphasis id="A00D52302DE0CABCA114F49E5E24B0FD" bold="true" pageId="0" pageNumber="13">D.</emphasis>
|
|
<taxonomicName id="F8CBD80174D7810891C9CDD52056111E" lsidName="L. zamorae" pageId="0" pageNumber="13" rank="species" species="zamorae">
|
|
<emphasis id="31C7BE4806BE68E257EE2F9D870B5BE9" italics="true" pageId="0" pageNumber="13">L. zamorae</emphasis>
|
|
</taxonomicName>
|
|
;
|
|
<emphasis id="33CA566AF53E2807322E9E8761A5AD69" bold="true" pageId="0" pageNumber="13">E.</emphasis>
|
|
<taxonomicName id="BBB2E6EA69418EC6ACD967D41C31527F" authorityName="Irmler" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Somoleptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Somoleptus alajuelae" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="alajuelae">
|
|
<emphasis id="F9E5F01C441991E2B8CA8F2AEA737857" italics="true" pageId="0" pageNumber="13">Somoleptus alajuelae</emphasis>
|
|
</taxonomicName>
|
|
;
|
|
<emphasis id="A07A8D7F55F0105749856EF199DF4784" bold="true" pageId="0" pageNumber="13">F.</emphasis>
|
|
<taxonomicName id="0569A01684F83FE0864A8ADF4036A15C" lsidName="S. curtioculatus" pageId="0" pageNumber="13" rank="species" species="curtioculatus">
|
|
<emphasis id="84ECEAE276D7B02828EEE27FDDF35D57" italics="true" pageId="0" pageNumber="13">S. curtioculatus</emphasis>
|
|
</taxonomicName>
|
|
. Scale bars: 0.5 mm.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="659C31606E069E5B1B39F63C5CAFA04B" doi="10.3897/contrib.entomol.74.e114543.figure21" httpUri="https://binary.pensoft.net/fig/975722" pageId="0" pageNumber="13" start="Figure 21" startId="F8">
|
|
<paragraph id="40FBBB2C56C1FD644EE63315482CB7B7" pageId="0" pageNumber="13">
|
|
<emphasis id="11E130355299652E62D105E5E7750C01" bold="true" pageId="0" pageNumber="13">Figure 21.</emphasis>
|
|
<emphasis id="CE2FA1597F96C2B01BCB577606575BF9" bold="true" pageId="0" pageNumber="13">A.</emphasis>
|
|
Fore-body of
|
|
<taxonomicName id="278BC4AC8F53F04F85C2ECDD24BE03D6" authorityName="Irmler" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Somoleptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Somoleptus densiceps" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="densiceps">
|
|
<emphasis id="AEFAD06AE5DE3EBC9F4E43CD32DD108F" italics="true" pageId="0" pageNumber="13">Somoleptus densiceps</emphasis>
|
|
</taxonomicName>
|
|
;
|
|
<emphasis id="54357C8DC3A1DA9EA72647E3A0479AD7" bold="true" pageId="0" pageNumber="13">B.</emphasis>
|
|
<taxonomicName id="719F1594F82B7DD76657D205834C1E05" lsidName="S. guianensis" pageId="0" pageNumber="13" rank="species" species="guianensis">
|
|
<emphasis id="0F2CB4064985DF66F4B860777DDDBDFE" italics="true" pageId="0" pageNumber="13">S. guianensis</emphasis>
|
|
</taxonomicName>
|
|
;
|
|
<emphasis id="A855175B17C36B0805345160D01D98A3" bold="true" pageId="0" pageNumber="13">C.</emphasis>
|
|
<taxonomicName id="4CF3B17790F3EEC7EA71615651CA4A5B" lsidName="S. luteicornis" pageId="0" pageNumber="13" rank="species" species="luteicornis">
|
|
<emphasis id="8E8CEE55499938E912870A5652D692FE" italics="true" pageId="0" pageNumber="13">S. luteicornis</emphasis>
|
|
</taxonomicName>
|
|
;
|
|
<emphasis id="18D5845247D97A28F7FDCCFEA835EFA7" bold="true" pageId="0" pageNumber="13">D.</emphasis>
|
|
<taxonomicName id="616DC6FE8B22586FE0FED9BD17DECB4F" lsidName="S. paramocola" pageId="0" pageNumber="13" rank="species" species="paramocola">
|
|
<emphasis id="3C1A28BE1DC787ED317A9F41032AF5E8" italics="true" pageId="0" pageNumber="13">S. paramocola</emphasis>
|
|
</taxonomicName>
|
|
;
|
|
<emphasis id="25DAEC3A803F0D174CA6F5DC3285C4D6" bold="true" pageId="0" pageNumber="13">E.</emphasis>
|
|
<taxonomicName id="E1A7AA277F69244B8DF0441A3A345C27" lsidName="S. puntarenae" pageId="0" pageNumber="13" rank="species" species="puntarenae">
|
|
<emphasis id="68CE41CDDE2C68275467746515ACB10A" italics="true" pageId="0" pageNumber="13">S. puntarenae</emphasis>
|
|
</taxonomicName>
|
|
;
|
|
<emphasis id="F12654753E6D9AF32BC024588B3A00E4" bold="true" pageId="0" pageNumber="13">F.</emphasis>
|
|
<taxonomicName id="BA44B4D3F48ED949361F2E0848E1CFF1" lsidName="S. umicola" pageId="0" pageNumber="13" rank="species" species="umicola">
|
|
<emphasis id="26847336DFFCE531116BDDBBE72FF4C3" italics="true" pageId="0" pageNumber="13">S. umicola</emphasis>
|
|
</taxonomicName>
|
|
. Scale bars: 0.5 mm.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection id="5E073EE8D871F1FEC877E34B7C3EB73B" pageId="0" pageNumber="13" type="etymology">
|
|
<paragraph id="7D79AE5074A03D5D937411216FCA8D1A" pageId="0" pageNumber="13">Etymology.</paragraph>
|
|
<paragraph id="365CADEF26B41C39B93529120C7742CE" pageId="0" pageNumber="13">
|
|
The species name is a combination of the Latin words
|
|
<emphasis id="8D8D4B036362C6F31FC6959749CE7D2B" italics="true" pageId="0" pageNumber="13">umidus</emphasis>
|
|
meaning
|
|
<normalizedToken id="FC67C2A9A18419FF977C1A0B9F639524" originalValue="“wet”">"wet"</normalizedToken>
|
|
and
|
|
<emphasis id="D79B85AA0671E37C96E20A32720D50AB" italics="true" pageId="0" pageNumber="13">colo</emphasis>
|
|
meaning
|
|
<normalizedToken id="141EFCDC8A63677B93906BD166148EE7" originalValue="“dwelling”">"dwelling"</normalizedToken>
|
|
and refers to the wet habitat, where it was found.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="03A1047C9BE6932D58557F90BADA6346" pageId="0" pageNumber="13" type="geography">
|
|
<paragraph id="E3B2604DB738C5287B0E99806D3CC183" pageId="0" pageNumber="13">Geography.</paragraph>
|
|
<paragraph id="F5A900C38E18C86205C91EF45988AA81" pageId="0" pageNumber="13">Northern Costa Rica.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="05593B896CA793D6486E398E783F8610" pageId="0" pageNumber="13" type="ecology">
|
|
<paragraph id="5F550941C124DE70258FA76E0506CF16" pageId="0" pageNumber="13">Ecology.</paragraph>
|
|
<paragraph id="31E62C8C6D63EBB2FF9AE73E1525FBD6" pageId="0" pageNumber="13">Under moss of river bank.</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |