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<document id="FAE44BEFE8B2530E254B5FEBB85F80CA" ID-DOI="10.11646/zootaxa.3736.5.8" ID-GBIF-Dataset="c231e1f1-ac35-406c-871b-a5ad87995af9" ID-ISSN="1175-5326" ID-Zenodo-Dep="249099" ID-ZooBank="D961FDF8-B2FB-4B63-A01A-45A1978FC6D4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460699311845" checkinUser="plazi" docAuthor="Giłka, Wojciech, Zakrzewska, Marta, Dominiak, Patrycja &amp; Urbanek, Aleksandra" docDate="2013" docId="03E687CEEA14FFF1FF0BFE7A4932FEE5" docLanguage="en" docName="zt03736p586.pdf" docOrigin="Zootaxa 3736 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Archistempellina bifurca Gilka et Zakrzewska, sp. nov." docType="treatment" docVersion="9" lastPageNumber="573" masterDocId="FFDFFFB6EA16FFF5FF9CFFA6482BFFF9" masterDocTitle="Non-biting midges of the tribe Tanytarsini in Eocene amber from the Rovno region (Ukraine): a pioneer systematic study with notes on the phylogeny (Diptera: Chironomidae)" masterLastPageNumber="586" masterPageNumber="569" pageNumber="571" updateTime="1698354172026" updateUser="plazi">
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<mods:title id="365EC831778347440135E5E48143809F">Non-biting midges of the tribe Tanytarsini in Eocene amber from the Rovno region (Ukraine): a pioneer systematic study with notes on the phylogeny (Diptera: Chironomidae)</mods:title>
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<mods:namePart id="4723DC3C2A9D45EF6B5799253D383E81">Giłka, Wojciech</mods:namePart>
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<mods:namePart id="8740DE30BD85CEB4B3A8339BEBE57D8B">Zakrzewska, Marta</mods:namePart>
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<mods:namePart id="4A4AFD3CF5A044FA4F139517B80AF643">Dominiak, Patrycja</mods:namePart>
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<mods:namePart id="000FEB3CE3EBCAA718F429F2C5F5EF27">Urbanek, Aleksandra</mods:namePart>
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<heading id="D0B881B4EA14FFF7FF0BFE7A4B10FE0F" bold="true" box="[151,827,476,502]" fontSize="11" level="1" pageId="2" pageNumber="571" reason="1">
<emphasis id="B93BEACAEA14FFF7FF0BFE7A4B10FE0F" bold="true" box="[151,827,476,502]" pageId="2" pageNumber="571">
<taxonomicName id="4C4F4D5BEA14FFF7FF0BFE7A4AFBFE0F" ID-CoL="8B3HQ" authority="Gilka et Zakrzewska" authorityName="Gilka et Zakrzewska" box="[151,720,476,502]" class="Insecta" family="Chironomidae" genus="Archistempellina" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="571" phylum="Arthropoda" rank="species" species="bifurca" status="sp. nov.">
<emphasis id="B93BEACAEA14FFF7FF0BFE7A49EEFE0F" bold="true" box="[151,453,476,502]" italics="true" pageId="2" pageNumber="571">Archistempellina bifurca</emphasis>
Giłka
<emphasis id="B93BEACAEA14FFF7FD85FE7A4A05FE0F" bold="true" box="[537,558,476,502]" italics="true" pageId="2" pageNumber="571">et</emphasis>
Zakrzewska
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,
<taxonomicNameLabel id="A20857B1EA14FFF7FD41FE7A4B10FE0F" box="[733,827,476,502]" pageId="2" pageNumber="571" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
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<subSubSection id="C3556553EA14FFF7FF0BFD8449DEFDA6" pageId="2" pageNumber="571" type="materials_examined">
<paragraph id="8BF036D8EA14FFF7FF0BFD8449DEFDA6" blockId="2.[151,1437,546,1375]" pageId="2" pageNumber="571">
<emphasis id="B93BEACAEA14FFF7FF0BFD84496DFDC2" bold="true" box="[151,326,546,571]" pageId="2" pageNumber="571">
<typeStatus id="54F4887AEA14FFF7FF0BFD8448F9FDC2" box="[151,210,546,571]" pageId="2" pageNumber="571">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54F4887AEA14FFF7FED2FD844991FDC2" box="[334,442,546,571]" pageId="2" pageNumber="571" type="holotype">Holotype</typeStatus>
. Adult male (tarsus of left fore leg and right mid leg missing) in
<date id="FFF11018EA14FFF7FB02FD854CC4FDC3" box="[1182,1263,546,571]" pageId="2" pageNumber="571">8 x 6 x</date>
<quantity id="4CB79B3DEA14FFF7FB6BFD854D12FDC3" box="[1271,1337,547,571]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="571" unit="mm" value="4.0">4 mm</quantity>
piece of amber (SIZ K-26075,
<figureCitation id="13742A5DEA14FFF7FE0CFDE049E5FDA6" box="[400,462,582,607]" captionStart="FIGURE 1" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[248,1338,157,1913]" captionTargetId="figure@3.[248,1339,157,1913]" captionTargetPageId="3" captionText="FIGURE 1. Archistempellina bifurca sp. nov., male. A— holotype (inclusion); B— habitus; C, D— antenna (D— photographed using Nomarski DIC); E— wing; F K— hypopygium: F, I— dorsal aspect; G— ventral aspect; H— lateral aspect; J— superior volsella; K— median volsella (J, K— magnified three times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/249100/files/figure.png" pageId="2" pageNumber="571">Fig. 1</figureCitation>
A).
</paragraph>
</subSubSection>
<subSubSection id="C3556553EA14FFF7FF5BFDCD4A27FD5E" pageId="2" pageNumber="571" type="etymology">
<paragraph id="8BF036D8EA14FFF7FF5BFDCD4A27FD5E" blockId="2.[151,1437,546,1375]" pageId="2" pageNumber="571">
<emphasis id="B93BEACAEA14FFF7FF5BFDCD498FFD7D" bold="true" box="[199,420,619,644]" pageId="2" pageNumber="571">Derivatio nominis.</emphasis>
From Latin, meaning forked, in reference to the shape of the hypopygial median volsella armed with two slender lamellae.
</paragraph>
</subSubSection>
<subSubSection id="C3556553EA14FFF7FF5BFD154CD3FCED" pageId="2" pageNumber="571" type="diagnosis">
<paragraph id="8BF036D8EA14FFF7FF5BFD154CD3FCED" blockId="2.[151,1437,546,1375]" pageId="2" pageNumber="571">
<emphasis id="B93BEACAEA14FFF7FF5BFD15496AFD35" bold="true" box="[199,321,691,716]" pageId="2" pageNumber="571">Diagnosis.</emphasis>
Tibia of fore leg with small spur; femur of mid leg longer than femur of hind leg. Anal point tapering to pointed apex, reaching over superior and median volsellae and over half length of inferior volsella. Superior volsella very small, cylindrical. Stem of median volsella longer than its falciform lamellae.
</paragraph>
</subSubSection>
<subSubSection id="C3556553EA14FFF7FF5BFCB84C8BF828" pageId="2" pageNumber="571" type="description">
<paragraph id="8BF036D8EA14FFF7FF5BFCB84C61FCCE" blockId="2.[151,1437,546,1375]" box="[199,1098,798,823]" pageId="2" pageNumber="571">
<emphasis id="B93BEACAEA14FFF7FF5BFCB849CEFCCE" bold="true" box="[199,485,798,823]" pageId="2" pageNumber="571">Description. Adult male</emphasis>
(n = 1). Total length
<quantity id="4CB79B3DEA14FFF7FD4FFCB94B07FCCE" box="[723,812,799,823]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="571" unit="mm" value="2.7">2.7 mm</quantity>
; wing length c. 1400 Μm.
</paragraph>
<paragraph id="8BF036D8EA14FFF7FF5BFCE54C9DFC5D" blockId="2.[151,1437,546,1375]" pageId="2" pageNumber="571">
<emphasis id="B93BEACAEA14FFF7FF5BFCE5492FFCA5" box="[199,260,835,860]" italics="true" pageId="2" pageNumber="571">Head</emphasis>
(
<figureCitation id="13742A5DEA14FFF7FE89FCE5497DFCA2" box="[277,342,835,860]" captionStart="FIGURE 1" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[248,1338,157,1913]" captionTargetId="figure@3.[248,1339,157,1913]" captionTargetPageId="3" captionText="FIGURE 1. Archistempellina bifurca sp. nov., male. A— holotype (inclusion); B— habitus; C, D— antenna (D— photographed using Nomarski DIC); E— wing; F K— hypopygium: F, I— dorsal aspect; G— ventral aspect; H— lateral aspect; J— superior volsella; K— median volsella (J, K— magnified three times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/249100/files/figure.png" pageId="2" pageNumber="571">Fig. 1</figureCitation>
BD). Eyes bare, with well developed dorsomedian extensions. Frontal tubercles not observed. Antenna with 13 well discernible flagellomeres, AR 1.121.15, plume fully developed (
<figureCitation id="13742A5DEA14FFF7FBE8FCC04C9FFC86" box="[1140,1204,870,895]" captionStart="FIGURE 1" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[248,1338,157,1913]" captionTargetId="figure@3.[248,1339,157,1913]" captionTargetPageId="3" captionText="FIGURE 1. Archistempellina bifurca sp. nov., male. A— holotype (inclusion); B— habitus; C, D— antenna (D— photographed using Nomarski DIC); E— wing; F K— hypopygium: F, I— dorsal aspect; G— ventral aspect; H— lateral aspect; J— superior volsella; K— median volsella (J, K— magnified three times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/249100/files/figure.png" pageId="2" pageNumber="571">Fig. 1</figureCitation>
C, D). Palps slightly asymmetrical, length of palpomeres 25 (Μm): 48, 123, 147, 205245. Clypeals unobservable.
</paragraph>
<paragraph id="8BF036D8EA14FFF7FF5BFC084BAFFC15" blockId="2.[151,1437,546,1375]" pageId="2" pageNumber="571">
<emphasis id="B93BEACAEA14FFF7FF5BFC084933FC3E" box="[199,280,942,967]" italics="true" pageId="2" pageNumber="571">Thorax</emphasis>
(
<figureCitation id="13742A5DEA14FFF7FEB4FC084942FC3E" box="[296,361,942,967]" captionStart="FIGURE 1" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[248,1338,157,1913]" captionTargetId="figure@3.[248,1339,157,1913]" captionTargetPageId="3" captionText="FIGURE 1. Archistempellina bifurca sp. nov., male. A— holotype (inclusion); B— habitus; C, D— antenna (D— photographed using Nomarski DIC); E— wing; F K— hypopygium: F, I— dorsal aspect; G— ventral aspect; H— lateral aspect; J— superior volsella; K— median volsella (J, K— magnified three times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/249100/files/figure.png" pageId="2" pageNumber="571">Fig. 1</figureCitation>
B). Antepronotum relatively well developed, scutum not overreaching antepronotum. Tubercle on scutum absent. Most of setae unobservable, at least 8 scutellars.
</paragraph>
<paragraph id="8BF036D8EA14FFF7FF5BFC5E4994FB51" blockId="2.[151,1437,546,1375]" pageId="2" pageNumber="571">
<emphasis id="B93BEACAEA14FFF7FF5BFC5E48D4FBF6" box="[199,255,1016,1039]" italics="true" pageId="2" pageNumber="571">Wing</emphasis>
(
<figureCitation id="13742A5DEA14FFF7FE8DFC504979FBF6" box="[273,338,1014,1039]" captionStart="FIGURE 1" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[248,1338,157,1913]" captionTargetId="figure@3.[248,1339,157,1913]" captionTargetPageId="3" captionText="FIGURE 1. Archistempellina bifurca sp. nov., male. A— holotype (inclusion); B— habitus; C, D— antenna (D— photographed using Nomarski DIC); E— wing; F K— hypopygium: F, I— dorsal aspect; G— ventral aspect; H— lateral aspect; J— superior volsella; K— median volsella (J, K— magnified three times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/249100/files/figure.png" pageId="2" pageNumber="571">Fig. 1</figureCitation>
E). Slender, with anal lobe weak, broadest at 2/3 length, width: 387 Μm, length/width ratio 3.62. RM slightly oblique relative to R. Sc unobservable. R1 and R2+3 arcuate. FCu placed distally of RM; VR Cu 1.25. Veins ending as follows (in order from base to tip): An, Cu1, R1, R2+3, M3+4, R4+5, M1+2; distances between ends of R1R2+3 and R2+3R4+5 unequal (VR C c. 4); An reaching FCu at least. Wing membrane covered with dense macrotrichia in distal half.
</paragraph>
<paragraph id="8BF036D8EA14FFF7FF5BFB124992FAA6" blockId="2.[151,1437,546,1375]" pageId="2" pageNumber="571">
<emphasis id="B93BEACAEA14FFF7FF5BFB1248D7FB32" box="[199,252,1204,1227]" italics="true" pageId="2" pageNumber="571">Legs</emphasis>
. Tibia of fore leg with small spur, c. 10 Μm long. Single tibial comb of mid leg fan-shaped, teeth up to 20 Μm long, with single slender c. 4045 Μm long spur; combs of hind leg broadly separated, fan-shaped, one comb broad (composed of c. 15 teeth) up to 20 Μm long, without spur, second comb narrow, teeth up to 25 Μm long, bearing c. 50 Μm long spur. Sensilla chaetica on ta1 of p2 not observed. Pulvilli absent. For length of leg segments and leg ratios see
<tableCitation id="C6CD0363EA14FFF7FEC3FAE04998FAA6" box="[351,435,1350,1375]" captionStart="TABLE 1" captionStartId="2.[151,239,1418,1441]" captionTargetBox="[159,1318,1468,1626]" captionTargetPageId="2" captionText="TABLE 1. Leg segment lengths (µm) and leg ratios of male Archistempellina bifurca sp. nov." httpUri="http://table.plazi.org/id/DF306650EA14FFF7FF0BFA2C4CAAFA58" pageId="2" pageNumber="571" tableUuid="DF306650EA14FFF7FF0BFA2C4CAAFA58">Table 1</tableCitation>
.
</paragraph>
<caption id="DF306650EA14FFF7FF0BFA2C4CAAFA58" ID-Table-UUID="DF306650EA14FFF7FF0BFA2C4CAAFA58" box="[151,1153,1418,1441]" httpUri="http://table.plazi.org/id/DF306650EA14FFF7FF0BFA2C4CAAFA58" pageId="2" pageNumber="571" targetBox="[159,1318,1468,1626]" targetIsTable="true" targetPageId="2">
<paragraph id="8BF036D8EA14FFF7FF0BFA2C4CAAFA58" blockId="2.[151,1153,1418,1441]" box="[151,1153,1418,1441]" pageId="2" pageNumber="571">
<emphasis id="B93BEACAEA14FFF7FF0BFA2C4929FA58" bold="true" box="[151,258,1418,1441]" pageId="2" pageNumber="571">TABLE 1</emphasis>
. Leg segment lengths (µm) and leg ratios of male
<taxonomicName id="4C4F4D5BEA14FFF7FC81FA2C4C0EFA58" box="[797,1061,1418,1441]" class="Insecta" family="Chironomidae" genus="Archistempellina" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="571" phylum="Arthropoda" rank="species" species="bifurca" status="sp. nov.">
<emphasis id="B93BEACAEA14FFF7FC81FA2C4C0EFA58" box="[797,1061,1418,1441]" italics="true" pageId="2" pageNumber="571">Archistempellina bifurca</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20857B1EA14FFF7FBB7FA2C4CAAFA58" box="[1067,1153,1418,1441]" pageId="2" pageNumber="571" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BF036D8EA14FFF7FF4AFA1A4D0DF9AF" pageId="2" pageNumber="571">
<table id="F94FC478EA14000AFF03FA1A4D0DF9A3" box="[159,1318,1468,1626]" gridcols="7" gridrows="4" pageId="2" pageNumber="571">
<tr id="357F349AEA14000AFF03FA1A4D0DFA2F" box="[159,1318,1468,1494]" gridrow="0" pageId="2" pageNumber="571">
<th id="76AE5DE6EA14000AFF03FA1A48D7FA2F" box="[159,252,1468,1494]" gridcol="0" gridrow="0" pageId="2" pageNumber="571">fe</th>
<th id="76AE5DE6EA14000AFEF2FA1A4A00FA2F" box="[366,555,1468,1494]" gridcol="1" gridrow="0" pageId="2" pageNumber="571">ti ta1</th>
<th id="76AE5DE6EA14000AFD01FA1A4AE8FA2F" box="[669,707,1468,1494]" gridcol="2" gridrow="0" pageId="2" pageNumber="571">ta2</th>
<th id="76AE5DE6EA14000AFCA8FA1A4B71FA2F" box="[820,858,1468,1494]" gridcol="3" gridrow="0" pageId="2" pageNumber="571">ta3</th>
<th id="76AE5DE6EA14000AFC57FA1A4BDAFA2F" box="[971,1009,1468,1494]" gridcol="4" gridrow="0" pageId="2" pageNumber="571">ta4</th>
<th id="76AE5DE6EA14000AFBFFFA1A4C56FA2F" box="[1123,1149,1468,1494]" gridcol="5" gridrow="0" pageId="2" pageNumber="571">ta5</th>
<th id="76AE5DE6EA14000AFB66FA1A4D0DFA2F" box="[1274,1318,1468,1494]" gridcol="6" gridrow="0" pageId="2" pageNumber="571">LR</th>
</tr>
<tr id="357F349AEA14000AFF03FA4E4D0DF9FB" box="[159,1318,1512,1538]" gridrow="1" pageId="2" pageNumber="571">
<th id="76AE5DE6EA14000AFF03FA4E48D7F9FB" box="[159,252,1512,1538]" gridcol="0" gridrow="1" pageId="2" pageNumber="571">P1 585</th>
<td id="76AE5DE6EA14000AFEF2FA4E4A00F9FB" box="[366,555,1512,1538]" gridcol="1" gridrow="1" pageId="2" pageNumber="571">395410 760</td>
<td id="76AE5DE6EA14000AFD01FA4E4AE8F9FB" box="[669,707,1512,1538]" gridcol="2" gridrow="1" pageId="2" pageNumber="571">-</td>
<td id="76AE5DE6EA14000AFCA8FA4E4B71F9FB" box="[820,858,1512,1538]" gridcol="3" gridrow="1" pageId="2" pageNumber="571">-</td>
<td id="76AE5DE6EA14000AFC57FA4E4BDAF9FB" box="[971,1009,1512,1538]" gridcol="4" gridrow="1" pageId="2" pageNumber="571">255</td>
<td id="76AE5DE6EA14000AFBFFFA4E4C56F9FB" box="[1123,1149,1512,1538]" gridcol="5" gridrow="1" pageId="2" pageNumber="571">85</td>
<td id="76AE5DE6EA14000AFB66FA4E4D0DF9FB" box="[1274,1318,1512,1538]" gridcol="6" gridrow="1" pageId="2" pageNumber="571">1.85</td>
</tr>
<tr id="357F349AEA14000AFF03F9B24D0DF9D7" box="[159,1318,1556,1582]" gridrow="2" pageId="2" pageNumber="571">
<th id="76AE5DE6EA14000AFF03F9B248D7F9D7" box="[159,252,1556,1582]" gridcol="0" gridrow="2" pageId="2" pageNumber="571">P2 685</th>
<td id="76AE5DE6EA14000AFEF2F9B24A00F9D7" box="[366,555,1556,1582]" gridcol="1" gridrow="2" pageId="2" pageNumber="571">560 400</td>
<td id="76AE5DE6EA14000AFD01F9B24AE8F9D7" box="[669,707,1556,1582]" gridcol="2" gridrow="2" pageId="2" pageNumber="571">195</td>
<td id="76AE5DE6EA14000AFCA8F9B24B71F9D7" box="[820,858,1556,1582]" gridcol="3" gridrow="2" pageId="2" pageNumber="571">140</td>
<td id="76AE5DE6EA14000AFC57F9B24BDAF9D7" box="[971,1009,1556,1582]" gridcol="4" gridrow="2" pageId="2" pageNumber="571">90</td>
<td id="76AE5DE6EA14000AFBFFF9B24C56F9D7" box="[1123,1149,1556,1582]" gridcol="5" gridrow="2" pageId="2" pageNumber="571">60</td>
<td id="76AE5DE6EA14000AFB66F9B24D0DF9D7" box="[1274,1318,1556,1582]" gridcol="6" gridrow="2" pageId="2" pageNumber="571">0.71</td>
</tr>
<tr id="357F349AEA14000AFF03F9E64D0DF9A3" box="[159,1318,1600,1626]" gridrow="3" pageId="2" pageNumber="571">
<th id="76AE5DE6EA14000AFF03F9E648D7F9A3" box="[159,252,1600,1626]" gridcol="0" gridrow="3" pageId="2" pageNumber="571">P3 645</th>
<td id="76AE5DE6EA14000AFEF2F9E64A00F9A3" box="[366,555,1600,1626]" gridcol="1" gridrow="3" pageId="2" pageNumber="571">595 520</td>
<td id="76AE5DE6EA14000AFD01F9E64AE8F9A3" box="[669,707,1600,1626]" gridcol="2" gridrow="3" pageId="2" pageNumber="571">285</td>
<td id="76AE5DE6EA14000AFCA8F9E64B71F9A3" box="[820,858,1600,1626]" gridcol="3" gridrow="3" pageId="2" pageNumber="571">240</td>
<td id="76AE5DE6EA14000AFC57F9E64BDAF9A3" box="[971,1009,1600,1626]" gridcol="4" gridrow="3" pageId="2" pageNumber="571">140</td>
<td id="76AE5DE6EA14000AFBFFF9E64C56F9A3" box="[1123,1149,1600,1626]" gridcol="5" gridrow="3" pageId="2" pageNumber="571">80</td>
<td id="76AE5DE6EA14000AFB66F9E64D0DF9A3" box="[1274,1318,1600,1626]" gridcol="6" gridrow="3" pageId="2" pageNumber="571">0.87</td>
</tr>
</table>
</paragraph>
<paragraph id="8BF036D8EA14FFF7FF5BF93C4C8BF828" blockId="2.[151,1436,1689,2002]" pageId="2" pageNumber="571">
<emphasis id="B93BEACAEA14FFF7FF5BF93C497DF948" box="[199,342,1690,1713]" italics="true" pageId="2" pageNumber="571">Hypopygium</emphasis>
(
<figureCitation id="13742A5DEA14FFF7FEFAF93F498DF94B" box="[358,422,1689,1714]" captionStart="FIGURE 1" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[248,1338,157,1913]" captionTargetId="figure@3.[248,1339,157,1913]" captionTargetPageId="3" captionText="FIGURE 1. Archistempellina bifurca sp. nov., male. A— holotype (inclusion); B— habitus; C, D— antenna (D— photographed using Nomarski DIC); E— wing; F K— hypopygium: F, I— dorsal aspect; G— ventral aspect; H— lateral aspect; J— superior volsella; K— median volsella (J, K— magnified three times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/249100/files/figure.png" pageId="2" pageNumber="571">Fig. 1</figureCitation>
FK). Gonostylus cylindrical, straight, c. 115 Μm long, distinctly longer than gonocoxite, with strong apical seta. Anal tergite subtriangular, bearing several strong median setae. Anal point slender, styliform, tapering to pointed apex, without spinulae and crests, bearing at least 4 posterolateral setae on each side; anal point reaching over superior and median volsellae and over half length of inferior volsella; upturned in lateral view with tip bent down (
<figureCitation id="13742A5DEA14FFF7FE27F88F49D0F8BB" box="[443,507,1833,1858]" captionStart="FIGURE 1" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[248,1338,157,1913]" captionTargetId="figure@3.[248,1339,157,1913]" captionTargetPageId="3" captionText="FIGURE 1. Archistempellina bifurca sp. nov., male. A— holotype (inclusion); B— habitus; C, D— antenna (D— photographed using Nomarski DIC); E— wing; F K— hypopygium: F, I— dorsal aspect; G— ventral aspect; H— lateral aspect; J— superior volsella; K— median volsella (J, K— magnified three times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/249100/files/figure.png" pageId="2" pageNumber="571">Fig. 1</figureCitation>
FI). Superior volsella extraordinarily small, cylindrical, slightly swollen in distal part, bearing at least 2 setae at rounded apex (
<figureCitation id="13742A5DEA14FFF7FD03F8EB4AF4F89F" box="[671,735,1869,1894]" captionStart="FIGURE 1" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[248,1338,157,1913]" captionTargetId="figure@3.[248,1339,157,1913]" captionTargetPageId="3" captionText="FIGURE 1. Archistempellina bifurca sp. nov., male. A— holotype (inclusion); B— habitus; C, D— antenna (D— photographed using Nomarski DIC); E— wing; F K— hypopygium: F, I— dorsal aspect; G— ventral aspect; H— lateral aspect; J— superior volsella; K— median volsella (J, K— magnified three times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/249100/files/figure.png" pageId="2" pageNumber="571">Fig. 1</figureCitation>
J). Digitus absent. Stem of median volsella about the size and shape of superior volsella, c. 20 Μm long, slightly curved, medially directed, bearing 2 slender falciform lamellae; stem distinctly longer than lamellae (
<figureCitation id="13742A5DEA14FFF7FDDEF8334AAFF857" box="[578,644,1941,1966]" captionStart="FIGURE 1" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[248,1338,157,1913]" captionTargetId="figure@3.[248,1339,157,1913]" captionTargetPageId="3" captionText="FIGURE 1. Archistempellina bifurca sp. nov., male. A— holotype (inclusion); B— habitus; C, D— antenna (D— photographed using Nomarski DIC); E— wing; F K— hypopygium: F, I— dorsal aspect; G— ventral aspect; H— lateral aspect; J— superior volsella; K— median volsella (J, K— magnified three times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/249100/files/figure.png" pageId="2" pageNumber="571">Fig. 1</figureCitation>
K). Inferior volsella reaching 1/3 length of gonostylus at most, clubshaped, with distinctly enlarged head-like apical half, armed with several stout curved setae.
</paragraph>
</subSubSection>
<caption id="DF306650EA15FFF6FF0BF83F4C12F815" httpUri="https://zenodo.org/record/249100/files/figure.png" pageId="3" pageNumber="572" targetBox="[248,1338,157,1913]" targetPageId="3">
<paragraph id="8BF036D8EA15FFF6FF0BF83F4C12F815" blockId="3.[151,1436,1945,2028]" pageId="3" pageNumber="572">
<emphasis id="B93BEACAEA15FFF6FF0BF83F4938F857" bold="true" box="[151,275,1945,1967]" pageId="3" pageNumber="572">FIGURE 1.</emphasis>
<taxonomicName id="4C4F4D5BEA15FFF6FE85F83F4A38F856" box="[281,531,1945,1967]" class="Insecta" family="Chironomidae" genus="Archistempellina" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="572" phylum="Arthropoda" rank="species" species="bifurca" status="sp. nov.">
<emphasis id="B93BEACAEA15FFF6FE85F83F4A38F856" box="[281,531,1945,1967]" italics="true" pageId="3" pageNumber="572">Archistempellina bifurca</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20857B1EA15FFF6FD86F83F4A40F857" box="[538,619,1945,1966]" pageId="3" pageNumber="572" rank="species">
<emphasis id="B93BEACAEA15FFF6FD86F83F4A40F857" bold="true" box="[538,619,1945,1966]" pageId="3" pageNumber="572">sp. nov.</emphasis>
</taxonomicNameLabel>
, male.
<emphasis id="B93BEACAEA15FFF6FD2AF83F4AC9F857" bold="true" box="[694,738,1945,1966]" pageId="3" pageNumber="572">A—</emphasis>
holotype (inclusion);
<emphasis id="B93BEACAEA15FFF6FC20F83F4BCDF857" bold="true" box="[956,998,1945,1966]" pageId="3" pageNumber="572">B—</emphasis>
habitus;
<emphasis id="B93BEACAEA15FFF6FBA1F83F4C64F856" bold="true" box="[1085,1103,1945,1967]" pageId="3" pageNumber="572">C</emphasis>
,
<emphasis id="B93BEACAEA15FFF6FBC0F83F4CA3F857" bold="true" box="[1116,1160,1945,1966]" pageId="3" pageNumber="572">D—</emphasis>
antenna (
<emphasis id="B93BEACAEA15FFF6FB79F83F4D3AF857" bold="true" box="[1253,1297,1945,1966]" pageId="3" pageNumber="572">D—</emphasis>
photographed using Nomarski DIC);
<emphasis id="B93BEACAEA15FFF6FE1FF81E4986F834" bold="true" box="[387,429,1976,1997]" pageId="3" pageNumber="572">E—</emphasis>
wing;
<emphasis id="B93BEACAEA15FFF6FE73F81E4A13F834" bold="true" box="[495,568,1976,1997]" pageId="3" pageNumber="572">FK—</emphasis>
hypopygium:
<emphasis id="B93BEACAEA15FFF6FD5AF81E4AF8F834" bold="true" box="[710,723,1976,1997]" pageId="3" pageNumber="572">F</emphasis>
,
<emphasis id="B93BEACAEA15FFF6FD7EF81E4B2EF834" bold="true" box="[738,773,1976,1997]" pageId="3" pageNumber="572">I—</emphasis>
dorsal aspect;
<emphasis id="B93BEACAEA15FFF6FC04F81E4BEEF837" bold="true" box="[920,965,1976,1998]" pageId="3" pageNumber="572">G—</emphasis>
ventral aspect;
<emphasis id="B93BEACAEA15FFF6FBFDF81E4CA5F834" bold="true" box="[1121,1166,1976,1997]" pageId="3" pageNumber="572">H—</emphasis>
lateral aspect;
<emphasis id="B93BEACAEA15FFF6FABFF81E4D62F834" bold="true" box="[1315,1353,1976,1997]" pageId="3" pageNumber="572">J—</emphasis>
superior volsella;
<emphasis id="B93BEACAEA15FFF6FF6FF871490BF815" bold="true" box="[243,288,2007,2028]" pageId="3" pageNumber="572">K—</emphasis>
median volsella (
<emphasis id="B93BEACAEA15FFF6FE52F87149F0F815" bold="true" box="[462,475,2007,2028]" pageId="3" pageNumber="572">J</emphasis>
,
<emphasis id="B93BEACAEA15FFF6FE74F8714A3EF815" bold="true" box="[488,533,2007,2028]" pageId="3" pageNumber="572">K—</emphasis>
magnified three times relative to hypopygium drawn).
</paragraph>
</caption>
<subSubSection id="C3556553EA12FFF1FF5BFF314932FEE5" pageId="4" pageNumber="573" type="discussion">
<paragraph id="8BF036D8EA12FFF1FF5BFF314932FEE5" blockId="4.[151,1436,151,285]" pageId="4" pageNumber="573">
<emphasis id="B93BEACAEA12FFF1FF5BFF314910FF49" bold="true" box="[199,315,151,176]" pageId="4" pageNumber="573">Remarks.</emphasis>
The best diagnostic characters that allow the separation of
<taxonomicName id="4C4F4D5BEA12FFF1FC4EFF314CC3FF49" box="[978,1256,151,176]" class="Insecta" family="Chironomidae" genus="Archistempellina" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="573" phylum="Arthropoda" rank="species" species="bifurca">
<emphasis id="B93BEACAEA12FFF1FC4EFF314CC3FF49" box="[978,1256,151,176]" italics="true" pageId="4" pageNumber="573">Archistempellina bifurca</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4F4D5BEA12FFF1FA82FF3E4DB7FF49" box="[1310,1436,151,176]" class="Insecta" family="Chironomidae" genus="Archistempellina" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="573" phylum="Arthropoda" rank="species" species="falcifera">
<emphasis id="B93BEACAEA12FFF1FA82FF3E4DB7FF49" box="[1310,1436,151,176]" italics="true" pageId="4" pageNumber="573">A. falcifera</emphasis>
</taxonomicName>
are the shape of the anal point, the arrangement/position of the anal point apex in relation to the hypopygial volsellae (
<emphasis id="B93BEACAEA12FFF1FE97FF46490CFF01" box="[267,295,224,248]" italics="true" pageId="4" pageNumber="573">cf.</emphasis>
<figureCitation id="13742A5DEA12FFF1FEB1FF794941FF01" box="[301,362,223,248]" captionStart="FIGURE 1" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[248,1338,157,1913]" captionTargetId="figure@3.[248,1339,157,1913]" captionTargetPageId="3" captionText="FIGURE 1. Archistempellina bifurca sp. nov., male. A— holotype (inclusion); B— habitus; C, D— antenna (D— photographed using Nomarski DIC); E— wing; F K— hypopygium: F, I— dorsal aspect; G— ventral aspect; H— lateral aspect; J— superior volsella; K— median volsella (J, K— magnified three times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/249100/files/figure.png" pageId="4" pageNumber="573">Fig. 1</figureCitation>
F, G, I and
<figureCitation id="13742A5DEA12FFF1FE74FF794A00FF01" box="[488,555,223,248]" captionStart="FIGURE 2" captionStartId="5.[151,250,1940,1962]" captionTargetBox="[250,1340,157,1913]" captionTargetId="figure@5.[250,1341,157,1913]" captionTargetPageId="5" captionText="FIGURE 2. Archistempellina falcifera sp. nov., male. A — holotype (inclusion); B — habitus; C — antenna; D — wing; E J — hypopygium: E, G — ventral aspect; F — lateral aspect; H — anal point; I — superior volsella, J — median volsella (H J — magnified c. three times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/249101/files/figure.png" pageId="4" pageNumber="573">Fig. 2</figureCitation>
E, G, H), and the lamellae/stem length proportions in median volsellae (
<emphasis id="B93BEACAEA12FFF1FAD1FF464D42FF01" box="[1357,1385,224,248]" italics="true" pageId="4" pageNumber="573">cf.</emphasis>
<figureCitation id="13742A5DEA12FFF1FAF3FF79488AFEE5" captionStart="FIGURE 1" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[248,1338,157,1913]" captionTargetId="figure@3.[248,1339,157,1913]" captionTargetPageId="3" captionText="FIGURE 1. Archistempellina bifurca sp. nov., male. A— holotype (inclusion); B— habitus; C, D— antenna (D— photographed using Nomarski DIC); E— wing; F K— hypopygium: F, I— dorsal aspect; G— ventral aspect; H— lateral aspect; J— superior volsella; K— median volsella (J, K— magnified three times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/249100/files/figure.png" pageId="4" pageNumber="573">Fig. 1</figureCitation>
K and 2J).
</paragraph>
</subSubSection>
</treatment>
</document>