treatments-xml/data/03/E6/87/03E68784FFA5C21285C2FA09FD70FD62.xml
2024-06-21 12:22:17 +02:00

504 lines
64 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="5AF7DE0FD50966EAA21F66C62E0BF371" ID-CLB-Dataset="36863" ID-DOI="10.11646/zootaxa.3794.1.1" ID-GBIF-Dataset="1e1241a0-cc91-44fc-818d-e561293e2017" ID-ISSN="1175-5326" ID-Zenodo-Dep="285981" ID-ZooBank="529185ED-26B1-4C8F-8649-8A030A793138" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460415134178" checkinUser="plazi" docAuthor="Geniez, Philippe, Sá-Sousa, Paulo, Guillaume, Claude P., Cluchier, Alexandre &amp; Crochet, Pierre-André" docDate="2014" docId="03E68784FFA5C21285C2FA09FD70FD62" docLanguage="en" docName="zt03794p051.pdf" docOrigin="Zootaxa 3794 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Podarcis guadarramae subsp. lusitanicus Geniez, Sá-Sousa, Guillaume, Cluchier &amp; Crochet, 2014, subsp. nov." docType="treatment" docVersion="8" lastPageNumber="20" masterDocId="FFDFFFFCFFA8C2018555FFC8FF8CFF92" masterDocTitle="Systematics of the Podarcis hispanicus complex (Sauria, Lacertidae) III: valid nomina of the western and central Iberian forms" masterLastPageNumber="51" masterPageNumber="1" pageNumber="14" updateTime="1698328581752" updateUser="ExternalLinkService">
<mods:mods id="63A854890DC091CBFEC4FE7CE1C4546E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5927BD5EB34D4BC612F7FA12A027D9C4">
<mods:title id="73C50F29294F0237A9CFAC7E210E115A">Systematics of the Podarcis hispanicus complex (Sauria, Lacertidae) III: valid nomina of the western and central Iberian forms</mods:title>
</mods:titleInfo>
<mods:name id="19E2B3B1E2FDDC195B06897F0EE710C9" type="personal">
<mods:role id="3725AA89A735B52B2631C68C13C39645">
<mods:roleTerm id="45189361B5E049C9AA9F0DE0F90C860B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2B0899E0D91A3E9E6561C889E0DEBF95">Geniez, Philippe</mods:namePart>
</mods:name>
<mods:name id="3C47E51F8FB03190543270BDB751AC03" type="personal">
<mods:role id="A51F9D0EC9FB8B85B94591D5B94AD599">
<mods:roleTerm id="881AD05325305CED811F97AA55854E32">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E67885802C93291EF9035347BBF9A90C">Sá-Sousa, Paulo</mods:namePart>
</mods:name>
<mods:name id="2FE10AAFEB38270019AF271B26D7D253" type="personal">
<mods:role id="FEC026C9043F2794A754212147A71D40">
<mods:roleTerm id="17D3281B9AAE0542CD16E5B94A5A77BE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3CD0EC058D593910A77B320C8F57D0EC">Guillaume, Claude P.</mods:namePart>
</mods:name>
<mods:name id="C332E3CFEE7DF5226691E94994BD65E3" type="personal">
<mods:role id="2998CB002A00EE254DC314188687D69E">
<mods:roleTerm id="17383BC20075CF8BDC8EF60394329AF0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4A598E24585730ADD89C60BD7A13ADD3">Cluchier, Alexandre</mods:namePart>
</mods:name>
<mods:name id="FBDC7756EA25C34CC6F3B97EA38E0B98" type="personal">
<mods:role id="2734AE22F73C5DFA9F42B8E7EAC9EDAC">
<mods:roleTerm id="F76B72448EAC77602971CA1A419A8501">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="530A34665E69A177830112007EBD5253">Crochet, Pierre-André</mods:namePart>
</mods:name>
<mods:typeOfResource id="21F8C4D7B278961FBEEE93E9219E21D2">text</mods:typeOfResource>
<mods:relatedItem id="71B85F520F1870AA70079CFD1E411DF6" type="host">
<mods:titleInfo id="0D96A261C5FAC24F33049C25AA136369">
<mods:title id="799E013302094995FACAD07491A1FE72">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="807586F608513720F693C423D69A4031">
<mods:date id="AD7DD302D80C148567065EB43486E89C">2014</mods:date>
<mods:detail id="1DC30D71E6E794009C6474CDB218B81F" type="volume">
<mods:number id="0BAD33E771F7CA38A54F20DE94139A2F">3794</mods:number>
</mods:detail>
<mods:detail id="5A6F86B4BF039560EAE7DBA1DE10C053" type="issue">
<mods:number id="2FA40E7FB50E3F74142F40080CB8B35C">1</mods:number>
</mods:detail>
<mods:extent id="5AB52B34664BBB540FFDA231DC2C96D9" unit="page">
<mods:start id="E2CDDCE480E636FA11E3255EBCB49902">1</mods:start>
<mods:end id="9588FFC41F441F1BA3DBC9280EF9CF97">51</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="D9B804821BB61ADEABA3FA4A604F9088">journal article</mods:classification>
<mods:identifier id="5AC6C6A5DEB7F88968038FD5A6DFAB52" type="CLB-Dataset">36863</mods:identifier>
<mods:identifier id="5AA202A39A22FE45F785C19FECC3C65D" type="DOI">10.11646/zootaxa.3794.1.1</mods:identifier>
<mods:identifier id="994795DE07199459EF489139F04389E2" type="GBIF-Dataset">1e1241a0-cc91-44fc-818d-e561293e2017</mods:identifier>
<mods:identifier id="CA4222C90771C886E5F052CCFEE03014" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="81BEEBCFFEDEBC5C9BF07F98B7DF8888" type="Zenodo-Dep">285981</mods:identifier>
<mods:identifier id="5BFFB7C75340AAB097A2C00A72607B31" type="ZooBank">529185ED-26B1-4C8F-8649-8A030A793138</mods:identifier>
</mods:mods>
<treatment id="03E68784FFA5C21285C2FA09FD70FD62" ID-DOI="http://doi.org/10.5281/zenodo.5694219" ID-GBIF-Taxon="127667738" ID-Zenodo-Dep="5694219" LSID="urn:lsid:plazi:treatment:03E68784FFA5C21285C2FA09FD70FD62" httpUri="http://treatment.plazi.org/id/03E68784FFA5C21285C2FA09FD70FD62" lastPageId="19" lastPageNumber="20" pageId="13" pageNumber="14">
<subSubSection id="C3556519FFA5C20C85C2FA09FD4AFA49" box="[151,710,1473,1499]" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="8BF03692FFA5C20C85C2FA09FD4AFA49" blockId="13.[151,710,1473,1499]" box="[151,710,1473,1499]" pageId="13" pageNumber="14">
<heading id="D0B881FEFFA5C20C85C2FA09FD4AFA49" bold="true" box="[151,710,1473,1499]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<emphasis id="B93BEA80FFA5C20C85C2FA09FD4AFA49" bold="true" box="[151,710,1473,1499]" pageId="13" pageNumber="14">
<taxonomicName id="4C4F4D11FFA5C20C85C2FA09FDBAFA49" box="[151,566,1473,1499]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="subSpecies" species="guadarramae" status="subsp. nov." subSpecies="lusitanicus">
<emphasis id="B93BEA80FFA5C20C85C2FA09FDBAFA49" bold="true" box="[151,566,1473,1499]" italics="true" pageId="13" pageNumber="14">Podarcis guadarramae lusitanicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20857FBFFA5C20C8768FA09FD4AFA49" box="[573,710,1473,1499]" pageId="13" pageNumber="14" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3556519FFA5C20C85C2F9CFFB5FF88E" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="8BF03692FFA5C20C85C2F9CFFB5FF88E" blockId="13.[151,1436,1543,2037]" pageId="13" pageNumber="14">
<emphasis id="B93BEA80FFA5C20C85C2F9CFFE81F9B2" bold="true" box="[151,269,1543,1568]" pageId="13" pageNumber="14">
<typeStatus id="54F48830FFA5C20C85C2F9CFFE84F9B2" box="[151,264,1543,1568]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
:
</emphasis>
Muséum national dHistoire naturelle
<collectionCode id="ED5EAE57FFA5C20C8787F9C0FCABF98D" LSID="urn:lsid:biocol.org:col:34988" box="[722,807,1544,1567]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="14">MNHN</collectionCode>
2012.0263 (formerly
<collectionCode id="ED5EAE57FFA5C20C817FF9C0FBE8F9B2" LSID="urn:lsid:biocol.org:col:12677" box="[1066,1124,1544,1568]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12677" name="Borough of Beverley Art Gallery and Museum" pageId="13" pageNumber="14">BEV</collectionCode>
.3987),
<typeStatus id="54F48830FFA5C20C81EDF9CFFA96F9B2" box="[1208,1306,1543,1568]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
by present designation; an adult male collected
<quantity id="4CB79B77FFA5C20C8714F9E4FDF1F9D7" box="[577,637,1580,1605]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="14" unit="km" value="1.0">1 km</quantity>
past Âncora towards Póvoa de Varzim (south of Caminha,
<collectingCountry id="F3587602FFA5C20C8063F9E4FA14F9D7" box="[1334,1432,1580,1605]" name="Portugal" pageId="13" pageNumber="14">Portugal</collectingCountry>
, district of Viana do Castelo) [
<geoCoordinate id="EE7B5055FFA5C20C84A6F998FDEEF9FA" box="[499,610,1616,1640]" direction="north" orientation="latitude" pageId="13" pageNumber="14" precision="55" value="41.794">41.794°N</geoCoordinate>
/
<geoCoordinate id="EE7B5055FFA5C20C872AF987FD65F9FA" box="[639,745,1615,1640]" direction="west" orientation="longitude" pageId="13" pageNumber="14" precision="55" value="-8.864">8.864°W</geoCoordinate>
],
<quantity id="4CB79B77FFA5C20C87AAF998FCB4F9F5" box="[767,824,1616,1640]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="14" unit="m" value="30.0">30 m</quantity>
a.s.l. in
<date id="FFF11052FFA5C20C86F7F998FB94F9FA" box="[930,1048,1615,1640]" pageId="13" pageNumber="14" value="1985-05">May 1985</date>
by C.P. Guillaume, P. Geniez, U. Mathis and J. Magraner (
<figureCitation id="13742A17FFA5C20C84E1F9BCFE74F91E" box="[436,504,1652,1677]" captionStart="FIGURE 7" captionStartId="14.[151,250,1909,1931]" captionTargetBox="[341,1255,193,1892]" captionTargetId="figure@14.[341,1256,193,1893]" captionTargetPageId="14" captionText="FIGURE 7. Holotype of Podarcis guadarramae lusitanicus, male MNHN 2012.0263. Portugal, 1 km past Âncora towards Póvoa do Varzim (district of Viana do Castelo), 14.05. 1985. Photos P. Geniez. Note the very flattened head, the very large black spots on the pileus, the lack of vertebral stripe, the very broad black supradorsolateral stripes invading a large part of the pale green dorsum and the light dorsolateral stripes made of weakly elongated spots." httpUri="https://zenodo.org/record/285988/files/figure.png" pageId="13" pageNumber="14">Fig. 7</figureCitation>
).
<typeStatus id="54F48830FFA5C20C875BF9BDFDF7F91F" box="[526,635,1653,1677]" pageId="13" pageNumber="14" type="paratype">Paratypes</typeStatus>
:
<collectionCode id="ED5EAE57FFA5C20C87DEF9BCFD6DF91E" LSID="urn:lsid:biocol.org:col:34992" box="[651,737,1652,1676]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="13" pageNumber="14">RMNH</collectionCode>
.
<collectionCode id="ED5EAE57FFA5C20C87B1F9BCFCBEF91E" box="[740,818,1652,1676]" pageId="13" pageNumber="14">RENA</collectionCode>
35253, 35297, 35317, males from a quarry in Coto de Caza San Martin near Ardia (
<collectingCountry id="F3587602FFA5C20C84BDF95FFDA0F922" box="[488,556,1687,1712]" name="Spain" pageId="13" pageNumber="14">Spain</collectingCountry>
, province of Pontevedra) [
<geoCoordinate id="EE7B5055FFA5C20C860CF95FFC5BF922" box="[857,983,1687,1712]" direction="north" orientation="latitude" pageId="13" pageNumber="14" precision="5" value="42.4553">42.4553°N</geoCoordinate>
/
<geoCoordinate id="EE7B5055FFA5C20C86A5F950FBE5F922" box="[1008,1129,1688,1712]" direction="west" orientation="longitude" pageId="13" pageNumber="14" precision="5" value="-8.8744">8.8744°W</geoCoordinate>
];
<collectionCode id="ED5EAE57FFA5C20C8129F95FFB34F922" box="[1148,1208,1687,1712]" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="13" pageNumber="14">EBD</collectionCode>
.16033, female from Hío, near Cangas (
<collectingCountry id="F3587602FFA5C20C843AF974FE3FF947" box="[367,435,1724,1749]" name="Spain" pageId="13" pageNumber="14">Spain</collectingCountry>
, province of Pontevedra) [
<geoCoordinate id="EE7B5055FFA5C20C87B0F974FCD8F946" box="[741,852,1724,1748]" direction="north" orientation="latitude" pageId="13" pageNumber="14" precision="55" value="42.271">42.271°N</geoCoordinate>
/ 8°.830°W];
<collectionCode id="ED5EAE57FFA5C20C86AFF974FBB9F946" box="[1018,1077,1724,1748]" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="13" pageNumber="14">EBD</collectionCode>
.9247, 9252, males,
<collectionCode id="ED5EAE57FFA5C20C8049F974FADAF946" box="[1308,1366,1724,1748]" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="13" pageNumber="14">EBD</collectionCode>
.9249- 9251, females, from Pouso da Serra, Donón (province of Pontevedra) [
<geoCoordinate id="EE7B5055FFA5C20C8686F917FBCFF96A" box="[979,1091,1759,1784]" direction="north" orientation="latitude" pageId="13" pageNumber="14" precision="55" value="42.275">42.275°N</geoCoordinate>
/
<geoCoordinate id="EE7B5055FFA5C20C8135F928FB46F96A" box="[1120,1226,1760,1784]" direction="west" orientation="longitude" pageId="13" pageNumber="14" precision="55" value="-8.847">8.847°W</geoCoordinate>
];
<collectionCode id="ED5EAE57FFA5C20C81B5F917FA95F96A" LSID="urn:lsid:biocol.org:col:12677" box="[1248,1305,1759,1784]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12677" name="Borough of Beverley Art Gallery and Museum" pageId="13" pageNumber="14">BEV</collectionCode>
.6299-6307, males,
<collectionCode id="ED5EAE57FFA5C20C85B3F8CCFE93F88F" LSID="urn:lsid:biocol.org:col:12677" box="[230,287,1796,1821]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12677" name="Borough of Beverley Art Gallery and Museum" pageId="13" pageNumber="14">BEV</collectionCode>
.6308, female, from Vila Real (
<collectingCountry id="F3587602FFA5C20C8722F8CCFD56F88F" box="[631,730,1796,1821]" name="Portugal" pageId="13" pageNumber="14">Portugal</collectingCountry>
, district of Vila Real) [
<geoCoordinate id="EE7B5055FFA5C20C868DF8CCFBCBF88E" box="[984,1095,1796,1820]" direction="north" orientation="latitude" pageId="13" pageNumber="14" precision="55" value="41.31">41.310°N</geoCoordinate>
/
<geoCoordinate id="EE7B5055FFA5C20C8108F8CCFB44F88E" box="[1117,1224,1796,1820]" direction="west" orientation="longitude" pageId="13" pageNumber="14" precision="55" value="-7.839">7.839°W</geoCoordinate>
].
</paragraph>
</subSubSection>
<subSubSection id="C3556519FFA5C20C8592F8EFFB0BF8F7" pageId="13" pageNumber="14" type="etymology">
<paragraph id="8BF03692FFA5C20C8592F8EFFB0BF8F7" blockId="13.[151,1436,1543,2037]" pageId="13" pageNumber="14">
<emphasis id="B93BEA80FFA5C20C8592F8EFFEC2F8D2" bold="true" box="[199,334,1831,1856]" pageId="13" pageNumber="14">Etymology.</emphasis>
the adjectival epithet
<emphasis id="B93BEA80FFA5C20C871EF8EFFD49F8D2" box="[587,709,1831,1856]" italics="true" pageId="13" pageNumber="14">lusitanicus</emphasis>
refers to the Lusitanians (latin
<emphasis id="B93BEA80FFA5C20C8179F8E1FB04F8D2" box="[1068,1160,1833,1856]" italics="true" pageId="13" pageNumber="14">Lusitani</emphasis>
), an ancient pre-Roman people inhabiting the centre and east of present-day
<collectingCountry id="F3587602FFA5C20C878AF884FCB2F8F7" box="[735,830,1868,1893]" name="Portugal" pageId="13" pageNumber="14">Portugal</collectingCountry>
(and adjacent area of
<collectingCountry id="F3587602FFA5C20C8162F884FBF0F8F7" box="[1079,1148,1868,1893]" name="Spain" pageId="13" pageNumber="14">Spain</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C3556519FFA5C2128592F8A7FB97FE8E" lastPageId="19" lastPageNumber="20" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="8BF03692FFA5C2118592F8A7FC65FE8F" blockId="13.[151,1436,1543,2037]" lastBlockId="16.[151,1436,151,285]" lastPageId="16" lastPageNumber="17" pageId="13" pageNumber="14">
<emphasis id="B93BEA80FFA5C20C8592F8A7FECDF81A" bold="true" box="[199,321,1903,1928]" pageId="13" pageNumber="14">Diagnosis.</emphasis>
This is the lineage referred to as
<taxonomicName id="4C4F4D11FFA5C20C8781F8A7FC30F81A" box="[724,956,1903,1928]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="hispanicus">
<emphasis id="B93BEA80FFA5C20C8781F8A7FC30F81A" box="[724,956,1903,1928]" italics="true" pageId="13" pageNumber="14">Podarcis hispanicus</emphasis>
</taxonomicName>
<typeStatus id="54F48830FFA5C20C8685F8B8FB8EF81A" box="[976,1026,1904,1928]" pageId="13" pageNumber="14">type</typeStatus>
1A” by
<bibRefCitation id="EFDE4B63FFA5C20C813BF8A7FAC8F81A" author="Pinho" box="[1134,1348,1903,1928]" pageId="13" pageNumber="14" refString="Pinho, C., Ferrand, N. &amp; Harris, D. J. (2006) Reexamination of the Iberian and North African Podarcis (Squamata: Lacertidae) phylogeny based on increased mitochondrial DNA sequencing. Molecular Phylogenetics and Evolution, 38, 266 - 273. http: // dx. doi. org / 10.1016 / j. ympev. 2005.06.012" type="journal article" year="2006">
Pinho
<emphasis id="B93BEA80FFA5C20C81EFF8B9FB7AF81A" box="[1210,1270,1903,1928]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
(2006
</bibRefCitation>
, 2007),
<bibRefCitation id="EFDE4B63FFA5C20C85C2F85CFEDFF83E" author="Carretero" box="[151,339,1940,1964]" pageId="13" pageNumber="14" refString="Carretero, M. A. (2008) An integrated assessment of a group with complex systematics: the Iberomaghrebian lizards genus Podarcis (Squamata, Lacertidae). Integrative Zoology, 4, 247 - 266. http: // dx. doi. org / 10.1111 / j. 1749 - 4877.2008.00102. x" type="journal article" year="2008">Carretero (2008)</bibRefCitation>
and
<bibRefCitation id="EFDE4B63FFA5C20C84DFF85CFD5CF83E" author="Kaliontzopoulou" box="[394,720,1940,1965]" pageId="13" pageNumber="14" refString="Kaliontzopoulou, A., Pinho, C., Harris, D. J. &amp; Carretero, M. A. (2011) When cryptic diversity blurs the picture: a cautionary tale from Iberian and North African Podarcis wall lizards. Biological Journal of the Linnean Society, 103, 779 - 800. http: // dx. doi. org / 10.1111 / j. 1095 - 8312.2011.01703. x" type="journal article" year="2011">
Kaliontzopoulou
<emphasis id="B93BEA80FFA5C20C8718F85DFD0AF83F" box="[589,646,1940,1965]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
(2011
</bibRefCitation>
,
<bibRefCitation id="EFDE4B63FFA5C20C878EF85CFC94F83E" author="Kaliontzopoulou" box="[731,792,1940,1964]" pageId="13" pageNumber="14" refString="Kaliontzopoulou, A., Carretero, M. A. &amp; Llorente, G. A. (2012) Morphology of the Podarcis wall lizards (Squamata: Lacertidae) from the Iberian Peninsula and North Africa: patterns of variation in a putative cryptic species complex. Zoological Journal of the Linnean Society, 164, 173 - 193. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2011.00760. x" type="journal article" year="2012">2012</bibRefCitation>
). A typical wall lizard of moderate size (adult males
<quantity id="4CB79B77FFA5C20C803EF85CFF4FF842" metricMagnitude="-2" metricUnit="m" metricValue="4.15" pageId="13" pageNumber="14" unit="mm" value="41.5">41.5 mm</quantity>
to
<quantity id="4CB79B77FFA5C20C85A3F870FEE8F842" box="[246,356,1976,2001]" metricMagnitude="-2" metricUnit="m" metricValue="6.25" pageId="13" pageNumber="14" unit="mm" value="62.5">62.5 mm</quantity>
, mean 51.5, adult females 40.0 mm to 60.0 mm, mean 48.7), very similar to
<taxonomicName id="4C4F4D11FFA5C20C806DF870FE43F867" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="subSpecies" species="guadarramae" subSpecies="guadarramae">
<emphasis id="B93BEA80FFA5C20C806DF870FE43F867" italics="true" pageId="13" pageNumber="14">Podarcis guadarramae guadarramae</emphasis>
</taxonomicName>
(see
<figureCitation id="13742A17FFA5C20C875CF814FDC7F866" box="[521,587,2012,2037]" captionStart="FIGURE 8" captionStartId="15.[151,250,1434,1456]" captionTargetBox="[151,1436,193,1412]" captionTargetId="figure@15.[151,1436,193,1413]" captionTargetPageId="15" captionText="FIGURE 8. Variation in males of Podarcis guadarramae lusitanicus. A) Portugal, Serra da Estrela, just north of the Lagoa de Viriato (district of Castelo Branco), 1579 m, 21.08. 2008 (PGe. 796). Photo O. Buisson. B) and C) Portugal, Serra da Estrela, top of the Vale Glacirio do Zêzere, crossroad N 338 x N 339, 1800 m (district of Guarda), 10.05. 1985 (respectively BEV. 3982 and 3981). Photos P. Geniez. D). Spain, 200 m E. Castro Baroña (42 km SW. Santiago, province of Pontevedra), 12.10. 2006 (PGe. 1035). Photo J. Speybroeck. E) and F). Spain, 1 km ESE. Viana do Bolo (province of Orense), 734 m, 1.05. 2012 (respectively PGe. 1134 and 1132). Photos J. Speybroeck (E) and W. Beukema (F)." httpUri="https://zenodo.org/record/285989/files/figure.png" pageId="13" pageNumber="14">Fig. 8</figureCitation>
) and often not safely identifiable based on present knowledge. On average has a slightly more flattened appearance with a flatter head (
<figureCitation id="13742A17FFA7C20E8612F9BEFC07F91D" box="[839,907,1654,1679]" captionStart="FIGURE 9" captionStartId="16.[151,250,1746,1768]" captionTargetBox="[455,1137,383,1708]" captionTargetId="figure@16.[455,1137,383,1708]" captionTargetPageId="16" captionText="FIGURE 9. Portraits of males of A) Podarcis guadarramae guadarramae. Spain, El Berrueco, in the village, 940 m (province of Ávila), 14.06. 2010 (BEV. 10937). Photo P. Geniez &amp; P. - A. Crochet. B) Podarcis guadarramae lusitanicus. Portugal, Serra da Estrela, Lagoa Comprida (district of Guarda) (PGe. 1024). Photo P. Geniez. C) Podarcis virescens. Spain, 1 km past Villanueva de los Escuderos towards Cuenca (province of Cuenca), 23.06. 2001 (holotype MNHN 2012.026). Photo P. Geniez." httpUri="https://zenodo.org/record/285990/files/figure.png" pageId="15" pageNumber="16">Fig. 9</figureCitation>
B); a lower number of femoral pores (13 to 21 for males, average 16.5, 12 to 19 for females, average 15.6) and lower number of dorsal scales (47 to 66 for males, average 56.9, 46 to 61 for females, average 53.4); “
<taxonomicName id="4C4F4D11FFA7C20E8787F976FCE9F945" box="[722,869,1726,1751]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="guadarramae">guadarramae</taxonomicName>
striped pattern” rare in females, when present pale dorsolateral stripes are narrower and less obvious than in
<taxonomicName id="4C4F4D11FFA7C20E864DF92BFC6BF969" box="[792,999,1762,1787]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="subSpecies" species="guadarramae" subSpecies="guadarramae">
<emphasis id="B93BEA80FFA7C20E864DF92BFC6BF969" box="[792,999,1762,1787]" italics="true" pageId="15" pageNumber="16">P. g. guadarramae</emphasis>
</taxonomicName>
(compare
<figureCitation id="13742A17FFA7C20E810BF92AFB22F969" box="[1118,1198,1762,1787]" captionStart="FIGURE 10" captionStartId="17.[151,250,1448,1470]" captionTargetBox="[151,1436,215,1415]" captionTargetId="figure@17.[151,1436,215,1416]" captionTargetPageId="17" captionText="FIGURE 10. Variation in females and juveniles of Podarcis guadarramae lusitanicus. A) and B) Females. Portugal, Serra da Estrela, top of the Vale Glaciário do Zêzere, crossroad N 338 x N 339, 1,800 m (district of Guarda), 10.05. 1985 (respectively BEV. 3986 and 3984). Photos P. Geniez. C) Female. Spain, Sierra de la Cabrera, Lago de Sanabria (province of Zamora), 23.10. 2009 (PGe. 1105). Photo M. Berroneau. D) Subadulte female. Spain, 1 km ESE. Viana do Bolo (province of Orense), 734 m, 1.05. 2012 (PGe. 1133). Photo W. Beukema. E) Juvenile. Spain, Sierra de la Cabrera, Lago de Sanabria (province of Zamora), 23.10. 2009 (PGe. 1117). Photo M. Berroneau. F) Newborn individual. Portugal, small hydroelectric power, 1 km above Montesinho, 1,000 m (district of Bragança), 14.08. 2003 (PGe. 782). Photo P. Geniez." httpUri="https://zenodo.org/record/285991/files/figure.png" pageId="15" pageNumber="16">Fig. 10</figureCitation>
B with
<figureCitation id="13742A17FFA7C20E81AAF92AFACDF969" box="[1279,1345,1762,1787]" captionStart="FIGURE 5" captionStartId="11.[151,250,1782,1804]" captionTargetBox="[151,1436,517,1760]" captionTargetId="figure@11.[151,1436,517,1761]" captionTargetPageId="11" captionText="FIGURE 5. Variation in females, undersides and juveniles of Podarcis guadarramae guadarramae. A) Female. Spain, El Tiemblo (province of Ávila). 27.06. 2001 (BEV. 2007). Photo P. Geniez. B) Female with the “ guadarramae striped pattern ”. Spain, Navalmoral (province of Ávila), 27.6. 2001 (BEV. 1999). Photo P. Geniez. C) Typical underside of a male, Spain, Navalilla, in the village (province of Segovia), 13.05. 1996 (BEV. 4487). Photo P. Geniez. D) Male with a white underside. Spain, San Martin de Valdeiglesias (province of Madrid), 27.06. 2001 (BEV. 2018). Photo P. Geniez. E) Juvenile male. Spain, San Martin de Valdeiglesias (province of Madrid), 27.06. 2001 (BEV. 2025). Photo P. Geniez. F) Juvenile female. Spain, La Alberca, Iglesia de Nuestra Señora de la Asunción (province of Salamanca), 18.09. 2007 (PGe. 790). Photo P. - A. Crochet." httpUri="https://zenodo.org/record/285986/files/figure.png" pageId="15" pageNumber="16">Fig. 5</figureCitation>
B), dark supradorsolateral stripes usually wider, leaving a narrower area of pale coloration in the middle of the back, in many adult males the dark supra-dorsolateral stripes can be very wide and very fragmented, invading the dorsal region, where they create a reticulated, dappled or ocellated pattern of light green, white or creamy spots on black back devoid of stripes (this pattern is typical to
<taxonomicName id="4C4F4D11FFA7C20E8790F8BBFCF0F819" box="[709,892,1906,1931]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="subSpecies" species="guadarramae" subSpecies="lusitanicus">
<emphasis id="B93BEA80FFA7C20E8790F8BBFCF0F819" box="[709,892,1906,1931]" italics="true" pageId="15" pageNumber="16">P. g. lusitanicus</emphasis>
</taxonomicName>
and seems to be unknown in other taxa of the
<taxonomicName id="4C4F4D11FFA7C20E85C2F85EFEF0F83D" box="[151,380,1942,1967]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="hispanicus">
<emphasis id="B93BEA80FFA7C20E85C2F85EFEF0F83D" box="[151,380,1942,1967]" italics="true" pageId="15" pageNumber="16">Podarcis hispanicus</emphasis>
</taxonomicName>
complex, see
<figureCitation id="13742A17FFA7C20E8774F85EFDFCF83C" box="[545,624,1942,1967]" captionStart="FIGURE 7" captionStartId="14.[151,250,1909,1931]" captionTargetBox="[341,1255,193,1892]" captionTargetId="figure@14.[341,1256,193,1893]" captionTargetPageId="14" captionText="FIGURE 7. Holotype of Podarcis guadarramae lusitanicus, male MNHN 2012.0263. Portugal, 1 km past Âncora towards Póvoa do Varzim (district of Viana do Castelo), 14.05. 1985. Photos P. Geniez. Note the very flattened head, the very large black spots on the pileus, the lack of vertebral stripe, the very broad black supradorsolateral stripes invading a large part of the pale green dorsum and the light dorsolateral stripes made of weakly elongated spots." httpUri="https://zenodo.org/record/285988/files/figure.png" pageId="15" pageNumber="16">Figs. 7</figureCitation>
&amp;
<figureCitation id="13742A17FFA7C20E87C2F85FFD29F83D" box="[663,677,1943,1967]" captionStart="FIGURE 8" captionStartId="15.[151,250,1434,1456]" captionTargetBox="[151,1436,193,1412]" captionTargetId="figure@15.[151,1436,193,1413]" captionTargetPageId="15" captionText="FIGURE 8. Variation in males of Podarcis guadarramae lusitanicus. A) Portugal, Serra da Estrela, just north of the Lagoa de Viriato (district of Castelo Branco), 1579 m, 21.08. 2008 (PGe. 796). Photo O. Buisson. B) and C) Portugal, Serra da Estrela, top of the Vale Glacirio do Zêzere, crossroad N 338 x N 339, 1800 m (district of Guarda), 10.05. 1985 (respectively BEV. 3982 and 3981). Photos P. Geniez. D). Spain, 200 m E. Castro Baroña (42 km SW. Santiago, province of Pontevedra), 12.10. 2006 (PGe. 1035). Photo J. Speybroeck. E) and F). Spain, 1 km ESE. Viana do Bolo (province of Orense), 734 m, 1.05. 2012 (respectively PGe. 1134 and 1132). Photos J. Speybroeck (E) and W. Beukema (F)." httpUri="https://zenodo.org/record/285989/files/figure.png" pageId="15" pageNumber="16">8</figureCitation>
B), pale spots inside the light dorsolateral stripes more contrasting and isolated as they are often positioned on a darker background, green-backed individuals not rare especially in mountains but also along the Atlantic coast. Diagnostic positions in the DNA sequence of the mitochondrial NADH dehydrogenase subunit 4 (ND4) gene relative to other lineages of the
<taxonomicName id="4C4F4D11FFB8C21186A1FF75FB0AFF47" box="[1012,1158,188,213]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="hispanicus">
<emphasis id="B93BEA80FFB8C21186A1FF75FB0AFF47" box="[1012,1158,188,213]" italics="true" pageId="16" pageNumber="17">P. hispanicus</emphasis>
</taxonomicName>
complex include an A at position 10905, C at position 11395 and A at position 11448 (positions numbered according to the
<taxonomicName id="4C4F4D11FFB8C2118072FF29FA17FF6A" box="[1319,1435,223,248]" class="Reptilia" family="Lacertidae" genus="Lacerta" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="muralis">
<emphasis id="B93BEA80FFB8C2118072FF29FA17FF6A" box="[1319,1435,223,248]" italics="true" pageId="16" pageNumber="17">P. muralis</emphasis>
</taxonomicName>
mitochondrion complete genome GenBank accession number NC_
<date id="FFF11052FFB8C21186D3FECCFC51FE8F" box="[902,989,260,285]" pageId="16" pageNumber="17" value="07-01-16">011607</date>
).
</paragraph>
<caption id="DF30661AFFA6C20F85C2F8BDFC32F875" httpUri="https://zenodo.org/record/285988/files/figure.png" pageId="14" pageNumber="15" targetBox="[341,1255,193,1892]" targetPageId="14">
<paragraph id="8BF03692FFA6C20F85C2F8BDFC32F875" blockId="14.[151,1436,1909,2023]" pageId="14" pageNumber="15">
<emphasis id="B93BEA80FFA6C20F85C2F8BDFE99F819" bold="true" box="[151,277,1909,1931]" pageId="14" pageNumber="15">FIGURE 7.</emphasis>
Holotype of
<taxonomicName id="4C4F4D11FFA6C20F84F1F8BDFC8BF819" box="[420,775,1909,1931]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="subSpecies" species="guadarramae" subSpecies="lusitanicus">
<emphasis id="B93BEA80FFA6C20F84F1F8BDFC8BF819" box="[420,775,1909,1931]" italics="true" pageId="14" pageNumber="15">Podarcis guadarramae lusitanicus</emphasis>
</taxonomicName>
, male MNHN 2012.0263. Portugal, 1 km past Âncora towards Póvoa do Varzim (district of Viana do Castelo), 14.05.1985. Photos P. Geniez. Note the very flattened head, the very large black spots on the pileus, the lack of vertebral stripe, the very broad black supradorsolateral stripes invading a large part of the pale green dorsum and the light dorsolateral stripes made of weakly elongated spots.
</paragraph>
</caption>
<caption id="DF30661AFFA7C20E85C2FA52FC56F9D8" httpUri="https://zenodo.org/record/285989/files/figure.png" pageId="15" pageNumber="16" targetBox="[151,1436,193,1412]" targetPageId="15">
<paragraph id="8BF03692FFA7C20E85C2FA52FC56F9D8" blockId="15.[151,1436,1434,1610]" pageId="15" pageNumber="16">
<emphasis id="B93BEA80FFA7C20E85C2FA52FE9FFA22" bold="true" box="[151,275,1434,1456]" pageId="15" pageNumber="16">FIGURE 8.</emphasis>
Variation in males of
<taxonomicName id="4C4F4D11FFA7C20E84A0FA52FCD9FA22" box="[501,853,1434,1456]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="subSpecies" species="guadarramae" subSpecies="lusitanicus">
<emphasis id="B93BEA80FFA7C20E84A0FA52FCD9FA22" box="[501,853,1434,1456]" italics="true" pageId="15" pageNumber="16">Podarcis guadarramae lusitanicus</emphasis>
</taxonomicName>
.
<emphasis id="B93BEA80FFA7C20E8637FA52FCF0FA22" bold="true" box="[866,892,1434,1456]" pageId="15" pageNumber="16">A)</emphasis>
Portugal, Serra da Estrela, just north of the Lagoa de Viriato (district of Castelo Branco), 1579 m, 21.08.2008 (PGe.796). Photo O. Buisson.
<emphasis id="B93BEA80FFA7C20E8151FA71FB92FA5D" bold="true" box="[1028,1054,1465,1487]" pageId="15" pageNumber="16">B)</emphasis>
and
<emphasis id="B93BEA80FFA7C20E811BFA71FBE5FA5D" bold="true" box="[1102,1129,1465,1487]" pageId="15" pageNumber="16">C)</emphasis>
Portugal, Serra da Estrela, top of the Vale Glacirio do Zêzere, crossroad N 338 x N 339, 1800 m (district of Guarda), 10.05.1985 (respectively BEV.3982 and 3981). Photos P. Geniez.
<emphasis id="B93BEA80FFA7C20E84F7FA3EFE4FF99E" bold="true" box="[418,451,1526,1548]" pageId="15" pageNumber="16">D).</emphasis>
Spain, 200 m E. Castro Baroña (42 km SW. Santiago, province of Pontevedra), 12.10.2006 (PGe.1035). Photo J. Speybroeck.
<emphasis id="B93BEA80FFA7C20E8752F9DDFDACF9B9" bold="true" box="[519,544,1557,1579]" pageId="15" pageNumber="16">E)</emphasis>
and
<emphasis id="B93BEA80FFA7C20E870FF9DDFDF4F9B9" bold="true" box="[602,632,1557,1579]" pageId="15" pageNumber="16">F).</emphasis>
Spain, 1 km ESE. Viana do Bolo (province of Orense), 734 m, 1.05.2012 (respectively PGe.1134 and 1132). Photos J. Speybroeck (E) and W. Beukema (F).
</paragraph>
</caption>
<caption id="DF30661AFFB8C21185C2F91AFA91F8D6" httpUri="https://zenodo.org/record/285990/files/figure.png" pageId="16" pageNumber="17" targetBox="[455,1137,383,1708]" targetPageId="16">
<paragraph id="8BF03692FFB8C21185C2F91AFA91F8D6" blockId="16.[151,1435,1746,1860]" pageId="16" pageNumber="17">
<emphasis id="B93BEA80FFB8C21185C2F91AFE9EF97A" bold="true" box="[151,274,1746,1768]" pageId="16" pageNumber="17">FIGURE 9.</emphasis>
Portraits of males of
<emphasis id="B93BEA80FFB8C21184BBF91AFD84F97A" bold="true" box="[494,520,1746,1768]" pageId="16" pageNumber="17">A)</emphasis>
<taxonomicName id="4C4F4D11FFB8C211875AF91AFC06F97A" box="[527,906,1746,1768]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="subSpecies" species="guadarramae" subSpecies="guadarramae">
<emphasis id="B93BEA80FFB8C211875AF91AFC06F97A" box="[527,906,1746,1768]" italics="true" pageId="16" pageNumber="17">Podarcis guadarramae guadarramae</emphasis>
</taxonomicName>
. Spain, El Berrueco, in the village, 940 m (province of Ávila), 14.06.2010 (BEV.10937). Photo P. Geniez &amp; P.-A. Crochet.
<emphasis id="B93BEA80FFB8C2118608F938FCFBF894" bold="true" box="[861,887,1776,1798]" pageId="16" pageNumber="17">B)</emphasis>
<taxonomicName id="4C4F4D11FFB8C2118629F938FB57F894" box="[892,1243,1776,1798]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="subSpecies" species="guadarramae" subSpecies="lusitanicus">
<emphasis id="B93BEA80FFB8C2118629F938FB57F894" box="[892,1243,1776,1798]" italics="true" pageId="16" pageNumber="17">Podarcis guadarramae lusitanicus</emphasis>
</taxonomicName>
. Portugal, Serra da Estrela, Lagoa Comprida (district of Guarda) (PGe.1024). Photo P. Geniez.
<emphasis id="B93BEA80FFB8C21186C1F8C7FC23F8B7" bold="true" box="[916,943,1807,1829]" pageId="16" pageNumber="17">C)</emphasis>
<taxonomicName id="4C4F4D11FFB8C21186E0F8C7FBFEF8B7" box="[949,1138,1807,1829]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="virescens">
<emphasis id="B93BEA80FFB8C21186E0F8C7FBFEF8B7" box="[949,1138,1807,1829]" italics="true" pageId="16" pageNumber="17">Podarcis virescens</emphasis>
</taxonomicName>
. Spain, 1 km past Villanueva de los Escuderos towards Cuenca (province of Cuenca), 23.06.2001 (holotype MNHN 2012.026). Photo P.Geniez.
</paragraph>
</caption>
<caption id="DF30661AFFB9C21085C2FA60FBBFF9E4" httpUri="https://zenodo.org/record/285991/files/figure.png" pageId="17" pageNumber="18" targetBox="[151,1436,215,1415]" targetPageId="17">
<paragraph id="8BF03692FFB9C21085C2FA60FBBFF9E4" blockId="17.[151,1436,1448,1654]" pageId="17" pageNumber="18">
<emphasis id="B93BEA80FFB9C21085C2FA60FEACFA2C" bold="true" box="[151,288,1448,1470]" pageId="17" pageNumber="18">FIGURE 10.</emphasis>
Variation in females and juveniles of
<taxonomicName id="4C4F4D11FFB9C21087F3FA60FB8BFA2C" box="[678,1031,1448,1470]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="subSpecies" species="guadarramae" subSpecies="lusitanicus">
<emphasis id="B93BEA80FFB9C21087F3FA60FB8BFA2C" box="[678,1031,1448,1470]" italics="true" pageId="17" pageNumber="18">Podarcis guadarramae lusitanicus</emphasis>
</taxonomicName>
.
<emphasis id="B93BEA80FFB9C2108141FA60FBA2FA2C" bold="true" box="[1044,1070,1448,1470]" pageId="17" pageNumber="18">A)</emphasis>
and
<emphasis id="B93BEA80FFB9C2108137FA60FBF6FA2C" bold="true" box="[1122,1146,1448,1470]" pageId="17" pageNumber="18">B)</emphasis>
Females. Portugal, Serra da Estrela, top of the Vale Glaciário do Zêzere, crossroad N 338 x N 339, 1,800 m (district of Guarda), 10.05.1985 (respectively BEV.3986 and 3984). Photos P. Geniez.
<emphasis id="B93BEA80FFB9C2108717FA2DFDD1FA69" bold="true" box="[578,605,1509,1531]" pageId="17" pageNumber="18">C)</emphasis>
Female. Spain, Sierra de la Cabrera, Lago de Sanabria (province of Zamora), 23.10.2009 (PGe.1105). Photo M. Berroneau.
<emphasis id="B93BEA80FFB9C210873CF9CCFD0FF988" bold="true" box="[617,643,1540,1562]" pageId="17" pageNumber="18">D)</emphasis>
Subadulte female. Spain, 1 km ESE. Viana do Bolo (province of Orense), 734 m, 1.05.2012 (PGe.1133). Photo W. Beukema.
<emphasis id="B93BEA80FFB9C21087C7F9EAFD27F9AA" bold="true" box="[658,683,1570,1592]" pageId="17" pageNumber="18">E)</emphasis>
Juvenile. Spain, Sierra de la Cabrera, Lago de Sanabria (province of Zamora), 23.10.2009 (PGe.1117). Photo M. Berroneau.
<emphasis id="B93BEA80FFB9C21087B7F989FD75F9C5" bold="true" box="[738,761,1601,1623]" pageId="17" pageNumber="18">F)</emphasis>
Newborn individual. Portugal, small hydroelectric power, 1 km above Montesinho, 1,000 m (district of Bragança), 14.08.2003 (PGe.782). Photo P. Geniez.
</paragraph>
</caption>
<paragraph id="8BF03692FFB9C2138592F96AFAC5FE8E" blockId="17.[151,1436,1698,2011]" lastBlockId="18.[151,1437,151,608]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<emphasis id="B93BEA80FFB9C2108592F96AFE21F929" bold="true" box="[199,429,1698,1723]" pageId="17" pageNumber="18">Range and ecology</emphasis>
(
<figureCitation id="13742A17FFB9C21084EBF96AFD88F929" box="[446,516,1698,1723]" captionStart="FIGURE 2" captionStartId="7.[151,250,1713,1735]" captionTargetBox="[151,1436,473,1701]" captionTargetId="figure@7.[151,1436,473,1703]" captionTargetPageId="7" captionText="FIGURE 2. Distribution of Podarcis types 1 A, 1 B and 2 based on specimens examined by the authors or on published or unpublished genetic data. See appendix 1 for list of records. Dashed line identifies the south-western limit of type 2 in areas where we have few records, based on published maps (see text). “ 1 A ”, “ 1 B ” and “ 2 ” indicate records supported by genetic data. Arrows identify type localities." httpUri="https://zenodo.org/record/285983/files/figure.png" pageId="17" pageNumber="18">Fig. 2</figureCitation>
): only in north-western
<collectingCountry id="F3587602FFB9C2108649F96AFCEEF929" box="[796,866,1698,1723]" name="Georgia" pageId="17" pageNumber="18">Iberia</collectingCountry>
: northern third of
<collectingCountry id="F3587602FFB9C2108162F96AFB15F929" box="[1079,1177,1698,1723]" name="Portugal" pageId="17" pageNumber="18">Portugal</collectingCountry>
, reaching the Serra da Estrela southwards (
<emphasis id="B93BEA80FFB9C210842EF90FFE1BF94D" box="[379,407,1735,1759]" italics="true" pageId="17" pageNumber="18">cf.</emphasis>
also “
<taxonomicName id="4C4F4D11FFB9C2108489F90EFD33F94D" box="[476,703,1734,1759]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="hispanicus">
<emphasis id="B93BEA80FFB9C2108489F90EFD33F94D" box="[476,703,1734,1759]" italics="true" pageId="17" pageNumber="18">Podarcis hispanicus</emphasis>
</taxonomicName>
1” in
<bibRefCitation id="EFDE4B63FFB9C2108656F90FFC21F94D" author="Sa-Sousa" box="[771,941,1735,1759]" pageId="17" pageNumber="18" refString="Sa-Sousa, P. (2000) A predictive distribution model for the Iberian wall lizard (Podarcis hispanicus) in Portugal. Herpetological Journal, 10, 1 - 11." type="journal article" year="2000">Sá-Sousa 2000</bibRefCitation>
), whole part of the Galicia region except the north, and north-western part of Castilla-y-León, reaching its northern limits near Ezaro (E of Cape Finisterre) along the Galician coast. Eastern limits poorly known but currently include the western slopes of the Puerto de Manzanal near Manzanal del Puerto (Léon,
<collectingCountry id="F3587602FFB9C21087DEF8FAFD40F8D9" box="[651,716,1842,1867]" name="Spain" pageId="17" pageNumber="18">Spain</collectingCountry>
) in the south and Oriñon (Santander,
<collectingCountry id="F3587602FFB9C210812FF8FAFB30F8D9" box="[1146,1212,1842,1867]" name="Spain" pageId="17" pageNumber="18">Spain</collectingCountry>
) along the northern coast of
<collectingCountry id="F3587602FFB9C21085AFF89EFEB1F8FD" box="[250,317,1878,1903]" name="Spain" pageId="17" pageNumber="18">Spain</collectingCountry>
. The north-western limit of the range of “
<taxonomicName id="4C4F4D11FFB9C210864AF89EFB88F8FD" box="[799,1028,1878,1903]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="hispanicus">
<emphasis id="B93BEA80FFB9C210864AF89EFB88F8FD" box="[799,1028,1878,1903]" italics="true" pageId="17" pageNumber="18">Podarcis hispanicus</emphasis>
</taxonomicName>
” (sensu lato) in Sá-Sousa &amp; Pérez- Mellado (2002) coincides with the limit of the range of
<taxonomicName id="4C4F4D11FFB9C2108657F8B2FB09F801" box="[770,1157,1914,1939]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="subSpecies" species="guadarramae" subSpecies="lusitanicus">
<emphasis id="B93BEA80FFB9C2108657F8B2FB09F801" box="[770,1157,1914,1939]" italics="true" pageId="17" pageNumber="18">Podarcis guadarramae lusitanicus</emphasis>
</taxonomicName>
(see also
<bibRefCitation id="EFDE4B63FFB9C21081A6F8B2FAF9F801" author="Galan" box="[1267,1397,1914,1939]" pageId="17" pageNumber="18" refString="Galan, P. (1986) Morfologia y distribucion del genero Podarcis Wagler, 1830 (Sauria, Lacertidae) en el noroeste de la Peninsula Iberica. Revista Espanola de Herpetologia., 1, 87 - 132." type="journal article" year="1986">Galán 1986</bibRefCitation>
for details on the northern-western limit in
<collectingCountry id="F3587602FFB9C21087D0F856FD46F825" box="[645,714,1950,1975]" name="Spain" pageId="17" pageNumber="18">Spain</collectingCountry>
).
<bibRefCitation id="EFDE4B63FFB9C21087B0F856FC68F825" author="Perez-Mellado" box="[741,996,1950,1975]" pageId="17" pageNumber="18" refString="Perez-Mellado, V. (2010) Podarcis hispanica (Steindachner, 1870). Lagartixa-iberica. In: Loureiro, A., Ferrand de Almeida, N., Carretero, M. A. &amp; Paulo, O. S. (Eds.), Atlas dos Anfibios e Repteis de Portugal, 2 nd Edition. Esfera do Caos, Lisboa, pp. 150 - 151." type="book chapter" year="2010">Pérez-Mellado (2010)</bibRefCitation>
provides a detailed account on the distribution in
<collectingCountry id="F3587602FFB9C2108469F80AFE17F849" box="[316,411,1986,2011]" name="Portugal" pageId="17" pageNumber="18">Portugal</collectingCountry>
(as
<taxonomicName id="4C4F4D11FFB9C210849EF80AFD22F849" box="[459,686,1986,2011]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="hispanicus">
<emphasis id="B93BEA80FFB9C210849EF80AFD22F849" box="[459,686,1986,2011]" italics="true" pageId="17" pageNumber="18">Podarcis hispanicus</emphasis>
</taxonomicName>
“morfotipo 1”: all Portuguese populations of
<typeStatus id="54F48830FFB9C21081E7F80BFB6EF849" box="[1202,1250,1987,2011]" pageId="17" pageNumber="18">type</typeStatus>
1 should be this taxon). Its ecology is very similar to the ecology of
<taxonomicName id="4C4F4D11FFBAC21387A5FF51FC48FF22" box="[752,964,151,176]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="subSpecies" species="guadarramae" subSpecies="guadarramae">
<emphasis id="B93BEA80FFBAC21387A5FF51FC48FF22" box="[752,964,151,176]" italics="true" pageId="18" pageNumber="19">P. g. guadarramae</emphasis>
</taxonomicName>
but it reaches sea level along the coasts. Competition with
<taxonomicName id="4C4F4D11FFBAC2138430FF75FE56FF47" box="[357,474,188,213]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="bocagei">
<emphasis id="B93BEA80FFBAC2138430FF75FE56FF47" box="[357,474,188,213]" italics="true" pageId="18" pageNumber="19">P. bocagei</emphasis>
</taxonomicName>
in the northwestern part of its range often excludes it from terrestrial and man-made habitats (stone walls for example), restricting it to natural rocky habitats in these areas (pers. obs., see also
<bibRefCitation id="EFDE4B63FFBAC213800CFF17FF5FFE8F" author="Galan" pageId="18" pageNumber="19" refString="Galan, P. (1986) Morfologia y distribucion del genero Podarcis Wagler, 1830 (Sauria, Lacertidae) en el noroeste de la Peninsula Iberica. Revista Espanola de Herpetologia., 1, 87 - 132." type="journal article" year="1986">Galán 1986</bibRefCitation>
). On small islands off Galicia the two species usually exclude each other (
<bibRefCitation id="EFDE4B63FFBAC2138144FECCFAB1FE8E" author="Arntzen" box="[1041,1341,260,284]" pageId="18" pageNumber="19" refString="Arntzen, J. W. &amp; Sa-Sousa, P. (2007) Morphological and genetical differentiation of lizards (Podarcis bocagei and P. hispanica) in the Ria de Arosa Archipelago (Galicia, Spain) resulting from vicariance and occasional dispersal. In: Renema, W. (Ed.), Biogeography, time, and place: distributions, barriers, and islands. Springer, New York &amp; Heidelberg, pp. 365 - 401." type="book chapter" year="2007">Arntzen &amp; Sá-Sousa 2007</bibRefCitation>
).
</paragraph>
<paragraph id="8BF03692FFBAC2138592FEEFFE2FFDF2" blockId="18.[151,1437,151,608]" pageId="18" pageNumber="19">
<emphasis id="B93BEA80FFBAC2138592FEEFFE60FED2" bold="true" box="[199,492,295,320]" pageId="18" pageNumber="19">Geographical variation.</emphasis>
Populations in the north-eastern part of the range of
<emphasis id="B93BEA80FFBAC21381C5FEEFFE9BFEF7" italics="true" pageId="18" pageNumber="19">
<taxonomicName id="4C4F4D11FFBAC21381C5FEEFFE9FFEF7" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="subSpecies" species="guadarramae" subSpecies="lusitanicus">Podarcis guadarramae lusitanicus</taxonomicName>
,
</emphasis>
including most of the Cantabrian Mountains area, roughly from Medulas and Carrucedo in
<collectingCountry id="F3587602FFBAC2138079FE84FAE0FEF7" box="[1324,1388,332,357]" name="Spain" pageId="18" pageNumber="19">Spain</collectingCountry>
and eastern piedmonts of the Serra de Montesinho in north-east
<collectingCountry id="F3587602FFBAC213866FFEA7FC15FE1A" box="[826,921,367,392]" name="Portugal" pageId="18" pageNumber="19">Portugal</collectingCountry>
to Picos de Europas, especially north of Caín in the Ruta del Cares, in
<collectingCountry id="F3587602FFBAC21384E0FE5CFE74FE3F" box="[437,504,404,429]" name="Spain" pageId="18" pageNumber="19">Spain</collectingCountry>
, are on average smaller (
<quantity id="4CB79B77FFBAC2138640FE5CFC32FE3E" box="[789,958,404,429]" metricMagnitude="-2" metricUnit="m" metricValue="4.975" metricValueMax="5.8" metricValueMin="4.15" pageId="18" pageNumber="19" unit="mm" value="49.75" valueMax="58.0" valueMin="41.5">41.5 to 58 mm</quantity>
of SVL for males, average 49.6,
<quantity id="4CB79B77FFBAC2138068FE5CFF4FFE5D" metricMagnitude="-2" metricUnit="m" metricValue="4.85" metricValueMax="5.7" metricValueMin="4.0" pageId="18" pageNumber="19" unit="mm" value="48.5" valueMax="57.0" valueMin="40.0">40 to 57 mm</quantity>
for females, average 47.2), with a lower number of dorsal scales (47 to 63 for males, average 54.5, 46 to 58 for females, average 52.2) and a larger number of transverse rows of ventral plates (25 to 30 for males, average 27.3, 29 to 35 for females, average 30.9); coloration is often quite distinctive, with a more finely dappled or punctuated and less contrasted dorsal pattern with reduced dark supradorsolateral stripes, pale dorsolateral stripes less contrasted (see
<figureCitation id="13742A17FFBAC2138410FD8FFE19FDF2" box="[325,405,583,608]" captionStart="FIGURE 11" captionStartId="18.[151,250,1595,1617]" captionTargetBox="[151,1436,665,1550]" captionTargetId="figure@18.[151,1436,665,1551]" captionTargetPageId="18" captionText="FIGURE 11. Podarcis guadarramae lusitanicus, Cantabric form. A), B) and D) Respectively two males and a juvenile. Spain, Picos de Europa, Desfiladero de Cares (= Senda del Cares), in the province of Santander, 25.04. 2013 (respectively PGe. 1088, 1087 and 1090). Photos M. Ricordel. C) Male. Portugal, eastern piedmonts of the Serra de Montesinho, 8 km past Rio de Onor towards Varge (district of Bragança), 15.8. 2003 (BEV. 8331). Photo P. Geniez." httpUri="https://zenodo.org/record/285992/files/figure.png" pageId="18" pageNumber="19">Fig. 11</figureCitation>
).
</paragraph>
<caption id="DF30661AFFBAC21385C2F9F3FC21F93E" httpUri="https://zenodo.org/record/285992/files/figure.png" pageId="18" pageNumber="19" targetBox="[151,1436,665,1550]" targetPageId="18">
<paragraph id="8BF03692FFBAC21385C2F9F3FC21F93E" blockId="18.[151,1436,1595,1708]" pageId="18" pageNumber="19">
<emphasis id="B93BEA80FFBAC21385C2F9F3FE92F9C3" bold="true" box="[151,286,1595,1617]" pageId="18" pageNumber="19">FIGURE 11.</emphasis>
<taxonomicName id="4C4F4D11FFBAC2138471F9F3FD7EF9C3" authority="Cantabric" authorityName="Cantabric" box="[292,754,1595,1617]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="subSpecies" species="guadarramae" subSpecies="lusitanicus">
<emphasis id="B93BEA80FFBAC2138471F9F3FD0FF9C3" box="[292,643,1595,1617]" italics="true" pageId="18" pageNumber="19">Podarcis guadarramae lusitanicus</emphasis>
, Cantabric
</taxonomicName>
form.
<emphasis id="B93BEA80FFBAC2138662F9F3FCDDF9C3" bold="true" box="[823,849,1595,1617]" pageId="18" pageNumber="19">A)</emphasis>
,
<emphasis id="B93BEA80FFBAC213860BF9F3FCF4F9C3" bold="true" box="[862,888,1595,1617]" pageId="18" pageNumber="19">B)</emphasis>
and
<emphasis id="B93BEA80FFBAC21386FCF9F3FC48F9C3" bold="true" box="[937,964,1595,1617]" pageId="18" pageNumber="19">D)</emphasis>
Respectively two males and a juvenile. Spain, Picos de Europa, Desfiladero de Cares (= Senda del Cares), in the province of Santander, 25.04.2013 (respectively PGe.1088, 1087 and 1090). Photos M. Ricordel.
<emphasis id="B93BEA80FFBAC2138741F9B0FDA3F91C" bold="true" box="[532,559,1656,1678]" pageId="18" pageNumber="19">C)</emphasis>
Male. Portugal, eastern piedmonts of the Serra de Montesinho, 8 km past Rio de Onor towards Varge (district of Bragança), 15.8.2003 (BEV.8331). Photo P. Geniez.
</paragraph>
</caption>
<paragraph id="8BF03692FFBAC2128592F91CFE46FF65" blockId="18.[151,1436,1748,2025]" lastBlockId="19.[151,1436,151,752]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<emphasis id="B93BEA80FFBAC2138592F91CFD32F97F" bold="true" box="[199,702,1748,1773]" pageId="18" pageNumber="19">Situation in contact zones with other taxa.</emphasis>
Largely sympatric with
<taxonomicName id="4C4F4D11FFBAC2138687F91CFB19F97F" box="[978,1173,1748,1773]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="bocagei">
<emphasis id="B93BEA80FFBAC2138687F91CFB19F97F" box="[978,1173,1748,1773]" italics="true" pageId="18" pageNumber="19">Podarcis bocagei</emphasis>
</taxonomicName>
in the north-west of its range (see
<bibRefCitation id="EFDE4B63FFBAC213845AF930FE61F882" author="Loureiro" box="[271,493,1784,1809]" pageId="18" pageNumber="19" refString="Loureiro, A., Ferrand de Almeida, N., Carretero, M. A. &amp; Paulo, O. S. (Eds.) (2010) Atlas dos Anfibios e Repteis de Portugal, 2 nd Edition. Esfera do Caos: Lisboa, 252 pp." type="book" year="2010">
Loureiro
<emphasis id="B93BEA80FFBAC2138422F931FE23F883" box="[375,431,1784,1809]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
2010
</bibRefCitation>
for
<collectingCountry id="F3587602FFBAC213874EF930FDF6F883" box="[539,634,1784,1809]" name="Portugal" pageId="18" pageNumber="19">Portugal</collectingCountry>
and Pleguezuelos
<emphasis id="B93BEA80FFBAC213861FF931FC0DF883" box="[842,897,1784,1809]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
2002 for
<collectingCountry id="F3587602FFBAC21386B8F930FBBEF883" box="[1005,1074,1784,1809]" name="Spain" pageId="18" pageNumber="19">Spain</collectingCountry>
). Reproductive isolation relative to this species is nearly complete but a few individuals of recent mixed ancestry have been detected, as well as some evidence of ancient introgression (
<bibRefCitation id="EFDE4B63FFBAC2138734F888FC1CF8CA" author="Arntzen" box="[609,912,1856,1880]" pageId="18" pageNumber="19" refString="Arntzen, J. W. &amp; Sa-Sousa, P. (2007) Morphological and genetical differentiation of lizards (Podarcis bocagei and P. hispanica) in the Ria de Arosa Archipelago (Galicia, Spain) resulting from vicariance and occasional dispersal. In: Renema, W. (Ed.), Biogeography, time, and place: distributions, barriers, and islands. Springer, New York &amp; Heidelberg, pp. 365 - 401." type="book chapter" year="2007">Arntzen &amp; Sá-Sousa 2007</bibRefCitation>
; see also Pinho
<emphasis id="B93BEA80FFBAC213811DF889FB0EF8CB" box="[1096,1154,1856,1881]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
2007, 2008). Sympatric, sometimes syntopic, with
<taxonomicName id="4C4F4D11FFBAC21384E9F8ADFDC6F8EF" box="[444,586,1892,1917]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="carbonelli">
<emphasis id="B93BEA80FFBAC21384E9F8ADFDC6F8EF" box="[444,586,1892,1917]" italics="true" pageId="18" pageNumber="19">P. carbonelli</emphasis>
</taxonomicName>
in the north of
<collectingCountry id="F3587602FFBAC21387AFF8ACFCD5F8EF" box="[762,857,1892,1917]" name="Portugal" pageId="18" pageNumber="19">Portugal</collectingCountry>
(pers. obs., see also
<bibRefCitation id="EFDE4B63FFBAC2138117F8ACFAABF8EE" author="Loureiro" box="[1090,1319,1892,1917]" pageId="18" pageNumber="19" refString="Loureiro, A., Ferrand de Almeida, N., Carretero, M. A. &amp; Paulo, O. S. (Eds.) (2010) Atlas dos Anfibios e Repteis de Portugal, 2 nd Edition. Esfera do Caos: Lisboa, 252 pp." type="book" year="2010">
Loureiro
<emphasis id="B93BEA80FFBAC21381FEF8ADFB68F8EF" box="[1195,1252,1892,1917]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
2010
</bibRefCitation>
). Globally parapatric with
<emphasis id="B93BEA80FFBAC213841DF841FE4BF832" box="[328,455,1929,1952]" italics="true" pageId="18" pageNumber="19">P. v i re s c en s</emphasis>
in
<collectingCountry id="F3587602FFBAC21384A4F840FDDCF833" box="[497,592,1928,1953]" name="Portugal" pageId="18" pageNumber="19">Portugal</collectingCountry>
(
<bibRefCitation id="EFDE4B63FFBAC213870AF840FC87F832" author="Sa-Sousa" box="[607,779,1928,1952]" pageId="18" pageNumber="19" refString="Sa-Sousa, P. (2000) A predictive distribution model for the Iberian wall lizard (Podarcis hispanicus) in Portugal. Herpetological Journal, 10, 1 - 11." type="journal article" year="2000">Sá-Sousa 2000</bibRefCitation>
,
<bibRefCitation id="EFDE4B63FFBAC2138642F840FBA1F832" author="Harris" box="[791,1069,1928,1953]" pageId="18" pageNumber="19" refString="Harris, D. J. &amp; Sa-Sousa, P. (2001) Species distinction and relationships of the Western Iberian Podarcis lizards (Reptilia, Lacertidae) based on morphology and mitochondrial DNA sequences. Herpetological Journal, 11, 129 - 136." type="journal article" year="2001">Harris &amp; Sá-Sousa 2001</bibRefCitation>
, Sá-Sousa
<emphasis id="B93BEA80FFBAC21381FCF841FB6EF833" box="[1193,1250,1928,1953]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
2002) but a few syntopic populations have been described (see
<bibRefCitation id="EFDE4B63FFBAC21387F5F864FC04F856" author="Perez-Mellado" box="[672,904,1964,1989]" pageId="18" pageNumber="19" refString="Perez-Mellado, V. (2010) Podarcis hispanica (Steindachner, 1870). Lagartixa-iberica. In: Loureiro, A., Ferrand de Almeida, N., Carretero, M. A. &amp; Paulo, O. S. (Eds.), Atlas dos Anfibios e Repteis de Portugal, 2 nd Edition. Esfera do Caos, Lisboa, pp. 150 - 151." type="book chapter" year="2010">Pérez-Mellado 2010</bibRefCitation>
). In spite of this,
<bibRefCitation id="EFDE4B63FFBAC2138113F864FA9AF856" box="[1094,1302,1964,1989]" pageId="18" pageNumber="19" refString="Pinho, C., Harris, D. J. &amp; Ferrand, N. (2008) Non-equilibrium estimates of gene flow inferred from nuclear genealogies suggest that Iberian and North African wall lizards (Podarcis spp.) are an assemblage of incipient species. BMC Evolutionary Biology, 8, 63. http: // dx. doi. org / 10.1186 / 1471 - 2148 - 8 - 63" type="journal article">
Pinho
<emphasis id="B93BEA80FFBAC21381DBF865FB49F857" box="[1166,1221,1964,1989]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
(2008)
</bibRefCitation>
detected no trace of nuclear gene flow between
<taxonomicName id="4C4F4D11FFBAC213877CF819FD57F87B" box="[553,731,2000,2025]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="subSpecies" species="guadarramae" subSpecies="lusitanicus">
<emphasis id="B93BEA80FFBAC213877CF819FD57F87B" box="[553,731,2000,2025]" italics="true" pageId="18" pageNumber="19">P. g. lusitanicus</emphasis>
</taxonomicName>
and
<emphasis id="B93BEA80FFBAC2138647F819FC1DF87A" box="[786,913,2001,2024]" italics="true" pageId="18" pageNumber="19">P. v i re s c en s</emphasis>
. Sá-Sousa
<emphasis id="B93BEA80FFBAC2138146F819FBC0F87B" box="[1043,1100,2000,2025]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
(2002) found specimens with
<typeStatus id="54F48830FFBBC21285C2FF50FF4BFF22" box="[151,199,152,176]" pageId="19" pageNumber="20">type</typeStatus>
2 morphological characters in the Serra da Estrela but as this population is far from the contact zone with this species, it is unlikely that this results from introgression. We have no information on contact zones with
<taxonomicName id="4C4F4D11FFBBC212803DFF75FEA3FF6A" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="subSpecies" species="guadarramae" subSpecies="guadarramae">
<emphasis id="B93BEA80FFBBC212803DFF75FEA3FF6A" italics="true" pageId="19" pageNumber="20">P. g. guadarramae</emphasis>
</taxonomicName>
or
<taxonomicName id="4C4F4D11FFBBC2128401FF29FE4FFF6A" box="[340,451,223,248]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="liolepis">
<emphasis id="B93BEA80FFBBC2128401FF29FE4FFF6A" box="[340,451,223,248]" italics="true" pageId="19" pageNumber="20">P. liolepis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF03692FFBBC2128592FECCFB97FE8E" blockId="19.[151,1436,151,752]" box="[199,1051,260,285]" pageId="19" pageNumber="20">
<emphasis id="B93BEA80FFBBC2128592FECCFDB2FE8F" bold="true" box="[199,574,260,285]" pageId="19" pageNumber="20">Comparison with other species.</emphasis>
See
<taxonomicName id="4C4F4D11FFBBC2128721FECCFC80FE8F" box="[628,780,260,285]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="guadarramae">
<emphasis id="B93BEA80FFBBC2128721FECCFC80FE8F" box="[628,780,260,285]" italics="true" pageId="19" pageNumber="20">guadarramae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4F4D11FFBBC2128616FECDFC26FE8E" box="[835,938,261,284]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="virescens">
<emphasis id="B93BEA80FFBBC2128616FECDFC26FE8E" box="[835,938,261,284]" italics="true" pageId="19" pageNumber="20">virescens</emphasis>
</taxonomicName>
accounts.
</paragraph>
</subSubSection>
<subSubSection id="C3556519FFBBC2128592FEEFFD70FD62" pageId="19" pageNumber="20" type="description">
<paragraph id="8BF03692FFBBC2128592FEEFFD70FD62" blockId="19.[151,1436,151,752]" pageId="19" pageNumber="20">
<emphasis id="B93BEA80FFBBC2128592FEEFFD83FED2" bold="true" box="[199,527,295,320]" pageId="19" pageNumber="20">
Description of the
<typeStatus id="54F48830FFBBC21284F7FEEFFD87FED2" box="[418,523,295,320]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
.
</emphasis>
An adult male (
<figureCitation id="13742A17FFBBC2128797FEEFFC8AFED2" box="[706,774,295,320]" captionStart="FIGURE 7" captionStartId="14.[151,250,1909,1931]" captionTargetBox="[341,1255,193,1892]" captionTargetId="figure@14.[341,1256,193,1893]" captionTargetPageId="14" captionText="FIGURE 7. Holotype of Podarcis guadarramae lusitanicus, male MNHN 2012.0263. Portugal, 1 km past Âncora towards Póvoa do Varzim (district of Viana do Castelo), 14.05. 1985. Photos P. Geniez. Note the very flattened head, the very large black spots on the pileus, the lack of vertebral stripe, the very broad black supradorsolateral stripes invading a large part of the pale green dorsum and the light dorsolateral stripes made of weakly elongated spots." httpUri="https://zenodo.org/record/285988/files/figure.png" pageId="19" pageNumber="20">Fig. 7</figureCitation>
) measuring
<quantity id="4CB79B77FFBBC21286C5FEEFFC52FEAD" box="[912,990,295,320]" metricMagnitude="-2" metricUnit="m" metricValue="5.2" pageId="19" pageNumber="20" unit="mm" value="52.0">52 mm</quantity>
of snout-vent length,
<quantity id="4CB79B77FFBBC2128186FEE0FAB9FEAD" box="[1235,1333,296,320]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="19" pageNumber="20" unit="mm" value="11.4">11.4 mm</quantity>
of pileus length,
<quantity id="4CB79B77FFBBC21285BEFE84FECDFEF6" box="[235,321,332,357]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="19" pageNumber="20" unit="mm" value="5.5">5.5 mm</quantity>
of pileus width,
<quantity id="4CB79B77FFBBC21284AAFE84FDD9FEF6" box="[511,597,332,357]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="19" pageNumber="20" unit="mm" value="5.1">5.1 mm</quantity>
of head height (giving a height / length head ratio of 0.93), and having the following scalation features: 60 longitudinal rows of dorsal scales at mid-body, 32 gular scales counted along a line from the contact between the fourth pair of maxillary scales to the collar, 27 transversal rows of ventral plates from the collar to the anal plates, 16 femoral pores on each side, 23 subdigital lamellae beneath the fourth toe, 93 scales on the temporal area, one small but well-defined masseteric shield on each side. Coloration on the live animal: iris whitish; pileus strongly black-marked; dorsum mainly black with a vestigial pale central area showing as a row of pale green spots along mid-dorsum, a row of pale cream spots along the dorsolateral area and flanks black with numerous large creamy spots; tail mainly regenerated, with basis not strongly widened and coloration similar to the body for the original tail, mainly pale brown with a black longitudinal lateral stripe for the regenerated part; central and median ventral plates uniform, marginal ventral plates with triangular black marks. Ventral coloration on the live specimen has not been photographed. Coloration of the preserved specimen is similar but pale areas on the back and flanks now have various shades of light grey.
</paragraph>
</subSubSection>
</treatment>
</document>