219 lines
30 KiB
XML
219 lines
30 KiB
XML
<document id="D11F53281EE23C0A9151A5286ECCF327" ID-DOI="10.5252/zoosystema2018v40a12" ID-GBIF-Dataset="6c3d39df-7484-4878-8952-9c6a0d926465" ID-ISSN="1638-9387" ID-Zenodo-Dep="3741548" ID-ZooBank="urn:lsid:zoobank.org:pub:6CBF9039-7E4D-4938-909A-2DB5113C8D35" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1586173524968" checkinUser="felipe" docAuthor="Salazar-Vallejo, Sergio I." docDate="2018" docId="C17687BDFFFFFFD7FC3EFAB6DBEFFD5B" docLanguage="en" docName="zoosystema.40.12.227-325.pdf.imf" docOrigin="Zoosystema 40 (12)" docStyle="DocumentStyle:AFAF64E709DD7C8340B70A24A5804402.4:Zoosystema.2018-2019.journal_article" docStyleId="AFAF64E709DD7C8340B70A24A5804402" docStyleName="Zoosystema.2018-2019.journal_article" docStyleVersion="4" docTitle="Hesione helenensis Salazar-Vallejo 2018, n. sp." docType="treatment" docVersion="5" lastPageNumber="263" masterDocId="3D4FFFC5FFDBFFF1FFD4FFA3DA42FFE9" masterDocTitle="Revision of Hesione Savigny in Lamarck, 1818 (Annelida, Errantia, Hesionidae)" masterLastPageNumber="325" masterPageNumber="227" pageNumber="261" updateTime="1698735891220" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:mods id="9C6D5F08C4F635055A7DF30D5B811223" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="4DE7C59B568DD9FAD19B0AF0092C0491">
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<mods:title id="A54C102943CB4AB87067C7E3EFB9C7AE">Revision of Hesione Savigny in Lamarck, 1818 (Annelida, Errantia, Hesionidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="065358952F0FCBEE52E5ABFCED1939FA" type="personal">
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<mods:role id="1FDA6DD64749973703423AFA92A1ACC5">
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<mods:roleTerm id="95D8D2AE6ABFAAD2DEE629892F84447E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B4BF290271FB77419BFB77B7F9D57721">Salazar-Vallejo, Sergio I.</mods:namePart>
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<mods:typeOfResource id="A037C6767FE21D3F56031B4FC899D041">text</mods:typeOfResource>
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<mods:titleInfo id="FBB8FDDB127829D7C3B06AA388BAB6F4">
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<mods:title id="F8638A691AA180FB02E022DE44F1B976">Zoosystema</mods:title>
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</mods:titleInfo>
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<mods:part id="17D7DB01C1A917A5827B155386521CA1">
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<mods:date id="0B0A113365282F7546064DF82A262C21">2018</mods:date>
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<mods:detail id="A55ECF7255B1657FC4D38914E17A8FB5" type="pubDate">
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<mods:number id="E157FD43DF391C14EFC70F4964C64FCD">2018-06-19</mods:number>
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<mods:detail id="C4FB4ABA9B76A16E178008A3FA708550" type="volume">
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<mods:number id="650414339A6CE3F87A8AD3A126209E7E">40</mods:number>
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</mods:detail>
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<mods:detail id="4149C1617AA2291AB2CFCB7E77E4CADA" type="issue">
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<mods:number id="58163D117FB827C850674F407B9A9093">12</mods:number>
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<mods:end id="E050A894F0800A07741D61CF97E39B20">325</mods:end>
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<mods:classification id="61646B95A2826FD97F078D1F47E08E1C">journal article</mods:classification>
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<mods:identifier id="B86F03869B9822B3F0B852B782589607" type="DOI">10.5252/zoosystema2018v40a12</mods:identifier>
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<mods:identifier id="6F21C70ED86DD6F85ED31BF5605A251E" type="GBIF-Dataset">6c3d39df-7484-4878-8952-9c6a0d926465</mods:identifier>
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<mods:identifier id="EC4777591F31F5E47C72ABC76C86693B" type="ISSN">1638-9387</mods:identifier>
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<mods:identifier id="8035FD7321680EFA9904E01B06CDEE2D" type="Zenodo-Dep">3741548</mods:identifier>
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<mods:identifier id="6D53D60D44EB07386B69B2D913D8218F" type="ZooBank">urn:lsid:zoobank.org:pub:6CBF9039-7E4D-4938-909A-2DB5113C8D35</mods:identifier>
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</mods:mods>
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<treatment id="C17687BDFFFFFFD7FC3EFAB6DBEFFD5B" ID-DOI="http://doi.org/10.5281/zenodo.3810264" ID-GBIF-Taxon="163437897" ID-Zenodo-Dep="3810264" LSID="urn:lsid:plazi:treatment:C17687BDFFFFFFD7FC3EFAB6DBEFFD5B" httpUri="http://treatment.plazi.org/id/C17687BDFFFFFFD7FC3EFAB6DBEFFD5B" lastPageId="38" lastPageNumber="263" pageId="36" pageNumber="261">
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<subSubSection id="01C56520FFFFFFD5FC3EFAB6DEB0FAD9" box="[1002,1266,1301,1328]" pageId="36" pageNumber="261" type="nomenclature">
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<paragraph id="496036ABFFFFFFD5FC3EFAB6DEB0FAD9" blockId="36.[1002,1266,1301,1359]" box="[1002,1266,1301,1328]" pageId="36" pageNumber="261">
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<heading id="122881C7FFFFFFD5FC3EFAB6DEB0FAD9" box="[1002,1266,1301,1328]" centered="true" fontSize="11" level="2" pageId="36" pageNumber="261" reason="2">
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<taxonomicName id="8EDF4D28FFFFFFD5FC3EFAB6DEF2FAD9" ID-CoL="6LRM8" authority="Salazar-Vallejo, 2018" authorityName="Salazar-Vallejo" authorityYear="2018" box="[1002,1200,1301,1328]" class="Polychaeta" family="Hesionidae" genus="Hesione" kingdom="Animalia" order="Phyllodocida" pageId="36" pageNumber="261" phylum="Annelida" rank="species" species="helenensis" status="sp. nov.">
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<emphasis id="7BABEAB9FFFFFFD5FC3EFAB6DEF2FAD9" bold="true" box="[1002,1200,1301,1328]" italics="true" pageId="36" pageNumber="261">Hesione helenensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="609857C2FFFFFFD5FB63FAB5DEB0FAD9" box="[1207,1266,1302,1328]" pageId="36" pageNumber="261" rank="species">n. sp.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="01C56520FFFFFFD5FBEBFA96DFD0FA6D" pageId="36" pageNumber="261" type="description">
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<paragraph id="496036ABFFFFFFD5FBEBFA96DEDFFAA6" blockId="36.[1002,1266,1301,1359]" box="[1087,1181,1333,1359]" pageId="36" pageNumber="261">
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(
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<figureCitation id="D1E42A2EFFFFFFD5FB93FA96DED7FAA6" box="[1095,1173,1333,1359]" captionStart="FIG" captionStartId="37.[132,143,1300,1318]" captionTargetBox="[172,1414,215,1255]" captionTargetId="figure@37.[172,1415,214,1255]" captionTargetPageId="37" captionText="FIG. 20. — Hesione helenensis n. sp., holotype,USNM 33148:A, anterior region, dorsal view, pharynx partially exposed; B, prostomium, dorsal view (arrow points to left antenna);C, chaetiger 9, left parapodium, anterior view (insets:acicular lobe and bases of cirrostyles); D, same,neurochaetae (insets: blades); E, posterior region, dorsal view. Scale bars: A, 1.7 mm; B, 0.2 mm; C, 0.5 mm; D, 160 µm; E, 1.8 mm." figureDoi="http://doi.org/10.5281/zenodo.3741590" httpUri="https://zenodo.org/record/3741590/files/figure.png" pageId="36" pageNumber="261">Fig. 20</figureCitation>
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)
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</paragraph>
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<paragraph id="496036ABFFFFFFD5FC9EFAD1DFD0FA6D" blockId="36.[842,1426,1394,1412]" box="[842,1426,1394,1412]" pageId="36" pageNumber="261">
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<uri id="3D4E3AA9FFFFFFD5FC9EFAD1DFD0FA6D" box="[842,1426,1394,1412]" pageId="36" pageNumber="261">
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urn:lsid:zoobank.org:act:
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<uuid id="3D790C7EFFFFFFD5FBC6FAD1DFD0FA6D" box="[1042,1426,1394,1412]" pageId="36" pageNumber="261">F44B445E-71C8-493B-B59F-2B193435A522</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="01C56520FFFFFFD5FCF9FA00DFD5F99D" pageId="36" pageNumber="261" type="materials_examined">
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<paragraph id="496036ABFFFFFFD5FCF9FA00DFD5F99D" blockId="36.[813,1456,1443,1652]" pageId="36" pageNumber="261">
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<materialsCitation id="F9B73CF6FFFFFFD5FCF9FA00DFD5F99D" ID-GBIF-Occurrence="2595817423" collectingDate="1964-04" collectionCode="A" collectorName="Loveridge" country="Saint Helena Island" location="South Atlantic Ocean." municipality="James Bay" pageId="36" pageNumber="261" specimenCode="USNM 33148, USNM 1479052" specimenCount="2" typeStatus="holotype">
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TYPE MATERIAL. —
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<emphasis id="7BABEAB9FFFFFFD5FC03FA00DEDDFA52" bold="true" box="[983,1183,1443,1467]" pageId="36" pageNumber="261">
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<location id="4C006070FFFFFFD5FC03FA00DEDDFA52" LSID="urn:lsid:plazi:treatment:C17687BDFFFFFFD7FC3EFAB6DBEFFD5B:4C006070FFFFFFD5FC03FA00DEDDFA52" box="[983,1183,1443,1467]" country="Saint Helena Island" municipality="James Bay" name="South Atlantic Ocean." pageId="36" pageNumber="261">South Atlantic Ocean.</location>
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</emphasis>
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<typeStatus id="96648809FFFFFFD5FB77FA00DEB8FA52" box="[1187,1274,1443,1467]" pageId="36" pageNumber="261" type="holotype">Holotype</typeStatus>
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,
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<specimenCode id="19799ED0FFFFFFD5FAD6FA00DFC4FA52" box="[1282,1414,1443,1467]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="36" pageNumber="261" type="Museum">USNM 33148</specimenCode>
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, and one
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<typeStatus id="96648809FFFFFFD5FC85FA1DD9E2FA3F" box="[849,928,1470,1494]" pageId="36" pageNumber="261" type="paratype">paratype</typeStatus>
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,
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<specimenCode id="19799ED0FFFFFFD5FC73FA1DDE01FA3C" box="[935,1091,1469,1494]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="36" pageNumber="261" type="Museum">USNM 1479052</specimenCode>
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,
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<collectingCountry id="31C8763BFFFFFFD5FB9FFA1EDEBAFA3C" box="[1099,1272,1469,1493]" name="Saint Helena Island" pageId="36" pageNumber="261">Saint Helena Island</collectingCountry>
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,
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<collectingMunicipality id="A904ACD1FFFFFFD5FAD4FA1DDF1BFA3F" box="[1280,1369,1470,1494]" pageId="36" pageNumber="261">James Bay</collectingMunicipality>
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,
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<date id="3D61106BFFFFFFD5FAB5FA1EDFEEFA3C" box="[1377,1452,1469,1493]" pageId="36" pageNumber="261" value="1964-04">
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<collectingDate id="2D25E983FFFFFFD5FAB5FA1EDFEEFA3C" box="[1377,1452,1469,1493]" pageId="36" pageNumber="261" value="1964-04">IV.1964</collectingDate>
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</date>
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,
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<collectionCode id="2FCEAE6EFFFFFFD5FCF9FA7BD97DFA19" box="[813,831,1496,1520]" pageId="36" pageNumber="261">A</collectionCode>
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.
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<collectorName id="E42A537DFFFFFFD5FC92FA7BD9DDFA19" box="[838,927,1496,1520]" pageId="36" pageNumber="261">Loveridge</collectorName>
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coll. [paratype
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<quantity id="8E279B4EFFFFFFD5FBFDFA7BDE28FA06" box="[1065,1130,1496,1520]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" pageId="36" pageNumber="261" unit="mm" value="39.0">39 mm</quantity>
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long,
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<quantity id="8E279B4EFFFFFFD5FB76FA7BDE9AFA06" box="[1186,1240,1496,1520]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="36" pageNumber="261" unit="mm" value="4.0">4 mm</quantity>
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wide; slightly distorted; anterior end compressed; dorsal cirrostyles swollen basally, multiarticulated; chaetal lobes markedly contracted, invaginated into neuropodia; acicular lobes barely seen, rounded, upper tine 3-4 times longer than lower one; ventral cirri basally swollen, longer than neuropodia, slightly articulated; posterior end truncate; anus with about 6 anal papillae].
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="01C56520FFFFFFD5FCF9F929DE27F91B" pageId="36" pageNumber="261" type="etymology">
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<paragraph id="496036ABFFFFFFD5FCF9F929DE27F91B" blockId="36.[813,1455,1674,1778]" pageId="36" pageNumber="261">
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ETYMOLOGY. — The species name is an adjective derived from
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<collectingCountry id="31C8763BFFFFFFD5FAAAF929D931F954" name="Saint Helena, Ascension and Tristan da Cunha" pageId="36" pageNumber="261">Saint Helena</collectingCountry>
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island, and as in some other similar names, the last vowel in the proper name is suppressed, and the suffix -
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<emphasis id="7BABEAB9FFFFFFD5FAC6F963DF7EF93E" box="[1298,1340,1728,1751]" italics="true" pageId="36" pageNumber="261">ensis</emphasis>
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is added to indicate its current distribution.
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</paragraph>
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</subSubSection>
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<subSubSection id="01C56520FFFFFFD5FCF9F8A4D983F8D3" pageId="36" pageNumber="261" type="distribution">
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<paragraph id="496036ABFFFFFFD5FCF9F8A4D983F8D3" blockId="36.[813,1455,1799,1850]" pageId="36" pageNumber="261">
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DISTRIBUTION. — Only known from
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<collectingCountry id="31C8763BFFFFFFD5FB59F8ABDF44F8C9" box="[1165,1286,1800,1824]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="36" pageNumber="261">Saint Helena</collectingCountry>
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island, in subtidal rocky bottoms.
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</paragraph>
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</subSubSection>
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<subSubSection id="01C56520FFFFFFD5FCF9F8F3DEF9F804" pageId="36" pageNumber="261" type="diagnosis">
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<paragraph id="496036ABFFFFFFD5FCF9F8F3DEF9F804" blockId="36.[813,1456,1872,2029]" pageId="36" pageNumber="261">
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DIAGNOSIS. —
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<taxonomicName id="8EDF4D28FFFFFFD5FC11F8F2DE4CF881" authorityName="Savigny in Lamarck" authorityYear="1818" box="[965,1038,1873,1896]" class="Polychaeta" family="Hesionidae" genus="Hesione" kingdom="Animalia" order="Phyllodocida" pageId="36" pageNumber="261" phylum="Annelida" rank="genus">
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<emphasis id="7BABEAB9FFFFFFD5FC11F8F2DE4CF881" box="[965,1038,1873,1896]" italics="true" pageId="36" pageNumber="261">Hesione</emphasis>
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</taxonomicName>
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with prostomium slightly curved laterally; parapodia with dorsal cirri basally swollen, dorsal cirrophore twice as long as wide; larger acicula blackish; acicular lobe single, short, massive, blunt, upper tine 3 times longer than lower one; neurochaetal blades bidentate, 4-6 times as long as wide; subdistal tooth smaller; guards approaching distal tooth.
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</paragraph>
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</subSubSection>
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<caption id="1DA06623FFFEFFD4FF50FAB7D928FABA" ID-DOI="http://doi.org/10.5281/zenodo.3741590" ID-Zenodo-Dep="3741590" httpUri="https://zenodo.org/record/3741590/files/figure.png" pageId="37" pageNumber="262" startId="37.[132,143,1300,1318]" targetBox="[172,1414,215,1255]" targetPageId="37">
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<paragraph id="496036ABFFFEFFD4FF50FAB7D928FABA" blockId="37.[132,1456,1300,1363]" pageId="37" pageNumber="262">
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FIG. 20. —
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<taxonomicName id="8EDF4D28FFFEFFD4FF09FAB7DB3EFACF" authority="Salazar-Vallejo, 2018" authorityName="Salazar-Vallejo" authorityYear="2018" box="[221,380,1300,1318]" class="Polychaeta" family="Hesionidae" genus="Hesione" kingdom="Animalia" order="Phyllodocida" pageId="37" pageNumber="262" phylum="Annelida" rank="species" species="helenensis" status="sp. nov.">
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<emphasis id="7BABEAB9FFFEFFD4FF09FAB7DB3EFACF" box="[221,380,1300,1318]" italics="true" pageId="37" pageNumber="262">Hesione helenensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="609857C2FFFEFFD4FE55FAB7DBF0FACF" box="[385,434,1300,1318]" pageId="37" pageNumber="262" rank="species">n. sp.</taxonomicNameLabel>
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, holotype,USNM 33148:
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<emphasis id="7BABEAB9FFFEFFD4FD56FAB7D8CDFACF" bold="true" box="[642,655,1300,1318]" pageId="37" pageNumber="262">A</emphasis>
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, anterior region, dorsal view, pharynx partially exposed;
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<emphasis id="7BABEAB9FFFEFFD4FB8AFAB7DE29FACF" bold="true" box="[1118,1131,1300,1318]" pageId="37" pageNumber="262">B</emphasis>
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, prostomium, dorsal view (arrow points to left antenna);
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<emphasis id="7BABEAB9FFFEFFD4FEDDFA88DB55FAD4" bold="true" box="[265,279,1323,1341]" pageId="37" pageNumber="262">C</emphasis>
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, chaetiger 9, left parapodium, anterior view (insets:acicular lobe and bases of cirrostyles);
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<emphasis id="7BABEAB9FFFEFFD4FBD0FA88DE50FAD4" bold="true" box="[1028,1042,1323,1341]" pageId="37" pageNumber="262">D</emphasis>
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, same,neurochaetae (insets: blades);
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<emphasis id="7BABEAB9FFFEFFD4FA9BFA88DF19FAD4" bold="true" box="[1359,1371,1323,1341]" pageId="37" pageNumber="262">E</emphasis>
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, posterior region, dorsal view. Scale bars: A, 1.7 mm; B, 0.2 mm; C, 0.5 mm; D, 160 µm; E, 1.8 mm.
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</paragraph>
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</caption>
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<subSubSection id="01C56520FFFEFFD7FF50FA77D801FEFD" lastPageId="38" lastPageNumber="263" pageId="37" pageNumber="262" type="description">
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<paragraph id="496036ABFFFEFFD4FF50FA77DB57FA04" blockId="37.[132,775,1492,2030]" box="[132,277,1492,1518]" pageId="37" pageNumber="262">DESCRIPTION</paragraph>
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<paragraph id="496036ABFFFEFFD4FF50FA56DB07F967" blockId="37.[132,775,1492,2030]" pageId="37" pageNumber="262">
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<typeStatus id="96648809FFFEFFD4FF50FA56DAAFF9E6" box="[132,237,1525,1551]" pageId="37" pageNumber="262" type="holotype">Holotype</typeStatus>
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,
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<specimenCode id="19799ED0FFFEFFD4FF2CFA56DBDBF9E6" box="[248,409,1525,1551]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="37" pageNumber="262" type="Museum">USNM 33148</specimenCode>
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, complete, subcylindrical, tapered posteriorly, without pigmentation (
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<figureCitation id="D1E42A2EFFFEFFD4FDC1F9B7D83EF9C6" box="[533,636,1556,1583]" captionStart="FIG" captionStartId="37.[132,143,1300,1318]" captionTargetBox="[172,1414,215,1255]" captionTargetId="figure@37.[172,1415,214,1255]" captionTargetPageId="37" captionText="FIG. 20. — Hesione helenensis n. sp., holotype,USNM 33148:A, anterior region, dorsal view, pharynx partially exposed; B, prostomium, dorsal view (arrow points to left antenna);C, chaetiger 9, left parapodium, anterior view (insets:acicular lobe and bases of cirrostyles); D, same,neurochaetae (insets: blades); E, posterior region, dorsal view. Scale bars: A, 1.7 mm; B, 0.2 mm; C, 0.5 mm; D, 160 µm; E, 1.8 mm." figureDoi="http://doi.org/10.5281/zenodo.3741590" httpUri="https://zenodo.org/record/3741590/files/figure.png" pageId="37" pageNumber="262">Fig. 20A</figureCitation>
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) in ethanol; most cirri and many neurochaetal blades broken; left parapodium of chaetiger 9 removed (kept in vial). Body
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<quantity id="8E279B4EFFFEFFD4FD6CF9F7D945F986" box="[696,775,1620,1647]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" pageId="37" pageNumber="262" unit="mm" value="39.0">39 mm</quantity>
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long,
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<quantity id="8E279B4EFFFEFFD4FF16F9D7DB41F966" box="[194,259,1652,1679]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="37" pageNumber="262" unit="mm" value="4.0">4 mm</quantity>
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wide.
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</paragraph>
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<paragraph id="496036ABFFFEFFD4FF4FF937DB6FF867" blockId="37.[132,775,1492,2030]" pageId="37" pageNumber="262">
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Prostomium slightly as wide as long, anterior margin projected anteriorly, lateral margins rounded, posterior margin with a shallow cleft, about 1/5 as long as prostomial length, longitudinal depression slight (
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<figureCitation id="D1E42A2EFFFEFFD4FD5AF957D8B8F8E7" box="[654,762,1780,1806]" captionStart="FIG" captionStartId="37.[132,143,1300,1318]" captionTargetBox="[172,1414,215,1255]" captionTargetId="figure@37.[172,1415,214,1255]" captionTargetPageId="37" captionText="FIG. 20. — Hesione helenensis n. sp., holotype,USNM 33148:A, anterior region, dorsal view, pharynx partially exposed; B, prostomium, dorsal view (arrow points to left antenna);C, chaetiger 9, left parapodium, anterior view (insets:acicular lobe and bases of cirrostyles); D, same,neurochaetae (insets: blades); E, posterior region, dorsal view. Scale bars: A, 1.7 mm; B, 0.2 mm; C, 0.5 mm; D, 160 µm; E, 1.8 mm." figureDoi="http://doi.org/10.5281/zenodo.3741590" httpUri="https://zenodo.org/record/3741590/files/figure.png" pageId="37" pageNumber="262">Fig. 20B</figureCitation>
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). Antennae minute, left one visible, fusiform, 2-3 times as long as wide, about as long as interocular distance. Eyes brownish, anterior ones larger and more separated than posterior ones.
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</paragraph>
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<paragraph id="496036ABFFFEFFD4FF4FF830D818F804" blockId="37.[132,775,1492,2030]" pageId="37" pageNumber="262">Tentacular cirri long, thin, broken, longest one (incomplete) reaches chaetiger 3. Lateral cushions low, smooth, barely projected, most divided into 2 or 3 regions.</paragraph>
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<paragraph id="496036ABFFFEFFD4FC97FA76D981F946" blockId="37.[813,1456,1493,2029]" pageId="37" pageNumber="262">
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Parapodia with chaetal lobes about as long as wide, truncate; dorsal cirri with cirrophores about twice as long as wide (
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<figureCitation id="D1E42A2EFFFEFFD4FCE0F9B7D9D5F9C6" box="[820,919,1556,1583]" captionStart="FIG" captionStartId="37.[132,143,1300,1318]" captionTargetBox="[172,1414,215,1255]" captionTargetId="figure@37.[172,1415,214,1255]" captionTargetPageId="37" captionText="FIG. 20. — Hesione helenensis n. sp., holotype,USNM 33148:A, anterior region, dorsal view, pharynx partially exposed; B, prostomium, dorsal view (arrow points to left antenna);C, chaetiger 9, left parapodium, anterior view (insets:acicular lobe and bases of cirrostyles); D, same,neurochaetae (insets: blades); E, posterior region, dorsal view. Scale bars: A, 1.7 mm; B, 0.2 mm; C, 0.5 mm; D, 160 µm; E, 1.8 mm." figureDoi="http://doi.org/10.5281/zenodo.3741590" httpUri="https://zenodo.org/record/3741590/files/figure.png" pageId="37" pageNumber="262">Fig. 20C</figureCitation>
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); cirrostyles basally swollen, annulated, articulated medially and distally, as long as body width (tips lost). Ventral cirri with cirrostyles basally swollen, surpassing chaetal lobe, darker than parapodia (if observed in compound microscope preparations).
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</paragraph>
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<paragraph id="496036ABFFFEFFD4FC97F917DEA4F8E7" blockId="37.[813,1456,1493,2029]" pageId="37" pageNumber="262">
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Neuraciculae blackish, tapered, only one visible. Acicular lobe double, upper tine blunt, as long as wide, 4 times longer than lower tine blunt (
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<figureCitation id="D1E42A2EFFFEFFD4FBF4F957DEC3F8E7" box="[1056,1153,1780,1806]" captionStart="FIG" captionStartId="37.[132,143,1300,1318]" captionTargetBox="[172,1414,215,1255]" captionTargetId="figure@37.[172,1415,214,1255]" captionTargetPageId="37" captionText="FIG. 20. — Hesione helenensis n. sp., holotype,USNM 33148:A, anterior region, dorsal view, pharynx partially exposed; B, prostomium, dorsal view (arrow points to left antenna);C, chaetiger 9, left parapodium, anterior view (insets:acicular lobe and bases of cirrostyles); D, same,neurochaetae (insets: blades); E, posterior region, dorsal view. Scale bars: A, 1.7 mm; B, 0.2 mm; C, 0.5 mm; D, 160 µm; E, 1.8 mm." figureDoi="http://doi.org/10.5281/zenodo.3741590" httpUri="https://zenodo.org/record/3741590/files/figure.png" pageId="37" pageNumber="262">Fig. 20C</figureCitation>
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[insets]).
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</paragraph>
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<paragraph id="496036ABFFFEFFD4FC97F8B7DF43F867" blockId="37.[813,1456,1493,2029]" pageId="37" pageNumber="262">
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Neurochaetae about 20 per bundle (
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<figureCitation id="D1E42A2EFFFEFFD4FB18F8B7DF71F8C7" box="[1228,1331,1812,1838]" captionStart="FIG" captionStartId="37.[132,143,1300,1318]" captionTargetBox="[172,1414,215,1255]" captionTargetId="figure@37.[172,1415,214,1255]" captionTargetPageId="37" captionText="FIG. 20. — Hesione helenensis n. sp., holotype,USNM 33148:A, anterior region, dorsal view, pharynx partially exposed; B, prostomium, dorsal view (arrow points to left antenna);C, chaetiger 9, left parapodium, anterior view (insets:acicular lobe and bases of cirrostyles); D, same,neurochaetae (insets: blades); E, posterior region, dorsal view. Scale bars: A, 1.7 mm; B, 0.2 mm; C, 0.5 mm; D, 160 µm; E, 1.8 mm." figureDoi="http://doi.org/10.5281/zenodo.3741590" httpUri="https://zenodo.org/record/3741590/files/figure.png" pageId="37" pageNumber="262">Fig. 20D</figureCitation>
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), blades bidentate, 4-6 times as long as wide, decreasing in size ventrally, each with smaller subdistal teeth, and guards (most broken) reaching subdistal teeth (
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<figureCitation id="D1E42A2EFFFEFFD4FBEDF8D0DEDEF864" box="[1081,1180,1907,1933]" captionStart="FIG" captionStartId="37.[132,143,1300,1318]" captionTargetBox="[172,1414,215,1255]" captionTargetId="figure@37.[172,1415,214,1255]" captionTargetPageId="37" captionText="FIG. 20. — Hesione helenensis n. sp., holotype,USNM 33148:A, anterior region, dorsal view, pharynx partially exposed; B, prostomium, dorsal view (arrow points to left antenna);C, chaetiger 9, left parapodium, anterior view (insets:acicular lobe and bases of cirrostyles); D, same,neurochaetae (insets: blades); E, posterior region, dorsal view. Scale bars: A, 1.7 mm; B, 0.2 mm; C, 0.5 mm; D, 160 µm; E, 1.8 mm." figureDoi="http://doi.org/10.5281/zenodo.3741590" httpUri="https://zenodo.org/record/3741590/files/figure.png" pageId="37" pageNumber="262">Fig. 20D</figureCitation>
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[insets]).
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</paragraph>
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<paragraph id="496036ABFFFEFFD4FC97F830D9CBF804" blockId="37.[813,1456,1493,2029]" pageId="37" pageNumber="262">
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Posterior region tapered into a blunt cone, dorsal surface rugose (
|
|
<figureCitation id="D1E42A2EFFFEFFD4FC53F810D9ADF824" box="[903,1007,1971,1997]" captionStart="FIG" captionStartId="37.[132,143,1300,1318]" captionTargetBox="[172,1414,215,1255]" captionTargetId="figure@37.[172,1415,214,1255]" captionTargetPageId="37" captionText="FIG. 20. — Hesione helenensis n. sp., holotype,USNM 33148:A, anterior region, dorsal view, pharynx partially exposed; B, prostomium, dorsal view (arrow points to left antenna);C, chaetiger 9, left parapodium, anterior view (insets:acicular lobe and bases of cirrostyles); D, same,neurochaetae (insets: blades); E, posterior region, dorsal view. Scale bars: A, 1.7 mm; B, 0.2 mm; C, 0.5 mm; D, 160 µm; E, 1.8 mm." figureDoi="http://doi.org/10.5281/zenodo.3741590" httpUri="https://zenodo.org/record/3741590/files/figure.png" pageId="37" pageNumber="262">Fig. 20E</figureCitation>
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); pygidium smooth, anus with 7 blunt papillae.
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</paragraph>
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<paragraph id="496036ABFFFDFFD7FF4FFF7AD801FEFD" blockId="38.[132,775,217,276]" pageId="38" pageNumber="263">Pharynx partially exposed, anterior margin slightly eroded, dorsal papilla not seen. Oocytes not seen.</paragraph>
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</subSubSection>
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<subSubSection id="01C56520FFFDFFD7FF50FE9BDBEFFD5B" pageId="38" pageNumber="263" type="discussion">
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<paragraph id="496036ABFFFDFFD7FF50FE9BDAAAFEB8" blockId="38.[132,775,312,690]" box="[132,232,312,338]" pageId="38" pageNumber="263">REMARKS</paragraph>
|
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<paragraph id="496036ABFFFDFFD7FF50FEFADBEFFD5B" blockId="38.[132,775,312,690]" pageId="38" pageNumber="263">
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|
<taxonomicName id="8EDF4D28FFFDFFD7FF50FEFADB03FE9A" authority="Salazar-Vallejo, 2018" authorityName="Salazar-Vallejo" authorityYear="2018" box="[132,321,345,371]" class="Polychaeta" family="Hesionidae" genus="Hesione" kingdom="Animalia" order="Phyllodocida" pageId="38" pageNumber="263" phylum="Annelida" rank="species" species="helenensis" status="sp. nov.">
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|
<emphasis id="7BABEAB9FFFDFFD7FF50FEFADB03FE9A" box="[132,321,345,371]" italics="true" pageId="38" pageNumber="263">Hesione helenensis</emphasis>
|
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</taxonomicName>
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|
<taxonomicNameLabel id="609857C2FFFDFFD7FE9DFEFADBC7FE9A" box="[329,389,345,371]" pageId="38" pageNumber="263" rank="species">n. sp.</taxonomicNameLabel>
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|
resembles Northeastern Atlantic or Mediterranean specimens of
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|
<taxonomicName id="8EDF4D28FFFDFFD7FE6DFEDAD896FE7A" authority="Risso, 1826" authorityName=", Risso" authorityYear="1826" box="[441,724,376,403]" class="Polychaeta" family="Hesionidae" genus="Hesione" kingdom="Animalia" order="Phyllodocida" pageId="38" pageNumber="263" phylum="Annelida" rank="species" species="pantherina">
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|
<emphasis id="7BABEAB9FFFDFFD7FE6DFEDAD80FFE7B" box="[441,589,376,402]" italics="true" pageId="38" pageNumber="263">H. pantherina</emphasis>
|
|
<bibRefCitation id="2D4E4B5AFFFDFFD7FD87FEDAD896FE7A" author="Risso" box="[595,724,377,403]" firstAuthor="Risso" pageId="38" pageNumber="263" refId="ref97434" refString="RISSO A. 1826. - Histoire naturelle des principales productions de l'Europe Meridionale et particulierement de celles des environs de Nice et des Alpes Maritimes. Volume 4, Paris: Livrault Libraire, 439 p. https: // biodiversitylibrary. org / page / 50455192" type="book" year="1826">Risso, 1826</bibRefCitation>
|
|
</taxonomicName>
|
|
, and it could be confused with it. However, as indicated in the key below,
|
|
<taxonomicName id="8EDF4D28FFFDFFD7FF2FFE1ADBC0FE3B" authority="Salazar-Vallejo, 2018" authorityName="Salazar-Vallejo" authorityYear="2018" box="[251,386,440,466]" class="Polychaeta" family="Hesionidae" genus="Hesione" kingdom="Animalia" order="Phyllodocida" pageId="38" pageNumber="263" phylum="Annelida" rank="species" species="helenensis" status="sp. nov.">
|
|
<emphasis id="7BABEAB9FFFDFFD7FF2FFE1ADBC0FE3B" box="[251,386,440,466]" italics="true" pageId="38" pageNumber="263">H. helenensis</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="609857C2FFFDFFD7FE5CFE1ADB80FE3A" box="[392,450,441,467]" pageId="38" pageNumber="263" rank="species">n. sp.</taxonomicNameLabel>
|
|
resembles
|
|
<taxonomicName id="8EDF4D28FFFDFFD7FDE3FE1ADAFEFE1B" authority="Ehlers, 1887" authorityName="Ehlers" authorityYear="1887" class="Polychaeta" family="Hesionidae" genus="Hesione" kingdom="Animalia" order="Phyllodocida" pageId="38" pageNumber="263" phylum="Annelida" rank="species" species="praetexta">
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|
<emphasis id="7BABEAB9FFFDFFD7FDE3FE1AD8F4FE3B" box="[567,694,441,466]" italics="true" pageId="38" pageNumber="263">H. praetexta</emphasis>
|
|
<bibRefCitation id="2D4E4B5AFFFDFFD7FD69FE1BDAFEFE1B" author="Ehlers" firstAuthor="Ehlers" pageId="38" pageNumber="263" pagination="1 - 335" refId="ref92380" refString="EHLERS E. 1887. - Reports on the results of dredging under th direction of L. P. Pourtales, during the years 1868 - 1870, and of Alexander Agassiz in the Gulf of Mexico (1877 - 78), and in the Caribbean Sea (1878 - 79) in the U. S. Coast Survey Steamer ' Blake', 31. Reports on the annelids. Memoirs of the Museum of Comparative Zoology, Harvard College 15: 1 - 335, pls 1 - 60. https: // biodiversitylibrary. org / page / 30734338" type="journal article" year="1887">Ehlers, 1887</bibRefCitation>
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|
</taxonomicName>
|
|
reinstated. The main difference is that in
|
|
<taxonomicName id="8EDF4D28FFFDFFD7FD54FE7AD945FE1B" authority="Salazar-Vallejo, 2018" authorityName="Salazar-Vallejo" authorityYear="2018" box="[640,775,472,498]" class="Polychaeta" family="Hesionidae" genus="Hesione" kingdom="Animalia" order="Phyllodocida" pageId="38" pageNumber="263" phylum="Annelida" rank="species" species="helenensis" status="sp. nov.">
|
|
<emphasis id="7BABEAB9FFFDFFD7FD54FE7AD945FE1B" box="[640,775,472,498]" italics="true" pageId="38" pageNumber="263">H. helenensis</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="609857C2FFFDFFD7FF50FE5ADA82FDFA" box="[132,192,505,531]" pageId="38" pageNumber="263" rank="species">n. sp.</taxonomicNameLabel>
|
|
dorsal and ventral cirrostyles are basally swollen, and this feature is unique for the genus, and present throughout the body, although in posterior chaetigers ventral cirrostyles are less markedly swollen. Further, these swollen areas appear darker when parapodia are mounted and observed in the compound microscope.
|
|
</paragraph>
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|
</subSubSection>
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|
</treatment>
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|
</document> |