200 lines
28 KiB
XML
200 lines
28 KiB
XML
<document id="DF669063B3BBE3F583BCF1F3A3E61348" ID-DOI="10.37520/aemnp.2021.006" ID-ISSN="1804-6487" ID-Zenodo-Dep="5037544" ID-ZooBank="7A927D20-F3B3-4611-BDC3-7EC99A131B69" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624896857361" checkinUser="carolina" docAuthor="Anichtchenko, Alexander, Minkina, Łukasz, Vasiljeva, Anastasija & Medina, Milton Norman D." docDate="2021" docId="03AA81215339FFD853BE6A15BEB2F8A7" docLanguage="en" docName="ActaEntMusNatPra.61.1.99-111.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 61 (1)" docSource="http://dx.doi.org/10.37520/aemnp.2021.006" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Rhyparus flavohirtus Anichtchenko & Minkina & Vasiljeva & Medina 2021, sp. nov." docType="treatment" docVersion="3" lastPageNumber="102" masterDocId="FF93F9595338FFDB50746D24BA16FFBC" masterDocTitle="A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao (Coleoptera: Scarabaeidae: Aphodiinae)" masterLastPageNumber="111" masterPageNumber="99" pageNumber="100" updateTime="1698977438183" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="26130AE6C44B62C018100319C8EFD6AD">A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao (Coleoptera: Scarabaeidae: Aphodiinae)</mods:title>
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<mods:namePart id="A343E0C81837F3EED5186B5BA77623D6">Anichtchenko, Alexander</mods:namePart>
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<mods:namePart id="3B264738AD6A67589F783C955C6043CB">Minkina, Łukasz</mods:namePart>
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<mods:affiliation id="8DB3096D5AA2B1A0615E366458A18E36">os. Polana Szalfarska</mods:affiliation>
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<mods:namePart id="A7A49144EA5141408B08BF64E80CCE66">Vasiljeva, Anastasija</mods:namePart>
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<mods:affiliation id="00107AC241C7784F76B395153B0DE2C4">Institute of Life Sciences and Technologies, Daugavpils University, Daugavpils, Latvia</mods:affiliation>
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<mods:namePart id="31436A356D0B7E386E71B7D2E0830416">Medina, Milton Norman D.</mods:namePart>
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<mods:affiliation id="88FD259642E4DAC1D381ABC3BFFAC569">Institute of Biodiversity and Environment, Coleoptera Research Center, University of Mindanao, Davao City, Philippines</mods:affiliation>
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<taxonomicName id="4C034BB45339FFDA53BE6A15BEA4F8F5" authority="Anichtchenko & Minkina & Vasiljeva & Medina, 2021" authorityName="Anichtchenko & Minkina & Vasiljeva & Medina" authorityYear="2021" box="[970,1202,1841,1865]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="100" phylum="Arthropoda" rank="species" species="flavohirtus" status="sp. nov.">
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<emphasis id="B977EC255339FFDA53BE6A15BEA4F8F5" bold="true" box="[970,1202,1841,1865]" italics="true" pageId="1" pageNumber="100">Rhyparus flavohirtus</emphasis>
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<subSubSection id="C31963BC5339FFDA53866A75BEF0F8D8" box="[1010,1254,1873,1892]" pageId="1" pageNumber="100" type="description">
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<paragraph id="8BBC30375339FFDA53866A75BEF0F8D8" blockId="1.[828,1437,1873,1960]" box="[1010,1254,1873,1892]" pageId="1" pageNumber="100">
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(
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<figureCitation id="13382CB25339FFDA538C6A75BE5BF8D8" box="[1016,1101,1873,1892]" captionStart="Fig" captionStartId="3.[150,181,1195,1214]" captionTargetBox="[165,1407,196,1168]" captionTargetId="figure-396@3.[150,1435,166,1175]" captionTargetPageId="3" captionText="Fig. 2. Habitus of Rhyparus flavohirtus sp. nov., male, holotype: a – dorsal view; b – right dorsolateral view; c – right lateral view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5037548" httpUri="https://zenodo.org/record/5037548/files/figure.png" pageId="1" pageNumber="100">Figs 2a–c</figureCitation>
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,
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<figureCitation id="13382CB25339FFDA54226A75BE74F8D8" box="[1110,1122,1873,1892]" captionStart="Figs 7–12" captionStartId="8.[151,188,1611,1630]" captionTargetBox="[165,1410,204,1585]" captionTargetId="figure-191@8.[151,1436,189,1595]" captionTargetPageId="8" captionText="Figs 7–12. Head of Rhyparus species, frontal view: 7 – R. barclayi Masumoto, Ochi & Ho, 2019; 8 – R. flavohirtus sp. nov.; 9 – R. gracilis Arrow, 1905; 10 – R. helophoroides Fairmaire, 1893; 11 – R. mindanaoensis sp. nov.; 12 – R. philippinensis Arrow, 1905. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5037560" httpUri="https://zenodo.org/record/5037560/files/figure.png" pageId="1" pageNumber="100">8</figureCitation>
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,
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<figureCitation id="13382CB25339FFDA541F6A75BE94F8D8" box="[1131,1154,1873,1892]" captionStart="Figs 13–18" captionStartId="9.[150,187,1526,1545]" captionTargetBox="[158,1411,197,1485]" captionTargetId="figure-259@9.[150,1435,161,1515]" captionTargetPageId="9" captionText="Figs 13–18. Epipharynx of Rhyparus species: 13 – R. barclayi Masumoto, Ochi & Ho, 2019; 14 – R. flavohirtus sp. nov.; 15 – R. gracilis Arrow, 1905; 16 – R. helophoroides Fairmaire, 1893; 17– R. mindanaoensis sp. nov.; 18 – R. philippinensis Arrow, 1905. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5037562" httpUri="https://zenodo.org/record/5037562/files/figure.png" pageId="1" pageNumber="100">14</figureCitation>
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,
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<figureCitation id="13382CB25339FFDA54FF6A75BEB4F8D8" box="[1163,1186,1873,1892]" captionStart="Figs 19–24" captionStartId="10.[151,188,1324,1343]" captionTargetBox="[164,1415,209,1277]" captionTargetId="figure-243@10.[151,1436,189,1299]" captionTargetPageId="10" captionText="Figs 19–24. Elytra of Rhyparus species, oblique apico-dorsolateral view: 19 – R. barclayi Masumoto, Ochi & Ho, 2019; 20 – R. flavohirtus sp. nov.; 21 – R. gracilis Arrow, 1905; 22 – R. helophoroides Fairmaire, 1893; 23 – R. mindanaoensis sp. nov.; 24 – R. philippinensis Arrow, 1905. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5037564" httpUri="https://zenodo.org/record/5037564/files/figure.png" pageId="1" pageNumber="100">20</figureCitation>
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,
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<figureCitation id="13382CB25339FFDA54DF6A75BEF7F8D8" box="[1195,1249,1873,1892]" captionStart="Figs 25–30" captionStartId="11.[150,187,1486,1505]" captionTargetBox="[207,1392,193,1454]" captionTargetId="figure-302@11.[151,1436,180,1472]" captionTargetPageId="11" captionText="Figs 25–30.Aedeagus of Rhyparus species, a – lateral view, b – dorsal view. 25 – R.barclayi Masumoto, Ochi & Ho, 2019; 26 – R. flavohirtus sp. nov.; 27 – R. gracilis Arrow, 1905; 28 – R. helophoroides Fairmaire, 1893; 29 – R.mindanaoensis sp. nov.; 30 – R.philippinensis Arrow, 1905. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5037566" httpUri="https://zenodo.org/record/5037566/files/figure.png" pageId="1" pageNumber="100">26a–b</figureCitation>
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)
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</paragraph>
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<subSubSection id="C31963BC5339FFDA53486A5EB99AF814" pageId="1" pageNumber="100" type="materials_examined">
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<paragraph id="8BBC30375339FFDA53486A5EB99AF814" blockId="1.[828,1437,1873,1960]" pageId="1" pageNumber="100">
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<emphasis id="B977EC255339FFDA53486A5EB9ABF831" bold="true" box="[828,957,1914,1933]" pageId="1" pageNumber="100">
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<typeStatus id="54B88E955339FFDA53486A5EB97EF831" box="[828,872,1914,1933]" pageId="1" pageNumber="100">Type</typeStatus>
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material.
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</emphasis>
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Hඈඅඈඍඒඉൾ:J, labelled:ʻMindanao, Lake Holon,
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<date id="FFBD16F75339FFDA55226A5EBF8FF831" box="[1366,1433,1914,1933]" pageId="1" pageNumber="100" value="2019-06">vi.2019</date>
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ʼ (
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<collectionCode id="ED12A8F25339FFDA53366AB1B994F814" box="[834,898,1941,1960]" pageId="1" pageNumber="100">DUBC</collectionCode>
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).
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</paragraph>
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<subSubSection id="C31963BC5339FFD853486AE5BE69FA67" lastPageId="3" lastPageNumber="102" pageId="1" pageNumber="100" type="description">
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<paragraph id="8BBC30375339FFD953486AE5B886FA8B" blockId="1.[828,1436,1985,2072]" lastBlockId="2.[151,760,1280,2071]" lastPageId="2" lastPageNumber="101" pageId="1" pageNumber="100">
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<emphasis id="B977EC255339FFDA53486AE5B9D1F864" bold="true" box="[828,967,1985,2008]" pageId="1" pageNumber="100">Description.</emphasis>
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Large species, body length 7.0 mm. Body surface matt, only pygidium with slightly shiny elevated area; black, except for brown or reddish-black preapical glandular area, more densely covered with yellow setae dorsally compared to other species (
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<figureCitation id="13382CB2533AFFD9526C6804B892FA8B" box="[536,644,1312,1335]" captionStart="Fig" captionStartId="3.[150,181,1195,1214]" captionTargetBox="[165,1407,196,1168]" captionTargetId="figure-396@3.[150,1435,166,1175]" captionTargetPageId="3" captionText="Fig. 2. Habitus of Rhyparus flavohirtus sp. nov., male, holotype: a – dorsal view; b – right dorsolateral view; c – right lateral view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5037548" httpUri="https://zenodo.org/record/5037548/files/figure.png" pageId="2" pageNumber="101">Figs 2a–c</figureCitation>
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).
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</paragraph>
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<caption id="DF7C60BF533AFFD950E369ABBF88FB1E" ID-DOI="http://doi.org/10.5281/zenodo.5037546" ID-Zenodo-Dep="5037546" box="[151,1438,1167,1186]" httpUri="https://zenodo.org/record/5037546/files/figure.png" pageId="2" pageNumber="101" startId="2.[151,179,1167,1186]" targetBox="[154,1401,189,1126]" targetPageId="2">
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<paragraph id="8BBC3037533AFFD950E369ABBF88FB1E" blockId="2.[151,1438,1167,1186]" box="[151,1438,1167,1186]" pageId="2" pageNumber="101">
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Fig 1. Habitus of
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<taxonomicName id="4C034BB4533AFFD9515F69ABB8D0FB1E" authority="Masumoto, Ochi & Ho, 2019" authorityName="Masumoto, Ochi & Ho" authorityYear="2019" box="[299,710,1167,1186]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="101" phylum="Arthropoda" rank="species" species="barclayi">
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<emphasis id="B977EC25533AFFD9515F69ABBBD1FB1E" box="[299,455,1167,1186]" italics="true" pageId="2" pageNumber="101">Rhyparus barclayi</emphasis>
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<bibRefCitation id="EF924DC6533AFFD951B869ABB8D0FB1E" author="MASUMOTO K. & OCHI T. & HO B. H." box="[460,710,1167,1186]" pageId="2" pageNumber="101" pagination="13 - 18" refId="ref6667" refString="MASUMOTO K., OCHI T. & HO B. H. 2019: A new Rhyparus species from Taiwan (Scarabaeidae, Aphodiinae, Rhyparini). Kogane 22: 13 - 18." type="journal article" year="2019">Masumoto, Ochi & Ho, 2019</bibRefCitation>
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</taxonomicName>
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: a – dorsal view; b – right dorsolateral view; c – right lateral view. Scale bar = 1.0 mm.
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</paragraph>
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</caption>
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<paragraph id="8BBC3037533AFFD950C16864B8FBF96B" blockId="2.[151,760,1280,2071]" pageId="2" pageNumber="101">
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Head as observed dorsally and frontally (
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<figureCitation id="13382CB2533AFFD952036864B8DFFAEB" box="[631,713,1344,1367]" captionStart="Fig" captionStartId="3.[150,181,1195,1214]" captionTargetBox="[165,1407,196,1168]" captionTargetId="figure-396@3.[150,1435,166,1175]" captionTargetPageId="3" captionText="Fig. 2. Habitus of Rhyparus flavohirtus sp. nov., male, holotype: a – dorsal view; b – right dorsolateral view; c – right lateral view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5037548" httpUri="https://zenodo.org/record/5037548/files/figure.png" pageId="2" pageNumber="101">Figs 2c</figureCitation>
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and
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<figureCitation id="13382CB2533AFFD950E36844BAB0FACB" box="[151,166,1376,1399]" captionStart="Figs 7–12" captionStartId="8.[151,188,1611,1630]" captionTargetBox="[165,1410,204,1585]" captionTargetId="figure-191@8.[151,1436,189,1595]" captionTargetPageId="8" captionText="Figs 7–12. Head of Rhyparus species, frontal view: 7 – R. barclayi Masumoto, Ochi & Ho, 2019; 8 – R. flavohirtus sp. nov.; 9 – R. gracilis Arrow, 1905; 10 – R. helophoroides Fairmaire, 1893; 11 – R. mindanaoensis sp. nov.; 12 – R. philippinensis Arrow, 1905. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5037560" httpUri="https://zenodo.org/record/5037560/files/figure.png" pageId="2" pageNumber="101">8</figureCitation>
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, respectively): Clypeus anteriorly not sinuate, with indistinct teeth on sides, laterally distinctly emarginated and next with widely rounded angle, separated by rather small emargination from distinctly protruding, rounded genae; clypeal disc distinctly convex, ringed by deep groove, with pair of distinct, rounded tubercles. Frons with four short, blunt, posteriorly vanishing longitudinal costae, median pair shorter than lateral pair and separated one from another by narrow, deep furrow, distance between neighboring median and lateral costae twice as broad as that between two median costae. Head covered with irregularly spaced fine punctures bearing small setae. Epipharynx as in
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<figureCitation id="13382CB2533AFFD952EF6BE4B8FFF96B" box="[667,745,1728,1751]" captionStart="Figs 13–18" captionStartId="9.[150,187,1526,1545]" captionTargetBox="[158,1411,197,1485]" captionTargetId="figure-259@9.[150,1435,161,1515]" captionTargetPageId="9" captionText="Figs 13–18. Epipharynx of Rhyparus species: 13 – R. barclayi Masumoto, Ochi & Ho, 2019; 14 – R. flavohirtus sp. nov.; 15 – R. gracilis Arrow, 1905; 16 – R. helophoroides Fairmaire, 1893; 17– R. mindanaoensis sp. nov.; 18 – R. philippinensis Arrow, 1905. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5037562" httpUri="https://zenodo.org/record/5037562/files/figure.png" pageId="2" pageNumber="101">Fig. 14</figureCitation>
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.
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</paragraph>
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<paragraph id="8BBC3037533AFFD950C16BC4BE11FA0B" blockId="2.[151,760,1280,2071]" lastBlockId="2.[828,1437,1280,2071]" pageId="2" pageNumber="101">Pronotum subquadrate, 1.13 times as wide as long; with six distinctly (but not sharply) convex costae and seven longitudinal intercostae, surface densely covered with yellow setae. Lateral margin with moderately protruding two lobes, anterior lobe triangular, median lobe obtusely angled, slightly wider than anterior lobe. Costae on anterior part present anteriorly as rounded swellings with dense concentration of setae; costae 1 (paramedian pair), 3 and 4 (lateral pairs) – continuous, costae 2 widely interrupted by transverse depression with deep caviform fossae, and another fossa in middle of third furrow at base of costae. First (median) inter-costa covered with distinct medium-sized punctures, very densely concentrated in middle to form elongated depression; lateral intercostae 1–3 abundantly punctured only in anterior part, intercostae 2 and 3 smooth in basal two thirds.</paragraph>
|
||
<paragraph id="8BBC3037533AFFD9532E68E4BF8BF80B" blockId="2.[828,1437,1280,2071]" pageId="2" pageNumber="101">
|
||
Elytra (
|
||
<figureCitation id="13382CB2533AFFD953DD68E4BE03FA6B" box="[937,1045,1472,1495]" captionStart="Fig" captionStartId="3.[150,181,1195,1214]" captionTargetBox="[165,1407,196,1168]" captionTargetId="figure-396@3.[150,1435,166,1175]" captionTargetPageId="3" captionText="Fig. 2. Habitus of Rhyparus flavohirtus sp. nov., male, holotype: a – dorsal view; b – right dorsolateral view; c – right lateral view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5037548" httpUri="https://zenodo.org/record/5037548/files/figure.png" pageId="2" pageNumber="101">Figs 2a–c</figureCitation>
|
||
,
|
||
<figureCitation id="13382CB2533AFFD9545568E4BE28FA6B" box="[1057,1086,1472,1495]" captionStart="Figs 19–24" captionStartId="10.[151,188,1324,1343]" captionTargetBox="[164,1415,209,1277]" captionTargetId="figure-243@10.[151,1436,189,1299]" captionTargetPageId="10" captionText="Figs 19–24. Elytra of Rhyparus species, oblique apico-dorsolateral view: 19 – R. barclayi Masumoto, Ochi & Ho, 2019; 20 – R. flavohirtus sp. nov.; 21 – R. gracilis Arrow, 1905; 22 – R. helophoroides Fairmaire, 1893; 23 – R. mindanaoensis sp. nov.; 24 – R. philippinensis Arrow, 1905. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5037564" httpUri="https://zenodo.org/record/5037564/files/figure.png" pageId="2" pageNumber="101">20</figureCitation>
|
||
), elongate, 1.73 times as long as wide, each with five costae of similar height and width from elytral base to apex, and one very long, well defined accessory costa present between costae 3 and 4, terminating posteriorly before preapical depressed area; costae slightly shiny on top, covered with yellow setae: costae 1 (sutural) with one row of setae in middle, costae 2 and 3 bordered by row of setae on each side. Intercostae 1 and 2 flat with 3 rows of punctures, with punctures in lateral rows larger than punctures in median row, punctures of intercostae 3a, 3b and 4 arranged in two rows, distances between neighboring punctures not uniform, but larger than or approximately equal to puncture diameter. Apical bulbous area of second costa densely covered with yellow macrosetae. Caudal bulbs weakly developed. Internal and median protrusion of caudal bulbs connected; external protrusion moderately developed.
|
||
</paragraph>
|
||
<paragraph id="8BBC3037533AFFD9532E6AE4B9BAF7AB" blockId="2.[828,1437,1280,2071]" pageId="2" pageNumber="101">Pygidium with elevated area at base, extended as costa to pygidial apex, areas on each side of costa being strongly depressed.</paragraph>
|
||
<caption id="DF7C60BF533BFFD850E2698FBF8BFB02" ID-DOI="http://doi.org/10.5281/zenodo.5037548" ID-Zenodo-Dep="5037548" box="[150,1437,1195,1214]" httpUri="https://zenodo.org/record/5037548/files/figure.png" pageId="3" pageNumber="102" startId="3.[150,181,1195,1214]" targetBox="[165,1407,196,1168]" targetPageId="3">
|
||
<paragraph id="8BBC3037533BFFD850E2698FBF8BFB02" blockId="3.[150,1437,1195,1214]" box="[150,1437,1195,1214]" pageId="3" pageNumber="102">
|
||
Fig. 2. Habitus of
|
||
<taxonomicName id="4C034BB4533BFFD85145698FBBF2FB02" authority="Anichtchenko & Minkina & Vasiljeva & Medina, 2021" authorityName="Anichtchenko & Minkina & Vasiljeva & Medina" authorityYear="2021" box="[305,484,1195,1214]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="102" phylum="Arthropoda" rank="species" species="flavohirtus" status="sp. nov.">
|
||
<emphasis id="B977EC25533BFFD85145698FBBF2FB02" box="[305,484,1195,1214]" italics="true" pageId="3" pageNumber="102">Rhyparus flavohirtus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A244515E533BFFD8519D698FB83BFB02" box="[489,557,1195,1214]" pageId="3" pageNumber="102" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male, holotype: a – dorsal view; b – right dorsolateral view; c – right lateral view. Scale bar = 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBC3037533BFFD850C16800BBCCF8C7" blockId="3.[151,760,1316,2075]" pageId="3" pageNumber="102">All legs sturdy, matt, covered with small punctures and triangular short setae (except for simple and smooth area along outer margin), greatest quantity of setae on mesotibia. Profemora strongly expanded, widest near basal third, with deep groove anteriorly; surface with medium-sized punctures throughout. Protibiae slenderer than mesotibiae, only slightly dilated apically and noticeably incurved apically; apex of protibiae with three small teeth. Mesofemora slightly narrower than profemur, posterior margin bilobed, surface punctate throughout, punctures smaller compared to those on profemora. Mesotibiae narrow at base, gradually expanded from base to about one third of length, and then narrowed to apex; apex of mesotibiae with large projection and tooth (larger than protibiae apical teeth). Metafemora long, slightly narrower than mesofemora; narrow at base, gradually expanded, apical part narrowed; punctures as on mesofemora. Metatibiae larger than mesotibiae, but similar in shape, but not narrowed preapically, only slightly widened to apex.</paragraph>
|
||
<paragraph id="8BBC3037533BFFD850C16AA0BF05FA07" blockId="3.[151,760,1316,2075]" lastBlockId="3.[828,1437,1316,1819]" pageId="3" pageNumber="102">Venter matt, black. Prosternum with long anterior process with rounded apex and hastate posterior process. Mesoventrite with shallow setigerous punctures and small setae. Meso-metaventral plate with distinct, shallow medi- an impression; surface otherwise with irregular medium- sized setigerous punctures. Abdomen with six abdominal ventrites, all separated by impressed sutures; surface punctate-setose. Ventrites 3–5 triangularly grooved on each lateral broadened area, groove noticeably large on ventrite 5; ventrites 5–6 fused, medially with costae.</paragraph>
|
||
<paragraph id="8BBC3037533BFFD8532E68E0BE69FA67" blockId="3.[828,1437,1316,1819]" box="[858,1151,1476,1499]" pageId="3" pageNumber="102">
|
||
Aedeagus as in
|
||
<figureCitation id="13382CB2533BFFD8547668E0BE6DFA67" box="[1026,1147,1476,1499]" captionStart="Figs 25–30" captionStartId="11.[150,187,1486,1505]" captionTargetBox="[207,1392,193,1454]" captionTargetId="figure-302@11.[151,1436,180,1472]" captionTargetPageId="11" captionText="Figs 25–30.Aedeagus of Rhyparus species, a – lateral view, b – dorsal view. 25 – R.barclayi Masumoto, Ochi & Ho, 2019; 26 – R. flavohirtus sp. nov.; 27 – R. gracilis Arrow, 1905; 28 – R. helophoroides Fairmaire, 1893; 29 – R.mindanaoensis sp. nov.; 30 – R.philippinensis Arrow, 1905. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5037566" httpUri="https://zenodo.org/record/5037566/files/figure.png" pageId="3" pageNumber="102">Figs 26a–b</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31963BC533BFFD8534868C0B9A1F9C7" pageId="3" pageNumber="102" type="diagnosis">
|
||
<paragraph id="8BBC3037533BFFD8534868C0B9A1F9C7" blockId="3.[828,1437,1316,1819]" pageId="3" pageNumber="102">
|
||
<emphasis id="B977EC25533BFFD8534868C0BE22FA47" bold="true" box="[828,1076,1508,1531]" pageId="3" pageNumber="102">Differential diagnosis.</emphasis>
|
||
The new species can be distinguished from all
|
||
<taxonomicName id="4C034BB4533BFFD853AD6B20BE82F9A7" box="[985,1172,1540,1563]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="102" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B977EC25533BFFD853AD6B20BE29F9A7" box="[985,1087,1540,1563]" italics="true" pageId="3" pageNumber="102">Rhyparus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
worldwide based on the combination of very long accessory costa, dense setation of dorsal surface, and number of rows of punctures on intercostae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31963BC533BFFD853486BA0BF8BF967" pageId="3" pageNumber="102" type="etymology">
|
||
<paragraph id="8BBC3037533BFFD853486BA0BF8BF967" blockId="3.[828,1437,1316,1819]" pageId="3" pageNumber="102">
|
||
<emphasis id="B977EC25533BFFD853486BA0B9AAF927" bold="true" box="[828,956,1668,1691]" pageId="3" pageNumber="102">Etymology.</emphasis>
|
||
The specific epithet is composed of the Latin adjectives
|
||
<emphasis id="B977EC25533BFFD853DE6B80B9F1F907" box="[938,999,1700,1723]" italics="true" pageId="3" pageNumber="102">flavus</emphasis>
|
||
, -
|
||
<emphasis id="B977EC25533BFFD853886B80BE1FF907" box="[1020,1033,1700,1723]" italics="true" pageId="3" pageNumber="102">a</emphasis>
|
||
, -
|
||
<emphasis id="B977EC25533BFFD854696B80BE2BF907" box="[1053,1085,1700,1723]" italics="true" pageId="3" pageNumber="102">um</emphasis>
|
||
and
|
||
<emphasis id="B977EC25533BFFD8541A6B80BEBAF907" box="[1134,1196,1700,1723]" italics="true" pageId="3" pageNumber="102">hirtus</emphasis>
|
||
, -
|
||
<emphasis id="B977EC25533BFFD854B46B80BEDBF907" box="[1216,1229,1700,1723]" italics="true" pageId="3" pageNumber="102">a</emphasis>
|
||
, -
|
||
<emphasis id="B977EC25533BFFD854956B80BF17F907" box="[1249,1281,1700,1723]" italics="true" pageId="3" pageNumber="102">um</emphasis>
|
||
, and alludes to the dense covering of yellow setae on the dorsum of body.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31963BC533BFFD853486BC0BEB2F8A7" pageId="3" pageNumber="102" type="distribution">
|
||
<paragraph id="8BBC3037533BFFD853486BC0BEB2F8A7" blockId="3.[828,1437,1316,1819]" pageId="3" pageNumber="102">
|
||
<emphasis id="B977EC25533BFFD853486BC0B9D8F947" bold="true" box="[828,974,1764,1787]" pageId="3" pageNumber="102">Distribution.</emphasis>
|
||
Species known only from
|
||
<typeStatus id="54B88E95533BFFD8548C6BC0BF30F947" box="[1272,1318,1764,1787]" pageId="3" pageNumber="102">type</typeStatus>
|
||
locality in Mindanao Island, the
|
||
<collectingCountry id="F31470A7533BFFD854506A20BEB6F8A7" box="[1060,1184,1796,1819]" name="Philippines" pageId="3" pageNumber="102">Philippines</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |