165 lines
19 KiB
XML
165 lines
19 KiB
XML
<document id="A6EE3CB126344A5297BF9E2B707383D0" ID-ISSN="1175-5326" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629742108138" checkinUser="felipe" docAuthor="Murányi, Dávid" docDate="2011" docId="03903327FF8BFFF2FF6DFAA0F9B6D134" docLanguage="en" docName="zt03049p046.pdf" docOrigin="Zootaxa 3049" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Isoperla obliqua Zwick 1978" docType="treatment" docVersion="4" lastPageNumber="33" masterDocId="FFA94B5FFF94FFD2FFFAFF92FD0BD04B" masterDocTitle="Balkanian species of the genus Isoperla Banks, 1906 (Plecoptera: Perlodidae)" masterLastPageNumber="46" masterPageNumber="1" pageNumber="32" updateTime="1699105341794" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="48D84A2CB5A90B49930245C01E11DF34">Balkanian species of the genus Isoperla Banks, 1906 (Plecoptera: Perlodidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="C722C42F1F8F43B99E561CB47C6C3828" type="personal">
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<mods:namePart id="C6929F2DBAFFD349871A360EAC44A542">Murányi, Dávid</mods:namePart>
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<mods:typeOfResource id="52A6214A6A0650A02A013E7EC8353D78">text</mods:typeOfResource>
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<mods:title id="218392AD874CA8D7808EB9DAE6AD090E">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="241C56A66C5688449ACD4899CF69B7BC">
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<mods:date id="8A75AD5E4B344D1EC37783800AC2D53B">2011</mods:date>
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<mods:number id="86F979F25996807170EC94CE946170AC">2011-10-05</mods:number>
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<mods:detail id="2F7A0370843AE1E1589C9175FBEEA202" type="volume">
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<mods:number id="53CD39AB0555728CC695F7ACB32EB542">3049</mods:number>
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</mods:detail>
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<mods:start id="D548E19339EBBE50D7A458EC25205283">1</mods:start>
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<mods:classification id="3B23EECDD41420B1C1497C06FF29FDE8">journal article</mods:classification>
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<mods:identifier id="1158E874738775A36559085F9DA3B173" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<treatment id="03903327FF8BFFF2FF6DFAA0F9B6D134" ID-DOI="http://doi.org/10.5281/zenodo.5282971" ID-Zenodo-Dep="5282971" LSID="urn:lsid:plazi:treatment:03903327FF8BFFF2FF6DFAA0F9B6D134" httpUri="http://treatment.plazi.org/id/03903327FF8BFFF2FF6DFAA0F9B6D134" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
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<subSubSection id="C323D1BAFF8BFFCDFF6DFAA0FF5DD507" box="[151,598,1330,1356]" pageId="31" pageNumber="32" type="nomenclature">
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<paragraph id="8B868231FF8BFFCDFF6DFAA0FF5DD507" blockId="31.[151,598,1330,1356]" box="[151,598,1330,1356]" pageId="31" pageNumber="32">
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<heading id="D0CE355DFF8BFFCDFF6DFAA0FF5DD507" bold="true" box="[151,598,1330,1356]" fontSize="11" level="1" pageId="31" pageNumber="32" reason="1">
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<emphasis id="B94D5E23FF8BFFCDFF6DFAA0FF5DD507" bold="true" box="[151,598,1330,1356]" pageId="31" pageNumber="32">
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<taxonomicName id="4C39F9B2FF8BFFCDFF6DFAA0FCF0D507" ID-CoL="6MZLD" authority="Zwick, 1978" authorityName="Zwick" authorityYear="1978" box="[151,507,1330,1356]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="obliqua">
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<emphasis id="B94D5E23FF8BFFCDFF6DFAA0FC57D507" bold="true" box="[151,348,1330,1356]" italics="true" pageId="31" pageNumber="32">Isoperla obliqua</emphasis>
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<bibRefCitation id="EFA8FFC0FF8BFFCDFE9EFAA0FCF0D507" author="Zwick" box="[356,507,1330,1356]" firstAuthor="Zwick" pageId="31" pageNumber="32" pagination="213 - 239" refId="ref30884" refString="Zwick, P. (1978) Steinfliegen (Plecoptera) aus Griechenland und benachbarten Landern. Mitteilungen der Schweizerischen entomologischen Gesellschaft, 51, 21 - 38, 213 - 239." type="journal article" year="1978">Zwick, 1978</bibRefCitation>
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</taxonomicName>
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stat. n.
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C323D1BAFF8BFFCDFF6DFAEAF939D5C5" box="[151,1074,1400,1422]" pageId="31" pageNumber="32" type="reference_group">
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<paragraph id="8B868231FF8BFFCDFF6DFAEAF939D5C5" blockId="31.[151,1074,1400,1422]" box="[151,1074,1400,1422]" pageId="31" pageNumber="32">
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<treatmentCitationGroup id="AB29A51FFF8BFFCDFF6DFAEAF939D5C5" box="[151,1074,1400,1422]" pageId="31" pageNumber="32">
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<taxonomicName id="4C39F9B2FF8BFFCDFF6DFAEAFF23D5C5" ID-CoL="6MZLD" authority="Zwick, 1978" authorityName="Zwick" authorityYear="1978" box="[151,552,1400,1422]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="subSpecies" species="tripartita" subSpecies="obliqua">
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<emphasis id="B94D5E23FF8BFFCDFF6DFAEAFCA9D5C5" box="[151,418,1400,1422]" italics="true" pageId="31" pageNumber="32">Isoperla tripartita obliqua</emphasis>
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<bibRefCitation id="EFA8FFC0FF8BFFCDFE52FAEAFF23D5C5" author="Zwick" box="[424,552,1400,1422]" firstAuthor="Zwick" pageId="31" pageNumber="32" pagination="213 - 239" refId="ref30884" refString="Zwick, P. (1978) Steinfliegen (Plecoptera) aus Griechenland und benachbarten Landern. Mitteilungen der Schweizerischen entomologischen Gesellschaft, 51, 21 - 38, 213 - 239." type="journal article" year="1978">Zwick, 1978</bibRefCitation>
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</taxonomicName>
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—
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<treatmentCitation id="0A98A420FF8BFFCDFDB4FAEAFFF0D5C5" author="Zwick, P." box="[590,763,1400,1422]" page="230" pageId="31" pageNumber="32" year="1978">
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<bibRefCitation id="EFA8FFC0FF8BFFCDFDB4FAEAFFF0D5C5" author="Zwick" box="[590,763,1400,1422]" firstAuthor="Zwick" page="230" pageId="31" pageNumber="32" pagination="213 - 239" refId="ref30884" refString="Zwick, P. (1978) Steinfliegen (Plecoptera) aus Griechenland und benachbarten Landern. Mitteilungen der Schweizerischen entomologischen Gesellschaft, 51, 21 - 38, 213 - 239." type="journal article" year="1978">Zwick 1978: 230</bibRefCitation>
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</treatmentCitation>
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(original description of male).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C323D1BAFF8BFFCDFF6DFA2FF9F0D5BC" pageId="31" pageNumber="32" type="materials_examined">
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<paragraph id="8B868231FF8BFFCDFF6DFA2FF9F0D5BC" blockId="31.[151,1436,1469,2015]" pageId="31" pageNumber="32">
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<emphasis id="B94D5E23FF8BFFCDFF6DFA2FFC82D59D" bold="true" box="[151,393,1469,1494]" pageId="31" pageNumber="32">Material examined:</emphasis>
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<materialsCitation id="3B51886CFF8BFFCDFE6FFA2FF9FCD5BC" collectingDate="1992-05-25" collectionCode="HNHM" collectorName="H. Malicky" country="Greece" county="Selinous Stream" determinerName="G. Vincon" elevation="1000" location="Achaia" municipality="Mts Erimanthos" pageId="31" pageNumber="32" specimenCount="1">
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<collectingCountry id="F32EC2A1FF8BFFCDFE6FFA2FFF0CD59E" box="[405,519,1469,1493]" name="Greece" pageId="31" pageNumber="32">GREECE</collectingCountry>
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:
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<location id="8EE6D4EAFF8BFFCDFDEDFA2FFF6DD59E" LSID="urn:lsid:plazi:treatment:03903327FF8BFFF2FF6DFAA0F9B6D134:8EE6D4EAFF8BFFCDFDEDFA2FFF6DD59E" box="[535,614,1469,1493]" country="Greece" county="Selinous Stream" municipality="Mts Erimanthos" name="Achaia" pageId="31" pageNumber="32">Achaia</location>
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prefecture:
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<location id="8EE6D4EAFF8BFFCDFD03FA2FFE87D59E" LSID="urn:lsid:plazi:treatment:03903327FF8BFFF2FF6DFAA0F9B6D134:8EE6D4EAFF8BFFCDFD03FA2FFE87D59E" box="[761,908,1469,1493]" country="Greece" county="Selinous Stream" municipality="Mts Erimanthos" name="Ano Vlassia" pageId="31" pageNumber="32">Ano Vlassia</location>
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,
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<collectingMunicipality id="6BE2184BFF8BFFCDFC67FA2CF952D59E" box="[925,1113,1469,1493]" pageId="31" pageNumber="32">Mts Erimanthos</collectingMunicipality>
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,
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<collectingCounty id="62E7FABDFF8BFFCDFB90FA2FF822D59E" box="[1130,1321,1469,1493]" pageId="31" pageNumber="32">Selinous Stream</collectingCounty>
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,
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<quantity id="4CC12FD4FF8BFFCDFAC0FA2FF89CD59E" box="[1338,1431,1469,1493]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="31" pageNumber="32" unit="m" value="1000.0">
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<elevation id="00146502FF8BFFCDFAC0FA2FF89CD59E" box="[1338,1431,1469,1493]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="31" pageNumber="32" unit="m" value="1000.0">1000 m</elevation>
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</quantity>
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,
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<date id="FF87A4F1FF8BFFCDFF6DFA4DFC1CD5BC" box="[151,279,1503,1527]" pageId="31" pageNumber="32" value="1992-05-25">
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<collectingDate id="EFC35D19FF8BFFCDFF6DFA4DFC1CD5BC" box="[151,279,1503,1527]" pageId="31" pageNumber="32" value="1992-05-25">25.05.1992</collectingDate>
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</date>
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, leg.
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<collectorName id="26CCE7E7FF8BFFCDFEA9FA72FCDFD5BC" box="[339,468,1503,1527]" pageId="31" pageNumber="32">H. Malicky</collectorName>
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, det.
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<determinerName id="7E5A6EF5FF8BFFCDFDF5FA4DFF88D5BC" box="[527,643,1503,1527]" pageId="31" pageNumber="32">G. Vinçon</determinerName>
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||
:
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||
<quantity id="4CC12FD4FF8BFFCDFD77FA4DFFBBD5BC" box="[653,688,1503,1527]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="31" pageNumber="32" unit="m" value="2.0">
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||
<elevation id="00146502FF8BFFCDFD77FA4DFFBBD5BC" box="[653,688,1503,1527]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="31" pageNumber="32" unit="m" value="2.0">2m</elevation>
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</quantity>
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2f (
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<collectionCode id="ED281AF4FF8BFFCDFD24FA4DFE32D5BC" box="[734,825,1503,1527]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="31" pageNumber="32">HNHM</collectionCode>
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; one penial armature prepared on slide)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C323D1BAFF8BFFCDFF3DF990FFB9D795" pageId="31" pageNumber="32" type="diagnosis">
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||
<paragraph id="8B868231FF8BFFCDFF3DF990FC30D6C8" blockId="31.[151,1436,1469,2015]" pageId="31" pageNumber="32">
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<emphasis id="B94D5E23FF8BFFCDFF3DF990FC48D650" bold="true" box="[199,323,1538,1563]" pageId="31" pageNumber="32">Diagnosis:</emphasis>
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This species is characterized by a coloured medial penial armature that is divided into an upper and a lower part. Its upper medial armature is further divided into diverging, tear drop-shaped left and right arms with a lower medial armature that is weakly subdivided. All scales on armatures are spike-like and the lateral penial armatures vestigial.
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</paragraph>
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<paragraph id="8B868231FF8BFFCDFF3DF91CFFFFD6A7" blockId="31.[151,1436,1469,2015]" pageId="31" pageNumber="32">
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<emphasis id="B94D5E23FF8BFFCDFF3DF91CFF2AD6EC" bold="true" box="[199,545,1678,1703]" pageId="31" pageNumber="32">Complementary description:</emphasis>
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General appearence, dimensions, male and female abdomen: Similar to
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<taxonomicName id="4C39F9B2FF8BFFCDFAA5F91DFC58D682" authority="tripartita" authorityName="tripartita Illies" authorityYear="1954" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="subSpecies" species="tripartita" subSpecies="tripartita">
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<emphasis id="B94D5E23FF8BFFCDFAA5F91DFC58D682" italics="true" pageId="31" pageNumber="32">I. tripartita tripartita</emphasis>
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</taxonomicName>
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as redescribed above. Head and pronotal pattern of the examined specimens remind one of that depicted in
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<figureCitation id="13029EB4FF8BFFCDFEE3F946FC60D6A7" box="[281,363,1748,1772]" captionStart="FIGURES 94–98" captionStartId="29.[151,264,1941,1964]" captionTargetBox="[166,1414,919,1928]" captionTargetId="figure-404@29.[151,1436,901,1947]" captionTargetPageId="29" captionText="FIGURES 94–98. Isoperla tripartita tripartita Illies, 1954, variations of head and pronotal pattern — 94: Albania, Mts Prokletije; 95: Macedonia, Mts Jakupica; 96: Albania, Mts Prokletije; 97: Albania, Mts Gjalica e Lumës; 98: Albania, Mts Ostrovicë — scale 1 mm (for all figures)." pageId="31" pageNumber="32">Fig. 96</figureCitation>
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, but with rugosities on the occiput.
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</paragraph>
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<paragraph id="8B868231FF8BFFCDFF3DF964FC19D7D2" blockId="31.[151,1436,1469,2015]" pageId="31" pageNumber="32">
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Penis: Lobes and the distribution of non coloured scales, hairs and sensilla cannot be studied in detail because the specimens were collected without an everted penis. However, the narrow stripe of hydra-like scales, characteristic for the species group, can be detected. The medial penial armature is in agreement with the original description (
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<bibRefCitation id="EFA8FFC0FF8BFFCDFF28F8CCFC54D73D" author="Zwick" box="[210,351,1886,1910]" firstAuthor="Zwick" pageId="31" pageNumber="32" pagination="213 - 239" refId="ref30884" refString="Zwick, P. (1978) Steinfliegen (Plecoptera) aus Griechenland und benachbarten Landern. Mitteilungen der Schweizerischen entomologischen Gesellschaft, 51, 21 - 38, 213 - 239." type="journal article" year="1978">Zwick 1978</bibRefCitation>
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:
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<figureCitation id="13029EB4FF8BFFCDFE91F8CCFCE4D73D" box="[363,495,1886,1910]" captionStart="FIGURES 67–72" captionStartId="24.[151,263,1669,1692]" captionTargetBox="[169,1435,573,1643]" captionTargetId="figure-283@24.[151,1436,566,1648]" captionTargetPageId="24" captionText="FIGURES 67–72. Isoperla tripartita tripartita Illies, 1954 — 67: head and pronotum; 68: male terminalia, ventral view; 69: male terminalia, dorsal view; 70: meso- and metathorax, ventral view; 71: female terminalia, ventral view; 72: extruded penis, dorsolateral view (ML: medial lobe; LL: lateral lobe; VL: ventral lobe; BS: basal section) — 67–71: Austria, Wienerwald; 72: Hungary, Mts Börzsöny — scale 1 mm: 67–71; 0.5 mm: 72." pageId="31" pageNumber="32">Figs. 72j, k</figureCitation>
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). Lateral penial armatures are vestigial, consisting of only a few uncoloured spikelike scales.
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</paragraph>
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<paragraph id="8B868231FF8BFFCDFF3DF836FCB6D7F7" blockId="31.[151,1436,1469,2015]" box="[199,445,1956,1980]" pageId="31" pageNumber="32">Larva, egg: unknown.</paragraph>
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<paragraph id="8B868231FF8BFFCDFF3DF854FFB9D795" blockId="31.[151,1436,1469,2015]" box="[199,690,1990,2015]" pageId="31" pageNumber="32">
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<emphasis id="B94D5E23FF8BFFCDFF3DF854FC34D794" bold="true" box="[199,319,1990,2015]" pageId="31" pageNumber="32">Affinities:</emphasis>
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See
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<taxonomicName id="4C39F9B2FF8BFFCDFE8FF855FF6AD795" authority="tripartita" authorityName="tripartita Illies" authorityYear="1954" box="[373,609,1991,2014]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="subSpecies" species="tripartita" subSpecies="tripartita">
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<emphasis id="B94D5E23FF8BFFCDFE8FF855FF6AD795" box="[373,609,1991,2014]" italics="true" pageId="31" pageNumber="32">I. tripartita tripartita</emphasis>
|
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</taxonomicName>
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above.
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</paragraph>
|
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</subSubSection>
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||
<subSubSection id="C323D1BAFFB4FFF2FF3DFF0AFE01D15C" pageId="32" pageNumber="33" type="biology_ecology">
|
||
<paragraph id="8B868231FFB4FFF2FF3DFF0AFE01D15C" blockId="32.[151,1436,151,384]" pageId="32" pageNumber="33">
|
||
<emphasis id="B94D5E23FFB4FFF2FF3DFF0AFF08D0FB" bold="true" box="[199,515,151,176]" pageId="32" pageNumber="33">Ecology and distribution:</emphasis>
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||
<materialsCitation id="3B51886CFFB4FFF2FDF5FF05FE0ED15C" location="Andros Island" municipality="Greek Euboea Island" pageId="32" pageNumber="33" specimenCount="1" stateProvince="Peloponnisos">
|
||
The species was originally described from the
|
||
<collectingMunicipality id="6BE2184BFFB4FFF2FBBAFF05F833D0E4" box="[1088,1336,151,175]" pageId="32" pageNumber="33">Greek Euboea Island</collectingMunicipality>
|
||
,
|
||
<location id="8EE6D4EAFFB4FFF2FAB3FF05FDD5D099" LSID="urn:lsid:plazi:treatment:03903327FF8BFFF2FF6DFAA0F9B6D134:8EE6D4EAFFB4FFF2FAB3FF05FDD5D099" municipality="Greek Euboea Island" name="Andros Island" pageId="32" pageNumber="33" stateProvince="Peloponnisos">Andros Island</location>
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||
, the
|
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<collectingRegion id="49FD4CD3FFB4FFF2FEE6FF28FC96D099" box="[284,413,186,210]" country="Greece" name="Peloponnisos" pageId="32" pageNumber="33">Peloponnes</collectingRegion>
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and the Greek mainland (
|
||
<bibRefCitation id="EFA8FFC0FFB4FFF2FD22FF28FE61D099" author="Zwick" box="[728,874,186,210]" firstAuthor="Zwick" pageId="32" pageNumber="33" pagination="213 - 239" refId="ref30884" refString="Zwick, P. (1978) Steinfliegen (Plecoptera) aus Griechenland und benachbarten Landern. Mitteilungen der Schweizerischen entomologischen Gesellschaft, 51, 21 - 38, 213 - 239." type="journal article" year="1978">Zwick 1978</bibRefCitation>
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||
). The species has not been referenced since its description. The known localities are submontane streams and spring outlets, the adults being caught in May, usually in common with a diverse community of stoneflies
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</materialsCitation>
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.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C323D1BAFFB4FFF2FF3DFEB1F9B6D134" pageId="32" pageNumber="33" type="discussion">
|
||
<paragraph id="8B868231FFB4FFF2FF3DFEB1F9B6D134" blockId="32.[151,1436,151,384]" pageId="32" pageNumber="33">
|
||
<emphasis id="B94D5E23FFB4FFF2FF3DFEB1FC36D170" bold="true" box="[199,317,291,315]" pageId="32" pageNumber="33">Remarks:</emphasis>
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The area of the species completely overlaps with
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<taxonomicName id="4C39F9B2FFB4FFF2FC93FEB1F958D171" authority="tripartita" authorityName="tripartita Illies" authorityYear="1954" box="[873,1107,291,314]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="tripartita" subSpecies="tripartita">
|
||
<emphasis id="B94D5E23FFB4FFF2FC93FEB1F958D171" box="[873,1107,291,314]" italics="true" pageId="32" pageNumber="33">I. tripartita tripartita</emphasis>
|
||
</taxonomicName>
|
||
, so I propose full species status herein. Nevertheless, as it was already stated in the original description (
|
||
<bibRefCitation id="EFA8FFC0FFB4FFF2FBDDFED7F9BCD116" author="Zwick" box="[1063,1207,325,349]" firstAuthor="Zwick" pageId="32" pageNumber="33" pagination="213 - 239" refId="ref30884" refString="Zwick, P. (1978) Steinfliegen (Plecoptera) aus Griechenland und benachbarten Landern. Mitteilungen der Schweizerischen entomologischen Gesellschaft, 51, 21 - 38, 213 - 239." type="journal article" year="1978">Zwick 1978</bibRefCitation>
|
||
), transitional forms towards
|
||
<taxonomicName id="4C39F9B2FFB4FFF2FF0CFEFBFCE9D134" authority="tripartita" authorityName="tripartita Illies" authorityYear="1954" box="[246,482,360,384]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="tripartita" subSpecies="tripartita">
|
||
<emphasis id="B94D5E23FFB4FFF2FF0CFEFBFCE9D134" box="[246,482,360,384]" italics="true" pageId="32" pageNumber="33">I. tripartita tripartita</emphasis>
|
||
</taxonomicName>
|
||
occur and the validity of the taxon requires further investigation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |