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<mods:title id="2BA956D82B4FBE5B9157752CC0C0A5D4">Phylogenetic analysis of the circum-Antarctic Subfamily Migadopinae (Coleoptera, Carabidae) and assessment of the trans-Tasman Amarotypus clade</mods:title>
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<mods:namePart id="CB64C37BCFB26C49428253217B3FAEE6">Liebherr, James K.</mods:namePart>
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<mods:namePart id="F42286D8F3E115A8F04154575D38DE6A">Roig-Juñent, Sergio</mods:namePart>
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<mods:affiliation id="C989BE0E5A35CB7102CAA5577A80D44F">Laboratorio de Entomología, Instituto Argentina de Investigaciones de las Zonas Áridas (IADIZA, CCT CONICET, Mendoza), 5500 Mendoza, Argentina</mods:affiliation>
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<mods:namePart id="590C5CFED0A38DF34949BC39306AE6D9">Will, Kipling W.</mods:namePart>
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<mods:date id="9407A619968752E6D35D3AED0459009E">2024</mods:date>
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<subSubSection id="B23631271D0477B4CF0F27A95EE962ED" type="nomenclature">
<paragraph id="95729367785976A01BC8FCAF4C6A6CF9">
<treatmentCitationGroup id="EE5BB63886D7081D155347990AF484A7">
<treatmentCitation id="413816CAC0AA033B9B37340C6AC3F0D1" author="Larochelle A &amp; Larivière M-C" page="10" year="2022">
<taxonomicName id="D0C3D7F0A3EC22F20B4C99FEC70F9F43" authority="Larochelle &amp; Lariviere, 2022: 10" authorityName="Larochelle &amp; Lariviere" authorityPageNumber="10" authorityYear="2022" class="Insecta" family="Carabidae" genus="Amaroxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="EC1D59F6AF515B1669C216F90AED64A9" italics="true">Amaroxenus</emphasis>
<bibRefCitation id="BEDF73E138B1C305849E307A921329BA" DOI="10.5281/zenodo.7300590" author="Larochelle A &amp; Larivière M-C" journalOrPublisher="Insecta Mundi" pagination="1-30" refId="B28" refString="Larochelle A, Lariviere M-C (2022) Synopsis of the tribe Amarotypini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 0942: 1 - 30. https: // doi. org / 10.5281 / zenodo. 7300590" title="Synopsis of the tribe Amarotypini in New Zealand (Coleoptera: Carabidae)." volume="942" year="2022">Larochelle &amp; Larivière, 2022: 10</bibRefCitation>
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.
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</paragraph>
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<subSubSection id="SECID0EIJBG" type="type species">
<paragraph id="8C712B5AF268132BE5A764BF0EB0FA51">
<heading id="DE28FF3924D188CD68E35394EF66458F" reason="title">
<typeStatus id="C00D02FD98B26C4C2EDD2FEE52671E1D">Type</typeStatus>
species.
</heading>
</paragraph>
<paragraph id="19E8242897C5F73ABCB281F15CC7FEA3">
<treatmentCitationGroup id="CD24AFB45E8BDFCA47CF5F0725AA8ADD">
<treatmentCitation id="9E992953AB7DFAFEB489C161D0A82151" author="Larochelle A &amp; Larivière M-C" page="10" year="2022">
<taxonomicName id="C73A96D1C46975E902BAE7FF1895CCFB" authority="Larochelle &amp; Lariviere, 2022: 10" authorityName="Larochelle &amp; Lariviere" authorityPageNumber="10" authorityYear="2022" class="Insecta" family="Carabidae" genus="Amaroxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="kahurangiensis">
<emphasis id="67C6384402574DA131F23F4AED5FBB46" italics="true">Amaroxenus kahurangiensis</emphasis>
<bibRefCitation id="032884856A866B4F71CF958D7EE47E15" DOI="10.5281/zenodo.7300590" author="Larochelle A &amp; Larivière M-C" journalOrPublisher="Insecta Mundi" pagination="1-30" refId="B28" refString="Larochelle A, Lariviere M-C (2022) Synopsis of the tribe Amarotypini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 0942: 1 - 30. https: // doi. org / 10.5281 / zenodo. 7300590" title="Synopsis of the tribe Amarotypini in New Zealand (Coleoptera: Carabidae)." volume="942" year="2022">Larochelle &amp; Larivière, 2022: 10</bibRefCitation>
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(by original designation).
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</paragraph>
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<subSubSection id="SECID0EZJBG" type="diagnosis">
<paragraph id="3CD08137F66E951D8FD4490403F8E3C0">
<heading id="84502DB8D68B3E93E9E507539C073E80" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="C2BF389688FBF740AFA5E1A20E5AC026">
Within
<taxonomicName id="DFF7683C9FAE10D061EE263751A93271" authorityName="Chaudoir" authorityYear="1861" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="tribe" tribe="Migadopini">Migadopini</taxonomicName>
exhibiting an elongate parascutellar stria that extends beyond the elytral mid-length; mentum medially bidentate, the mentum tooth notched medially; submentum with six setae; pronotum cordate, with basal width subequal to apical width, the lateral margin sinuate anteriad distinct hind angle; pronotal anterior marginal bead usually incomplete medially — in
<taxonomicName id="0A326FB34509D83A002A083B50FD7C6B" authorityName="Liebherr &amp; Roig-Juñent &amp; Will" authorityYear="2024" class="Insecta" family="Carabidae" genus="Amaroxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="embersoni" status="sp. nov.">
<emphasis id="B181959CCF214A53CC6DA9CE6D4DC8B4" italics="true">A. embersoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9D5A80A10008AEF7CF1EFA6E7E81E664" rank="species">sp. nov.</taxonomicNameLabel>
polymorphically complete or incomplete medially — always present laterally; metathoracic wings vestigial; pro- and mesothoracic legs with tarsomeres 14 laterally expanded in both sexes; female gonocoxite unipartite, the narrower apical portion indistinctly defined relative to the equally sclerotized, broader basal portion, no microtrichia — or a single small seta — situated along the inner margin of the gonocoxite at the point of apical narrowing (Fig.
<figureCitation id="FC1F532D173778B142CC2E191EBE6574" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Female left gonocoxite, ventral view. A. Amarotypus edwardsii; B. Amaroxenus embersoni; C. Amaroxenus marrisi; D. Migadopiella octoguttata; E. Amarophilus otagoensis." figureDoi="10.3897/dez.71.134268.figure3" httpUri="https://binary.pensoft.net/fig/1177804">3 B, C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ETKBG" type="Diversity">
<paragraph id="4F397DDC84B87469CD709AFAB5C8F0D1">
<heading id="A2EC689F198232A506F45B1014BEC319" reason="title">Diversity.</heading>
</paragraph>
<paragraph id="AEF722A94B44407D8C296DBE15E13D51">
Four species were described by
<bibRefCitation id="B4E82AA61C42E4F66A416F6215263CF6" DOI="10.5281/zenodo.7300590" author="Larochelle A &amp; Larivière M-C" firstAuthor="Larochelle" journalOrPublisher="Insecta Mundi" pagination="1-30" refId="B28" refString="Larochelle A, Larivière M-C (2022) Synopsis of the tribe Amarotypini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 0942: 130. https://doi.org/10.5281/zenodo.7300590" title="Synopsis of the tribe Amarotypini in New Zealand (Coleoptera: Carabidae)." volume="942" year="2022">Larochelle and Larivière (2022)</bibRefCitation>
, to which we add two new species, with the six species determinable using the following key.
</paragraph>
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