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<document id="D8F92EA918216F3A21F40A9166A64192" ID-DOI="10.11646/zootaxa.4576.3.6" ID-GBIF-Dataset="7abdc62c-c350-49dd-8060-4ed64da07362" ID-ISSN="1175-5326" ID-Zenodo-Dep="3715772" ID-ZooBank="56BC8573-D4A1-4B18-9BF6-7AB5F7984BFD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1584603665960" checkinUser="plazi" docAuthor="Muona, Jyrki" docDate="2019" docId="03D387E9FFAB776AFF6FE904FD63FA2E" docLanguage="en" docName="zootaxa.4576.3.6.pdf" docOrigin="Zootaxa 4576 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pactopus burmensis Muona 2019, new species" docType="treatment" docVersion="5" lastPageNumber="541" masterDocId="FFEAFF91FFA7777EFFF8EE40FF81FFA7" masterDocTitle="Throscidae (Coleoptera) relationships, with descriptions of new fossil genera and species" masterLastPageNumber="543" masterPageNumber="521" pageNumber="533" updateTime="1698730058212" updateUser="plazi">
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<mods:titleInfo id="0637DDA68065FB5AC9D87354B7AF0450">
<mods:title id="CBEBDFD78E56F8DA4DBAE2239A0FA2CE">Throscidae (Coleoptera) relationships, with descriptions of new fossil genera and species</mods:title>
</mods:titleInfo>
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<mods:namePart id="8A50DA4CCFEDF1A267DC835B7CCCC89C">Muona, Jyrki</mods:namePart>
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<mods:typeOfResource id="C46CA166FD8E9F75AACBB900183E050D">text</mods:typeOfResource>
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<mods:title id="294966BDB895DD9A3E1B769175DF5B5C">Zootaxa</mods:title>
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<mods:part id="9F273BBC4B543D9A43ACD2BF60477F33">
<mods:date id="1ECFDA2737137F05FADC6BFD4FE08B74">2019</mods:date>
<mods:detail id="022978D2A385CFC0991DE08361090AFC" type="pubDate">
<mods:number id="83A46870358818C2792F0F16E86335BE">2019-04-03</mods:number>
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<mods:number id="5192107A501B7A941302D3F79FB6928D">4576</mods:number>
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<mods:classification id="C1F13AEB03B32F6F0250BAA8D8747516">journal article</mods:classification>
<mods:identifier id="022044CCDA34BC12820142B6BB455A39" type="DOI">10.11646/zootaxa.4576.3.6</mods:identifier>
<mods:identifier id="94F4E9B363E44E529CFD34D83D8CD388" type="GBIF-Dataset">7abdc62c-c350-49dd-8060-4ed64da07362</mods:identifier>
<mods:identifier id="EBDECEEEF43FBCE0819E41D72A8EB85C" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="146A76722085B6A3B9B9A16192AF34F0" type="Zenodo-Dep">3715772</mods:identifier>
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<treatment id="03D387E9FFAB776AFF6FE904FD63FA2E" ID-DOI="http://doi.org/10.5281/zenodo.3716782" ID-GBIF-Taxon="162639251" ID-Zenodo-Dep="3716782" LSID="urn:lsid:plazi:treatment:03D387E9FFAB776AFF6FE904FD63FA2E" httpUri="http://treatment.plazi.org/id/03D387E9FFAB776AFF6FE904FD63FA2E" lastPageId="20" lastPageNumber="541" pageId="12" pageNumber="533">
<subSubSection id="C3606574FFAB7772FF6FE904FDA0F8F9" box="[151,545,1860,1886]" pageId="12" pageNumber="533" type="nomenclature">
<paragraph id="8BC536FFFFAB7772FF6FE904FDA0F8F9" blockId="12.[151,545,1860,1919]" box="[151,545,1860,1886]" pageId="12" pageNumber="533">
<heading id="D08D8193FFAB7772FF6FE904FDA0F8F9" bold="true" box="[151,545,1860,1886]" fontSize="11" level="1" pageId="12" pageNumber="533" reason="1">
<emphasis id="B90EEAEDFFAB7772FF6FE904FDA0F8F9" bold="true" box="[151,545,1860,1886]" pageId="12" pageNumber="533">
<taxonomicName id="4C7A4D7CFFAB7772FF6FE904FE0BF8F9" ID-CoL="6C6B" ID-ENA="1710478" authority="Muona, 2019" authorityName="Muona" authorityYear="2019" box="[151,394,1860,1886]" class="Insecta" family="Throscidae" genus="Pactopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="533" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="B90EEAEDFFAB7772FF6FE904FE0BF8F9" bold="true" box="[151,394,1860,1886]" italics="true" pageId="12" pageNumber="533">Pactopus burmensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D5796FFAB7772FE6AE904FDA0F8F9" box="[402,545,1860,1886]" pageId="12" pageNumber="533" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3606574FFAB7772FF6FE926FEC4F8D8" box="[151,325,1894,1919]" pageId="12" pageNumber="533" type="description">
<paragraph id="8BC536FFFFAB7772FF6FE926FEC4F8D8" blockId="12.[151,545,1860,1919]" box="[151,325,1894,1919]" pageId="12" pageNumber="533">
<figureCitation id="13412A7AFFAB7772FF6FE926FE9CF8D8" box="[151,285,1894,1919]" captionStart-0="FIGURE 12" captionStart-1="FIGURE 13" captionStart-2="FIGURE 14" captionStartId-0="13.[151,250,1275,1297]" captionStartId-1="13.[151,250,1944,1966]" captionStartId-2="14.[151,250,1994,2016]" captionTargetBox-0="[151,1436,265,1253]" captionTargetBox-1="[217,1369,1347,1923]" captionTargetBox-2="[151,1436,589,1973]" captionTargetId-0="figure@13.[151,1436,265,1253]" captionTargetId-1="figure@13.[217,1369,1347,1923]" captionTargetId-2="figure@14.[151,1436,589,1973]" captionTargetPageId-0="13" captionTargetPageId-1="13" captionTargetPageId-2="14" captionText-0="FIGURE 12. Pactopus burmensis n. sp., front leg." captionText-1="FIGURE 13. Pactopus burmensis n. sp., ventral view. AtG = abdominal tarsal groove; MtG = metathoracic tarsal groove; PP = prosternal peg." captionText-2="FIGURE 14. Pactopus burmensis n. sp., abdomen, ventral view. AtG = abdominal tarsal groove." figureDoi-0="http://doi.org/10.5281/zenodo.3715798" figureDoi-1="http://doi.org/10.5281/zenodo.3715800" figureDoi-2="http://doi.org/10.5281/zenodo.3715802" httpUri-0="https://zenodo.org/record/3715798/files/figure.png" httpUri-1="https://zenodo.org/record/3715800/files/figure.png" httpUri-2="https://zenodo.org/record/3715802/files/figure.png" pageId="12" pageNumber="533">Figs. 12–14</figureCitation>
,
<figureCitation id="13412A7AFFAB7772FED1E926FEC4F8D8" box="[297,325,1894,1919]" captionStart="FIGURE 25" captionStartId="20.[151,250,1150,1172]" captionText="FIGURE 25. Pactopus burmensis n. sp., holotype." pageId="12" pageNumber="533">25</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3606574FFAB7770FF6FE9EEFEB2FF5F" lastPageId="14" lastPageNumber="535" pageId="12" pageNumber="533" type="materials_examined">
<paragraph id="8BC536FFFFAB7772FF6FE9EEFA1DF84D" blockId="12.[151,1436,1966,2027]" pageId="12" pageNumber="533">
<emphasis id="B90EEAEDFFAB7772FF6FE9EEFEC1F860" bold="true" box="[151,320,1966,1991]" pageId="12" pageNumber="533">Type material</emphasis>
.
<typeStatus id="54C1885DFFAB7772FEA9E9EEFE38F860" box="[337,441,1966,1991]" pageId="12" pageNumber="533" type="holotype">Holotype</typeStatus>
labeled:
<taxonomicName id="4C7A4D7CFFAB7772FDDEE9EFFC8AF860" authority="Muona, 2019" authorityName="Muona" authorityYear="2019" box="[550,779,1966,1991]" class="Insecta" family="Throscidae" genus="Pactopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="533" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="B90EEAEDFFAB7772FDDEE9EFFC8AF860" box="[550,779,1966,1991]" italics="true" pageId="12" pageNumber="533">Pactopus burmensis</emphasis>
</taxonomicName>
<emphasis id="B90EEAEDFFAB7772FCECE9EEFCD7F860" bold="true" box="[788,854,1966,1991]" pageId="12" pageNumber="533">
<taxonomicNameLabel id="A23D5796FFAB7772FCECE9EEFCD7F860" box="[788,854,1966,1991]" pageId="12" pageNumber="533" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
/
<typeStatus id="54C1885DFFAB7772FCA7E9EFFC77F860" box="[863,1014,1967,1991]" pageId="12" pageNumber="533" type="holotype">HOLOTYPE</typeStatus>
/J.Muona des. 10-2015. Second label with following data:
<date id="FFC4103FFFAB7772FE70E992FE7CF84C" box="[392,509,2002,2027]" pageId="12" pageNumber="533" value="1401" valueMax="1572">1401-1572</date>
/Burmese amber/LarvaAdult/Throscida
<quantity id="4C829B1AFFAB7772FC3FE992FBA1F84C" box="[967,1056,2002,2027]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="12" pageNumber="533" unit="mm" value="8.5">8.5 mm</quantity>
/98.79+-0.62 Ma. Embedded in a
</paragraph>
<paragraph id="8BC536FFFFAA7773FF6FEED7FC40FF72" blockId="13.[151,1436,151,213]" pageId="13" pageNumber="534">
cone-shaped, dark brown Burmese Amber piece,
<quantity id="4C829B1AFFAA7773FD29EED8FCA3FF08" box="[721,802,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="13" pageNumber="534" unit="mm" value="12.0">12 mm</quantity>
<date id="FFC4103FFFAA7773FCD3EED8FCDEFF17" box="[811,863,152,176]" pageId="13" pageNumber="534" value="2010-10">
x
<quantity id="4C829B1AFFAA7773FCBBEED8FCDEFF17" box="[835,863,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="534" unit="mm" value="10.0">10</quantity>
</date>
mm
<date id="FFC4103FFFAA7773FC66EED8FC53FF17" box="[926,978,152,176]" pageId="13" pageNumber="534" value="2010-10">
x
<quantity id="4C829B1AFFAA7773FC4EEED8FC53FF17" box="[950,978,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="534" unit="mm" value="10.0">10</quantity>
</date>
mm, sex unknown. Age 98.79 +/- 0.62 Mya. Numerous unidentified plant remains are present as well (
<figureCitation id="13412A7AFFAA7773FC98EEFCFC35FF72" box="[864,948,188,213]" captionStart="FIGURE 25" captionStartId="20.[151,250,1150,1172]" captionText="FIGURE 25. Pactopus burmensis n. sp., holotype." pageId="13" pageNumber="534">Fig. 25</figureCitation>
).
</paragraph>
<caption id="DF056677FFAA7773FF6FEABBFD1DFAB6" ID-DOI="http://doi.org/10.5281/zenodo.3715798" ID-Zenodo-Dep="3715798" box="[151,668,1275,1297]" httpUri="https://zenodo.org/record/3715798/files/figure.png" pageId="13" pageNumber="534" startId="13.[151,250,1275,1297]" targetBox="[151,1436,265,1253]" targetPageId="13">
<paragraph id="8BC536FFFFAA7773FF6FEABBFD1DFAB6" blockId="13.[151,668,1275,1297]" box="[151,668,1275,1297]" pageId="13" pageNumber="534">
<emphasis id="B90EEAEDFFAA7773FF6FEABBFE9EFAB6" bold="true" box="[151,287,1275,1297]" pageId="13" pageNumber="534">FIGURE 12.</emphasis>
<taxonomicName id="4C7A4D7CFFAA7773FEDDEABCFE73FAB6" authority="Muona, 2019" authorityName="Muona" authorityYear="2019" box="[293,498,1275,1297]" class="Insecta" family="Throscidae" genus="Pactopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="534" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="B90EEAEDFFAA7773FEDDEABCFE73FAB6" box="[293,498,1275,1297]" italics="true" pageId="13" pageNumber="534">Pactopus burmensis</emphasis>
</taxonomicName>
<emphasis id="B90EEAEDFFAA7773FE00EABBFDB0FAB6" bold="true" box="[504,561,1275,1297]" pageId="13" pageNumber="534">
<taxonomicNameLabel id="A23D5796FFAA7773FE00EABBFDB0FAB6" box="[504,561,1275,1297]" pageId="13" pageNumber="534" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, front leg.
</paragraph>
</caption>
<caption id="DF056677FFAA7773FF6FE9D8FEAFF876" ID-DOI="http://doi.org/10.5281/zenodo.3715800" ID-Zenodo-Dep="3715800" httpUri="https://zenodo.org/record/3715800/files/figure.png" pageId="13" pageNumber="534" startId="13.[151,250,1944,1966]" targetBox="[217,1369,1347,1923]" targetPageId="13">
<paragraph id="8BC536FFFFAA7773FF6FE9D8FEAFF876" blockId="13.[151,1435,1944,2001]" pageId="13" pageNumber="534">
<emphasis id="B90EEAEDFFAA7773FF6FE9D8FE9EF809" bold="true" box="[151,287,1944,1966]" pageId="13" pageNumber="534">FIGURE 13.</emphasis>
<taxonomicName id="4C7A4D7CFFAA7773FEDCE9DAFE70F809" authority="Muona, 2019" authorityName="Muona" authorityYear="2019" box="[292,497,1944,1967]" class="Insecta" family="Throscidae" genus="Pactopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="534" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="B90EEAEDFFAA7773FEDCE9DAFE70F809" box="[292,497,1944,1967]" italics="true" pageId="13" pageNumber="534">Pactopus burmensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D5796FFAA7773FE0EE9D8FDAEF809" box="[502,559,1944,1966]" pageId="13" pageNumber="534" rank="species">
<emphasis id="B90EEAEDFFAA7773FE0EE9D8FDA8F809" bold="true" box="[502,553,1944,1966]" pageId="13" pageNumber="534">n. sp</emphasis>
.
</taxonomicNameLabel>
, ventral view. AtG = abdominal tarsal groove; MtG = metathoracic tarsal groove; PP = prosternal peg.
</paragraph>
</caption>
<paragraph id="8BC536FFFFA97770FF3FEED8FEB2FF5F" blockId="14.[151,1436,151,536]" pageId="14" pageNumber="535">
<typeStatus id="54C1885DFFA97770FF3FEED8FEA8FF17" box="[199,297,152,176]" pageId="14" pageNumber="535" type="paratype">Paratype</typeStatus>
in a box-shaped piece of fairly lightly coloured Burmese Amber,
<quantity id="4C829B1AFFA97770FBC5EED8FB11FF08" box="[1085,1168,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="14" pageNumber="535" unit="mm" value="12.0">12 mm</quantity>
<date id="FFC4103FFFA97770FB63EED8FB42FF17" box="[1179,1219,151,176]" pageId="14" pageNumber="535">
x
<quantity id="4C829B1AFFA97770FB4DEED7FB42FF17" box="[1205,1219,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="14" pageNumber="535" unit="mm" value="6.0">6</quantity>
</date>
mm
<date id="FFC4103FFFA97770FAFEEED8FAACFF17" box="[1286,1325,151,176]" pageId="14" pageNumber="535">
x
<quantity id="4C829B1AFFA97770FAE7EED7FAACFF17" box="[1311,1325,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="14" pageNumber="535" unit="mm" value="6.0">6</quantity>
</date>
mm, sex unknown. Age 98.79 +/- 0.62 Mya. Some unidentified plant remains are present as well. Length
<quantity id="4C829B1AFFA97770FB1CEEFCFAC2FF73" box="[1252,1347,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="14" pageNumber="535" unit="mm" value="2.9">2. 9 mm</quantity>
(PT) to 3.0 mm (HT).
</paragraph>
</subSubSection>
<subSubSection id="C3606574FFA97770FF3FEF44FE2DFDBF" pageId="14" pageNumber="535" type="diagnosis">
<paragraph id="8BC536FFFFA97770FF3FEF44FE2DFDBF" blockId="14.[151,1436,151,536]" pageId="14" pageNumber="535">
<emphasis id="B90EEAEDFFA97770FF3FEF44FEBBFEBA" bold="true" box="[199,314,260,285]" pageId="14" pageNumber="535">Diagnosis</emphasis>
. The combination of
<taxonomicName id="4C7A4D7CFFA97770FDD7EF44FD1BFEBA" box="[559,666,260,285]" class="Insecta" family="Throscidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="535" phylum="Arthropoda" rank="tribe" tribe="Throscini">Throscini</taxonomicName>
<typeStatus id="54C1885DFFA97770FD5BEF45FD52FEBA" box="[675,723,261,285]" pageId="14" pageNumber="535">type</typeStatus>
protibiae (
<figureCitation id="13412A7AFFA97770FCB6EF44FC22FEBB" box="[846,931,260,285]" captionStart="FIGURE 12" captionStartId="13.[151,250,1275,1297]" captionTargetBox="[151,1436,265,1253]" captionTargetId="figure@13.[151,1436,265,1253]" captionTargetPageId="13" captionText="FIGURE 12. Pactopus burmensis n. sp., front leg." figureDoi="http://doi.org/10.5281/zenodo.3715798" httpUri="https://zenodo.org/record/3715798/files/figure.png" pageId="14" pageNumber="535">Fig. 12</figureCitation>
), metathoracic and abdominal tarsal grooves and unmodified head is unique among
<taxonomicName id="4C7A4D7CFFA97770FD97EF67FD6DFEE7" authorityName="Laporte" authorityYear="1840" box="[623,748,295,320]" class="Insecta" family="Throscidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="535" phylum="Arthropoda" rank="family">Throscidae</taxonomicName>
and places this species in
<taxonomicName id="4C7A4D7CFFA97770FBCFEF69FB1EFEE7" authorityName="LeConte" authorityYear="1868" box="[1079,1183,297,320]" class="Insecta" family="Throscidae" genus="Pactopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="535" phylum="Arthropoda" rank="genus">
<emphasis id="B90EEAEDFFA97770FBCFEF69FB1EFEE7" box="[1079,1183,297,320]" italics="true" pageId="14" pageNumber="535">Pactopus</emphasis>
</taxonomicName>
.
<taxonomicName id="4C7A4D7CFFA97770FB4CEF69FA1DFEE7" authorityName="Muona" authorityYear="2019" box="[1204,1436,295,320]" class="Insecta" family="Throscidae" genus="Pactopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="535" phylum="Arthropoda" rank="genus">
<emphasis id="B90EEAEDFFA97770FB4CEF69FA1DFEE7" box="[1204,1436,295,320]" italics="true" pageId="14" pageNumber="535">Pactopus burmensis</emphasis>
</taxonomicName>
resembles the extinct
<taxonomicName id="4C7A4D7CFFA97770FE70EF0DFD19FEC2" authority="Wickham" authorityName="Wickham" authorityYear="1914" box="[392,664,332,357]" class="Insecta" family="Throscidae" genus="Pactopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="535" phylum="Arthropoda" rank="species" species="americanus">
<emphasis id="B90EEAEDFFA97770FE70EF0DFDA4FEC3" box="[392,549,333,356]" italics="true" pageId="14" pageNumber="535">P. americanus</emphasis>
Wickham
</taxonomicName>
in being relatively wide in shape, having strongly bent tarsal grooves on metaventrite and long, straight abdominal tarsal grooves. The extant
<taxonomicName id="4C7A4D7CFFA97770FC3BEF31FBFFFE20" authority="Leconte" authorityName="Leconte" box="[963,1150,367,392]" class="Insecta" family="Throscidae" genus="Pactopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="535" phylum="Arthropoda" rank="species" species="horni">
<emphasis id="B90EEAEDFFA97770FC3BEF31FB9AFE2F" box="[963,1051,367,392]" italics="true" pageId="14" pageNumber="535">P. horni</emphasis>
Leconte
</taxonomicName>
and the Eocene
<taxonomicName id="4C7A4D7CFFA97770FAC2EF31FF67FE0B" authority="Muona" authorityName="Muona" authorityYear="1993" class="Insecta" family="Throscidae" genus="Pactopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="535" phylum="Arthropoda" rank="species" species="fasolti">
<emphasis id="B90EEAEDFFA97770FAC2EF31FA18FE2F" box="[1338,1433,367,392]" italics="true" pageId="14" pageNumber="535">P. fasolti</emphasis>
Muona
</taxonomicName>
and
<taxonomicName id="4C7A4D7CFFA97770FED8EFD5FE61FE0B" authority="Muona" authorityName="Muona" authorityYear="1993" box="[288,480,404,428]" class="Insecta" family="Throscidae" genus="Pactopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="535" phylum="Arthropoda" rank="species" species="fafneri">
<emphasis id="B90EEAEDFFA97770FED8EFD5FE08FE0B" box="[288,393,404,428]" italics="true" pageId="14" pageNumber="535">P. fafneri</emphasis>
Muona
</taxonomicName>
are more parallel-sided and much narrower than
<emphasis id="B90EEAEDFFA97770FBEFEFD5FA1DFE0A" box="[1047,1436,404,429]" italics="true" pageId="14" pageNumber="535">
<taxonomicName id="4C7A4D7CFFA97770FBEFEFD5FB2DFE0A" authorityName="Muona" authorityYear="2019" box="[1047,1196,404,429]" class="Insecta" family="Throscidae" genus="Pactopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="535" phylum="Arthropoda" rank="species" species="burmensis">P. burmensis</taxonomicName>
.
<taxonomicName id="4C7A4D7CFFA97770FB40EFD5FA1DFE0A" authorityName="Muona" authorityYear="2019" box="[1208,1436,404,429]" class="Insecta" family="Throscidae" genus="Pactopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="535" phylum="Arthropoda" rank="genus">Pactopus burmensis</taxonomicName>
</emphasis>
differs from
<taxonomicName id="4C7A4D7CFFA97770FEDCEFF9FE42FE77" authorityName="Wickham" authorityYear="1914" box="[292,451,441,464]" class="Insecta" family="Throscidae" genus="Pactopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="535" phylum="Arthropoda" rank="species" species="americanus">
<emphasis id="B90EEAEDFFA97770FEDCEFF9FE42FE77" box="[292,451,441,464]" italics="true" pageId="14" pageNumber="535">P. americanus</emphasis>
</taxonomicName>
in having the prosernal process widest in the apical third and narrowing both anteriorly and posteriorly and by having the tarsal gooves on metaventrite more abruptly bent and close to parallel to metanepisterna caudally.
</paragraph>
</subSubSection>
<caption id="DF056677FFA97770FF6FE98AFBF3F847" ID-DOI="http://doi.org/10.5281/zenodo.3715802" ID-Zenodo-Dep="3715802" box="[151,1138,1994,2016]" httpUri="https://zenodo.org/record/3715802/files/figure.png" pageId="14" pageNumber="535" startId="14.[151,250,1994,2016]" targetBox="[151,1436,589,1973]" targetPageId="14">
<paragraph id="8BC536FFFFA97770FF6FE98AFBF3F847" blockId="14.[151,1138,1994,2016]" box="[151,1138,1994,2016]" pageId="14" pageNumber="535">
<emphasis id="B90EEAEDFFA97770FF6FE98AFE9EF847" bold="true" box="[151,287,1994,2016]" pageId="14" pageNumber="535">FIGURE 14.</emphasis>
<taxonomicName id="4C7A4D7CFFA97770FEDDE98BFE73F847" authority="Muona, 2019" authorityName="Muona" authorityYear="2019" box="[293,498,1994,2016]" class="Insecta" family="Throscidae" genus="Pactopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="535" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="B90EEAEDFFA97770FEDDE98BFE73F847" box="[293,498,1994,2016]" italics="true" pageId="14" pageNumber="535">Pactopus burmensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D5796FFA97770FE00E98AFDB0F847" box="[504,561,1994,2016]" pageId="14" pageNumber="535" rank="species">
<emphasis id="B90EEAEDFFA97770FE00E98AFDAAF847" bold="true" box="[504,555,1994,2016]" pageId="14" pageNumber="535">n. sp</emphasis>
.
</taxonomicNameLabel>
, abdomen, ventral view. AtG = abdominal tarsal groove.
</paragraph>
</caption>
<caption id="DF056677FFA87771FF6FED8BFBB9FC46" ID-DOI="http://doi.org/10.5281/zenodo.3715804" ID-Zenodo-Dep="3715804" box="[151,1080,971,993]" httpUri="https://zenodo.org/record/3715804/files/figure.png" pageId="15" pageNumber="536" startId="15.[151,250,971,993]" targetBox="[151,1436,220,934]" targetPageId="15">
<paragraph id="8BC536FFFFA87771FF6FED8BFBB9FC46" blockId="15.[151,1080,971,993]" box="[151,1080,971,993]" pageId="15" pageNumber="536">
<emphasis id="B90EEAEDFFA87771FF6FED8BFE9EFC47" bold="true" box="[151,287,971,993]" pageId="15" pageNumber="536">FIGURE 15.</emphasis>
<taxonomicName id="4C7A4D7CFFA87771FEDDED8CFE5DFC46" authority="Muona, 2019" authorityName="Muona" authorityYear="2019" box="[293,476,971,993]" class="Insecta" family="Throscidae" genus="Trixagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="536" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="B90EEAEDFFA87771FEDDED8CFE5DFC46" box="[293,476,971,993]" italics="true" pageId="15" pageNumber="536">Trixagus parvulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D5796FFA87771FE1BED8BFD9DFC46" box="[483,540,971,993]" pageId="15" pageNumber="536" rank="species">
<emphasis id="B90EEAEDFFA87771FE1BED8BFD97FC46" bold="true" box="[483,534,971,993]" pageId="15" pageNumber="536">n. sp</emphasis>
.
</taxonomicNameLabel>
, head, frontal view. ExE = excised eyes; L = labrum.
</paragraph>
</caption>
<caption id="DF056677FFA87771FF6FE9F8FD67F869" ID-DOI="http://doi.org/10.5281/zenodo.3715806" ID-Zenodo-Dep="3715806" box="[151,742,1976,1998]" httpUri="https://zenodo.org/record/3715806/files/figure.png" pageId="15" pageNumber="536" startId="15.[151,250,1976,1998]" targetBox="[151,1436,1060,1917]" targetPageId="15">
<paragraph id="8BC536FFFFA87771FF6FE9F8FD67F869" blockId="15.[151,742,1976,1998]" box="[151,742,1976,1998]" pageId="15" pageNumber="536">
<emphasis id="B90EEAEDFFA87771FF6FE9F8FE9EF869" bold="true" box="[151,287,1976,1998]" pageId="15" pageNumber="536">FIGURE 16.</emphasis>
<taxonomicName id="4C7A4D7CFFA87771FEDDE9F9FE5DF869" authority="Muona, 2019" authorityName="Muona" authorityYear="2019" box="[293,476,1976,1998]" class="Insecta" family="Throscidae" genus="Trixagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="536" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="B90EEAEDFFA87771FEDDE9F9FE5DF869" box="[293,476,1976,1998]" italics="true" pageId="15" pageNumber="536">Trixagus parvulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D5796FFA87771FE1BE9F8FD9DF869" box="[483,540,1976,1998]" pageId="15" pageNumber="536" rank="species">
<emphasis id="B90EEAEDFFA87771FE1BE9F8FD97F869" bold="true" box="[483,534,1976,1998]" pageId="15" pageNumber="536">n. sp</emphasis>
.
</taxonomicNameLabel>
, ventrolateral view.
</paragraph>
</caption>
<caption id="DF056677FFB7776EFF6FEA13FD65FBCE" ID-DOI="http://doi.org/10.5281/zenodo.3715808" ID-Zenodo-Dep="3715808" box="[151,740,1107,1129]" httpUri="https://zenodo.org/record/3715808/files/figure.png" pageId="16" pageNumber="537" startId="16.[151,250,1107,1129]" targetBox="[171,1420,193,1097]" targetPageId="16">
<paragraph id="8BC536FFFFB7776EFF6FEA13FD65FBCE" blockId="16.[151,740,1107,1129]" box="[151,740,1107,1129]" pageId="16" pageNumber="537">
<emphasis id="B90EEAEDFFB7776EFF6FEA13FE9EFBCE" bold="true" box="[151,287,1107,1129]" pageId="16" pageNumber="537">FIGURE 17.</emphasis>
<taxonomicName id="4C7A4D7CFFB7776EFEDDEA14FE5DFBCE" authority="Muona, 2019" authorityName="Muona" authorityYear="2019" box="[293,476,1107,1129]" class="Insecta" family="Throscidae" genus="Trixagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="537" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="B90EEAEDFFB7776EFEDDEA14FE5DFBCE" box="[293,476,1107,1129]" italics="true" pageId="16" pageNumber="537">Trixagus parvulus</emphasis>
</taxonomicName>
<emphasis id="B90EEAEDFFB7776EFE1BEA13FD9DFBCE" bold="true" box="[483,540,1107,1129]" pageId="16" pageNumber="537">
<taxonomicNameLabel id="A23D5796FFB7776EFE1BEA13FD9DFBCE" box="[483,540,1107,1129]" pageId="16" pageNumber="537" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, dorsolateral view.
</paragraph>
</caption>
<caption id="DF056677FFB7776EFF6FE99EFC3DF853" ID-DOI="http://doi.org/10.5281/zenodo.3715810" ID-Zenodo-Dep="3715810" box="[151,956,2014,2036]" httpUri="https://zenodo.org/record/3715810/files/figure.png" pageId="16" pageNumber="537" startId="16.[151,250,2014,2036]" targetBox="[167,1426,1179,1999]" targetPageId="16">
<paragraph id="8BC536FFFFB7776EFF6FE99EFC3DF853" blockId="16.[151,956,2014,2036]" box="[151,956,2014,2036]" pageId="16" pageNumber="537">
<emphasis id="B90EEAEDFFB7776EFF6FE99EFE9EF853" bold="true" box="[151,287,2014,2036]" pageId="16" pageNumber="537">FIGURE 18.</emphasis>
<taxonomicName id="4C7A4D7CFFB7776EFEDDE99EFD70F853" authority="Kirejtshuk &amp; Kovalev" authorityName="Kirejtshuk &amp; Kovalev" box="[293,753,2014,2036]" class="Insecta" family="Throscidae" genus="Rhomboaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="537" phylum="Arthropoda" rank="species" species="gratiosa">
<emphasis id="B90EEAEDFFB7776EFEDDE99EFD88F853" box="[293,521,2014,2036]" italics="true" pageId="16" pageNumber="537">Rhomboaspis gratiosa</emphasis>
Kirejtshuk &amp; Kovalev
</taxonomicName>
, frontoventral view.
</paragraph>
</caption>
<caption id="DF056677FFB6776FFF6FEDEFFBDCFC62" ID-DOI="http://doi.org/10.5281/zenodo.3715812" ID-Zenodo-Dep="3715812" box="[151,1117,943,965]" httpUri="https://zenodo.org/record/3715812/files/figure.png" pageId="17" pageNumber="538" startId="17.[151,250,943,965]" targetBox="[267,1334,192,929]" targetPageId="17">
<paragraph id="8BC536FFFFB6776FFF6FEDEFFBDCFC62" blockId="17.[151,1117,943,965]" box="[151,1117,943,965]" pageId="17" pageNumber="538">
<emphasis id="B90EEAEDFFB6776FFF6FEDEFFE9EFC62" bold="true" box="[151,287,943,965]" pageId="17" pageNumber="538">FIGURE 19.</emphasis>
Cladogram 1. Black dots = unique synapomorphies, circles homoplastic changes.
</paragraph>
</caption>
<caption id="DF056677FFB6776FFF6FE8B4FBDCF8AD" ID-DOI="http://doi.org/10.5281/zenodo.3715814" ID-Zenodo-Dep="3715814" box="[151,1117,1780,1802]" httpUri="https://zenodo.org/record/3715814/files/figure.png" pageId="17" pageNumber="538" startId="17.[151,250,1780,1802]" targetBox="[241,1346,1046,1759]" targetPageId="17">
<paragraph id="8BC536FFFFB6776FFF6FE8B4FBDCF8AD" blockId="17.[151,1117,1780,1802]" box="[151,1117,1780,1802]" pageId="17" pageNumber="538">
<emphasis id="B90EEAEDFFB6776FFF6FE8B4FE9EF8AD" bold="true" box="[151,287,1780,1802]" pageId="17" pageNumber="538">FIGURE 20.</emphasis>
Cladogram 1. Black dots = unique synapomorphies, circles homoplastic changes.
</paragraph>
</caption>
<caption id="DF056677FFB5776CFF6FEDFAFBDCFC77" ID-DOI="http://doi.org/10.5281/zenodo.3715816" ID-Zenodo-Dep="3715816" box="[151,1117,954,976]" httpUri="https://zenodo.org/record/3715816/files/figure.png" pageId="18" pageNumber="539" startId="18.[151,250,954,976]" targetBox="[231,1355,197,930]" targetPageId="18">
<paragraph id="8BC536FFFFB5776CFF6FEDFAFBDCFC77" blockId="18.[151,1117,954,976]" box="[151,1117,954,976]" pageId="18" pageNumber="539">
<emphasis id="B90EEAEDFFB5776CFF6FEDFAFE9EFC77" bold="true" box="[151,287,954,976]" pageId="18" pageNumber="539">FIGURE 21.</emphasis>
Cladogram 3. Black dots = unique synapomorphies, circles homoplastic changes.
</paragraph>
</caption>
<caption id="DF056677FFB5776CFF6FE91FFD46F8D2" ID-DOI="http://doi.org/10.5281/zenodo.3715818" ID-Zenodo-Dep="3715818" box="[151,711,1887,1909]" httpUri="https://zenodo.org/record/3715818/files/figure.png" pageId="18" pageNumber="539" startId="18.[151,250,1887,1909]" targetBox="[178,1417,1018,1876]" targetPageId="18">
<paragraph id="8BC536FFFFB5776CFF6FE91FFD46F8D2" blockId="18.[151,711,1887,1909]" box="[151,711,1887,1909]" pageId="18" pageNumber="539">
<emphasis id="B90EEAEDFFB5776CFF6FE91FFE9EF8D2" bold="true" box="[151,287,1887,1909]" pageId="18" pageNumber="539">FIGURE 22.</emphasis>
<taxonomicName id="4C7A4D7CFFB5776CFEDDE91FFD9AF8D2" authority="Muona, 2019" authorityName="Muona" authorityYear="2019" box="[293,539,1887,1909]" class="Insecta" family="Throscidae" genus="Pseudothroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="539" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="B90EEAEDFFB5776CFEDDE91FFD9AF8D2" box="[293,539,1887,1909]" italics="true" pageId="18" pageNumber="539">Pseudothroscus balticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D5796FFB5776CFDDAE91FFDDBF8D2" box="[546,602,1887,1909]" pageId="18" pageNumber="539" rank="species">
<emphasis id="B90EEAEDFFB5776CFDDAE91FFDD5F8D2" bold="true" box="[546,596,1887,1909]" pageId="18" pageNumber="539">n. sp</emphasis>
.
</taxonomicNameLabel>
, holotype.
</paragraph>
</caption>
<caption id="DF056677FFB4776DFF6FEA67FD21FB9A" ID-DOI="http://doi.org/10.5281/zenodo.3715820" ID-Zenodo-Dep="3715820" box="[151,672,1063,1085]" httpUri="https://zenodo.org/record/3715820/files/figure.png" pageId="19" pageNumber="540" startId="19.[151,250,1063,1085]" targetBox="[216,1373,183,1042]" targetPageId="19">
<paragraph id="8BC536FFFFB4776DFF6FEA67FD21FB9A" blockId="19.[151,672,1063,1085]" box="[151,672,1063,1085]" pageId="19" pageNumber="540">
<emphasis id="B90EEAEDFFB4776DFF6FEA67FE9EFB9A" bold="true" box="[151,287,1063,1085]" pageId="19" pageNumber="540">FIGURE 23.</emphasis>
<taxonomicName id="4C7A4D7CFFB4776DFEDDEA67FE75FB9A" authority="Muona, 2019" authorityName="Muona" authorityYear="2019" box="[293,500,1063,1085]" class="Insecta" family="Throscidae" genus="Tyrannothroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="540" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="B90EEAEDFFB4776DFEDDEA67FE75FB9A" box="[293,500,1063,1085]" italics="true" pageId="19" pageNumber="540">Tyrannothroscus rex</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D5796FFB4776DFE02EA67FDB2FB9A" box="[506,563,1063,1085]" pageId="19" pageNumber="540" rank="species">
<emphasis id="B90EEAEDFFB4776DFE02EA67FDACFB9A" bold="true" box="[506,557,1063,1085]" pageId="19" pageNumber="540">n. sp</emphasis>
.
</taxonomicNameLabel>
, holotype.
</paragraph>
</caption>
<caption id="DF056677FFB4776DFF6FE9C5FD03F83C" ID-DOI="http://doi.org/10.5281/zenodo.3715822" ID-Zenodo-Dep="3715822" box="[151,642,1925,1947]" httpUri="https://zenodo.org/record/3715822/files/figure.png" pageId="19" pageNumber="540" startId="19.[151,250,1925,1947]" targetBox="[228,1352,1135,1910]" targetPageId="19">
<paragraph id="8BC536FFFFB4776DFF6FE9C5FD03F83C" blockId="19.[151,642,1925,1947]" box="[151,642,1925,1947]" pageId="19" pageNumber="540">
<emphasis id="B90EEAEDFFB4776DFF6FE9C5FEA1F83C" bold="true" box="[151,288,1925,1947]" pageId="19" pageNumber="540">FIGURE 24.</emphasis>
<taxonomicName id="4C7A4D7CFFB4776DFEDDE9C6FE57F83C" authority="Muona, 2019" authorityName="Muona" authorityYear="2019" box="[293,470,1925,1947]" class="Insecta" family="Throscidae" genus="Potergus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="540" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="B90EEAEDFFB4776DFEDDE9C6FE57F83C" box="[293,470,1925,1947]" italics="true" pageId="19" pageNumber="540">Potergus balticus</emphasis>
</taxonomicName>
<emphasis id="B90EEAEDFFB4776DFE25E9C5FD97F83C" bold="true" box="[477,534,1925,1947]" pageId="19" pageNumber="540">
<taxonomicNameLabel id="A23D5796FFB4776DFE25E9C5FD97F83C" box="[477,534,1925,1947]" pageId="19" pageNumber="540" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype.
</paragraph>
</caption>
<caption id="DF056677FFB3776AFF6FEA3EFD1FFB33" box="[151,670,1150,1173]" pageId="20" pageNumber="541">
<paragraph id="8BC536FFFFB3776AFF6FEA3EFD1FFB33" blockId="20.[151,670,1150,1173]" box="[151,670,1150,1173]" pageId="20" pageNumber="541">
<emphasis id="B90EEAEDFFB3776AFF6FEA3EFEA1FB33" bold="true" box="[151,288,1150,1172]" pageId="20" pageNumber="541">FIGURE 25.</emphasis>
<taxonomicName id="4C7A4D7CFFB3776AFEDDEAC0FE73FB33" authority="Muona, 2019" authorityName="Muona" authorityYear="2019" box="[293,498,1150,1173]" class="Insecta" family="Throscidae" genus="Pactopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="541" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="B90EEAEDFFB3776AFEDDEAC0FE73FB33" box="[293,498,1150,1173]" italics="true" pageId="20" pageNumber="541">Pactopus burmensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D5796FFB3776AFE00EA3EFDB0FB33" box="[504,561,1150,1172]" pageId="20" pageNumber="541" rank="species">
<emphasis id="B90EEAEDFFB3776AFE00EA3EFDAAFB33" bold="true" box="[504,555,1150,1172]" pageId="20" pageNumber="541">n. sp</emphasis>
.
</taxonomicNameLabel>
, holotype.
</paragraph>
</caption>
<subSubSection id="C3606574FFB3776AFF3FEAFCFCD5FAC3" pageId="20" pageNumber="541" type="description">
<paragraph id="8BC536FFFFB3776AFF3FEAFCFCD5FAC3" blockId="20.[151,1436,1212,1417]" pageId="20" pageNumber="541">
<emphasis id="B90EEAEDFFB3776AFF3FEAFCFED0FB72" bold="true" box="[199,337,1212,1237]" pageId="20" pageNumber="541">Description</emphasis>
. Dorsum poorly visible in the
<typeStatus id="54C1885DFFB3776AFD34EAFCFCB0FB72" box="[716,817,1212,1237]" pageId="20" pageNumber="541" type="holotype">holotype</typeStatus>
, partly clear on the
<typeStatus id="54C1885DFFB3776AFBD3EAFDFB0EFB72" box="[1067,1167,1213,1237]" pageId="20" pageNumber="541" type="paratype">paratype</typeStatus>
. Elytra without keeled humeri, with sharp striae and slightly convex interstices, punctation poorly visible, but apparently week. Ventrum moderately densely punctate, punctures small. Antennae with slender club, all three apical antennomeres longer than wide. Prosternal process widest in the apical third (
<figureCitation id="13412A7AFFB3776AFCF7EB68FCE3FAE7" box="[783,866,1320,1345]" captionStart="FIGURE 13" captionStartId="13.[151,250,1944,1966]" captionTargetBox="[217,1369,1347,1923]" captionTargetId="figure@13.[217,1369,1347,1923]" captionTargetPageId="13" captionText="FIGURE 13. Pactopus burmensis n. sp., ventral view. AtG = abdominal tarsal groove; MtG = metathoracic tarsal groove; PP = prosternal peg." figureDoi="http://doi.org/10.5281/zenodo.3715800" httpUri="https://zenodo.org/record/3715800/files/figure.png" pageId="20" pageNumber="541">Fig. 13</figureCitation>
). Metaventrite with strongly curved tarsal grooves (
<figureCitation id="13412A7AFFB3776AFF67EB0CFF73FAC3" box="[159,242,1356,1381]" captionStart="FIGURE 13" captionStartId="13.[151,250,1944,1966]" captionTargetBox="[217,1369,1347,1923]" captionTargetId="figure@13.[217,1369,1347,1923]" captionTargetPageId="13" captionText="FIGURE 13. Pactopus burmensis n. sp., ventral view. AtG = abdominal tarsal groove; MtG = metathoracic tarsal groove; PP = prosternal peg." figureDoi="http://doi.org/10.5281/zenodo.3715800" httpUri="https://zenodo.org/record/3715800/files/figure.png" pageId="20" pageNumber="541">Fig. 13</figureCitation>
), abdominal tarsal grooves long and straight (
<figureCitation id="13412A7AFFB3776AFD0BEB0CFCC7FAC3" box="[755,838,1356,1381]" captionStart="FIGURE 14" captionStartId="14.[151,250,1994,2016]" captionTargetBox="[151,1436,589,1973]" captionTargetId="figure@14.[151,1436,589,1973]" captionTargetPageId="14" captionText="FIGURE 14. Pactopus burmensis n. sp., abdomen, ventral view. AtG = abdominal tarsal groove." figureDoi="http://doi.org/10.5281/zenodo.3715802" httpUri="https://zenodo.org/record/3715802/files/figure.png" pageId="20" pageNumber="541">Fig. 14</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3606574FFB3776AFF3FEB30FD63FA2E" box="[199,738,1392,1417]" pageId="20" pageNumber="541" type="etymology">
<paragraph id="8BC536FFFFB3776AFF3FEB30FD63FA2E" blockId="20.[151,1436,1212,1417]" box="[199,738,1392,1417]" pageId="20" pageNumber="541">
<emphasis id="B90EEAEDFFB3776AFF3FEB30FEC7FA2E" bold="true" box="[199,326,1392,1417]" pageId="20" pageNumber="541">Etymology</emphasis>
. Known only from Burmese Amber.
</paragraph>
</subSubSection>
</treatment>
</document>
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