343 lines
39 KiB
XML
343 lines
39 KiB
XML
<document id="6C85C01764B66657164D5B0F95E118CF" ID-DOI="10.5281/zenodo.4531515" ID-ISSN="1942-1354" ID-Zenodo-Dep="4531515" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612980280246" checkinUser="felipe" docAuthor="Stebnicka, Z. Teresa & Skelley, Paul E." docDate="2009" docId="03D387539E1966534EA0FBB36CE8FD80" docLanguage="en" docName="InsectaMundi.2009.0062.1-16.pdf" docOrigin="Insecta Mundi 2009 (62)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Haroldiataenius Chalumeau 1981" docType="key" docVersion="6" lastPageNumber="4" masterDocId="FFEAFF2B9E1A66564E60FFB06948FFDB" masterDocTitle="A revision of the genus Haroldiataenius Chalumeau (Scarabaeidae: Aphodiinae: Eupariini)" masterLastPageNumber="16" masterPageNumber="1" pageNumber="2" updateTime="1698869595032" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:mods id="489CFDEB2FDD0BDDB89C368BDFC64C82" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="EEEBFB85EF7C8E7D58CB3ED3D414D508">
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<mods:title id="A1870FF2A65C1039E64F5869788F54D6">A revision of the genus Haroldiataenius Chalumeau (Scarabaeidae: Aphodiinae: Eupariini)</mods:title>
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<mods:name id="F83AECA66AE468C4067BC917F9BA7076" type="personal">
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<mods:namePart id="8642AAF83C171EBFDCE08BEFC9FA8C5A">Stebnicka, Z. Teresa</mods:namePart>
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<mods:name id="7C1CC462DDDE7CE150DC3A25F8D6B561" type="personal">
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<mods:roleTerm id="48F73AAB9F4521BCA4C144018413A150">Author</mods:roleTerm>
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<mods:namePart id="01DAF87C48194F621CB459F41099D0DB">Skelley, Paul E.</mods:namePart>
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<mods:typeOfResource id="BB142033AE7F4419DBF6574E4ED6CF25">text</mods:typeOfResource>
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<mods:title id="838C69FC836DDCAE2411A0F253361B89">Insecta Mundi</mods:title>
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<mods:part id="146B13376491C606578F18B444B81F8C">
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<mods:date id="96586B2C3BB42FE762ADE31ED9FA66CB">2009</mods:date>
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<mods:number id="6DEEDC2641483CB3E0E26586716DC4F1">2009-01-30</mods:number>
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<mods:number id="3036BAD2F2FF4503C5F187868203C81A">2009</mods:number>
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<mods:number id="BE304D56F064105C42A62F0D20A5D779">62</mods:number>
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<mods:classification id="206EFD19515D23BB09565D46EE0982CA">journal article</mods:classification>
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<mods:identifier id="74DF457FEE0725EF8C4C640910CE5A1A" type="DOI">10.5281/zenodo.4531515</mods:identifier>
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<mods:identifier id="034C67221F247D9CF4841ACC68D7927F" type="ISSN">1942-1354</mods:identifier>
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<mods:identifier id="001AFFB508CCFF342A4506B9921D374A" type="Zenodo-Dep">4531515</mods:identifier>
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<treatment id="03D387539E1966534EA0FBB36CE8FD80" ID-DOI="http://doi.org/10.5281/zenodo.4531870" ID-Zenodo-Dep="4531870" LSID="urn:lsid:plazi:treatment:03D387539E1966534EA0FBB36CE8FD80" httpUri="http://treatment.plazi.org/id/03D387539E1966534EA0FBB36CE8FD80" lastPageId="5" lastPageNumber="4" pageId="3" pageNumber="2">
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<subSubSection id="C36065CE9E1966554EA0FBB36BD8FBC7" box="[192,656,1027,1052]" pageId="3" pageNumber="2" type="nomenclature">
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<paragraph id="8BC536459E1966554EA0FBB36BD8FBC7" blockId="3.[192,656,1027,1052]" box="[192,656,1027,1052]" pageId="3" pageNumber="2">
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<heading id="D08D81299E1966554EA0FBB36BD8FBC7" bold="true" box="[192,656,1027,1052]" fontSize="10" level="2" pageId="3" pageNumber="2" reason="3">
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<emphasis id="B90EEA579E1966554EA0FBB36BD8FBC7" bold="true" box="[192,656,1027,1052]" pageId="3" pageNumber="2">
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Key to Species of
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<taxonomicName id="4C7A4DC69E1966554FCCFBB56BD8FBC7" authorityName="Chalumeau" authorityYear="1981" box="[428,656,1029,1052]" class="Insecta" family="Aphodiidae" genus="Haroldiataenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
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<emphasis id="B90EEA579E1966554FCCFBB56BD8FBC7" bold="true" box="[428,656,1029,1052]" italics="true" pageId="3" pageNumber="2">Haroldiataenius</emphasis>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36065CE9E1966534EA0FBF36CE8FD80" lastPageId="5" lastPageNumber="4" pageId="3" pageNumber="2" type="key">
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<keyStep id="308E2EE09E1966554EA0FBF36CE9FB20" pageId="3" pageNumber="2">
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<paragraph id="8BC536459E1966554EA0FBF36CE8FB40" blockId="3.[192,1441,1091,1276]" pageId="3" pageNumber="2">
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<keyLead id="308B95709E1966554EA0FBF36CE8FB40" pageId="3" pageNumber="2">
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1. Body psammodiiform (Fig. 12); lateral margins of pronotum and elytron with long setal fringe, setal length much greater than distance between setal bases (
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<figureCitation id="13412AC09E1966554A5AFBD36DC8FBA0" box="[1082,1152,1123,1147]" captionStart="Figures 1-7" captionStartId="4.[192,289,1434,1456]" captionTargetBox="[192,1440,193,1402]" captionTargetId="figure-142@4.[190,1442,191,1404]" captionTargetPageId="4" captionText="Figures 1-7. Haroldiataenius spp. 1) H. mariarum, lateral habitus. 2) H. hintoni lateral habitus. 3) H. lucanus, mesotibia. 4) H. semipilosus, mesotibia. 5) H. saramari, lateral habitus. 6) H. lucanus, lateral habitus. 7) H. limbatus, lateral habitus." figureDoi="http://doi.org/10.5281/zenodo.4531517" httpUri="https://zenodo.org/record/4531517/files/figure.png" pageId="3" pageNumber="2">Fig. 1</figureCitation>
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). Clypeus transversely rugose.
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<collectingCountry id="F36D76D59E1966554FF6FB3368A7FB40" box="[406,495,1155,1179]" name="Mexico" pageId="3" pageNumber="2">Mexico</collectingCountry>
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,
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<collectingCountry id="F36D76D59E1966554F98FB346B16FB40" box="[504,606,1156,1179]" name="Panama" pageId="3" pageNumber="2">Panama</collectingCountry>
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........................................................................
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<taxonomicName id="4C7A4DC69E1966554AE4FB356CE8FB40" authority="(Bates)" baseAuthorityName="Bates" box="[1156,1440,1155,1180]" class="Insecta" family="Aphodiidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="mariarum">
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<emphasis id="B90EEA579E1966554AE4FB356CE8FB40" bold="true" box="[1156,1440,1155,1180]" pageId="3" pageNumber="2">
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<emphasis id="B90EEA579E1966554AE4FB356C70FB47" bold="true" box="[1156,1336,1157,1180]" italics="true" pageId="3" pageNumber="2">H. mariarum</emphasis>
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(Bates)
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</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="8BC536459E1966554EA0FB146CE9FB20" blockId="3.[192,1441,1091,1276]" pageId="3" pageNumber="2">
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<keyLead id="308B95709E1966554EA0FB146CE9FB20" pageId="3" pageNumber="2">
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— Body psammodiiform to robustly elongate (Fig. 13-15); lateral margins of pronotum or elytron bare, or with fringe of short setae; if setae present, then length equal to or less than distance between setal bases (
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<figureCitation id="13412AC09E1966554C56FB536BF7FB20" box="[566,703,1251,1275]" captionStart-0="Figures 1-7" captionStart-1="Figures 8-11" captionStartId-0="4.[192,289,1434,1456]" captionStartId-1="6.[192,287,989,1011]" captionTargetBox-0="[192,1440,193,1402]" captionTargetBox-1="[192,1443,196,958]" captionTargetId-0="figure-142@4.[190,1442,191,1404]" captionTargetId-1="figure-305@6.[190,1445,194,960]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionText-0="Figures 1-7. Haroldiataenius spp. 1) H. mariarum, lateral habitus. 2) H. hintoni lateral habitus. 3) H. lucanus, mesotibia. 4) H. semipilosus, mesotibia. 5) H. saramari, lateral habitus. 6) H. lucanus, lateral habitus. 7) H. limbatus, lateral habitus." captionText-1="Figures 8-11. Haroldiataenius spp., lateral habitus. 8) H. semipilosus. 9) H. griffini. 10) H. buvexus, paratype. 11) H. convexus, holotype." figureDoi-0="http://doi.org/10.5281/zenodo.4531517" figureDoi-1="http://doi.org/10.5281/zenodo.4531519" httpUri-0="https://zenodo.org/record/4531517/files/figure.png" httpUri-1="https://zenodo.org/record/4531519/files/figure.png" pageId="3" pageNumber="2">Fig. 2, 5-11</figureCitation>
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). Clypeus punctate, smooth, or weakly rugose .....................
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<emphasis id="B90EEA579E1966554BF2FB536CE9FB20" bold="true" box="[1426,1441,1251,1275]" pageId="3" pageNumber="2">2</emphasis>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="308E2EE09E1966554EA0FA936CE9FA40" pageId="3" pageNumber="2">
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<paragraph id="8BC536459E1966554EA0FA936CE8FA80" blockId="3.[192,1441,1315,1435]" pageId="3" pageNumber="2">
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<keyLead id="308B95709E1966554EA0FA936CE8FA80" pageId="3" pageNumber="2">
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2(1). Meso- and metafemoral posterior margin with distinct, complete line from trochanter to apex (
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<figureCitation id="13412AC09E1966554F21FAF368CDFA80" box="[321,389,1347,1371]" captionStart="Figures 1-7" captionStartId="4.[192,289,1434,1456]" captionTargetBox="[192,1440,193,1402]" captionTargetId="figure-142@4.[190,1442,191,1404]" captionTargetPageId="4" captionText="Figures 1-7. Haroldiataenius spp. 1) H. mariarum, lateral habitus. 2) H. hintoni lateral habitus. 3) H. lucanus, mesotibia. 4) H. semipilosus, mesotibia. 5) H. saramari, lateral habitus. 6) H. lucanus, lateral habitus. 7) H. limbatus, lateral habitus." figureDoi="http://doi.org/10.5281/zenodo.4531517" httpUri="https://zenodo.org/record/4531517/files/figure.png" pageId="3" pageNumber="2">Fig. 3</figureCitation>
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) .......................................................................................................................................
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<emphasis id="B90EEA579E1966554BF1FAF36CE8FA80" bold="true" box="[1425,1440,1347,1371]" pageId="3" pageNumber="2">3</emphasis>
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</keyLead>
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</paragraph>
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<paragraph id="8BC536459E1966554EA0FAD46CE9FA40" blockId="3.[192,1441,1315,1435]" pageId="3" pageNumber="2">
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<keyLead id="308B95709E1966554EA0FAD46CE9FA40" pageId="3" pageNumber="2">
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— Meso- and metafemoral posterior margin lacking complete line, if present, then only in distal half (
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<figureCitation id="13412AC09E1966554F21FA3368CEFA40" box="[321,390,1411,1435]" captionStart="Figures 1-7" captionStartId="4.[192,289,1434,1456]" captionTargetBox="[192,1440,193,1402]" captionTargetId="figure-142@4.[190,1442,191,1404]" captionTargetPageId="4" captionText="Figures 1-7. Haroldiataenius spp. 1) H. mariarum, lateral habitus. 2) H. hintoni lateral habitus. 3) H. lucanus, mesotibia. 4) H. semipilosus, mesotibia. 5) H. saramari, lateral habitus. 6) H. lucanus, lateral habitus. 7) H. limbatus, lateral habitus." figureDoi="http://doi.org/10.5281/zenodo.4531517" httpUri="https://zenodo.org/record/4531517/files/figure.png" pageId="3" pageNumber="2">Fig. 4</figureCitation>
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)........................................................................................................................................
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<emphasis id="B90EEA579E1966554BF2FA336CE9FA40" bold="true" box="[1426,1441,1411,1435]" pageId="3" pageNumber="2">4</emphasis>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="308E2EE09E1966554EA0FA736CE8F9A0" pageId="3" pageNumber="2">
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<paragraph id="8BC536459E1966554EA0FA736CE8F9C0" blockId="3.[192,1440,1475,1660]" pageId="3" pageNumber="2">
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<keyLead id="308B95709E1966554EA0FA736CE8F9C0" pageId="3" pageNumber="2">
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3(2). Clypeus rounded each side of median emargination. Base of head lacking band of coarse punctures. Elytral intervals convex (
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<figureCitation id="13412AC09E1966554C15FA536BF5FA20" box="[629,701,1507,1531]" captionStart="Figures 1-7" captionStartId="4.[192,289,1434,1456]" captionTargetBox="[192,1440,193,1402]" captionTargetId="figure-142@4.[190,1442,191,1404]" captionTargetPageId="4" captionText="Figures 1-7. Haroldiataenius spp. 1) H. mariarum, lateral habitus. 2) H. hintoni lateral habitus. 3) H. lucanus, mesotibia. 4) H. semipilosus, mesotibia. 5) H. saramari, lateral habitus. 6) H. lucanus, lateral habitus. 7) H. limbatus, lateral habitus." figureDoi="http://doi.org/10.5281/zenodo.4531517" httpUri="https://zenodo.org/record/4531517/files/figure.png" pageId="3" pageNumber="2">Fig. 5</figureCitation>
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), lacking setae on disc, some setae may be visible on apical declivity.
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<collectingRegion id="49BEF8A79E1966554FCCF9B36B40F9C0" box="[428,520,1539,1563]" country="United States of America" name="Florida" pageId="3" pageNumber="2">Florida</collectingRegion>
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,
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<collectingCountry id="F36D76D59E1966554C72F9B36B04F9C0" box="[530,588,1539,1563]" name="United States of America" pageId="3" pageNumber="2">USA</collectingCountry>
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.................................................................
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<taxonomicName id="4C7A4DC69E1966554A20F9B56CE8F9C0" authority="(Cartwright)" baseAuthorityName="Cartwright" box="[1088,1440,1539,1564]" class="Insecta" family="Aphodiidae" genus="Haroldiataenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="saramari">
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<emphasis id="B90EEA579E1966554A20F9B56CE8F9C0" bold="true" box="[1088,1440,1539,1564]" pageId="3" pageNumber="2">
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<emphasis id="B90EEA579E1966554A20F9B56DAFF9C7" bold="true" box="[1088,1255,1541,1564]" italics="true" pageId="3" pageNumber="2">H. saramari</emphasis>
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(Cartwright)
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</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="8BC536459E1966554EA0F9946CE8F9A0" blockId="3.[192,1440,1475,1660]" pageId="3" pageNumber="2">
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<keyLead id="308B95709E1966554EA0F9946CE8F9A0" pageId="3" pageNumber="2">
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— Clypeus angulate to distinctly toothed each side of median emargination. Base of head with band of coarse punctures. Elytral intervals tectiform, with short setae (
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<figureCitation id="13412AC09E1966554A54F9F36D30F980" box="[1076,1144,1603,1627]" captionStart="Figures 1-7" captionStartId="4.[192,289,1434,1456]" captionTargetBox="[192,1440,193,1402]" captionTargetId="figure-142@4.[190,1442,191,1404]" captionTargetPageId="4" captionText="Figures 1-7. Haroldiataenius spp. 1) H. mariarum, lateral habitus. 2) H. hintoni lateral habitus. 3) H. lucanus, mesotibia. 4) H. semipilosus, mesotibia. 5) H. saramari, lateral habitus. 6) H. lucanus, lateral habitus. 7) H. limbatus, lateral habitus." figureDoi="http://doi.org/10.5281/zenodo.4531517" httpUri="https://zenodo.org/record/4531517/files/figure.png" pageId="3" pageNumber="2">Fig. 6</figureCitation>
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).
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<collectingRegion id="49BEF8A79E1966554AEBF9F36C0AF980" box="[1163,1346,1603,1627]" country="Mexico" name="Baja California" pageId="3" pageNumber="2">Baja California</collectingRegion>
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,
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<collectingCountry id="F36D76D59E1966554B2CF9F36CE8F980" box="[1356,1440,1603,1627]" name="Mexico" pageId="3" pageNumber="2">Mexico</collectingCountry>
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and
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<collectingRegion id="49BEF8A79E1966554F0CF9D36887F9A0" box="[364,463,1635,1659]" country="United States of America" name="Arizona" pageId="3" pageNumber="2">Arizona</collectingRegion>
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,
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<collectingCountry id="F36D76D59E1966554FB8F9D36B5AF9A0" box="[472,530,1635,1659]" name="United States of America" pageId="3" pageNumber="2">USA</collectingCountry>
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.......................................................................................
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<taxonomicName id="4C7A4DC69E1966554ACAF9D56CE8F9A0" authority="(Horn)" baseAuthorityName="Horn" box="[1194,1440,1635,1660]" class="Insecta" family="Aphodiidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="lucanus">
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<emphasis id="B90EEA579E1966554ACAF9D56CE8F9A0" bold="true" box="[1194,1440,1635,1660]" pageId="3" pageNumber="2">
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<emphasis id="B90EEA579E1966554ACAF9D56C75F9A7" bold="true" box="[1194,1341,1637,1660]" italics="true" pageId="3" pageNumber="2">H. lucanus</emphasis>
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(Horn)
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</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="308E2EE09E1966554EA0F9136CE9F8E0" pageId="3" pageNumber="2">
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<paragraph id="8BC536459E1966554EA0F9136CE8F920" blockId="3.[192,1441,1699,1851]" pageId="3" pageNumber="2">
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<keyLead id="308B95709E1966554EA0F9136CE8F920" pageId="3" pageNumber="2">
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4(2). Lateral pronotal margin lacking distinct setae; lateral bead of pronotum gradually thickened toward anterior angle (
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<figureCitation id="13412AC09E1966554C2FF9736BDDF900" box="[591,661,1731,1755]" captionStart="Figures 1-7" captionStartId="4.[192,289,1434,1456]" captionTargetBox="[192,1440,193,1402]" captionTargetId="figure-142@4.[190,1442,191,1404]" captionTargetPageId="4" captionText="Figures 1-7. Haroldiataenius spp. 1) H. mariarum, lateral habitus. 2) H. hintoni lateral habitus. 3) H. lucanus, mesotibia. 4) H. semipilosus, mesotibia. 5) H. saramari, lateral habitus. 6) H. lucanus, lateral habitus. 7) H. limbatus, lateral habitus." figureDoi="http://doi.org/10.5281/zenodo.4531517" httpUri="https://zenodo.org/record/4531517/files/figure.png" pageId="3" pageNumber="2">Fig. 7</figureCitation>
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).
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<collectingCountry id="F36D76D59E1966554CC9F9736A4BF900" box="[681,771,1731,1755]" name="Mexico" pageId="3" pageNumber="2">Mexico</collectingCountry>
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,
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<collectingCountry id="F36D76D59E1966554D6CF9736AD0F900" box="[780,920,1731,1755]" name="Guatemala" pageId="3" pageNumber="2">Guatemala</collectingCountry>
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,
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<collectingCountry id="F36D76D59E1966554DC2F9736D57F900" box="[930,1055,1731,1755]" name="Honduras" pageId="3" pageNumber="2">Honduras</collectingCountry>
|
|
,
|
|
<collectingCountry id="F36D76D59E1966554A49F9736DF0F900" box="[1065,1208,1731,1755]" name="El Salvador" pageId="3" pageNumber="2">El Salvador</collectingCountry>
|
|
............................. ..................................................................................................................
|
|
<taxonomicName id="4C7A4DC69E1966554AFAF9556CE8F920" authority="(Bates)" baseAuthorityName="Bates" box="[1178,1440,1763,1788]" class="Insecta" family="Aphodiidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="limbatus">
|
|
<emphasis id="B90EEA579E1966554AFAF9556CE8F920" bold="true" box="[1178,1440,1763,1788]" pageId="3" pageNumber="2">
|
|
<emphasis id="B90EEA579E1966554AFAF9556C70F927" bold="true" box="[1178,1336,1765,1788]" italics="true" pageId="3" pageNumber="2">H. limbatus</emphasis>
|
|
(Bates)
|
|
</emphasis>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8BC536459E1966554EA0F8B46CE9F8E0" blockId="3.[192,1441,1699,1851]" pageId="3" pageNumber="2">
|
|
<keyLead id="308B95709E1966554EA0F8B46CE9F8E0" pageId="3" pageNumber="2">
|
|
— Lateral pronotal margin with distinct setal fringe; lateral bead of pronotum fine throughout length, not distinctly thickened near anterior angles ...............................................................
|
|
<emphasis id="B90EEA579E1966554BF2F8936CE9F8E0" bold="true" box="[1426,1441,1827,1851]" pageId="3" pageNumber="2">5</emphasis>
|
|
</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<caption id="DF0566CD9E1E66524EA0FA2A68ABFA31" ID-DOI="http://doi.org/10.5281/zenodo.4531517" ID-Zenodo-Dep="4531517" httpUri="https://zenodo.org/record/4531517/files/figure.png" pageId="4" pageNumber="3" startId="4.[192,289,1434,1456]" targetBox="[192,1440,193,1402]" targetPageId="4">
|
|
<paragraph id="8BC536459E1E66524EA0FA2A68ABFA31" blockId="4.[192,1441,1434,1514]" pageId="4" pageNumber="3">
|
|
<emphasis id="B90EEA579E1E66524EA0FA2A681AFA74" bold="true" box="[192,338,1434,1456]" pageId="4" pageNumber="3">Figures 1-7.</emphasis>
|
|
<taxonomicName id="4C7A4DC69E1E66524F37FA2B6B58FA6B" authorityName="Chalumeau" authorityYear="1981" box="[343,528,1435,1456]" class="Insecta" family="Aphodiidae" genus="Haroldiataenius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B90EEA579E1E66524F37FA2B6B58FA6B" box="[343,528,1435,1456]" italics="true" pageId="4" pageNumber="3">Haroldiataenius</emphasis>
|
|
</taxonomicName>
|
|
spp.
|
|
<emphasis id="B90EEA579E1E66524C2AFA2A6B28FA6B" bold="true" box="[586,608,1434,1456]" pageId="4" pageNumber="3">1)</emphasis>
|
|
<taxonomicName id="4C7A4DC69E1E66524C07FA2B6BB4FA6B" box="[615,764,1435,1456]" class="Insecta" family="Aphodiidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="mariarum">
|
|
<emphasis id="B90EEA579E1E66524C07FA2B6BB4FA6B" box="[615,764,1435,1456]" italics="true" pageId="4" pageNumber="3">H. mariarum</emphasis>
|
|
</taxonomicName>
|
|
, lateral habitus.
|
|
<emphasis id="B90EEA579E1E66524DDBFA2A6A99FA6B" bold="true" box="[955,977,1434,1456]" pageId="4" pageNumber="3">2)</emphasis>
|
|
<taxonomicName id="4C7A4DC69E1E66524DB8FA2B6D03FA6B" authorityName="Saylor. Subsequently" authorityYear="1933" box="[984,1099,1435,1456]" class="Insecta" family="Aphodiidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="hintoni">
|
|
<emphasis id="B90EEA579E1E66524DB8FA2B6D03FA6B" box="[984,1099,1435,1456]" italics="true" pageId="4" pageNumber="3">H. hintoni</emphasis>
|
|
</taxonomicName>
|
|
lateral habitus.
|
|
<emphasis id="B90EEA579E1E66524B63FA2A6C51FA6B" bold="true" box="[1283,1305,1434,1456]" pageId="4" pageNumber="3">3)</emphasis>
|
|
<taxonomicName id="4C7A4DC69E1E66524B40FA2B6CD1FA6B" box="[1312,1433,1435,1456]" class="Insecta" family="Aphodiidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="lucanus">
|
|
<emphasis id="B90EEA579E1E66524B40FA2B6CD1FA6B" box="[1312,1433,1435,1456]" italics="true" pageId="4" pageNumber="3">H. lucanus</emphasis>
|
|
</taxonomicName>
|
|
, mesotibia.
|
|
<emphasis id="B90EEA579E1E66524F5CFA07681BFA16" bold="true" box="[316,339,1463,1485]" pageId="4" pageNumber="3">4)</emphasis>
|
|
<taxonomicName id="4C7A4DC69E1E66524F3CFA086B4AFA17" authorityName="Stebnicka & Skelley" authorityYear="2009" baseAuthorityName="Stebnicka & Skelley" baseAuthorityYear="2009" box="[348,514,1463,1485]" class="Insecta" family="Aphodiidae" genus="Haroldiataenius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="semipilosus">
|
|
<emphasis id="B90EEA579E1E66524F3CFA086B4AFA17" box="[348,514,1463,1485]" italics="true" pageId="4" pageNumber="3">H. semipilosus</emphasis>
|
|
</taxonomicName>
|
|
, mesotibia.
|
|
<emphasis id="B90EEA579E1E66524CEEFA076BEDFA16" bold="true" box="[654,677,1463,1485]" pageId="4" pageNumber="3">5)</emphasis>
|
|
<taxonomicName id="4C7A4DC69E1E66524CCEFA086A72FA16" authorityName="Stebnicka & Skelley" authorityYear="2009" baseAuthorityName="Stebnicka & Skelley" baseAuthorityYear="2009" box="[686,826,1464,1485]" class="Insecta" family="Aphodiidae" genus="Haroldiataenius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="saramari">
|
|
<emphasis id="B90EEA579E1E66524CCEFA086A72FA16" box="[686,826,1464,1485]" italics="true" pageId="4" pageNumber="3">H. saramari</emphasis>
|
|
</taxonomicName>
|
|
, lateral habitus.
|
|
<emphasis id="B90EEA579E1E66524A62FA076D52FA16" bold="true" box="[1026,1050,1463,1485]" pageId="4" pageNumber="3">6)</emphasis>
|
|
<taxonomicName id="4C7A4DC69E1E66524A42FA086DD5FA17" box="[1058,1181,1463,1485]" class="Insecta" family="Aphodiidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="lucanus">
|
|
<emphasis id="B90EEA579E1E66524A42FA086DD5FA17" box="[1058,1181,1463,1485]" italics="true" pageId="4" pageNumber="3">H. lucanus</emphasis>
|
|
</taxonomicName>
|
|
, lateral habitus.
|
|
<emphasis id="B90EEA579E1E66524B05FA076C34FA16" bold="true" box="[1381,1404,1463,1485]" pageId="4" pageNumber="3">7)</emphasis>
|
|
<taxonomicName id="4C7A4DC69E1E66524BE5FA08686CFA32" class="Insecta" family="Aphodiidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="limbatus">
|
|
<emphasis id="B90EEA579E1E66524BE5FA08686CFA32" italics="true" pageId="4" pageNumber="3">H. limbatus</emphasis>
|
|
</taxonomicName>
|
|
, lateral habitus.
|
|
</paragraph>
|
|
</caption>
|
|
<keyStep id="308E2EE09E1E66524EA0F9AA6CE9F969" pageId="4" pageNumber="3">
|
|
<paragraph id="8BC536459E1E66524EA0F9AA6CE8F9AA" blockId="4.[192,1441,1562,1714]" pageId="4" pageNumber="3">
|
|
<keyLead id="308B95709E1E66524EA0F9AA6CE8F9AA" pageId="4" pageNumber="3">
|
|
5(4). Pronotal punctures of uniform size, punctures dense, separated by less than one diameter over entire surface (
|
|
<figureCitation id="13412AC09E1E66524F8FF98A6B7EF989" box="[495,566,1594,1618]" captionStart="Figures 1-7" captionStartId="4.[192,289,1434,1456]" captionTargetBox="[192,1440,193,1402]" captionTargetId="figure-142@4.[190,1442,191,1404]" captionTargetPageId="4" captionText="Figures 1-7. Haroldiataenius spp. 1) H. mariarum, lateral habitus. 2) H. hintoni lateral habitus. 3) H. lucanus, mesotibia. 4) H. semipilosus, mesotibia. 5) H. saramari, lateral habitus. 6) H. lucanus, lateral habitus. 7) H. limbatus, lateral habitus." figureDoi="http://doi.org/10.5281/zenodo.4531517" httpUri="https://zenodo.org/record/4531517/files/figure.png" pageId="4" pageNumber="3">Fig. 2</figureCitation>
|
|
, 15); base of pronotum with marginal line obscured by coarse punctures.
|
|
<collectingCountry id="F36D76D59E1E66524F58F9EA68D9F9A9" box="[312,401,1626,1650]" name="Mexico" pageId="4" pageNumber="3">Mexico</collectingCountry>
|
|
,
|
|
<collectingCountry id="F36D76D59E1E66524FFAF9EA6B6DF9A9" box="[410,549,1626,1650]" name="Guatemala" pageId="4" pageNumber="3">Guatemala</collectingCountry>
|
|
,
|
|
<collectingCountry id="F36D76D59E1E66524C4FF9EA6BE0F9A9" box="[559,680,1626,1650]" name="Honduras" pageId="4" pageNumber="3">Honduras</collectingCountry>
|
|
..................................................................
|
|
<taxonomicName id="4C7A4DC69E1E66524AFEF9EB6CE8F9AA" authority="(Saylor)" baseAuthorityName="Saylor" box="[1182,1440,1625,1650]" class="Insecta" family="Aphodiidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="hintoni">
|
|
<emphasis id="B90EEA579E1E66524AFEF9EB6CE8F9AA" bold="true" box="[1182,1440,1625,1650]" pageId="4" pageNumber="3">
|
|
<emphasis id="B90EEA579E1E66524AFEF9EB6C62F9A9" bold="true" box="[1182,1322,1627,1650]" italics="true" pageId="4" pageNumber="3">H. hintoni</emphasis>
|
|
(Saylor)
|
|
</emphasis>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8BC536459E1E66524EA0F9CA6CE9F969" blockId="4.[192,1441,1562,1714]" pageId="4" pageNumber="3">
|
|
<keyLead id="308B95709E1E66524EA0F9CA6CE9F969" pageId="4" pageNumber="3">
|
|
— Pronotal punctures mixed fine and coarse; coarse pronotal punctures more widely scattered, separated by at least their own diameter (
|
|
<figureCitation id="13412AC09E1E66524CC3F92A6A43F969" box="[675,779,1690,1714]" captionStart="Figures 8-11" captionStartId="6.[192,287,989,1011]" captionTargetBox="[192,1443,196,958]" captionTargetId="figure-305@6.[190,1445,194,960]" captionTargetPageId="6" captionText="Figures 8-11. Haroldiataenius spp., lateral habitus. 8) H. semipilosus. 9) H. griffini. 10) H. buvexus, paratype. 11) H. convexus, holotype." figureDoi="http://doi.org/10.5281/zenodo.4531519" httpUri="https://zenodo.org/record/4531519/files/figure.png" pageId="4" pageNumber="3">Fig. 8-11</figureCitation>
|
|
); base of pronotum with distinct marginal line ............
|
|
<emphasis id="B90EEA579E1E66524BF2F92A6CE9F969" bold="true" box="[1426,1441,1690,1714]" pageId="4" pageNumber="3">6</emphasis>
|
|
</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="308E2EE09E1E66524EA0F96A6CE9F8A9" pageId="4" pageNumber="3">
|
|
<paragraph id="8BC536459E1E66524EA0F96A6CE8F8EA" blockId="4.[192,1441,1754,1906]" pageId="4" pageNumber="3">
|
|
<keyLead id="308B95709E1E66524EA0F96A6CE8F8EA" pageId="4" pageNumber="3">
|
|
6(5). Clypeus strongly angulate to dentate each side of median emargination. Elytral intervals with long scattered setae, most prominent laterally (
|
|
<figureCitation id="13412AC09E1E66524D16F94A6AF4F8C9" box="[886,956,1786,1810]" captionStart="Figures 8-11" captionStartId="6.[192,287,989,1011]" captionTargetBox="[192,1443,196,958]" captionTargetId="figure-305@6.[190,1445,194,960]" captionTargetPageId="6" captionText="Figures 8-11. Haroldiataenius spp., lateral habitus. 8) H. semipilosus. 9) H. griffini. 10) H. buvexus, paratype. 11) H. convexus, holotype." figureDoi="http://doi.org/10.5281/zenodo.4531519" httpUri="https://zenodo.org/record/4531519/files/figure.png" pageId="4" pageNumber="3">Fig. 8</figureCitation>
|
|
).
|
|
<collectingRegion id="49BEF8A79E1E66524DB1F94A6D52F8CA" box="[977,1050,1786,1809]" country="United States of America" name="Texas" pageId="4" pageNumber="3">Texas</collectingRegion>
|
|
and
|
|
<collectingRegion id="49BEF8A79E1E66524A33F94A6DFFF8C9" box="[1107,1207,1786,1810]" country="United States of America" name="Arizona" pageId="4" pageNumber="3">Arizona</collectingRegion>
|
|
,
|
|
<collectingCountry id="F36D76D59E1E66524AA1F94A6DB6F8C9" box="[1217,1278,1786,1810]" name="United States of America" pageId="4" pageNumber="3">USA</collectingCountry>
|
|
;
|
|
<collectingCountry id="F36D76D59E1E66524B68F94A6C17F8C9" box="[1288,1375,1786,1810]" name="Mexico" pageId="4" pageNumber="3">Mexico</collectingCountry>
|
|
....... ......................................................................................................
|
|
<taxonomicName id="4C7A4DC69E1E66524A20F8AB6CE8F8EA" authority="(Van Dyke)" baseAuthorityName="Van Dyke" box="[1088,1440,1817,1842]" class="Insecta" family="Aphodiidae" genus="Haroldiataenius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="semipilosus">
|
|
<emphasis id="B90EEA579E1E66524A20F8AB6CE8F8EA" bold="true" box="[1088,1440,1817,1842]" pageId="4" pageNumber="3">
|
|
<emphasis id="B90EEA579E1E66524A20F8AB6C4AF8E9" bold="true" box="[1088,1282,1819,1842]" italics="true" pageId="4" pageNumber="3">H. semipilosus</emphasis>
|
|
(Van Dyke)
|
|
</emphasis>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8BC536459E1E66524EA0F88A6CE9F8A9" blockId="4.[192,1441,1754,1906]" pageId="4" pageNumber="3">
|
|
<keyLead id="308B95709E1E66524EA0F88A6CE9F8A9" pageId="4" pageNumber="3">
|
|
— Clypeus rounded or angulate each side of median emargination. Elytral intervals lacking setae (
|
|
<figureCitation id="13412AC09E1E66524F21F8EA68E6F8A9" box="[321,430,1882,1906]" captionStart="Figures 8-11" captionStartId="6.[192,287,989,1011]" captionTargetBox="[192,1443,196,958]" captionTargetId="figure-305@6.[190,1445,194,960]" captionTargetPageId="6" captionText="Figures 8-11. Haroldiataenius spp., lateral habitus. 8) H. semipilosus. 9) H. griffini. 10) H. buvexus, paratype. 11) H. convexus, holotype." figureDoi="http://doi.org/10.5281/zenodo.4531519" httpUri="https://zenodo.org/record/4531519/files/figure.png" pageId="4" pageNumber="3">Fig. 9-11</figureCitation>
|
|
). Southern
|
|
<collectingRegion id="49BEF8A79E1E66524C5AF8EA6BCEF8AA" box="[570,646,1882,1905]" country="United States of America" name="Texas" pageId="4" pageNumber="3">Texas</collectingRegion>
|
|
,
|
|
<collectingCountry id="F36D76D59E1E66524CF0F8EA6B82F8A9" box="[656,714,1882,1906]" name="United States of America" pageId="4" pageNumber="3">USA</collectingCountry>
|
|
.............................................................................................
|
|
<emphasis id="B90EEA579E1E66524BF2F8EA6CE9F8A9" bold="true" box="[1426,1441,1882,1906]" pageId="4" pageNumber="3">7</emphasis>
|
|
</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="308E2EE09E1F66534EA0FF736CE9FEA0" pageId="5" pageNumber="4">
|
|
<paragraph id="8BC536459E1F66534EA0FF736CE8FEC0" blockId="5.[192,1441,195,380]" pageId="5" pageNumber="4">
|
|
<keyLead id="308B95709E1F66534EA0FF736CE8FEC0" pageId="5" pageNumber="4">
|
|
7(6). Lateral elytral margin with indistinct fringe of setae, if visible, setae distinctly shorter than distance between their bases (
|
|
<figureCitation id="13412AC09E1F66534CF5FF536B90FF20" box="[661,728,227,251]" captionStart="Figures 8-11" captionStartId="6.[192,287,989,1011]" captionTargetBox="[192,1443,196,958]" captionTargetId="figure-305@6.[190,1445,194,960]" captionTargetPageId="6" captionText="Figures 8-11. Haroldiataenius spp., lateral habitus. 8) H. semipilosus. 9) H. griffini. 10) H. buvexus, paratype. 11) H. convexus, holotype." figureDoi="http://doi.org/10.5281/zenodo.4531519" httpUri="https://zenodo.org/record/4531519/files/figure.png" pageId="5" pageNumber="4">Fig. 9</figureCitation>
|
|
). Coarse pronotal punctures more widely scattered, diameter equal to or less than width of basal pronotal margin ......................
|
|
<taxonomicName id="4C7A4DC69E1F66534A3EFEB56CE8FEC0" authority="(Cartwright)" baseAuthorityName="Cartwright" box="[1118,1440,259,284]" class="Insecta" family="Aphodiidae" genus="Haroldiataenius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="griffini">
|
|
<emphasis id="B90EEA579E1F66534A3EFEB56CE8FEC0" bold="true" box="[1118,1440,259,284]" pageId="5" pageNumber="4">
|
|
<emphasis id="B90EEA579E1F66534A3EFEB56DA0FEC7" bold="true" box="[1118,1256,261,284]" italics="true" pageId="5" pageNumber="4">H. griffini</emphasis>
|
|
(Cartwright)
|
|
</emphasis>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8BC536459E1F66534EA0FE946CE9FEA0" blockId="5.[192,1441,195,380]" pageId="5" pageNumber="4">
|
|
<keyLead id="308B95709E1F66534EA0FE946CE9FEA0" pageId="5" pageNumber="4">
|
|
— Lateral elytral margin with distinct fringe of short setae, setae as long as distance between their bases (
|
|
<figureCitation id="13412AC09E1F66534FE6FEF368B5FE80" box="[390,509,323,347]" captionStart="Figures 8-11" captionStartId="6.[192,287,989,1011]" captionTargetBox="[192,1443,196,958]" captionTargetId="figure-305@6.[190,1445,194,960]" captionTargetPageId="6" captionText="Figures 8-11. Haroldiataenius spp., lateral habitus. 8) H. semipilosus. 9) H. griffini. 10) H. buvexus, paratype. 11) H. convexus, holotype." figureDoi="http://doi.org/10.5281/zenodo.4531519" httpUri="https://zenodo.org/record/4531519/files/figure.png" pageId="5" pageNumber="4">Fig. 10-11</figureCitation>
|
|
). Coarse pronotal punctures evenly distributed basally, their diameter generally greater than width of broad basal pronotal margin ..................................................................
|
|
<emphasis id="B90EEA579E1F66534BF2FED36CE9FEA0" bold="true" box="[1426,1441,355,379]" pageId="5" pageNumber="4">8</emphasis>
|
|
</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="308E2EE09E1F66534EA0FE136CE8FD80" pageId="5" pageNumber="4">
|
|
<paragraph id="8BC536459E1F66534EA0FE136CE8FE20" blockId="5.[192,1440,419,604]" pageId="5" pageNumber="4">
|
|
<keyLead id="308B95709E1F66534EA0FE136CE8FE20" pageId="5" pageNumber="4">
|
|
8(7). Clypeal anterior margin evenly rounded each side of median emargination (
|
|
<figureCitation id="13412AC09E1F66534A1FFE136D87FE60" box="[1151,1231,419,443]" captionStart="Figures 8-11" captionStartId="6.[192,287,989,1011]" captionTargetBox="[192,1443,196,958]" captionTargetId="figure-305@6.[190,1445,194,960]" captionTargetPageId="6" captionText="Figures 8-11. Haroldiataenius spp., lateral habitus. 8) H. semipilosus. 9) H. griffini. 10) H. buvexus, paratype. 11) H. convexus, holotype." figureDoi="http://doi.org/10.5281/zenodo.4531519" httpUri="https://zenodo.org/record/4531519/files/figure.png" pageId="5" pageNumber="4">Fig. 10</figureCitation>
|
|
). Elytral intervals weakly convex on disc. Body dark brown, nearly black. Big Bend region,
|
|
<collectingRegion id="49BEF8A79E1F66534AC4FE746DB8FE00" box="[1188,1264,452,475]" country="United States of America" name="Texas" pageId="5" pageNumber="4">Texas</collectingRegion>
|
|
,
|
|
<collectingCountry id="F36D76D59E1F66534A9AFE736C7CFE00" box="[1274,1332,451,475]" name="United States of America" pageId="5" pageNumber="4">USA</collectingCountry>
|
|
............ ......................................................................................................................
|
|
<emphasis id="B90EEA579E1F66534AD4FE556CE8FE20" bold="true" box="[1204,1440,483,508]" pageId="5" pageNumber="4">
|
|
<taxonomicName id="4C7A4DC69E1F66534AD4FE556C01FE27" authority="Stebnicka & Skelley, 2009" authorityName="Stebnicka & Skelley" authorityYear="2009" box="[1204,1353,485,508]" class="Insecta" family="Aphodiidae" genus="Haroldiataenius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="buvexus" status="sp. nov.">
|
|
<emphasis id="B90EEA579E1F66534AD4FE556C01FE27" bold="true" box="[1204,1353,485,508]" italics="true" pageId="5" pageNumber="4">H. buvexus</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicNameLabel id="A23D572C9E1F66534B38FE536CE8FE20" box="[1368,1440,483,507]" pageId="5" pageNumber="4" rank="species">n. sp.</taxonomicNameLabel>
|
|
</emphasis>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="8BC536459E1F66534EA0FDB46CE8FD80" blockId="5.[192,1440,419,604]" pageId="5" pageNumber="4">
|
|
<keyLead id="308B95709E1F66534EA0FDB46CE8FD80" pageId="5" pageNumber="4">
|
|
— Clypeal anterior margin angulate each side of median emargination (
|
|
<figureCitation id="13412AC09E1F66534A0AFDB36DF7FDC0" box="[1130,1215,515,539]" captionStart="Figures 8-11" captionStartId="6.[192,287,989,1011]" captionTargetBox="[192,1443,196,958]" captionTargetId="figure-305@6.[190,1445,194,960]" captionTargetPageId="6" captionText="Figures 8-11. Haroldiataenius spp., lateral habitus. 8) H. semipilosus. 9) H. griffini. 10) H. buvexus, paratype. 11) H. convexus, holotype." figureDoi="http://doi.org/10.5281/zenodo.4531519" httpUri="https://zenodo.org/record/4531519/files/figure.png" pageId="5" pageNumber="4">Fig. 11</figureCitation>
|
|
). Elytral intervals strongly convex on disc. Body dark reddish brown. Southern coastal
|
|
<collectingRegion id="49BEF8A79E1F66534A11FD946DF5FDE0" box="[1137,1213,548,571]" country="United States of America" name="Texas" pageId="5" pageNumber="4">Texas</collectingRegion>
|
|
,
|
|
<collectingCountry id="F36D76D59E1F66534AA7FD936C49FDE0" box="[1223,1281,547,571]" name="United States of America" pageId="5" pageNumber="4">USA</collectingCountry>
|
|
................... ..........................................................................................................
|
|
<taxonomicName id="4C7A4DC69E1F66534A00FDF56CE8FD80" authority="(Robinson)" baseAuthorityName="Robinson" box="[1120,1440,579,604]" class="Insecta" family="Aphodiidae" genus="Haroldiataenius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="convexus">
|
|
<emphasis id="B90EEA579E1F66534A00FDF56CE8FD80" bold="true" box="[1120,1440,579,604]" pageId="5" pageNumber="4">
|
|
<emphasis id="B90EEA579E1F66534A00FDF56C49FD87" bold="true" box="[1120,1281,581,604]" italics="true" pageId="5" pageNumber="4">H. convexus</emphasis>
|
|
(Robinson)
|
|
</emphasis>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |