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<document id="A0E522BAC4B589EFFFB5DA31EDD8D41F" ID-DOI="10.5281/zenodo.5384590" ID-ISSN="2345-7600" ID-Zenodo-Dep="5384590" ID-ZooBank="40BCDD62-D35E-46D1-95A3-2CC0DF219DEE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630299679004" checkinUser="valdenar" docAuthor="Ng, Peter K. L. &amp; Forges, Bertrand Richer De" docDate="2015" docId="03A9654BFFA2077B54D9FF3E7CA3F8A7" docLanguage="en" docName="63rbz110-225.pdf" docOrigin="Raffles Bulletin of Zoology 63" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Alcomaja Ng &amp; Forges 2015, n. gen." docType="key" docUuid="C3EE8C3E-4F10-433C-B1FA-974B8F3E563E" docUuidSource="ZooBank" docVersion="5" lastPageNumber="145" masterDocId="FF901D33FF8107585767FF837962FFA3" masterDocTitle="Revision of the spider crab genus Maja Lamarck, 1801 (Crustacea: Brachyura: Majoidea: Majidae), with descriptions of seven new genera and 17 new species from the Atlantic and Indo-West Pacific" masterLastPageNumber="225" masterPageNumber="110" pageNumber="145" updateTime="1699220132769" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="4A244EEB8CCA9C0FE073DFF52D55B10E">Revision of the spider crab genus Maja Lamarck, 1801 (Crustacea: Brachyura: Majoidea: Majidae), with descriptions of seven new genera and 17 new species from the Atlantic and Indo-West Pacific</mods:title>
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<mods:namePart id="F97AC150E607FE39E31389CA9C3F470C">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="F0CB8A32A963645B369B7678C6F3483C">Forges, Bertrand Richer De</mods:namePart>
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<mods:title id="E9704713103529CF72A93E5E41564911">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="A9DABCCC5E8E3BB47CD0AF6AAE5EA586">2015</mods:date>
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<treatment id="03A9654BFFA2077B54D9FF3E7CA3F8A7" ID-DOI="http://doi.org/10.5281/zenodo.5466795" ID-Zenodo-Dep="5466795" LSID="urn:lsid:zoobank.org:act:C3EE8C3E-4F10-433C-B1FA-974B8F3E563E" httpUri="http://treatment.plazi.org/id/03A9654BFFA2077B54D9FF3E7CA3F8A7" lastPageNumber="145" pageId="35" pageNumber="145">
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<heading id="D0F76331FFA2077B54D9FF3E7C2AFF76" bold="true" box="[958,1352,188,213]" centered="true" fontSize="10" level="2" pageId="35" pageNumber="145" reason="2">
<emphasis id="B974084FFFA2077B54D9FF3E7C2AFF76" bold="true" box="[958,1352,188,213]" pageId="35" pageNumber="145">
Key to species of
<taxonomicName id="4C00AFDEFFA2077B53ECFF3F7D96FF77" ID-CoL="7NNHK" authority="Ng &amp; Forges, 2015" authorityName="Ng &amp; Forges" authorityYear="2015" box="[1163,1268,188,212]" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="145" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B974084FFFA2077B53ECFF3F7D96FF77" bold="true" box="[1163,1268,188,212]" italics="true" pageId="35" pageNumber="145">Alcomaja</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B534FFA2077B539CFF3E7C2AFF76" box="[1275,1352,189,213]" pageId="35" pageNumber="145" rank="genus">n. gen.</taxonomicNameLabel>
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<subSubSection id="C31A87D6FFA2077B5424FF7B7CA3F8A7" pageId="35" pageNumber="145" type="key">
<keyStep id="30F4CCF8FFA2077B5424FF7B7CA3FE19" pageId="35" pageNumber="145">
<paragraph id="8BBFD45DFFA2077B5424FF7B7CA3FEC7" blockId="35.[835,1475,248,443]" pageId="35" pageNumber="145">
<keyLead id="30F17768FFA2077B5424FF7B7CA3FEC7" pageId="35" pageNumber="145">
1. Dorsal surface of carapace and ambulatory legs densely covered with long, stiff setae (
<figureCitation id="133BC8D8FFA2077B5353FE967DEBFE88" box="[1076,1161,277,299]" captionStart="Fig" captionStartId="40.[113,149,1934,1956]" captionTargetBox="[113,1451,257,1918]" captionTargetId="figure-15@40.[113,1451,257,1920]" captionTargetPageId="40" captionText="Fig. 17. General habitus, Alcomaja species. A, A. nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; B, A. nagashimaensis, male (37.2 × 32.4 mm) (CBM-ZC9928), Japan; C, A. nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Japan; D, A. latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; E, A. miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; F, A. desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384630" httpUri="https://zenodo.org/record/5384630/files/figure.png" pageId="35" pageNumber="145">Fig. 17F</figureCitation>
); G1 almost straight with distinct ventral fold (
<figureCitation id="133BC8D8FFA2077B548FFEB17D3FFEEB" box="[1000,1117,306,328]" captionStart="Fig" captionStartId="43.[136,171,1934,1956]" captionTargetBox="[138,1474,203,1912]" captionTargetId="figure-8@43.[132,1494,190,1915]" captionTargetPageId="43" captionText="Fig. 20. Left G1s, Alcomaja species. A, B, A. nagashimaensis (as Maja nagashimaensis, after T. Sakai, 1976: text-fig. 129); C, D, A. nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; EG, A. nagashimaensis, male (37.2 × 32.4 mm) (CBM- ZC9928), Japan; HJ, A. nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; KN, A. latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; OQ, A. miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; RT, A. desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines. Scale bars = 1.0 mm [H, KR]; 2.0 mm [E]; 0.5 mm [C, F, G, I, J, S, T]; 0.25 mm [D]." figureDoi="http://doi.org/10.5281/zenodo.5384636" httpUri="https://zenodo.org/record/5384636/files/figure.png" pageId="35" pageNumber="145">Fig. 20RT</figureCitation>
);
<collectingCountry id="F31794CDFFA2077B5317FEB17DBEFEEB" box="[1136,1244,306,328]" name="Philippines" pageId="35" pageNumber="145">Philippines</collectingCountry>
..................................... .........................................................
<taxonomicName id="4C00AFDEFFA2077B53A3FECC7CE6FEC7" ID-CoL="BKVT" authority="Ng &amp; Forges, 2015" authorityName="Ng &amp; Forges" authorityYear="2015" box="[1220,1412,335,356]" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="145" phylum="Arthropoda" rank="species" species="desmondi" status="sp. nov.">
<emphasis id="B974084FFFA2077B53A3FECC7CE6FEC7" box="[1220,1412,335,356]" italics="true" pageId="35" pageNumber="145">Alcomaja desmondi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B534FFA2077B52ECFECC7CA3FEC7" box="[1419,1473,335,356]" pageId="35" pageNumber="145" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="8BBFD45DFFA2077B5424FEEF7CA3FE19" blockId="35.[835,1475,248,443]" pageId="35" pageNumber="145">
<keyLead id="30F17768FFA2077B5424FEEF7CA3FE19" pageId="35" pageNumber="145"> Dorsal surface of carapace and ambulatory legs covered with scattered short, hooked setae, sometimes dense; G1 curved... .................................................................................................2</keyLead>
</paragraph>
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<keyStep id="30F4CCF8FFA2077B5424FE5C7CA3FD78" pageId="35" pageNumber="145">
<paragraph id="8BBFD45DFFA2077B5424FE5C7CA3FDE9" blockId="35.[835,1476,479,731]" pageId="35" pageNumber="145">
<keyLead id="30F17768FFA2077B5424FE5C7CA3FDE9" pageId="35" pageNumber="145">
2. Dorsal surface of carapace covered with small rounded granules, lateral and branchial spines strong even as adults, with 2 hepatic spines; adult ambulatory legs relatively short, with first leg of adult males not substantially elongated (
<figureCitation id="133BC8D8FFA2077B5393FDB67C13FDE8" box="[1268,1393,565,587]" captionStart="Fig" captionStartId="40.[113,149,1934,1956]" captionTargetBox="[113,1451,257,1918]" captionTargetId="figure-15@40.[113,1451,257,1920]" captionTargetPageId="40" captionText="Fig. 17. General habitus, Alcomaja species. A, A. nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; B, A. nagashimaensis, male (37.2 × 32.4 mm) (CBM-ZC9928), Japan; C, A. nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Japan; D, A. latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; E, A. miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; F, A. desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384630" httpUri="https://zenodo.org/record/5384630/files/figure.png" pageId="35" pageNumber="145">Figs. 17AE</figureCitation>
) .........3
</keyLead>
</paragraph>
<paragraph id="8BBFD45DFFA2077B5424FDD17CA3FD78" blockId="35.[835,1476,479,731]" pageId="35" pageNumber="145">
<keyLead id="30F17768FFA2077B5424FDD17CA3FD78" pageId="35" pageNumber="145">
Dorsal surface of carapace covered with small and large granules, sometimes sharp, lateral and branchial spines distinct but low to absent in adults, with 2 hepatic spines; adult ambulatory legs long, with first leg of adult males prominently elongated (
<figureCitation id="133BC8D8FFA2077B5417FD467A8FFD78" box="[880,1005,709,731]" captionStart="Fig" captionStartId="39.[136,171,1902,1924]" captionTargetBox="[136,1474,291,1887]" captionTargetId="figure-8@39.[136,1474,291,1887]" captionTargetPageId="39" captionText="Fig. 16. General habitus, Alcomaja species. A, A. irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; B, A. gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea; C, A. gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; D, A. confragosa, female (21.5 × 18.2 mm) (NMV J53978), Australia; E, A. confragosa, holotype male (as Maja confragosa, after Griffin &amp; Tranter 1986: pl. 16a)." figureDoi="http://doi.org/10.5281/zenodo.5384628" httpUri="https://zenodo.org/record/5384628/files/figure.png" pageId="35" pageNumber="145">Figs. 16AE</figureCitation>
)..........................................................................5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F4CCF8FFA2077B5424FD7C7CA3FC07" pageId="35" pageNumber="145">
<paragraph id="8BBFD45DFFA2077B5424FD7C7CA3FCC8" blockId="35.[835,1474,767,933]" pageId="35" pageNumber="145">
<keyLead id="30F17768FFA2077B5424FD7C7CA3FCC8" pageId="35" pageNumber="145">
3. Carapace relatively more elongate; pseudorostral spines long (
<figureCitation id="133BC8D8FFA2077B5417FC987AB7FC92" box="[880,981,795,817]" captionStart="Fig" captionStartId="40.[113,149,1934,1956]" captionTargetBox="[113,1451,257,1918]" captionTargetId="figure-15@40.[113,1451,257,1920]" captionTargetPageId="40" captionText="Fig. 17. General habitus, Alcomaja species. A, A. nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; B, A. nagashimaensis, male (37.2 × 32.4 mm) (CBM-ZC9928), Japan; C, A. nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Japan; D, A. latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; E, A. miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; F, A. desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384630" httpUri="https://zenodo.org/record/5384630/files/figure.png" pageId="35" pageNumber="145">Figs. 17E</figureCitation>
,
<figureCitation id="133BC8D8FFA2077B5484FC987D73FC92" box="[995,1041,795,817]" captionStart="Fig" captionStartId="72.[113,148,1742,1764]" captionTargetBox="[113,1451,281,1727]" captionTargetId="figure-15@72.[113,1451,281,1728]" captionTargetPageId="72" captionText="Fig. 36. Frontal regions of carapaces. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja brachydactyla, male (161.2 × 140.1 mm) (ZRC 2008.0179), Spain; D, Maja cornuta, male (115.3 × 103.4 mm) (ZRC 2013.1184), South Africa; E, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; F, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; G, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; H, Paramaja kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; I, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; J, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; K, Paramaja turgida n. sp., paratype male (67.9 × 60.5 mm) (MNHN), Philippines; L, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; M, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; N, Alcomaja latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; O, Alcomaja miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar." figureDoi="http://doi.org/10.5281/zenodo.5384670" httpUri="https://zenodo.org/record/5384670/files/figure.png" pageId="35" pageNumber="145">36O</figureCitation>
); G1 gently curved with distal part gently curved downwards (
<figureCitation id="133BC8D8FFA2077B5357FCBB7DCBFCED" box="[1072,1193,824,846]" captionStart="Fig" captionStartId="43.[136,171,1934,1956]" captionTargetBox="[138,1474,203,1912]" captionTargetId="figure-8@43.[132,1494,190,1915]" captionTargetPageId="43" captionText="Fig. 20. Left G1s, Alcomaja species. A, B, A. nagashimaensis (as Maja nagashimaensis, after T. Sakai, 1976: text-fig. 129); C, D, A. nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; EG, A. nagashimaensis, male (37.2 × 32.4 mm) (CBM- ZC9928), Japan; HJ, A. nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; KN, A. latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; OQ, A. miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; RT, A. desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines. Scale bars = 1.0 mm [H, KR]; 2.0 mm [E]; 0.5 mm [C, F, G, I, J, S, T]; 0.25 mm [D]." figureDoi="http://doi.org/10.5281/zenodo.5384636" httpUri="https://zenodo.org/record/5384636/files/figure.png" pageId="35" pageNumber="145">Fig. 20OQ</figureCitation>
);
<collectingCountry id="F31794CDFFA2077B53DBFCBB7C52FCED" box="[1212,1328,824,846]" name="Madagascar" pageId="35" pageNumber="145">Madagascar</collectingCountry>
........................ ..............................................................
<taxonomicName id="4C00AFDEFFA2077B5384FCD67CE1FCC9" ID-CoL="BKVX" authority="Ng &amp; Forges, 2015" authorityName="Ng &amp; Forges" authorityYear="2015" box="[1251,1411,853,874]" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="145" phylum="Arthropoda" rank="species" species="miriky" status="sp. nov.">
<emphasis id="B974084FFFA2077B5384FCD67CE1FCC9" box="[1251,1411,853,874]" italics="true" pageId="35" pageNumber="145">Alcomaja miriky</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B534FFA2077B52ECFCD57CA3FCC8" box="[1419,1473,854,875]" pageId="35" pageNumber="145" rank="species">n. sp.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BBFD45DFFA2077B5424FCF17CA3FC07" blockId="35.[835,1474,767,933]" pageId="35" pageNumber="145">
<keyLead id="30F17768FFA2077B5424FCF17CA3FC07" pageId="35" pageNumber="145">
Carapace relatively broad; pseudorostral spines relatively shorter (
<figureCitation id="133BC8D8FFA2077B5408FC0C7A93FC07" box="[879,1009,911,933]" captionStart="Fig" captionStartId="40.[113,149,1934,1956]" captionTargetBox="[113,1451,257,1918]" captionTargetId="figure-15@40.[113,1451,257,1920]" captionTargetPageId="40" captionText="Fig. 17. General habitus, Alcomaja species. A, A. nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; B, A. nagashimaensis, male (37.2 × 32.4 mm) (CBM-ZC9928), Japan; C, A. nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Japan; D, A. latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; E, A. miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; F, A. desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384630" httpUri="https://zenodo.org/record/5384630/files/figure.png" pageId="35" pageNumber="145">Figs. 17AD</figureCitation>
,
<figureCitation id="133BC8D8FFA2077B549AFC0C7D2EFC07" box="[1021,1100,911,933]" captionStart="Fig" captionStartId="72.[113,148,1742,1764]" captionTargetBox="[113,1451,281,1727]" captionTargetId="figure-15@72.[113,1451,281,1728]" captionTargetPageId="72" captionText="Fig. 36. Frontal regions of carapaces. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja brachydactyla, male (161.2 × 140.1 mm) (ZRC 2008.0179), Spain; D, Maja cornuta, male (115.3 × 103.4 mm) (ZRC 2013.1184), South Africa; E, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; F, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; G, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; H, Paramaja kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; I, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; J, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; K, Paramaja turgida n. sp., paratype male (67.9 × 60.5 mm) (MNHN), Philippines; L, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; M, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; N, Alcomaja latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; O, Alcomaja miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar." figureDoi="http://doi.org/10.5281/zenodo.5384670" httpUri="https://zenodo.org/record/5384670/files/figure.png" pageId="35" pageNumber="145">36M, N</figureCitation>
); G1 similar to above or otherwise ..4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F4CCF8FFA2077B5424FC4B7CA3FB28" pageId="35" pageNumber="145">
<paragraph id="8BBFD45DFFA2077B5424FC4B7CA3FB96" blockId="35.[835,1474,968,1163]" pageId="35" pageNumber="145">
<keyLead id="30F17768FFA2077B5424FC4B7CA3FB96" pageId="35" pageNumber="145">
4. Intercalated spine acutely triangular (
<figureCitation id="133BC8D8FFA2077B53BEFC4B7C58FC7D" box="[1241,1338,968,990]" captionStart="Fig" captionStartId="72.[113,148,1742,1764]" captionTargetBox="[113,1451,281,1727]" captionTargetId="figure-15@72.[113,1451,281,1728]" captionTargetPageId="72" captionText="Fig. 36. Frontal regions of carapaces. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja brachydactyla, male (161.2 × 140.1 mm) (ZRC 2008.0179), Spain; D, Maja cornuta, male (115.3 × 103.4 mm) (ZRC 2013.1184), South Africa; E, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; F, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; G, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; H, Paramaja kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; I, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; J, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; K, Paramaja turgida n. sp., paratype male (67.9 × 60.5 mm) (MNHN), Philippines; L, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; M, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; N, Alcomaja latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; O, Alcomaja miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar." figureDoi="http://doi.org/10.5281/zenodo.5384670" httpUri="https://zenodo.org/record/5384670/files/figure.png" pageId="35" pageNumber="145">Fig. 36M</figureCitation>
); G1 strongly curved, with distal part gently curved downwards (
<figureCitation id="133BC8D8FFA2077B522EFC667CDAFC58" box="[1353,1464,997,1019]" captionStart="Fig" captionStartId="43.[136,171,1934,1956]" captionTargetBox="[138,1474,203,1912]" captionTargetId="figure-8@43.[132,1494,190,1915]" captionTargetPageId="43" captionText="Fig. 20. Left G1s, Alcomaja species. A, B, A. nagashimaensis (as Maja nagashimaensis, after T. Sakai, 1976: text-fig. 129); C, D, A. nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; EG, A. nagashimaensis, male (37.2 × 32.4 mm) (CBM- ZC9928), Japan; HJ, A. nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; KN, A. latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; OQ, A. miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; RT, A. desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines. Scale bars = 1.0 mm [H, KR]; 2.0 mm [E]; 0.5 mm [C, F, G, I, J, S, T]; 0.25 mm [D]." figureDoi="http://doi.org/10.5281/zenodo.5384636" httpUri="https://zenodo.org/record/5384636/files/figure.png" pageId="35" pageNumber="145">Fig. 20AJ</figureCitation>
);
<collectingCountry id="F31794CDFFA2077B540FFB817AFDFBB4" box="[872,927,1026,1047]" name="Japan" pageId="35" pageNumber="145">Japan</collectingCountry>
and
<collectingCountry id="F31794CDFFA2077B54B6FB817D7BFBBB" box="[977,1049,1026,1048]" name="Taiwan" pageId="35" pageNumber="145">Taiwan</collectingCountry>
...................................................................... .............................
<taxonomicName id="4C00AFDEFFA2077B537BFB9C7CA3FB96" ID-CoL="BKVY" authority="(T. Sakai, 1969)" baseAuthorityName="T. Sakai" baseAuthorityYear="1969" box="[1052,1473,1055,1077]" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="145" phylum="Arthropoda" rank="species" species="nagashimaensis">
<emphasis id="B974084FFFA2077B537BFB9C7C7BFB97" box="[1052,1305,1055,1076]" italics="true" pageId="35" pageNumber="145">Alcomaja nagashimaensis</emphasis>
(T.
<bibRefCitation id="EF91A9ACFFA2077B5221FB9C7CD8FB96" author="Sakai T" box="[1350,1466,1055,1077]" pageId="35" pageNumber="145" pagination="243 - 280" refId="ref88407" refString="Sakai T (1969) Two new genera and twenty-two new species of crabs from Japan. Proceedings of the Biological Society of Washington, 82: 243 - 280, figs. 1 - 19, pls. 1, 2." type="journal article" year="1969">Sakai, 1969</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBFD45DFFA2077B5424FBBF7CA3FB28" blockId="35.[835,1474,968,1163]" pageId="35" pageNumber="145">
<keyLead id="30F17768FFA2077B5424FBBF7CA3FB28" pageId="35" pageNumber="145">
Intercalated spine basally subquadrate (
<figureCitation id="133BC8D8FFA2077B5388FBB87C2EFBF2" box="[1263,1356,1083,1105]" captionStart="Fig" captionStartId="72.[113,148,1742,1764]" captionTargetBox="[113,1451,281,1727]" captionTargetId="figure-15@72.[113,1451,281,1728]" captionTargetPageId="72" captionText="Fig. 36. Frontal regions of carapaces. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja brachydactyla, male (161.2 × 140.1 mm) (ZRC 2008.0179), Spain; D, Maja cornuta, male (115.3 × 103.4 mm) (ZRC 2013.1184), South Africa; E, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; F, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; G, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; H, Paramaja kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; I, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; J, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; K, Paramaja turgida n. sp., paratype male (67.9 × 60.5 mm) (MNHN), Philippines; L, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; M, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; N, Alcomaja latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; O, Alcomaja miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar." figureDoi="http://doi.org/10.5281/zenodo.5384670" httpUri="https://zenodo.org/record/5384670/files/figure.png" pageId="35" pageNumber="145">Fig. 36N</figureCitation>
); G1 gently curved, with distal part gently curved upwards (
<figureCitation id="133BC8D8FFA2077B525CFBDB7CD5FBCD" box="[1339,1463,1112,1134]" captionStart="Fig" captionStartId="43.[136,171,1934,1956]" captionTargetBox="[138,1474,203,1912]" captionTargetId="figure-8@43.[132,1494,190,1915]" captionTargetPageId="43" captionText="Fig. 20. Left G1s, Alcomaja species. A, B, A. nagashimaensis (as Maja nagashimaensis, after T. Sakai, 1976: text-fig. 129); C, D, A. nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; EG, A. nagashimaensis, male (37.2 × 32.4 mm) (CBM- ZC9928), Japan; HJ, A. nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; KN, A. latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; OQ, A. miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; RT, A. desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines. Scale bars = 1.0 mm [H, KR]; 2.0 mm [E]; 0.5 mm [C, F, G, I, J, S, T]; 0.25 mm [D]." figureDoi="http://doi.org/10.5281/zenodo.5384636" httpUri="https://zenodo.org/record/5384636/files/figure.png" pageId="35" pageNumber="145">Fig. 20KM</figureCitation>
);
<collectingCountry id="F31794CDFFA2077B540DFBF67D6FFB28" box="[874,1037,1141,1163]" name="Solomon Islands" pageId="35" pageNumber="145">Solomon Islands</collectingCountry>
...................................
<taxonomicName id="4C00AFDEFFA2077B5380FBF67CE1FB29" ID-CoL="BKVW" authority="Ng &amp; Forges, 2015" authorityName="Ng &amp; Forges" authorityYear="2015" box="[1255,1411,1141,1162]" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="145" phylum="Arthropoda" rank="species" species="latens" status="sp. nov.">
<emphasis id="B974084FFFA2077B5380FBF67CE1FB29" box="[1255,1411,1141,1162]" italics="true" pageId="35" pageNumber="145">Alcomaja latens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B534FFA2077B52ECFBF57CA3FB28" box="[1419,1473,1142,1163]" pageId="35" pageNumber="145" rank="species">n. sp.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F4CCF8FFA2077B5424FB2C7CA3FA2D" pageId="35" pageNumber="145">
<paragraph id="8BBFD45DFFA2077B5424FB2C7CA3FA9B" blockId="35.[835,1475,1199,1422]" pageId="35" pageNumber="145">
<keyLead id="30F17768FFA2077B5424FB2C7CA3FA9B" pageId="35" pageNumber="145">
5. Dorsal surface of carapace with distinct small and large sharp and rounded granules; intercalated tooth triangular (
<figureCitation id="133BC8D8FFA2077B5239FB487A18FB5E" captionStart="Fig" captionStartId="39.[136,171,1902,1924]" captionTargetBox="[136,1474,291,1887]" captionTargetId="figure-8@39.[136,1474,291,1887]" captionTargetPageId="39" captionText="Fig. 16. General habitus, Alcomaja species. A, A. irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; B, A. gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea; C, A. gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; D, A. confragosa, female (21.5 × 18.2 mm) (NMV J53978), Australia; E, A. confragosa, holotype male (as Maja confragosa, after Griffin &amp; Tranter 1986: pl. 16a)." figureDoi="http://doi.org/10.5281/zenodo.5384628" httpUri="https://zenodo.org/record/5384628/files/figure.png" pageId="35" pageNumber="145">Figs. 16D, E</figureCitation>
,
<figureCitation id="133BC8D8FFA2077B5419FB6B7AA5FB5D" box="[894,967,1256,1278]" captionStart="Fig" captionStartId="41.[136,171,1906,1928]" captionTargetBox="[136,1474,257,1891]" captionTargetId="figure-8@41.[136,1474,257,1891]" captionTargetPageId="41" captionText="Fig. 18. Frontal regions of carapace showing different structures of intercalated teeth, Alcomaja species. A, A. confragosa, holotype male (as Maja confragosa, after Griffin &amp; Tranter, 1986: pl. 16a); B, A. confragosa, female (21.5 × 18.2 mm) (NMV J53978), Australia; C, A. gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea; D, A. gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; E, A. irrorata n. sp., paratype male (44.7 × 39.1 mm) (ZRC 2013.1229), Philippines; F, A. irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384632" httpUri="https://zenodo.org/record/5384632/files/figure.png" pageId="35" pageNumber="145">18A, B</figureCitation>
); G1 gently curved, distal part relatively short (
<figureCitation id="133BC8D8FFA2077B52FCFB6B7AF5FAB8" captionStart="Fig" captionStartId="42.[113,148,794,816]" captionTargetBox="[114,1451,189,779]" captionTargetId="figure-15@42.[104,1473,179,782]" captionTargetPageId="42" captionText="Fig. 19. Left G1s, Alcomaja species. AC, A. irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; DF, A. gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea; G, A. confragosa (as Maja confragosa, after Griffin &amp; Tranter, 1986: fig. 73a). Scale bars = 1.0 mm [AC]." figureDoi="http://doi.org/10.5281/zenodo.5384634" httpUri="https://zenodo.org/record/5384634/files/figure.png" pageId="35" pageNumber="145">Fig. 19G</figureCitation>
);
<collectingCountry id="F31794CDFFA2077B54CDFA867D65FAB8" box="[938,1031,1285,1307]" name="Indonesia" pageId="35" pageNumber="145">Moluccas</collectingCountry>
and Western Australia ................................... ......................
<taxonomicName id="4C00AFDEFFA2077B5496FAA17CA3FA9B" ID-CoL="BKVS" authority="(Griffin &amp; Tranter, 1986)" baseAuthorityName="Griffin &amp; Tranter" baseAuthorityYear="1986" box="[1009,1473,1314,1336]" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="145" phylum="Arthropoda" rank="species" species="confragosa">
<emphasis id="B974084FFFA2077B5496FAA17DA3FA94" box="[1009,1217,1314,1335]" italics="true" pageId="35" pageNumber="145">Alcomaja confragosa</emphasis>
(
<bibRefCitation id="EF91A9ACFFA2077B53A8FAA17CD8FA9B" author="Griffin DJG &amp; Tranter HA" box="[1231,1466,1314,1336]" pageId="35" pageNumber="145" pagination="1 - 335" refId="ref84094" refString="Griffin DJG &amp; Tranter HA (1986) The Decapoda Brachyura of the Siboga Expedition. Part VIII. Majidae. Siboga-Expeditie, 39 C 4: 1 - 335, 22 pls." type="journal article" year="1986">Griffin &amp; Tranter, 1986</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBFD45DFFA2077B5424FABC7CA3FA2D" blockId="35.[835,1475,1199,1422]" pageId="35" pageNumber="145">
<keyLead id="30F17768FFA2077B5424FABC7CA3FA2D" pageId="35" pageNumber="145">
Dorsal surface of carapace as above or otherwise (
<figureCitation id="133BC8D8FFA2077B5225FABC7CDCFAF6" box="[1346,1470,1343,1365]" captionStart="Fig" captionStartId="39.[136,171,1902,1924]" captionTargetBox="[136,1474,291,1887]" captionTargetId="figure-8@39.[136,1474,291,1887]" captionTargetPageId="39" captionText="Fig. 16. General habitus, Alcomaja species. A, A. irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; B, A. gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea; C, A. gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; D, A. confragosa, female (21.5 × 18.2 mm) (NMV J53978), Australia; E, A. confragosa, holotype male (as Maja confragosa, after Griffin &amp; Tranter 1986: pl. 16a)." figureDoi="http://doi.org/10.5281/zenodo.5384628" httpUri="https://zenodo.org/record/5384628/files/figure.png" pageId="35" pageNumber="145">Figs. 16AC</figureCitation>
,
<figureCitation id="133BC8D8FFA2077B540EFADF7ACCFAD1" box="[873,942,1372,1394]" captionStart="Fig" captionStartId="41.[136,171,1906,1928]" captionTargetBox="[136,1474,257,1891]" captionTargetId="figure-8@41.[136,1474,257,1891]" captionTargetPageId="41" captionText="Fig. 18. Frontal regions of carapace showing different structures of intercalated teeth, Alcomaja species. A, A. confragosa, holotype male (as Maja confragosa, after Griffin &amp; Tranter, 1986: pl. 16a); B, A. confragosa, female (21.5 × 18.2 mm) (NMV J53978), Australia; C, A. gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea; D, A. gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; E, A. irrorata n. sp., paratype male (44.7 × 39.1 mm) (ZRC 2013.1229), Philippines; F, A. irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384632" httpUri="https://zenodo.org/record/5384632/files/figure.png" pageId="35" pageNumber="145">18CF</figureCitation>
); intercalated tooth subtruncate basally (
<figureCitation id="133BC8D8FFA2077B5225FAD87CDAFAD1" box="[1346,1464,1371,1394]" captionStart="Fig" captionStartId="41.[136,171,1906,1928]" captionTargetBox="[136,1474,257,1891]" captionTargetId="figure-8@41.[136,1474,257,1891]" captionTargetPageId="41" captionText="Fig. 18. Frontal regions of carapace showing different structures of intercalated teeth, Alcomaja species. A, A. confragosa, holotype male (as Maja confragosa, after Griffin &amp; Tranter, 1986: pl. 16a); B, A. confragosa, female (21.5 × 18.2 mm) (NMV J53978), Australia; C, A. gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea; D, A. gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; E, A. irrorata n. sp., paratype male (44.7 × 39.1 mm) (ZRC 2013.1229), Philippines; F, A. irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384632" httpUri="https://zenodo.org/record/5384632/files/figure.png" pageId="35" pageNumber="145">Fig. 18CF</figureCitation>
): G1 gently or strongly curved..................................................6
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<keyStep id="30F4CCF8FFA2077B5424FA317CA3F8A7" pageId="35" pageNumber="145">
<paragraph id="8BBFD45DFFA2077B5424FA317CA3F9FB" blockId="35.[835,1475,1458,1796]" pageId="35" pageNumber="145">
<keyLead id="30F17768FFA2077B5424FA317CA3F9FB" pageId="35" pageNumber="145">
6. Dorsal surface of carapace with distinct small and large sharp, rounded granules (
<figureCitation id="133BC8D8FFA2077B5339FA4C7DB9FA47" box="[1118,1243,1487,1509]" captionStart="Fig" captionStartId="39.[136,171,1902,1924]" captionTargetBox="[136,1474,291,1887]" captionTargetId="figure-8@39.[136,1474,291,1887]" captionTargetPageId="39" captionText="Fig. 16. General habitus, Alcomaja species. A, A. irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; B, A. gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea; C, A. gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; D, A. confragosa, female (21.5 × 18.2 mm) (NMV J53978), Australia; E, A. confragosa, holotype male (as Maja confragosa, after Griffin &amp; Tranter 1986: pl. 16a)." figureDoi="http://doi.org/10.5281/zenodo.5384628" httpUri="https://zenodo.org/record/5384628/files/figure.png" pageId="35" pageNumber="145">Figs. 16B, C</figureCitation>
,
<figureCitation id="133BC8D8FFA2077B5381FA4C7C4CFA47" box="[1254,1326,1487,1508]" captionStart="Fig" captionStartId="41.[136,171,1906,1928]" captionTargetBox="[136,1474,257,1891]" captionTargetId="figure-8@41.[136,1474,257,1891]" captionTargetPageId="41" captionText="Fig. 18. Frontal regions of carapace showing different structures of intercalated teeth, Alcomaja species. A, A. confragosa, holotype male (as Maja confragosa, after Griffin &amp; Tranter, 1986: pl. 16a); B, A. confragosa, female (21.5 × 18.2 mm) (NMV J53978), Australia; C, A. gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea; D, A. gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; E, A. irrorata n. sp., paratype male (44.7 × 39.1 mm) (ZRC 2013.1229), Philippines; F, A. irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384632" httpUri="https://zenodo.org/record/5384632/files/figure.png" pageId="35" pageNumber="145">18C, D</figureCitation>
); pseudorostral spines with rounded cross-section (
<figureCitation id="133BC8D8FFA2077B53AEFA687C26F9A2" box="[1225,1348,1515,1537]" captionStart="Fig" captionStartId="41.[136,171,1906,1928]" captionTargetBox="[136,1474,257,1891]" captionTargetId="figure-8@41.[136,1474,257,1891]" captionTargetPageId="41" captionText="Fig. 18. Frontal regions of carapace showing different structures of intercalated teeth, Alcomaja species. A, A. confragosa, holotype male (as Maja confragosa, after Griffin &amp; Tranter, 1986: pl. 16a); B, A. confragosa, female (21.5 × 18.2 mm) (NMV J53978), Australia; C, A. gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea; D, A. gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; E, A. irrorata n. sp., paratype male (44.7 × 39.1 mm) (ZRC 2013.1229), Philippines; F, A. irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384632" httpUri="https://zenodo.org/record/5384632/files/figure.png" pageId="35" pageNumber="145">Fig. 18C, D</figureCitation>
); ischium of third maxilliped distinctly granular (
<figureCitation id="133BC8D8FFA2077B53B6F98B7C29F9BD" box="[1233,1355,1544,1566]" captionStart="Fig" captionStartId="81.[136,171,1723,1745]" captionTargetBox="[137,1475,328,1708]" captionTargetId="figure-8@81.[136,1475,328,1708]" captionTargetPageId="81" captionText="Fig. 45. Third maxillipeds. A, Alcomaja irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; B, Alcomaja gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea [third max laterally inverted to match]; C, Alcomaja gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; D, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; E, Alcomaja nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; F, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; G, Alcomaja latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; H, Alcomaja miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; I, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; J, Paramaya ouch n. sp., holotype male (76.8 × 60.0 mm) (NMCR), Philippines; K, Paramaya ouch n. sp., paratype male (39.7 × 29.6 mm) (ZRC 2011.0045), Philippines; L, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.5384688" httpUri="https://zenodo.org/record/5384688/files/figure.png" pageId="35" pageNumber="145">Fig. 45B, C</figureCitation>
); G1 gently curved, distal part straight (
<figureCitation id="133BC8D8FFA2077B5314F9A67D85F998" box="[1139,1255,1573,1595]" captionStart="Fig" captionStartId="42.[113,148,794,816]" captionTargetBox="[114,1451,189,779]" captionTargetId="figure-15@42.[104,1473,179,782]" captionTargetPageId="42" captionText="Fig. 19. Left G1s, Alcomaja species. AC, A. irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; DF, A. gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea; G, A. confragosa (as Maja confragosa, after Griffin &amp; Tranter, 1986: fig. 73a). Scale bars = 1.0 mm [AC]." figureDoi="http://doi.org/10.5281/zenodo.5384634" httpUri="https://zenodo.org/record/5384634/files/figure.png" pageId="35" pageNumber="145">Fig. 19DF</figureCitation>
); South
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Sea and
<collectingCountry id="F31794CDFFA2077B540DF9C17AB4F9FB" box="[874,982,1602,1624]" name="Philippines" pageId="35" pageNumber="145">Philippines</collectingCountry>
...............
<taxonomicName id="4C00AFDEFFA2077B5352F9C17CA3F9FB" ID-CoL="8W5SN" authority="(Chen &amp; Ng, 1999)" baseAuthorityName="Chen &amp; Ng" baseAuthorityYear="1999" box="[1077,1473,1602,1624]" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="145" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B974084FFFA2077B5352F9C17D9BF9F4" box="[1077,1273,1602,1623]" italics="true" pageId="35" pageNumber="145">Alcomaja gracilipes</emphasis>
(
<bibRefCitation id="EF91A9ACFFA2077B5260F9C17CD8F9FB" author="Chen HL &amp; Ng PKL" box="[1287,1466,1602,1624]" pageId="35" pageNumber="145" pagination="754 - 758" refId="ref83546" refString="Chen HL &amp; Ng PKL (1999) Description of a new spider crab, Maja gracilipes, from the South China Sea, with notes on the taxonomic validity of M. brevispinosis Dai, 1981 (Crustacea: Decapoda: Brachyura: Majidae). Proceedings of the Biological Society of Washington, 112 (4): 754 - 758, Figs. 1, 2." type="journal article" year="1999">Chen &amp; Ng, 1999</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBFD45DFFA2077B5424F9DC7CA3F8A7" blockId="35.[835,1475,1458,1796]" pageId="35" pageNumber="145">
<keyLead id="30F17768FFA2077B5424F9DC7CA3F8A7" pageId="35" pageNumber="145">
Dorsal surface of carapace with small and large rounded granules, some of which merge basally (
<figureCitation id="133BC8D8FFA2077B5398F9F87C01F932" box="[1279,1379,1659,1681]" captionStart="Fig" captionStartId="39.[136,171,1902,1924]" captionTargetBox="[136,1474,291,1887]" captionTargetId="figure-8@39.[136,1474,291,1887]" captionTargetPageId="39" captionText="Fig. 16. General habitus, Alcomaja species. A, A. irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; B, A. gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea; C, A. gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; D, A. confragosa, female (21.5 × 18.2 mm) (NMV J53978), Australia; E, A. confragosa, holotype male (as Maja confragosa, after Griffin &amp; Tranter 1986: pl. 16a)." figureDoi="http://doi.org/10.5281/zenodo.5384628" httpUri="https://zenodo.org/record/5384628/files/figure.png" pageId="35" pageNumber="145">Figs. 16A</figureCitation>
,
<figureCitation id="133BC8D8FFA2077B5217F9FF7CD4F932" box="[1392,1462,1660,1681]" captionStart="Fig" captionStartId="41.[136,171,1906,1928]" captionTargetBox="[136,1474,257,1891]" captionTargetId="figure-8@41.[136,1474,257,1891]" captionTargetPageId="41" captionText="Fig. 18. Frontal regions of carapace showing different structures of intercalated teeth, Alcomaja species. A, A. confragosa, holotype male (as Maja confragosa, after Griffin &amp; Tranter, 1986: pl. 16a); B, A. confragosa, female (21.5 × 18.2 mm) (NMV J53978), Australia; C, A. gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea; D, A. gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; E, A. irrorata n. sp., paratype male (44.7 × 39.1 mm) (ZRC 2013.1229), Philippines; F, A. irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384632" httpUri="https://zenodo.org/record/5384632/files/figure.png" pageId="35" pageNumber="145">18E, F</figureCitation>
); pseudorostral spines dorsoventrally flattened (
<figureCitation id="133BC8D8FFA2077B525AF91B7CD4F90D" box="[1341,1462,1688,1710]" captionStart="Fig" captionStartId="41.[136,171,1906,1928]" captionTargetBox="[136,1474,257,1891]" captionTargetId="figure-8@41.[136,1474,257,1891]" captionTargetPageId="41" captionText="Fig. 18. Frontal regions of carapace showing different structures of intercalated teeth, Alcomaja species. A, A. confragosa, holotype male (as Maja confragosa, after Griffin &amp; Tranter, 1986: pl. 16a); B, A. confragosa, female (21.5 × 18.2 mm) (NMV J53978), Australia; C, A. gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea; D, A. gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; E, A. irrorata n. sp., paratype male (44.7 × 39.1 mm) (ZRC 2013.1229), Philippines; F, A. irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384632" httpUri="https://zenodo.org/record/5384632/files/figure.png" pageId="35" pageNumber="145">Fig. 18E, F</figureCitation>
); ischium of third maxilliped smooth (
<figureCitation id="133BC8D8FFA2077B53BCF9367C5AF968" box="[1243,1336,1717,1739]" captionStart="Fig" captionStartId="81.[136,171,1723,1745]" captionTargetBox="[137,1475,328,1708]" captionTargetId="figure-8@81.[136,1475,328,1708]" captionTargetPageId="81" captionText="Fig. 45. Third maxillipeds. A, Alcomaja irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; B, Alcomaja gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea [third max laterally inverted to match]; C, Alcomaja gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; D, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; E, Alcomaja nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; F, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; G, Alcomaja latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; H, Alcomaja miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; I, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; J, Paramaya ouch n. sp., holotype male (76.8 × 60.0 mm) (NMCR), Philippines; K, Paramaya ouch n. sp., paratype male (39.7 × 29.6 mm) (ZRC 2011.0045), Philippines; L, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.5384688" httpUri="https://zenodo.org/record/5384688/files/figure.png" pageId="35" pageNumber="145">Fig. 45A</figureCitation>
); G1 strongly curved, distal part curved upwards (
<figureCitation id="133BC8D8FFA2077B53ADF9517C20F94B" box="[1226,1346,1746,1768]" captionStart="Fig" captionStartId="42.[113,148,794,816]" captionTargetBox="[114,1451,189,779]" captionTargetId="figure-15@42.[104,1473,179,782]" captionTargetPageId="42" captionText="Fig. 19. Left G1s, Alcomaja species. AC, A. irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; DF, A. gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea; G, A. confragosa (as Maja confragosa, after Griffin &amp; Tranter, 1986: fig. 73a). Scale bars = 1.0 mm [AC]." figureDoi="http://doi.org/10.5281/zenodo.5384634" httpUri="https://zenodo.org/record/5384634/files/figure.png" pageId="35" pageNumber="145">Fig. 19AC</figureCitation>
);
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...........................................................
<taxonomicName id="4C00AFDEFFA2077B53B4F96C7CE1F8A7" ID-CoL="BKVV" authority="Ng &amp; Forges, 2015" authorityName="Ng &amp; Forges" authorityYear="2015" box="[1235,1411,1775,1796]" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="145" phylum="Arthropoda" rank="species" species="irrorata" status="sp. nov.">
<emphasis id="B974084FFFA2077B53B4F96C7CE1F8A7" box="[1235,1411,1775,1796]" italics="true" pageId="35" pageNumber="145">Alcomaja irrorata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B534FFA2077B52ECF96C7CA3F8A7" box="[1419,1473,1775,1796]" pageId="35" pageNumber="145" rank="species">n. sp.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>