540 lines
56 KiB
XML
540 lines
56 KiB
XML
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<mods:title id="9450914B493E9126A63128F2F2F3D323">A taxonomic revision of the Palaearctic members of the subgenus Lasius s. str. (Hymenoptera, Formicidae)</mods:title>
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<mods:namePart id="22B678D335284DBB30B6A5212DB338BE">Seifert, Bernhard</mods:namePart>
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<mods:title id="FB1032F9FC7A29D1303E4BBC98A14D47">Soil Organisms</mods:title>
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<mods:part id="E7AEFDA657DED05AF4E3E318DFACB717">
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<mods:date id="CE1A5B1A8696009CF425D881055D3324">2020</mods:date>
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<mods:number id="6D30B2F705928ECDC77A3F6A868A1439">2020-05-26</mods:number>
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<mods:number id="704CD4D3BD16665E73573E50215DAC5F">92</mods:number>
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<mods:identifier id="9028188127AF297DDBB7B368A90244FC" type="ISSN">2509-9523</mods:identifier>
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</mods:mods>
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<treatment id="153287B6FD1DFFE2FF0BFED95989FC6A" LSID="urn:lsid:plazi:treatment:153287B6FD1DFFE2FF0BFED95989FC6A" httpUri="http://treatment.plazi.org/id/153287B6FD1DFFE2FF0BFED95989FC6A" lastPageNumber="48" pageId="32" pageNumber="47">
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<subSubSection id="D581652BFD1DFFE2FF0BFED95889FEBC" box="[174,708,305,333]" pageId="32" pageNumber="47" type="nomenclature">
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<paragraph id="9D2436A0FD1DFFE2FF0BFED95889FEBC" blockId="32.[174,708,305,333]" box="[174,708,305,333]" pageId="32" pageNumber="47">
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<heading id="C66C81CCFD1DFFE2FF0BFED95889FEBC" bold="true" box="[174,708,305,333]" fontSize="10" level="2" pageId="32" pageNumber="47" reason="3">
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<emphasis id="AFEFEAB2FD1DFFE2FF0BFED95889FEBC" bold="true" box="[174,708,305,333]" pageId="32" pageNumber="47">
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4.4.1
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<taxonomicName id="5A9B4D23FD1DFFE2FF5AFED95889FEBC" ID-CoL="6P3BM" authority="(Latreille 1798)" baseAuthorityName="Latreille" baseAuthorityYear="1798" box="[255,708,305,333]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="47" phylum="Arthropoda" rank="species" species="brunneus">
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<emphasis id="AFEFEAB2FD1DFFE2FF5AFED95BA7FEBD" bold="true" box="[255,490,305,332]" italics="true" pageId="32" pageNumber="47">Lasius brunneus</emphasis>
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(Latreille 1798)
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="D581652BFD1DFFE2FF0BFEBD5842FD07" pageId="32" pageNumber="47" type="reference_group">
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<paragraph id="9D2436A0FD1DFFE2FF0BFEBD5B61FD27" blockId="32.[151,766,340,1975]" pageId="32" pageNumber="47">
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<taxonomicName id="5A9B4D23FD1DFFE2FF0BFEBD585CFE9D" authority="Latreille 1798" authorityName="Latreille" authorityYear="1798" box="[174,529,340,364]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="47" phylum="Arthropoda" rank="species" species="brunnea">
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<emphasis id="AFEFEAB2FD1DFFE2FF0BFEBD5B22FE9D" box="[174,367,340,364]" italics="true" pageId="32" pageNumber="47">Formica brunnea</emphasis>
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Latreille 1798
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</taxonomicName>
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[conception of
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<bibRefCitation id="F90A4B51FD1DFFE2FD64FEBD5A9AFE7C" author="Mayr, G." pageId="32" pageNumber="47" refId="ref51698" refString="Mayr, G. (1861): Die Europaischen Formiciden (Ameisen). Wien, 80 pp." type="book" year="1861">Mayr (1861)</bibRefCitation>
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and
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<typeStatus id="42208802FD1DFFE2FEAEFE9E5B2FFE7F" box="[267,354,374,398]" pageId="32" pageNumber="47" type="neotype">neotype</typeStatus>
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fixation] Both the original description and that of Latreille (1802) do not allow a species identification. The locus typicus is Brive/
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<collectingCountry id="E58C7630FD1DFFE2FDD0FE5F588CFE3E" box="[629,705,439,463]" name="France" pageId="32" pageNumber="47">France</collectingCountry>
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. The first unambiguous description was presented by
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<bibRefCitation id="F90A4B51FD1DFFE2FD64FE305A9AFDE1" author="Mayr, G." pageId="32" pageNumber="47" refId="ref51698" refString="Mayr, G. (1861): Die Europaischen Formiciden (Ameisen). Wien, 80 pp." type="book" year="1861">Mayr (1861)</bibRefCitation>
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that has been adopted by most of the European authors since then. A
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<typeStatus id="42208802FD1DFFE2FE2DFDF25B92FDC3" box="[392,479,538,562]" pageId="32" pageNumber="47" type="neotype">neotype</typeStatus>
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is fixed herewith in
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<specimenCount id="8B9DFD29FD1DFFE2FD61FDF15AAFFDA3" count="3" pageId="32" pageNumber="47" type="worker">three worker</specimenCount>
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specimens collected near Brive, labelled ”FRA:
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<geoCoordinate id="F8AF5067FD1DFFE2FF32FDB35B46FD82" box="[151,267,603,627]" degrees="44.9125" direction="north" orientation="latitude" pageId="32" pageNumber="47" precision="5" value="44.9125">44.9125°N</geoCoordinate>
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,
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<geoCoordinate id="F8AF5067FD1DFFE2FEB1FDB35B35FD82" box="[276,376,603,627]" degrees="1.4769" direction="east" orientation="longitude" pageId="32" pageNumber="47" precision="5" value="1.4769">1.4769°E</geoCoordinate>
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, Souillac–
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<quantity id="5A639B45FD1DFFE2FE42FDB35863FD82" box="[487,558,603,627]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="32" pageNumber="47" unit="km" value="1.7">1.7 km</quantity>
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N,
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<quantity id="5A639B45FD1DFFE2FDF7FDB358D8FD82" box="[594,661,603,627]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="32" pageNumber="47" unit="m" value="120.0">120 m</quantity>
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, tree row, leg. Galkowski
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<date id="E9251060FD1DFFE2FEE2FD945B8DFD65" box="[327,448,636,660]" pageId="32" pageNumber="47" value="2008-09-07">2008.09.07</date>
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“ and ”
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<typeStatus id="42208802FD1DFFE2FDAAFD945823FD65" box="[527,622,636,660]" pageId="32" pageNumber="47" type="neotype">Neotype</typeStatus>
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(top)
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<taxonomicName id="5A9B4D23FD1DFFE2FD11FD955BE6FD44" ID-CoL="6P3BM" authority="(Latreille 1798)" baseAuthorityName="Latreille" baseAuthorityYear="1798" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="47" phylum="Arthropoda" rank="species" species="brunneus">
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<emphasis id="AFEFEAB2FD1DFFE2FD11FD955AB0FD44" italics="true" pageId="32" pageNumber="47">Lasius brunneus</emphasis>
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(Latreille 1798)
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</taxonomicName>
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des. Seifert 2019“; depository:
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<collectionCode id="FB8AAE65FD1DFFE2FF32FD565A9FFD27" box="[151,210,702,726]" pageId="32" pageNumber="47">SMN</collectionCode>
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Görlitz.
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</paragraph>
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<paragraph id="9D2436A0FD1DFFE2FF0BFD375842FD07" blockId="32.[151,766,340,1975]" box="[174,527,734,759]" pageId="32" pageNumber="47">
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<taxonomicName id="5A9B4D23FD1DFFE2FF0BFD375842FD07" authority="(Latreille 1798)" baseAuthorityName="Latreille" baseAuthorityYear="1798" box="[174,527,734,759]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="47" phylum="Arthropoda" rank="species" species="pallidus">
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<emphasis id="AFEFEAB2FD1DFFE2FF0BFD375842FD07" bold="true" box="[174,527,734,759]" pageId="32" pageNumber="47">
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<emphasis id="AFEFEAB2FD1DFFE2FF0BFD375B15FD07" bold="true" box="[174,344,734,759]" italics="true" pageId="32" pageNumber="47">Lasius pallidus</emphasis>
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(Latreille 1798)
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</emphasis>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="D581652BFD1DFFE2FF0BFCE858B1FA79" pageId="32" pageNumber="47" type="description">
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<paragraph id="9D2436A0FD1DFFE2FF0BFCE85B95FC6A" blockId="32.[151,766,340,1975]" pageId="32" pageNumber="47">
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||
<taxonomicName id="5A9B4D23FD1DFFE2FF0BFCE85842FCE9" authority="Latreille 1798" authorityName="Latreille" authorityYear="1798" box="[174,527,768,792]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="47" phylum="Arthropoda" rank="species" species="pallida">
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<emphasis id="AFEFEAB2FD1DFFE2FF0BFCE85B29FCE9" box="[174,356,768,792]" italics="true" pageId="32" pageNumber="47">Formica pallida</emphasis>
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Latreille 1798
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</taxonomicName>
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[indirect indication] This taxon is not identifiable from data in the original description. Yet, as the synonymization with
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<taxonomicName id="5A9B4D23FD1DFFE2FD11FCAA58B6FCA8" box="[692,763,834,857]" pageId="32" pageNumber="47">
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<emphasis id="AFEFEAB2FD1DFFE2FD11FCAA58B6FCA8" box="[692,763,834,857]" italics="true" pageId="32" pageNumber="47">Lasius</emphasis>
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</taxonomicName>
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<emphasis id="AFEFEAB2FD1DFFE2FF0BFC8A5B59FC8B" box="[174,276,866,890]" italics="true" pageId="32" pageNumber="47">brunneus</emphasis>
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was established by Latreille himself (Latreille 1802), I follow his decision.
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</paragraph>
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<paragraph id="9D2436A0FD1DFFE2FF0BFC4C584BFC4D" blockId="32.[151,766,340,1975]" box="[174,518,932,956]" pageId="32" pageNumber="47">
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<taxonomicName id="5A9B4D23FD1DFFE2FF0BFC4C584BFC4D" authority="(Foerster 1850)" baseAuthorityName="Foerster" baseAuthorityYear="1850" box="[174,518,932,956]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="47" phylum="Arthropoda" rank="species" species="timidus">
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<emphasis id="AFEFEAB2FD1DFFE2FF0BFC4C584BFC4D" bold="true" box="[174,518,932,956]" pageId="32" pageNumber="47">
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<emphasis id="AFEFEAB2FD1DFFE2FF0BFC4C5B1FFC4D" bold="true" box="[174,338,932,956]" italics="true" pageId="32" pageNumber="47">Lasius timidus</emphasis>
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(Foerster 1850)
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</emphasis>
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</taxonomicName>
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</paragraph>
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<paragraph id="9D2436A0FD1DFFE2FF0BFC2E58B1FB91" blockId="32.[151,766,340,1975]" pageId="32" pageNumber="47">
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<taxonomicName id="5A9B4D23FD1DFFE2FF0BFC2E5BB0FC2C" authority="Foerster 1850" authorityName="Foerster" authorityYear="1850" box="[174,509,965,989]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="47" phylum="Arthropoda" rank="species" species="timida">
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<emphasis id="AFEFEAB2FD1DFFE2FF0BFC2E5B10FC2C" box="[174,349,965,989]" italics="true" pageId="32" pageNumber="47">Formica timida</emphasis>
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Foerster 1850
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</taxonomicName>
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[original description] This taxon was described from near Aachen /
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<collectingCountry id="E58C7630FD1DFFE2FD37FC0E58B5FC0F" box="[658,760,998,1022]" name="Germany" pageId="32" pageNumber="47">Germany</collectingCountry>
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. The reported morphology (coloration, length of frontal line, pilosity) in combination with zoogeography strongly suggests a synonymy with
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<taxonomicName id="5A9B4D23FD1DFFE2FDC2FBA158B8FB91" baseAuthorityName="Latreille" baseAuthorityYear="1798" box="[615,757,1096,1120]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="47" phylum="Arthropoda" rank="species" species="brunneus">
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<emphasis id="AFEFEAB2FD1DFFE2FDC2FBA158B8FB91" box="[615,757,1096,1120]" italics="true" pageId="32" pageNumber="47">L. brunneus</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="9D2436A0FD1DFFE2FF0BFB815866FB70" blockId="32.[151,766,340,1975]" box="[174,555,1129,1153]" pageId="32" pageNumber="47">
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<taxonomicName id="5A9B4D23FD1DFFE2FF0BFB815866FB70" authority="Forel 1874" authorityName="Forel" authorityYear="1874" box="[174,555,1129,1153]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="47" phylum="Arthropoda" rank="species" species="alienobrunneus">
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<emphasis id="AFEFEAB2FD1DFFE2FF0BFB815866FB70" bold="true" box="[174,555,1129,1153]" pageId="32" pageNumber="47">
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<emphasis id="AFEFEAB2FD1DFFE2FF0BFB815BE2FB70" bold="true" box="[174,431,1129,1153]" italics="true" pageId="32" pageNumber="47">Lasius alienobrunneus</emphasis>
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Forel 1874
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</emphasis>
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</taxonomicName>
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</paragraph>
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<paragraph id="9D2436A0FD1DFFE2FF0BFB6358C6FB53" blockId="32.[151,766,340,1975]" box="[174,651,1162,1186]" pageId="32" pageNumber="47">
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<taxonomicName id="5A9B4D23FD1DFFE2FF0BFB6358C6FB53" authority="Forel 1874" authorityName="Forel" authorityYear="1874" box="[174,651,1162,1186]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="47" phylum="Arthropoda" rank="variety" species="niger" variety="alienobrunneus">
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<emphasis id="AFEFEAB2FD1DFFE2FF0BFB635859FB53" box="[174,532,1162,1186]" italics="true" pageId="32" pageNumber="47">Lasius niger var. alienobrunneus</emphasis>
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Forel 1874
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</taxonomicName>
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</paragraph>
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<paragraph id="9D2436A0FD1DFFE2FF0BFB445B28FB32" blockId="32.[151,766,340,1975]" box="[174,357,1195,1220]" pageId="32" pageNumber="47">
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[
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<typeStatus id="42208802FD1DFFE2FF11FB445AA8FB35" box="[180,229,1196,1220]" pageId="32" pageNumber="47">type</typeStatus>
|
||
specimens]
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</paragraph>
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<paragraph id="9D2436A0FD1DFFE2FF0BFB2458B1FA79" blockId="32.[151,766,340,1975]" pageId="32" pageNumber="47">
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Forel did not mention a locus typicus.
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<specimenCount id="8B9DFD29FD1DFFE2FDC9FB2458B6FB15" box="[620,763,1228,1252]" count="5" pageId="32" pageNumber="47" type="worker">Five workers</specimenCount>
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stored in
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<collectionCode id="FB8AAE65FD1DFFE2FEBCFB055B14FAF5" box="[281,345,1261,1284]" pageId="32" pageNumber="47">MHN</collectionCode>
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<collectingRegion id="5F5FF842FD1DFFE2FEC6FB055BFAFAF4" box="[355,439,1261,1285]" country="Switzerland" name="Geneve" pageId="32" pageNumber="47">Genéve</collectingRegion>
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, labelled as ”
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<typeStatus id="42208802FD1DFFE2FDEBFB0558D9FAF4" box="[590,660,1261,1285]" pageId="32" pageNumber="47">Typus</typeStatus>
|
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“ and ”L.
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<taxonomicName id="5A9B4D23FD1DFFE2FF0BFAE65BC5FAD4" authority="For." authorityName="Forel" authorityYear="1874" box="[174,392,1294,1318]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="47" phylum="Arthropoda" rank="species" species="alienobrunneus">alienobrunneus For.</taxonomicName>
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/ Vaux“ do not show any notable deviation from the normal morphology of
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<taxonomicName id="5A9B4D23FD1DFFE2FDDDFAC758B1FAB7" baseAuthorityName="Latreille" baseAuthorityYear="1798" box="[632,764,1326,1350]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="47" phylum="Arthropoda" rank="species" species="brunneus">
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<emphasis id="AFEFEAB2FD1DFFE2FDDDFAC758B1FAB7" box="[632,764,1326,1350]" italics="true" pageId="32" pageNumber="47">L. brunneus</emphasis>
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</taxonomicName>
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both by NUMOBAT data and subjective impression (pictures in www.antweb.org under CASENT0911044).
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</paragraph>
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</subSubSection>
|
||
<subSubSection id="D581652BFD1DFFE2FF0BFA795B28F8E3" pageId="32" pageNumber="47" type="materials_examined">
|
||
<paragraph id="9D2436A0FD1DFFE2FF0BFA7958C2F9BF" blockId="32.[151,766,340,1975]" pageId="32" pageNumber="47">
|
||
<emphasis id="AFEFEAB2FD1DFFE2FF0BFA795BE1FA58" bold="true" box="[174,428,1425,1449]" pageId="32" pageNumber="47">All material examined</emphasis>
|
||
. A total of 32 nest samples with
|
||
<specimenCount id="8B9DFD29FD1DFFE2FF32FA5A5B5EFA3B" box="[151,275,1458,1482]" count="67" pageId="32" pageNumber="47" type="worker">67 workers</specimenCount>
|
||
were subject to NUMOBAT investigation.
|
||
<location id="9844607BFD1DFFE2FF32FA3B5A95FA1A" LSID="urn:lsid:plazi:treatment:153287B6FD1DFFE2FF0BFED95989FC6A:9844607BFD1DFFE2FF32FA3B5A95FA1A" box="[151,216,1491,1515]" country="Austria" municipality="For" name="These" pageId="32" pageNumber="47">These</location>
|
||
originated from
|
||
<collectingCountry id="E58C7630FD1DFFE2FE3DFA3B5BA5FA1A" box="[408,488,1491,1515]" name="Austria" pageId="32" pageNumber="47">Austria</collectingCountry>
|
||
(1 sample),
|
||
<collectingCountry id="E58C7630FD1DFFE2FDD7FA3B5882FA1A" box="[626,719,1491,1515]" name="Bulgaria" pageId="32" pageNumber="47">Bulgaria</collectingCountry>
|
||
(1), England (1),
|
||
<collectingCountry id="E58C7630FD1DFFE2FE80FA1C5B23F9FA" box="[293,366,1524,1547]" name="France" pageId="32" pageNumber="47">France</collectingCountry>
|
||
(3),
|
||
<collectingCountry id="E58C7630FD1DFFE2FE01FA1C5844F9FD" box="[420,521,1524,1548]" name="Germany" pageId="32" pageNumber="47">Germany</collectingCountry>
|
||
(11),
|
||
<collectingCountry id="E58C7630FD1DFFE2FDECFA1C58DBF9FD" box="[585,662,1524,1548]" name="Greece" pageId="32" pageNumber="47">Greece</collectingCountry>
|
||
(1),
|
||
<collectingCountry id="E58C7630FD1DFFE2FD6FFA1C58B6F9FD" box="[714,763,1524,1548]" name="Italy" pageId="32" pageNumber="47">Italy</collectingCountry>
|
||
(1),
|
||
<collectingCountry id="E58C7630FD1DFFE2FF1AF9FD5AB0F9DC" box="[191,253,1557,1581]" name="Spain" pageId="32" pageNumber="47">Spain</collectingCountry>
|
||
(2),
|
||
<collectingCountry id="E58C7630FD1DFFE2FE8FF9FD5B33F9DC" box="[298,382,1557,1581]" name="Sweden" pageId="32" pageNumber="47">Sweden</collectingCountry>
|
||
(1),
|
||
<collectingCountry id="E58C7630FD1DFFE2FE09F9FD5860F9DC" box="[428,557,1557,1581]" name="Switzerland" pageId="32" pageNumber="47">Switzerland</collectingCountry>
|
||
(2), and
|
||
<collectingCountry id="E58C7630FD1DFFE2FD22F9FD589FF9DC" box="[647,722,1557,1581]" name="Turkey" pageId="32" pageNumber="47">Turkey</collectingCountry>
|
||
(8).
|
||
<collectingMunicipality id="7D40ACDAFD1DFFE2FF32F9DE5AF6F9BC" box="[151,187,1590,1613]" pageId="32" pageNumber="47">For</collectingMunicipality>
|
||
details see supplementary information SI1.
|
||
</paragraph>
|
||
<paragraph id="9D2436A0FD1DFFE2FF0BF9BE5B28F8E3" blockId="32.[151,766,340,1975]" pageId="32" pageNumber="47">
|
||
<emphasis id="AFEFEAB2FD1DFFE2FF0BF9BE5B31F99E" bold="true" box="[174,380,1622,1647]" pageId="32" pageNumber="47">Geographic range</emphasis>
|
||
.
|
||
<location id="9844607BFD1DFFE2FE2CF9BE5867F99F" LSID="urn:lsid:plazi:treatment:153287B6FD1DFFE2FF0BFED95989FC6A:9844607BFD1DFFE2FE2CF9BE5867F99F" box="[393,554,1622,1646]" country="United Kingdom" county="Caucasus. In Scandinavia" latitude="60.0" municipality="Balkans" name="Eurocaucasian" pageId="32" pageNumber="47">Eurocaucasian</location>
|
||
, submeridional and temperate.
|
||
<collectorName id="306E5376FD1DFFE2FEB0F9905B27F97E" box="[277,362,1655,1679]" pageId="32" pageNumber="47">From S</collectorName>
|
||
<collectingCountry id="E58C7630FD1DFFE2FED3F99F5B9DF97E" box="[374,464,1655,1679]" name="United Kingdom" pageId="32" pageNumber="47">England</collectingCountry>
|
||
and Iberia across
|
||
<location id="9844607BFD1DFFE2FD0EF99F5AA5F941" LSID="urn:lsid:plazi:treatment:153287B6FD1DFFE2FF0BFED95989FC6A:9844607BFD1DFFE2FD0EF99F5AA5F941" country="United Kingdom" county="Caucasus. In Scandinavia" latitude="60.0" municipality="Balkans" name="Central Europe" pageId="32" pageNumber="47">Central Europe</location>
|
||
, the
|
||
<location id="9844607BFD1DFFE2FEBDF9705BCFF941" LSID="urn:lsid:plazi:treatment:153287B6FD1DFFE2FF0BFED95989FC6A:9844607BFD1DFFE2FEBDF9705BCFF941" box="[280,386,1688,1712]" country="United Kingdom" county="Caucasus. In Scandinavia" latitude="60.0" municipality="Balkans" name="Apennine" pageId="32" pageNumber="47">Apennine</location>
|
||
and the
|
||
<collectingMunicipality id="7D40ACDAFD1DFFE2FE79F9705879F941" box="[476,564,1688,1712]" pageId="32" pageNumber="47">Balkans</collectingMunicipality>
|
||
to
|
||
<collectingCountry id="E58C7630FD1DFFE2FDF1F9705882F941" box="[596,719,1688,1712]" name="Turkey" pageId="32" pageNumber="47">Asia Minor</collectingCountry>
|
||
and the
|
||
<collectingCounty id="74454E2CFD1DFFE2FF1AF9515B9EF920" box="[191,467,1721,1745]" pageId="32" pageNumber="47">Caucasus. In Scandinavia</collectingCounty>
|
||
north to
|
||
<geoCoordinate id="F8AF5067FD1DFFE2FD90F951583DF920" box="[565,624,1721,1745]" degrees="60" direction="north" orientation="latitude" pageId="32" pageNumber="47" precision="55555" value="60.0">60°N</geoCoordinate>
|
||
, absent from
|
||
<collectingCountry id="E58C7630FD1DFFE2FF32F9325AA0F903" box="[151,237,1754,1778]" name="Finland" pageId="32" pageNumber="47">Finland</collectingCountry>
|
||
.
|
||
<collectorName id="306E5376FD1DFFE2FF5DF9325B24F903" box="[248,361,1754,1778]" pageId="32" pageNumber="47">In N Tyrol</collectorName>
|
||
ascending to
|
||
<quantity id="5A639B45FD1DFFE2FDA5F9325800F900" box="[512,589,1754,1778]" metricMagnitude="3" metricUnit="m" metricValue="1.41" pageId="32" pageNumber="47" unit="m" value="1410.0">
|
||
<elevation id="16B6D193FD1DFFE2FDA5F9325800F900" box="[512,589,1754,1778]" metricMagnitude="3" metricUnit="m" metricValue="1.41" pageId="32" pageNumber="47" unit="m" value="1410.0">1410 m</elevation>
|
||
</quantity>
|
||
and in
|
||
<collectingCountry id="E58C7630FD1DFFE2FD3BF93258B6F903" box="[670,763,1754,1778]" name="Turkey" pageId="32" pageNumber="47">Anatolia</collectingCountry>
|
||
at
|
||
<geoCoordinate id="F8AF5067FD1DFFE2FF14F9135AA4F8E2" box="[177,233,1787,1811]" degrees="37" direction="north" orientation="latitude" pageId="32" pageNumber="47" precision="55555" value="37.0">37°N</geoCoordinate>
|
||
to
|
||
<quantity id="5A639B45FD1DFFE2FEAEF9135B2CF8E3" box="[267,353,1787,1811]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="32" pageNumber="47" unit="m" value="2000.0">
|
||
<elevation id="16B6D193FD1DFFE2FEAEF9135B2CF8E3" box="[267,353,1787,1811]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="32" pageNumber="47" unit="m" value="2000.0">2000 m</elevation>
|
||
</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D581652BFD1DFFE2FF0BF8F35F43FCE9" pageId="32" pageNumber="47" type="diagnosis">
|
||
<paragraph id="9D2436A0FD1DFFE2FF0BF8F358B8F867" blockId="32.[151,766,340,1975]" pageId="32" pageNumber="47">
|
||
<emphasis id="AFEFEAB2FD1DFFE2FF0BF8F35B50F8C2" bold="true" box="[174,285,1819,1843]" pageId="32" pageNumber="47">Diagnosis</emphasis>
|
||
(
|
||
<tableCitation id="D019031BFD1DFFE2FE88F8F45B3FF8C5" box="[301,370,1820,1844]" captionStart="Tab" captionStartId="69.[193,241,376,402]" captionTargetBox="[193,1313,480,1696]" captionTargetId="graphics-66@69.[188,1328,471,1701]" captionTargetPageId="69" captionText="Tab. 1*: Members of the Lasius brunneus species complex (nOcc+nGu+nSt <10). Data are given as arithmetic mean ± standard deviation [lower extreme, upper extreme] number of individuals." pageId="32" pageNumber="47">Tab. 1</tableCitation>
|
||
,
|
||
<figureCitation id="05A02A25FD1DFFE2FE25F8F45B88F8C5" box="[384,453,1820,1844]" captionStart="Figs" captionStartId="14.[151,193,1864,1886]" captionTargetBox="[151,1399,1329,1838]" captionTargetId="figure-490@14.[677,1402,1329,1705]" captionTargetPageId="14" captionText="Figs. 1 and 2: Lasius brunneus; neotype" figureDoi="http://doi.org/10.5281/zenodo.10832218" httpUri="https://zenodo.org/record/10832218/files/figure.png" pageId="32" pageNumber="47">Figs. 1</figureCitation>
|
||
–2; images in www.antWeb. org with specimen identifiers CASENT0172717, CASENT0172745, CASENT0172746, CASENT0179886, CASENT0179917, CASENT0179917, CASENT0911044:
|
||
</paragraph>
|
||
<paragraph id="9D2436A0FD1DFFE2FF0BF8775F43FCE9" blockId="32.[151,766,340,1975]" lastBlockId="32.[786,1399,241,923]" pageId="32" pageNumber="47">
|
||
The species is rather easily identified by a combination of short scape (SL/CS
|
||
<subScript id="011F34E5FD1DFFE2FBAFFEEA5E6FFEE1" attach="left" box="[1034,1058,258,272]" fontSize="6" pageId="32" pageNumber="47">900</subScript>
|
||
0.874), broad head (CL/CW
|
||
<subScript id="011F34E5FD1DFFE2FAFBFEEA5F3BFEE1" attach="left" box="[1374,1398,258,272]" fontSize="6" pageId="32" pageNumber="47">900</subScript>
|
||
1.041), reduced setae numbers on all body parts (nOcc
|
||
<subScript id="011F34E5FD1DFFE2FAFBFECA5F3BFEC1" attach="left" box="[1374,1398,290,304]" fontSize="6" pageId="32" pageNumber="47">900</subScript>
|
||
2.0, nGen
|
||
<subScript id="011F34E5FD1DFFE2FCDEFEAB59DEFEA0" attach="left" box="[891,915,323,337]" fontSize="6" pageId="32" pageNumber="47">900</subScript>
|
||
0.1, nGu
|
||
<subScript id="011F34E5FD1DFFE2FC50FEAB5E40FEA0" attach="left" box="[1013,1037,323,337]" fontSize="6" pageId="32" pageNumber="47">900</subScript>
|
||
1.3, nSC
|
||
<subScript id="011F34E5FD1DFFE2FBD5FEAB5EC5FEA0" attach="left" box="[1136,1160,323,337]" fontSize="6" pageId="32" pageNumber="47">900</subScript>
|
||
0.0, nHT
|
||
<subScript id="011F34E5FD1DFFE2FB4AFEAB5F4AFEA0" attach="left" box="[1263,1287,323,337]" fontSize="6" pageId="32" pageNumber="47">900</subScript>
|
||
0.1, nSt
|
||
<subScript id="011F34E5FD1DFFE2FAFBFEAB5F3BFEA0" attach="left" box="[1374,1398,323,337]" fontSize="6" pageId="32" pageNumber="47">900</subScript>
|
||
1.1) a very smooth pubescence surface on scape and tibiae, and reduced number of mandibular dents (MaDe
|
||
<subScript id="011F34E5FD1DFFE2FAFBFE6D5F3BFE62" attach="left" box="[1374,1398,389,403]" fontSize="6" pageId="32" pageNumber="47">900</subScript>
|
||
7.06). The dorsal and posterior profile of propodeum are linear and form a distinct, obtuse angle. The petiole is in anterocaudal view rather rectangular or slightly converging dorsad but typically with straight sides and forming a sharp, weakly emarginate dorsal crest. A typical coloration of medium-sized to large specimens is mesosoma, petiole and appendages light yellowish brown, head a little darker bronze brown and gaster dark to blackish brown. However, small workers from initial colonies may show a homogenously dark coloration as seen in
|
||
<taxonomicName id="5A9B4D23FD1DFFE2FCCAFD3759BCFD06" baseAuthorityName="Emery" baseAuthorityYear="1869" box="[879,1009,735,759]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="47" phylum="Arthropoda" rank="species" species="lasioides">
|
||
<emphasis id="AFEFEAB2FD1DFFE2FCCAFD3759BCFD06" box="[879,1009,735,759]" italics="true" pageId="32" pageNumber="47">L. lasioides</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="5A9B4D23FD1DFFE2FBBAFD375EEAFD06" box="[1055,1191,735,759]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="47" phylum="Arthropoda" rank="species" species="neglectus">
|
||
<emphasis id="AFEFEAB2FD1DFFE2FBBAFD375EEAFD06" box="[1055,1191,735,759]" italics="true" pageId="32" pageNumber="47">L. neglectus</emphasis>
|
||
</taxonomicName>
|
||
but can easily be separated by RAV-corrected NUMOBAT data.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D581652BFD1DFFE2FC8CFCC85E1CFCC8" box="[809,1105,800,825]" pageId="32" pageNumber="47" type="biology_ecology">
|
||
<paragraph id="9D2436A0FD1DFFE2FC8CFCC85E1CFCC8" blockId="32.[786,1399,241,923]" box="[809,1105,800,825]" pageId="32" pageNumber="47">
|
||
<emphasis id="AFEFEAB2FD1DFFE2FC8CFCC85933FCC9" bold="true" box="[809,894,800,824]" pageId="32" pageNumber="47">Biology</emphasis>
|
||
. See Seifert (2018).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D581652BFD1DFFE2FC8CFCA95989FC6A" pageId="32" pageNumber="47" type="discussion">
|
||
<paragraph id="9D2436A0FD1DFFE2FC8CFCA95989FC6A" blockId="32.[786,1399,241,923]" pageId="32" pageNumber="47">
|
||
<emphasis id="AFEFEAB2FD1DFFE2FC8CFCA959E9FCA8" bold="true" box="[809,932,833,857]" pageId="32" pageNumber="47">Comments</emphasis>
|
||
. For separation from the eastern sister species
|
||
<taxonomicName id="5A9B4D23FD1DFFE2FCC8FC8B59BBFC8B" authority="Seifert, 2020" authorityName="Seifert" authorityYear="2020" box="[877,1014,866,890]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="47" phylum="Arthropoda" rank="species" species="silvaticus" status="sp. nov.">
|
||
<emphasis id="AFEFEAB2FD1DFFE2FCC8FC8B59BBFC8B" box="[877,1014,866,890]" italics="true" pageId="32" pageNumber="47">L. silvaticus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B4DC57C9FD1DFFE2FBA6FC8B5E16FC8B" box="[1027,1115,867,890]" pageId="32" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="5A9B4D23FD1DFFE2FB3FFC8B5F0AFC8B" authorityName="Bingham" authorityYear="1903" box="[1178,1351,866,890]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="47" phylum="Arthropoda" rank="species" species="himalayanus">
|
||
<emphasis id="AFEFEAB2FD1DFFE2FB3FFC8B5F0AFC8B" box="[1178,1351,866,890]" italics="true" pageId="32" pageNumber="47">L. himalayanus</emphasis>
|
||
</taxonomicName>
|
||
see there and
|
||
<tableCitation id="D019031BFD1DFFE2FCD8FC6B5939FBF1" captionStart-0="Tab" captionStart-1="Tab" captionStart-2="Tab" captionStart-3="Tab" captionStartId-0="69.[193,241,376,402]" captionStartId-1="70.[96,144,374,400]" captionStartId-2="71.[192,240,392,418]" captionStartId-3="72.[224,272,393,419]" captionTargetBox-0="[193,1313,480,1696]" captionTargetBox-1="[96,1387,445,1688]" captionTargetBox-2="[198,1387,463,1680]" captionTargetBox-3="[228,1378,465,1682]" captionTargetId-0="graphics-66@69.[188,1328,471,1701]" captionTargetId-1="graphics-66@70.[92,1400,436,1692]" captionTargetId-2="graphics-49@71.[188,1404,454,1684]" captionTargetId-3="graphics-56@72.[220,1399,456,1686]" captionTargetPageId-0="69" captionTargetPageId-1="70" captionTargetPageId-2="71" captionTargetPageId-3="72" captionText-0="Tab. 1*: Members of the Lasius brunneus species complex (nOcc+nGu+nSt <10). Data are given as arithmetic mean ± standard deviation [lower extreme, upper extreme] number of individuals." captionText-1="Tab. 2: Members of the Lasius turcicus species complex (nOcc+nGu+nSt> 10)." captionText-2="Tab. 3: The Lasius obscuratus species complex & L. brevipalpus n.sp." captionText-3="Tab. 4: The Lasius paralienus species complex" pageId="32" pageNumber="47">
|
||
Tab. 1.
|
||
<subSubSection id="D581652BFD1DFFE2FC8CFC0C5E93FBF1" box="[809,1246,995,1024]" pageId="32" pageNumber="47" type="nomenclature">
|
||
<paragraph id="9D2436A0FD1DFFE2FC8CFC0C5E93FBF1" blockId="32.[809,1246,995,1024]" box="[809,1246,995,1024]" pageId="32" pageNumber="47">
|
||
<heading id="C66C81CCFD1DFFE2FC8CFC0C5E93FBF1" bold="true" box="[809,1246,995,1024]" fontSize="10" level="2" pageId="32" pageNumber="47" reason="3">
|
||
<emphasis id="AFEFEAB2FD1DFFE2FC8CFC0C5E93FBF1" bold="true" box="[809,1246,995,1024]" pageId="32" pageNumber="47">
|
||
4.4.2
|
||
<taxonomicName id="5A9B4D23FD1DFFE2FCD8FC0C5E25FC0F" authority="Seifert, 2020" authorityName="Seifert" authorityYear="2020" box="[893,1128,995,1022]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="47" phylum="Arthropoda" rank="species" species="silvaticus" status="sp. nov.">
|
||
<emphasis id="AFEFEAB2FD1DFFE2FCD8FC0C5E25FC0F" bold="true" box="[893,1128,995,1022]" italics="true" pageId="32" pageNumber="47">Lasius silvaticus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B4DC57C9FD1DFFE2FBD4FC0C5E93FBF1" box="[1137,1246,996,1024]" pageId="32" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D581652BFD1DFFE2FC8CFBEF592EFBCE" pageId="32" pageNumber="47" type="etymology">
|
||
<paragraph id="9D2436A0FD1DFFE2FC8CFBEF592EFBCE" blockId="32.[786,1400,1031,1981]" pageId="32" pageNumber="47">
|
||
<emphasis id="AFEFEAB2FD1DFFE2FC8CFBEF59E9FBEE" bold="true" box="[809,932,1031,1055]" pageId="32" pageNumber="47">Etymology</emphasis>
|
||
. The species name refers to the woodland habitat.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D581652BFD1DFFE2FC8CFBA05E17F920" pageId="32" pageNumber="47" type="materials_examined">
|
||
<paragraph id="9D2436A0FD1DFFE2FC8CFBA05F12FA1A" blockId="32.[786,1400,1031,1981]" pageId="32" pageNumber="47">
|
||
<emphasis id="AFEFEAB2FD1DFFE2FC8CFBA0599FFB91" bold="true" box="[809,978,1096,1120]" pageId="32" pageNumber="47">Type material</emphasis>
|
||
.
|
||
<typeStatus id="42208802FD1DFFE2FC4DFBA05E01FB91" box="[1000,1100,1096,1120]" pageId="32" pageNumber="47" type="holotype">Holotype</typeStatus>
|
||
plus
|
||
<specimenCount id="8B9DFD29FD1DFFE2FB33FBA05F3BFB91" box="[1174,1398,1096,1121]" count="2" pageId="32" pageNumber="47" type="worker" typeStatus="paratype">2 paratype workers</specimenCount>
|
||
labelled ”
|
||
<collectingCountry id="E58C7630FD1DFFE2FCDBFB81598AFB70" box="[894,967,1129,1153]" name="Iran" pageId="32" pageNumber="47">IRAN</collectingCountry>
|
||
:
|
||
<geoCoordinate id="F8AF5067FD1DFFE2FC73FB815E05FB70" box="[982,1096,1129,1153]" degrees="36.7198" direction="north" orientation="latitude" pageId="32" pageNumber="47" precision="5" value="36.7198">36.7198°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="F8AF5067FD1DFFE2FBFCFB815E84FB70" box="[1113,1225,1129,1153]" degrees="54.5813" direction="east" orientation="longitude" pageId="32" pageNumber="47" precision="5" value="54.5813">54.5813°E</geoCoordinate>
|
||
, Gorgan 19SE,
|
||
<quantity id="5A639B45FD1DFFE2FCB7FB62591AFB53" box="[786,855,1162,1186]" metricMagnitude="2" metricUnit="m" metricValue="8.53" pageId="32" pageNumber="47" unit="m" value="853.0">
|
||
<elevation id="16B6D193FD1DFFE2FCB7FB62591AFB53" box="[786,855,1162,1186]" metricMagnitude="2" metricUnit="m" metricValue="8.53" pageId="32" pageNumber="47" unit="m" value="853.0">853 m</elevation>
|
||
</quantity>
|
||
, Caspian mild and wet,
|
||
<collectorName id="306E5376FD1DFFE2FBF9FB625E99FB53" box="[1116,1236,1162,1186]" pageId="32" pageNumber="47">A.B. Yazdi</collectorName>
|
||
<date id="E9251060FD1DFFE2FB7EFB625F00FB53" box="[1243,1357,1162,1186]" pageId="32" pageNumber="47" value="2016-06-15">
|
||
<collectingDate id="F961E988FD1DFFE2FB7EFB625F00FB53" box="[1243,1357,1162,1186]" pageId="32" pageNumber="47" value="2016-06-15">2016.06.15</collectingDate>
|
||
</date>
|
||
-9“ and ”
|
||
<typeStatus id="42208802FD1DFFE2FCEEFB4359FFFB32" box="[843,946,1195,1219]" pageId="32" pageNumber="47" type="holotype">Holotype</typeStatus>
|
||
(bottom) and
|
||
<typeStatus id="42208802FD1DFFE2FBEAFB445EF4FB35" box="[1103,1209,1196,1220]" pageId="32" pageNumber="47" type="paratype">paratypes</typeStatus>
|
||
<taxonomicName id="5A9B4D23FD1DFFE2FB64FB44592DFB15" authority="Seifert" authorityName="Seifert" authorityYear="2020" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="47" phylum="Arthropoda" rank="species" species="silvaticus">
|
||
<emphasis id="AFEFEAB2FD1DFFE2FB64FB445F3AFB32" box="[1217,1399,1195,1219]" italics="true" pageId="32" pageNumber="47">Lasius silvaticus</emphasis>
|
||
Seifert
|
||
</taxonomicName>
|
||
“;
|
||
<specimenCount id="8B9DFD29FD1DFFE2FC23FB245E34FB15" box="[902,1145,1228,1252]" count="3" pageId="32" pageNumber="47" type="worker" typeStatus="paratype">3 paratype workers</specimenCount>
|
||
”
|
||
<collectingCountry id="E58C7630FD1DFFE2FB3EFB245EA9FB15" box="[1179,1252,1228,1252]" name="Iran" pageId="32" pageNumber="47">IRAN</collectingCountry>
|
||
:
|
||
<geoCoordinate id="F8AF5067FD1DFFE2FAA5FB245F3FFB15" box="[1280,1394,1228,1252]" degrees="36.7198" direction="north" orientation="latitude" pageId="32" pageNumber="47" precision="5" value="36.7198">36.7198°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="F8AF5067FD1DFFE2FCB7FB0559CEFAF4" box="[786,899,1261,1285]" degrees="54.5813" direction="east" orientation="longitude" pageId="32" pageNumber="47" precision="5" value="54.5813">54.5813°E</geoCoordinate>
|
||
, Gorgan 19SE,
|
||
<quantity id="5A639B45FD1DFFE2FBE3FB055EDEFAF5" box="[1094,1171,1261,1285]" metricMagnitude="2" metricUnit="m" metricValue="8.53" pageId="32" pageNumber="47" unit="m" value="853.0">
|
||
<elevation id="16B6D193FD1DFFE2FBE3FB055EDEFAF5" box="[1094,1171,1261,1285]" metricMagnitude="2" metricUnit="m" metricValue="8.53" pageId="32" pageNumber="47" unit="m" value="853.0">853 m</elevation>
|
||
</quantity>
|
||
, Caspian mild and wet,
|
||
<collectorName id="306E5376FD1DFFE2FCECFAE6598FFAD7" box="[841,962,1294,1318]" pageId="32" pageNumber="47">A.B. Yazdi</collectorName>
|
||
<date id="E9251060FD1DFFE2FC69FAE55E73FAD4" box="[972,1086,1293,1317]" pageId="32" pageNumber="47" value="2016-06-15">
|
||
<collectingDate id="F961E988FD1DFFE2FC69FAE55E73FAD4" box="[972,1086,1293,1317]" pageId="32" pageNumber="47" value="2016-06-15">2016.06.15</collectingDate>
|
||
</date>
|
||
-24“ and ”
|
||
<typeStatus id="42208802FD1DFFE2FB1CFAE65F6BFAD7" box="[1209,1318,1294,1318]" pageId="32" pageNumber="47" type="paratype">Paratypes</typeStatus>
|
||
<taxonomicName id="5A9B4D23FD1DFFE2FA95FAE65986FAB7" authority="Seifert" authorityName="Seifert" authorityYear="2020" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="47" phylum="Arthropoda" rank="species" species="silvaticus">Lasius silvaticus Seifert</taxonomicName>
|
||
“;
|
||
<specimenCount id="8B9DFD29FD1DFFE2FC7BFAC75EE6FAB7" box="[990,1195,1326,1351]" count="3" pageId="32" pageNumber="47" type="worker" typeStatus="paratype">3 paratype workers</specimenCount>
|
||
”
|
||
<collectingCountry id="E58C7630FD1DFFE2FB1EFAC75F49FAB6" box="[1211,1284,1327,1351]" name="Iran" pageId="32" pageNumber="47">IRAN</collectingCountry>
|
||
:
|
||
<geoCoordinate id="F8AF5067FD1DFFE2FAA9FAC65F3EFAB7" box="[1292,1395,1326,1350]" degrees="37.367" direction="north" orientation="latitude" pageId="32" pageNumber="47" precision="55" value="37.367">37.367°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="F8AF5067FD1DFFE2FCB7FAA75939FA96" box="[786,884,1359,1383]" degrees="55.817" direction="east" orientation="longitude" pageId="32" pageNumber="47" precision="55" value="55.817">55.817°E</geoCoordinate>
|
||
,
|
||
<collectingRegion id="5F5FF842FD1DFFE2FC27FAA759AFFA96" box="[898,994,1359,1383]" country="Iran" name="Golestan" pageId="32" pageNumber="47">Golestan</collectingRegion>
|
||
NP,
|
||
<collectingCounty id="74454E2CFD1DFFE2FBB8FAA75E39FA96" box="[1053,1140,1359,1383]" pageId="32" pageNumber="47">Galesha</collectingCounty>
|
||
canyon,
|
||
<quantity id="5A639B45FD1DFFE2FB7EFAA75F69FA96" box="[1243,1316,1359,1383]" metricMagnitude="2" metricUnit="m" metricValue="5.9399999999999995" pageId="32" pageNumber="47" unit="m" value="594.0">
|
||
<elevation id="16B6D193FD1DFFE2FB7EFAA75F69FA96" box="[1243,1316,1359,1383]" metricMagnitude="2" metricUnit="m" metricValue="5.9399999999999995" pageId="32" pageNumber="47" unit="m" value="594.0">594 m</elevation>
|
||
</quantity>
|
||
, forest, nest on tree,
|
||
<collectingMunicipality id="7D40ACDAFD1DFFE2FC1FFA985E49FA79" box="[954,1028,1392,1416]" pageId="32" pageNumber="47">Paknia</collectingMunicipality>
|
||
<date id="E9251060FD1DFFE2FBB1FA985EC1FA79" box="[1044,1164,1392,1416]" pageId="32" pageNumber="47" value="2007-05-23">
|
||
<collectingDate id="F961E988FD1DFFE2FBB1FA985EC1FA79" box="[1044,1164,1392,1416]" pageId="32" pageNumber="47" value="2007-05-23">2007.05.23</collectingDate>
|
||
</date>
|
||
- 1420“;
|
||
<specimenCount id="8B9DFD29FD1DFFE2FB5FFA985925FA58" count="3" pageId="32" pageNumber="47" type="worker" typeStatus="paratype">3 paratype workers</specimenCount>
|
||
”
|
||
<collectingCountry id="E58C7630FD1DFFE2FC23FA795982FA58" box="[902,975,1425,1449]" name="Iran" pageId="32" pageNumber="47">IRAN</collectingCountry>
|
||
:
|
||
<geoCoordinate id="F8AF5067FD1DFFE2FC40FA795E01FA58" box="[997,1100,1425,1449]" degrees="37.367" direction="north" orientation="latitude" pageId="32" pageNumber="47" precision="55" value="37.367">37.367°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="F8AF5067FD1DFFE2FBC6FA795E89FA58" box="[1123,1220,1425,1449]" degrees="55.817" direction="east" orientation="longitude" pageId="32" pageNumber="47" precision="55" value="55.817">55.817°E</geoCoordinate>
|
||
,
|
||
<collectingRegion id="5F5FF842FD1DFFE2FB79FA795F71FA58" box="[1244,1340,1425,1449]" country="Iran" name="Golestan" pageId="32" pageNumber="47">Golestan</collectingRegion>
|
||
NP,
|
||
<location id="9844607BFD1DFFE2FCB7FA5A5924FA3B" LSID="urn:lsid:plazi:treatment:153287B6FD1DFFE2FF0BFED95989FC6A:9844607BFD1DFFE2FCB7FA5A5924FA3B" box="[786,873,1458,1482]" country="Iran" county="Galesha" latitude="36.7198" longLatPrecision="6" longitude="54.5813" municipality="Paknia" name="Galesha" pageId="32" pageNumber="47" stateProvince="Golestan">Galesha</location>
|
||
canyon,
|
||
<quantity id="5A639B45FD1DFFE2FC68FA5A5E5EFA3B" box="[973,1043,1458,1482]" metricMagnitude="2" metricUnit="m" metricValue="5.9399999999999995" pageId="32" pageNumber="47" unit="m" value="594.0">
|
||
<elevation id="16B6D193FD1DFFE2FC68FA5A5E5EFA3B" box="[973,1043,1458,1482]" metricMagnitude="2" metricUnit="m" metricValue="5.9399999999999995" pageId="32" pageNumber="47" unit="m" value="594.0">594 m</elevation>
|
||
</quantity>
|
||
, forest, nest under stone,
|
||
<location id="9844607BFD1DFFE2FA89FA5A5F3BFA3B" LSID="urn:lsid:plazi:treatment:153287B6FD1DFFE2FF0BFED95989FC6A:9844607BFD1DFFE2FA89FA5A5F3BFA3B" box="[1324,1398,1458,1482]" country="Iran" county="Galesha" latitude="36.7198" longLatPrecision="6" longitude="54.5813" municipality="Paknia" name="Paknia" pageId="32" pageNumber="47" stateProvince="Golestan">Paknia</location>
|
||
<date id="E9251060FD1DFFE2FCB7FA3B59C7FA1A" box="[786,906,1491,1515]" pageId="32" pageNumber="47" value="2007-05-23">
|
||
<collectingDate id="F961E988FD1DFFE2FCB7FA3B59C7FA1A" box="[786,906,1491,1515]" pageId="32" pageNumber="47" value="2007-05-23">2007.05.23</collectingDate>
|
||
</date>
|
||
- 1427“; all material stored in
|
||
<collectionCode id="FB8AAE65FD1DFFE2FB6FFA3B5F48FA1A" box="[1226,1285,1491,1515]" pageId="32" pageNumber="47">SMN</collectionCode>
|
||
Görlitz.
|
||
</paragraph>
|
||
<paragraph id="9D2436A0FD1DFFE2FC8CFA1C5E24F99E" blockId="32.[786,1400,1031,1981]" pageId="32" pageNumber="47">
|
||
<emphasis id="AFEFEAB2FD1DFFE2FC8CFA1C5E75F9FD" bold="true" box="[809,1080,1524,1548]" pageId="32" pageNumber="47">All material examined</emphasis>
|
||
. A total of 9 nest samples with
|
||
<specimenCount id="8B9DFD29FD1DFFE2FCE9F9FD598AF9DC" box="[844,967,1557,1581]" count="24" pageId="32" pageNumber="47" type="worker">24 workers</specimenCount>
|
||
from seven localities in the
|
||
<collectingCountry id="E58C7630FD1DFFE2FAA9F9FD5F74F9DD" box="[1292,1337,1557,1580]" name="Iran" pageId="32" pageNumber="47">Iran</collectingCountry>
|
||
were subject to NUMOBAT investigation.
|
||
<location id="9844607BFD1DFFE2FB6CF9DE5EA0F9BC" LSID="urn:lsid:plazi:treatment:153287B6FD1DFFE2FF0BFED95989FC6A:9844607BFD1DFFE2FB6CF9DE5EA0F9BC" box="[1225,1261,1590,1613]" country="Iran" name="For" pageId="32" pageNumber="47">For</location>
|
||
details see supplementary information SI1.
|
||
</paragraph>
|
||
<paragraph id="9D2436A0FD1DFFE2FC8CF99F5E17F920" blockId="32.[786,1400,1031,1981]" pageId="32" pageNumber="47">
|
||
<emphasis id="AFEFEAB2FD1DFFE2FC8CF99F59BAF97E" bold="true" box="[809,1015,1655,1679]" pageId="32" pageNumber="47">Geographic range</emphasis>
|
||
. S Caspian; known so far only from a rather small area between 36.7 and
|
||
<geoCoordinate id="F8AF5067FD1DFFE2FB10F9705F47F941" box="[1205,1290,1688,1712]" degrees="37.4" direction="north" orientation="latitude" pageId="32" pageNumber="47" precision="5555" value="37.4">37.4° N</geoCoordinate>
|
||
, 54.4 and
|
||
<geoCoordinate id="F8AF5067FD1DFFE2FCB7F9515911F920" box="[786,860,1721,1745]" degrees="55.8" direction="east" orientation="longitude" pageId="32" pageNumber="47" precision="5555" value="55.8">55.8°E</geoCoordinate>
|
||
, and
|
||
<quantity id="5A639B45FD1DFFE2FC30F9515E6FF920" box="[917,1058,1721,1745]" metricMagnitude="2" metricUnit="m" metricValue="5.3" metricValueMax="9.0" metricValueMin="1.6" pageId="32" pageNumber="47" unit="m" value="530.0" valueMax="900.0" valueMin="160.0">160 to 900 m</quantity>
|
||
a.s.l.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D581652BFD1DFFE3FC8CF93258AEFEBA" lastPageId="33" lastPageNumber="48" pageId="32" pageNumber="47" type="diagnosis">
|
||
<paragraph id="9D2436A0FD1DFFE3FC8CF93258AEFEBA" blockId="32.[786,1400,1031,1981]" lastBlockId="33.[189,803,241,1384]" lastPageId="33" lastPageNumber="48" pageId="32" pageNumber="47">
|
||
<emphasis id="AFEFEAB2FD1DFFE2FC8CF93259D5F903" bold="true" box="[809,920,1754,1778]" pageId="32" pageNumber="47">Diagnosis</emphasis>
|
||
(
|
||
<tableCitation id="D019031BFD1DFFE2FC08F93259BAF903" box="[941,1015,1754,1778]" captionStart="Tab" captionStartId="69.[193,241,376,402]" captionTargetBox="[193,1313,480,1696]" captionTargetId="graphics-66@69.[188,1328,471,1701]" captionTargetPageId="69" captionText="Tab. 1*: Members of the Lasius brunneus species complex (nOcc+nGu+nSt <10). Data are given as arithmetic mean ± standard deviation [lower extreme, upper extreme] number of individuals." pageId="32" pageNumber="47">Tab. 1</tableCitation>
|
||
,
|
||
<figureCitation id="05A02A25FD1DFFE2FBAFF9325E15F903" box="[1034,1112,1754,1778]" captionStart="Figs" captionStartId="15.[189,229,644,666]" captionTargetBox="[189,1359,227,628]" captionTargetId="figure-13@15.[756,1361,224,632]" captionTargetPageId="15" captionText="Figs. 3 and 4: Lasius silvaticus sp. nov.; holotype" figureDoi="http://doi.org/10.5281/zenodo.10832222" httpUri="https://zenodo.org/record/10832222/files/figure.png" pageId="32" pageNumber="47">Figs. 3</figureCitation>
|
||
–4; key; image in www. antWeb.org with specimen identifier CFH000052): Showing all diagnostic characters of the
|
||
<taxonomicName id="5A9B4D23FD1DFFE2FB4BF8F45F3AF8C5" baseAuthorityName="Latreille" baseAuthorityYear="1798" box="[1262,1399,1820,1844]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="47" phylum="Arthropoda" rank="species" species="brunneus">
|
||
<emphasis id="AFEFEAB2FD1DFFE2FB4BF8F45F3AF8C5" box="[1262,1399,1820,1844]" italics="true" pageId="32" pageNumber="47">L. brunneus</emphasis>
|
||
</taxonomicName>
|
||
species complex. The main difference to
|
||
<taxonomicName id="5A9B4D23FD1DFFE2FB72F8D55F11F8A4" baseAuthorityName="Latreille" baseAuthorityYear="1798" box="[1239,1372,1853,1877]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="47" phylum="Arthropoda" rank="species" species="brunneus">
|
||
<emphasis id="AFEFEAB2FD1DFFE2FB72F8D55F11F8A4" box="[1239,1372,1853,1877]" italics="true" pageId="32" pageNumber="47">L. brunneus</emphasis>
|
||
</taxonomicName>
|
||
is the less sparse and longer pilosity on pronotum (PnHL/ CS
|
||
<subScript id="011F34E5FD1DFFE2FC97F8665907F86D" attach="left" box="[818,842,1934,1948]" fontSize="6" pageId="32" pageNumber="47">900</subScript>
|
||
0.130), underside of head (nGu
|
||
<subScript id="011F34E5FD1DFFE2FB07F8665EF7F86D" attach="left" box="[1186,1210,1934,1948]" fontSize="6" pageId="32" pageNumber="47">900</subScript>
|
||
3.6, GuHL/CS
|
||
<subScript id="011F34E5FD1DFFE2FAFBF8665F3BF86D" attach="left" box="[1374,1398,1934,1948]" fontSize="6" pageId="32" pageNumber="47">900</subScript>
|
||
0.125), propodeum and hind tibia (nHT
|
||
<subScript id="011F34E5FD1DFFE2FB1EF8475E9EF84C" attach="left" box="[1211,1235,1967,1981]" fontSize="6" pageId="32" pageNumber="47">900</subScript>
|
||
1.6, nSt
|
||
<subScript id="011F34E5FD1DFFE2FA8EF8475F0EF84C" attach="left" box="[1323,1347,1967,1981]" fontSize="6" pageId="32" pageNumber="47">900</subScript>
|
||
2.9). Accessory differences are the longer head and smaller postocular distance (CL/CW
|
||
<subScript id="011F34E5FD1CFFE3FE56FECA5846FEC1" attach="left" box="[499,523,290,304]" fontSize="6" pageId="33" pageNumber="48">900</subScript>
|
||
1.059, PoOc/CS
|
||
<subScript id="011F34E5FD1CFFE3FD1EFECA589EFEC1" attach="left" box="[699,723,290,304]" fontSize="6" pageId="33" pageNumber="48">900</subScript>
|
||
0.235). The coloration is fully comparable to
|
||
<taxonomicName id="5A9B4D23FD1CFFE3FDFFFEDC5891FEBA" baseAuthorityName="Latreille" baseAuthorityYear="1798" box="[602,732,307,331]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="48" phylum="Arthropoda" rank="species" species="brunneus">
|
||
<emphasis id="AFEFEAB2FD1CFFE3FDFFFEDC5891FEBA" box="[602,732,307,331]" italics="true" pageId="33" pageNumber="48">L. brunneus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D581652BFD1CFFE3FF71FEBC5BDBFDE1" pageId="33" pageNumber="48" type="biology_ecology">
|
||
<paragraph id="9D2436A0FD1CFFE3FF71FEBC5BDBFDE1" blockId="33.[189,803,241,1384]" pageId="33" pageNumber="48">
|
||
<emphasis id="AFEFEAB2FD1CFFE3FF71FEBC5B64FE9D" bold="true" box="[212,297,340,364]" pageId="33" pageNumber="48">Biology</emphasis>
|
||
. Habitat selection appears to be similar to its sibling species
|
||
<taxonomicName id="5A9B4D23FD1CFFE3FECFFE9E586CFE7C" baseAuthorityName="Latreille" baseAuthorityYear="1798" box="[362,545,373,397]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="48" phylum="Arthropoda" rank="species" species="brunneus">
|
||
<emphasis id="AFEFEAB2FD1CFFE3FECFFE9E586CFE7C" box="[362,545,373,397]" italics="true" pageId="33" pageNumber="48">Lasius brunneus</emphasis>
|
||
</taxonomicName>
|
||
: Seven samples were found in wet and humid Caspian broad-leafed forest and one sample in a city park with trees. The nests were both in the wood of trees, under stones, in litter and (in a clearcutting) in soil.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D581652BFD1CFFE3FF71FDF1595BFA96" pageId="33" pageNumber="48" type="discussion">
|
||
<paragraph id="9D2436A0FD1CFFE3FF71FDF1595BFA96" blockId="33.[189,803,241,1384]" pageId="33" pageNumber="48">
|
||
<emphasis id="AFEFEAB2FD1CFFE3FF71FDF15B02FDC0" bold="true" box="[212,335,537,561]" pageId="33" pageNumber="48">Comments</emphasis>
|
||
.
|
||
<emphasis id="AFEFEAB2FD1CFFE3FEFBFDF2583AFDC0" box="[350,631,537,561]" italics="true" pageId="33" pageNumber="48">
|
||
<taxonomicName id="5A9B4D23FD1CFFE3FEFBFDF25BA8FDC0" baseAuthorityName="Latreille" baseAuthorityYear="1798" box="[350,485,537,561]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="48" phylum="Arthropoda" rank="species" species="brunneus">L. brunneus</taxonomicName>
|
||
,
|
||
<taxonomicName id="5A9B4D23FD1CFFE3FE57FDF2583AFDC0" authority="Seifert, 2020" authorityName="Seifert" authorityYear="2020" box="[498,631,537,561]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="48" phylum="Arthropoda" rank="species" species="silvaticus" status="sp. nov.">L. silvaticus</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="B4DC57C9FD1CFFE3FD25FDF2589EFDC0" box="[640,723,538,561]" pageId="33" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="5A9B4D23FD1CFFE3FCAEFDF25B0AFDA3" authorityName="Bingham" authorityYear="1903" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="48" phylum="Arthropoda" rank="species" species="himalayanus">
|
||
<emphasis id="AFEFEAB2FD1CFFE3FCAEFDF25B0AFDA3" italics="true" pageId="33" pageNumber="48">L. himalayanus</emphasis>
|
||
</taxonomicName>
|
||
are hypothesized here to represent three cryptic species with different geographic distribution. They are clearly clustered on the sample level as different entities by exploratory data analyses using the 16 standard NUMOBAT characters unselectively. The classification error relative to the controlling LDA is 0% in a PCA and a TSCA using the first four principal components. NC-Ward, NC-part.kmeans, NC-part.hclust and NC-NMDS-kmeans misclassify
|
||
<specimenCount id="8B9DFD29FD1CFFE3FDC7FCAA58E0FCA8" box="[610,685,834,857]" count="1" pageId="33" pageNumber="48" type="generic">
|
||
one
|
||
<taxonomicName id="5A9B4D23FD1CFFE3FD32FCAA596CFCA8" baseAuthorityName="Latreille" baseAuthorityYear="1798" box="[663,801,833,857]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="48" phylum="Arthropoda" rank="species" species="brunneus">
|
||
<emphasis id="AFEFEAB2FD1CFFE3FD32FCAA596CFCA8" box="[663,801,833,857]" italics="true" pageId="33" pageNumber="48">L. brunneus</emphasis>
|
||
</taxonomicName>
|
||
</specimenCount>
|
||
sample from Aksehir /
|
||
<collectingCountry id="E58C7630FD1CFFE3FE11FC8A5BB2FC8B" box="[436,511,866,890]" name="Turkey" pageId="33" pageNumber="48">Turkey</collectingCountry>
|
||
as
|
||
<taxonomicName id="5A9B4D23FD1CFFE3FD8DFC8B5882FC8B" authorityName="Bingham" authorityYear="1903" box="[552,719,866,890]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="48" phylum="Arthropoda" rank="species" species="himalayanus">
|
||
<emphasis id="AFEFEAB2FD1CFFE3FD8DFC8B5882FC8B" box="[552,719,866,890]" italics="true" pageId="33" pageNumber="48">L. himalayanus</emphasis>
|
||
</taxonomicName>
|
||
(=error 2.9% in 34 samples). A mean classification error of 1.9% within six exploratory data analyses is in agreement with the heterospecificity threshold of the GAGE species concept. The three cryptic species seem to be parapatric or allopatric according to the poor information currently available and it will be interesting to see if the current taxonomic assessment would change after investigations in the putative contact zone of
|
||
<taxonomicName id="5A9B4D23FD1CFFE3FE56FB82583FFB70" baseAuthorityName="Latreille" baseAuthorityYear="1798" box="[499,626,1129,1153]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="48" phylum="Arthropoda" rank="species" species="brunneus">
|
||
<emphasis id="AFEFEAB2FD1CFFE3FE56FB82583FFB70" box="[499,626,1129,1153]" italics="true" pageId="33" pageNumber="48">L.brunneus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="5A9B4D23FD1CFFE3FD07FB82596CFB70" authority="Seifert, 2020" authorityName="Seifert" authorityYear="2020" box="[674,801,1129,1153]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="48" phylum="Arthropoda" rank="species" species="silvaticus" status="sp. nov.">
|
||
<emphasis id="AFEFEAB2FD1CFFE3FD07FB82596CFB70" box="[674,801,1129,1153]" italics="true" pageId="33" pageNumber="48">L. silvaticus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B4DC57C9FD1CFFE3FF18FB635B5CFB53" box="[189,273,1163,1186]" pageId="33" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
|
||
near the Turkish-Iranian border will have been done. The allopatric data currently available provide a strong signal for species separation: the discriminant D(8) = 40.81*SL–38.9*CW +71.2*PnHL+0.866*nHT– 0.674*sqPDCL+0.246*nSt–2.392 is D(8)= –1.885 ±0.956 [–4.75,0.00] in
|
||
<specimenCount id="8B9DFD29FD1CFFE3FEC0FAC65BADFAB7" box="[357,480,1326,1350]" count="56" pageId="33" pageNumber="48" type="worker">56 workers</specimenCount>
|
||
of
|
||
<taxonomicName id="5A9B4D23FD1CFFE3FDAEFAC758DCFAB7" baseAuthorityName="Latreille" baseAuthorityYear="1798" box="[523,657,1326,1350]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="48" phylum="Arthropoda" rank="species" species="brunneus">
|
||
<emphasis id="AFEFEAB2FD1CFFE3FDAEFAC758DCFAB7" box="[523,657,1326,1350]" italics="true" pageId="33" pageNumber="48">L. brunneus</emphasis>
|
||
</taxonomicName>
|
||
and D(8) = 4.390 ± 1.098 [2.63, 6.56] in
|
||
<specimenCount id="8B9DFD29FD1CFFE3FE57FAB85824FA96" box="[498,617,1359,1384]" count="24" pageId="33" pageNumber="48" type="worker">24 workers</specimenCount>
|
||
of
|
||
<taxonomicName id="5A9B4D23FD1CFFE3FD29FAB85942FA96" authorityName="Seifert" authorityYear="2020" box="[652,783,1359,1383]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="48" phylum="Arthropoda" rank="species" species="silvaticus">
|
||
<emphasis id="AFEFEAB2FD1CFFE3FD29FAB85942FA96" box="[652,783,1359,1383]" italics="true" pageId="33" pageNumber="48">L. silvaticus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</tableCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |