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<mods:title id="A14CE36F0DC93058E0AF0E4BB6B9B9EA">A taxonomic revision of the Palaearctic members of the subgenus Lasius s. str. (Hymenoptera, Formicidae)</mods:title>
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<treatment id="153287B6FD06FFFEFCEAFA3958BCFC4D" LSID="urn:lsid:plazi:treatment:153287B6FD06FFFEFCEAFA3958BCFC4D" httpUri="http://treatment.plazi.org/id/153287B6FD06FFFEFCEAFA3958BCFC4D" lastPageId="60" lastPageNumber="75" pageId="59" pageNumber="74">
<subSubSection id="D581652BFD06FFF9FCEAFA395F0EFA1C" box="[847,1347,1489,1517]" pageId="59" pageNumber="74" type="nomenclature">
<paragraph id="9D2436A0FD06FFF9FCEAFA395F0EFA1C" blockId="59.[847,1347,1489,1517]" box="[847,1347,1489,1517]" pageId="59" pageNumber="74">
<heading id="C66C81CCFD06FFF9FCEAFA395F0EFA1C" bold="true" box="[847,1347,1489,1517]" fontSize="10" level="2" pageId="59" pageNumber="74" reason="3">
<emphasis id="AFEFEAB2FD06FFF9FCEAFA395F0EFA1C" bold="true" box="[847,1347,1489,1517]" pageId="59" pageNumber="74">
4.4.46
<taxonomicName id="5A9B4D23FD06FFF9FC13FA395F0EFA1C" ID-CoL="6P3DC" authority="Seifert 1992" authorityName="Seifert" authorityYear="1992" box="[950,1347,1489,1517]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="59" pageNumber="74" phylum="Arthropoda" rank="species" species="wittmeri">
<emphasis id="AFEFEAB2FD06FFF9FC13FA395EC0FA1A" bold="true" box="[950,1165,1489,1515]" italics="true" pageId="59" pageNumber="74">Lasius wittmeri</emphasis>
<bibRefCitation id="F90A4B51FD06FFF9FB33FA395F0EFA1C" author="Seifert, B." box="[1174,1347,1489,1517]" pageId="59" pageNumber="74" pagination="1 - 67" refId="ref51794" refString="Seifert, B. (1992): A taxonomic revision of the Palaearctic members of the ant subgenus Lasius s. str. (Hymenoptera: Formicidae). - Abhandlungen und Berichte des Naturkundemuseums Gorlitz 66 / 5: 1 - 67." type="journal article" year="1992">Seifert 1992</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D581652BFD06FFF9FCEAFA1D5F14F9FD" box="[847,1369,1524,1548]" pageId="59" pageNumber="74" type="reference_group">
<paragraph id="9D2436A0FD06FFF9FCEAFA1D5F14F9FD" blockId="59.[824,1443,1524,1975]" box="[847,1369,1524,1548]" pageId="59" pageNumber="74">
<taxonomicName id="5A9B4D23FD06FFF9FCEAFA1D5ECCF9FD" ID-CoL="6P3DC" authority="Seifert 1992" authorityName="Seifert" authorityYear="1992" box="[847,1153,1524,1548]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="59" pageNumber="74" phylum="Arthropoda" rank="species" species="wittmeri">
<emphasis id="AFEFEAB2FD06FFF9FCEAFA1D59BAF9FD" box="[847,1015,1525,1548]" italics="true" pageId="59" pageNumber="74">Lasius wittmeri</emphasis>
<bibRefCitation id="F90A4B51FD06FFF9FC5BFA1C5ECCF9FD" author="Seifert, B." box="[1022,1153,1524,1548]" pageId="59" pageNumber="74" pagination="1 - 67" refId="ref51794" refString="Seifert, B. (1992): A taxonomic revision of the Palaearctic members of the ant subgenus Lasius s. str. (Hymenoptera: Formicidae). - Abhandlungen und Berichte des Naturkundemuseums Gorlitz 66 / 5: 1 - 67." type="journal article" year="1992">Seifert 1992</bibRefCitation>
</taxonomicName>
[
<typeStatus id="42208802FD06FFF9FB2BFA1C5EF2F9FD" box="[1166,1215,1524,1548]" pageId="59" pageNumber="74">type</typeStatus>
investigation]
</paragraph>
</subSubSection>
<subSubSection id="D581652BFD06FFF9FCEAF9FD5F71F867" pageId="59" pageNumber="74" type="materials_examined">
<paragraph id="9D2436A0FD06FFF9FCEAF9FD5FD1F9DC" blockId="59.[824,1443,1524,1975]" box="[847,1436,1557,1581]" pageId="59" pageNumber="74">
Type material:
<typeStatus id="42208802FD06FFF9FBADF9FD5E21F9DC" box="[1032,1132,1557,1581]" pageId="59" pageNumber="74" type="holotype">Holotype</typeStatus>
and
<specimenCount id="8B9DFD29FD06FFF9FB12F9FD5FD1F9DC" box="[1207,1436,1557,1581]" count="5" pageId="59" pageNumber="74" type="worker" typeStatus="paratype">5 paratype workers</specimenCount>
</paragraph>
<paragraph id="9D2436A0FD06FFF9FCEAF9DE5FD1F9BF" blockId="59.[824,1443,1524,1975]" box="[847,1436,1589,1614]" pageId="59" pageNumber="74">labelled ”Kashmir, 1976, W. Wittmer“ and ”Pahalgam</paragraph>
<paragraph id="9D2436A0FD06FFF9FCEAF9BF5FD1F99F" blockId="59.[824,1443,1524,1975]" box="[847,1436,1622,1647]" pageId="59" pageNumber="74">
<date id="E9251060FD06FFF9FCEAF9BF59F8F99E" box="[847,949,1623,1647]" pageId="59" pageNumber="74" value="2200-07-07">
7. 7.
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</date>
“;
<specimenCount id="8B9DFD29FD06FFF9FB80F9BE5EB9F99F" box="[1061,1268,1622,1647]" count="6" pageId="59" pageNumber="74" type="worker" typeStatus="paratype">6 paratype workers</specimenCount>
labelled ”Paki-
</paragraph>
<paragraph id="9D2436A0FD06FFF9FCEAF9905FD1F961" blockId="59.[824,1443,1524,1975]" box="[847,1436,1655,1680]" pageId="59" pageNumber="74">stan 1974, C. Baroni Urbani“ and ”Naran 7900 Kagan</paragraph>
<paragraph id="9D2436A0FD06FFF9FCEAF9705FEEF941" blockId="59.[824,1443,1524,1975]" box="[847,1443,1688,1712]" pageId="59" pageNumber="74">
Valley 25.V“; both samples deposited in
<collectionCode id="FB8AAE65FD06FFF9FAB7F9715F1FF941" box="[1298,1362,1689,1712]" pageId="59" pageNumber="74">NHM</collectionCode>
Basel.
</paragraph>
<paragraph id="9D2436A0FD06FFF9FCEAF9515EC3F8C5" blockId="59.[824,1443,1524,1975]" pageId="59" pageNumber="74">
<emphasis id="AFEFEAB2FD06FFF9FCEAF9515E1DF920" bold="true" box="[847,1104,1721,1745]" pageId="59" pageNumber="74">All material examined</emphasis>
.
<materialsCitation id="2DF33CFDFD06FFF9FBF8F9515EC6F8C5" country="Pakistan" location="These" municipality="For" pageId="59" pageNumber="74" specimenCount="12" specimenCount-worker="12">
A total of 2 nest samples with
<specimenCount id="8B9DFD29FD06FFF9FC9DF93259F9F903" box="[824,948,1754,1778]" count="12" pageId="59" pageNumber="74" type="worker">12 workers</specimenCount>
were subject to NUMOBAT investigation.
<location id="9844607BFD06FFF9FC9DF9135934F8E2" LSID="urn:lsid:plazi:treatment:153287B6FD06FFFEFCEAFA3958BCFC4D:9844607BFD06FFF9FC9DF9135934F8E2" box="[824,889,1787,1811]" country="Pakistan" municipality="For" name="These" pageId="59" pageNumber="74">These</location>
originated from
<collectingCountry id="E58C7630FD06FFF9FB8BF9135EC6F8E2" box="[1070,1163,1787,1811]" name="Pakistan" pageId="59" pageNumber="74">Pakistan</collectingCountry>
and
<collectingCountry id="E58C7630FD06FFF9FB1BF9135EB7F8E2" box="[1214,1274,1787,1811]" name="India" pageId="59" pageNumber="74">India</collectingCountry>
.
<collectingMunicipality id="7D40ACDAFD06FFF9FAA6F9135F6AF8E3" box="[1283,1319,1787,1810]" pageId="59" pageNumber="74">For</collectingMunicipality>
details see supplementary information SI1
</materialsCitation>
.
</paragraph>
<paragraph id="9D2436A0FD06FFF9FCEAF8D45F71F867" blockId="59.[824,1443,1524,1975]" pageId="59" pageNumber="74">
<emphasis id="AFEFEAB2FD06FFF9FCEAF8D45E52F8A4" bold="true" box="[847,1055,1852,1877]" pageId="59" pageNumber="74">Geographic range</emphasis>
. The two known sites are situated at the southwestern flank of the Himalayas at
<geoCoordinate id="F8AF5067FD06FFF9FA99F8B65FD5F887" box="[1340,1432,1886,1910]" degrees="34.90" direction="north" orientation="latitude" pageId="59" pageNumber="74" precision="555" value="34.9">34.90°N</geoCoordinate>
,
<geoCoordinate id="F8AF5067FD06FFF9FC9DF89659C0F867" box="[824,909,1918,1942]" degrees="73.75" direction="east" orientation="longitude" pageId="59" pageNumber="74" precision="555" value="73.75">73.75°E</geoCoordinate>
,
<quantity id="5A639B45FD06FFF9FC3DF89659A4F867" box="[920,1001,1918,1942]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="59" pageNumber="74" unit="m" value="2400.0">2400 m</quantity>
and
<geoCoordinate id="F8AF5067FD06FFF9FBB8F8965E34F867" box="[1053,1145,1918,1942]" degrees="34.04" direction="north" orientation="latitude" pageId="59" pageNumber="74" precision="555" value="34.04">34.04°N</geoCoordinate>
,
<geoCoordinate id="F8AF5067FD06FFF9FB21F8965E97F867" box="[1156,1242,1918,1942]" degrees="75.33" direction="east" orientation="longitude" pageId="59" pageNumber="74" precision="555" value="75.33">75.33°E</geoCoordinate>
,
<quantity id="5A639B45FD06FFF9FB40F8965F75F867" box="[1253,1336,1918,1942]" metricMagnitude="3" metricUnit="m" metricValue="2.65" pageId="59" pageNumber="74" unit="m" value="2650.0">2650 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="D581652BFD06FFFEFCEAF877583AFCE9" lastPageId="60" lastPageNumber="75" pageId="59" pageNumber="74" type="diagnosis">
<paragraph id="9D2436A0FD06FFF9FCEAF8775FD0F846" blockId="59.[824,1443,1524,1975]" box="[847,1437,1951,1975]" pageId="59" pageNumber="74">
<emphasis id="AFEFEAB2FD06FFF9FCEAF87759F3F846" bold="true" box="[847,958,1951,1975]" pageId="59" pageNumber="74">Diagnosis</emphasis>
(Tab. 9,
<figureCitation id="05A02A25FD06FFF9FB86F8775E36F846" box="[1059,1147,1951,1975]" captionStart="Figs" captionStartId="28.[151,191,1767,1789]" captionTargetBox="[151,1285,1344,1750]" captionTargetPageId="28" captionText="Figs. 89 and 90: Lasius wittmeri; holotype" figureDoi="http://doi.org/10.5281/zenodo.10832318" httpUri="https://zenodo.org/record/10832318/files/figure.png" pageId="59" pageNumber="74">Figs. 89</figureCitation>
90, key, images in www.
</paragraph>
<paragraph id="9D2436A0FD01FFFEFF32FF19583AFCE9" blockId="60.[151,764,241,956]" pageId="60" pageNumber="75">
antWeb.org with specimen identifiers CASENT0912298): The most similar Himalayan species is
<taxonomicName id="5A9B4D23FD01FFFEFDEEFEFB58F3FEDB" authorityName="Seifert" authorityYear="1992" box="[587,702,274,298]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="75" phylum="Arthropoda" rank="species" species="lawarai">
<emphasis id="AFEFEAB2FD01FFFEFDEEFEFB58F3FEDB" box="[587,702,274,298]" italics="true" pageId="60" pageNumber="75">L. lawarai</emphasis>
</taxonomicName>
from which it differs by larger eyes (EYE/CS
<subScript id="011F34E5FD01FFFEFD9BFEAB581BFEA0" attach="left" box="[574,598,323,337]" fontSize="6" pageId="60" pageNumber="75">900</subScript>
0.240), smaller postocular index (PoOc/CL
<subScript id="011F34E5FD01FFFEFE63FE8C5B93FE83" attach="left" box="[454,478,356,370]" fontSize="6" pageId="60" pageNumber="75">900</subScript>
0.241), longer scape (SL/ CS
<subScript id="011F34E5FD01FFFEFF12FE6D5A82FE62" attach="left" box="[183,207,389,403]" fontSize="6" pageId="60" pageNumber="75">900</subScript>
0.979) and smaller torulo-clypeal distance (dClAn/ CS
<subScript id="011F34E5FD01FFFEFF12FE4E5A82FE45" attach="left" box="[183,207,422,436]" fontSize="6" pageId="60" pageNumber="75">900</subScript>
3.60%). The Tibetan
<taxonomicName id="5A9B4D23FD01FFFEFE71FE7F5817FE5F" authorityName="Seifert" authorityYear="1992" box="[468,602,406,430]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="75" phylum="Arthropoda" rank="species" species="schaeferi">
<emphasis id="AFEFEAB2FD01FFFEFE71FE7F5817FE5F" box="[468,602,406,430]" italics="true" pageId="60" pageNumber="75">L. schaeferi</emphasis>
</taxonomicName>
differs by a much shorter frontal pubescence (PLF
<subScript id="011F34E5FD01FFFEFDEDFE2F582DFE24" attach="left" box="[584,608,455,469]" fontSize="6" pageId="60" pageNumber="75">900</subScript>
29.6 vs. 38.8 µm) and the presence of very distinct standing setae on hind tibia, the morphology of which differs clearly from neighboring pubescence hairs. Seta counts in
<taxonomicName id="5A9B4D23FD01FFFEFD21FDF258B6FDC0" authorityName="Seifert" authorityYear="1992" box="[644,763,538,561]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="75" phylum="Arthropoda" rank="species" species="wittmeri">
<emphasis id="AFEFEAB2FD01FFFEFD21FDF258B6FDC0" box="[644,763,538,561]" italics="true" pageId="60" pageNumber="75">L. wittmeri</emphasis>
</taxonomicName>
are not clearly reproducible because of unclear thickness differences between elongated semierect pubescence hairs and semierect setae. Yet, this missing differentiation may be used as accessory character to distinguish
<taxonomicName id="5A9B4D23FD01FFFEFD2FFD7658B1FD44" authorityName="Seifert" authorityYear="1992" box="[650,764,669,693]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="75" phylum="Arthropoda" rank="species" species="lawarai">
<emphasis id="AFEFEAB2FD01FFFEFD2FFD7658B1FD44" box="[650,764,669,693]" italics="true" pageId="60" pageNumber="75">L. lawarai</emphasis>
</taxonomicName>
also from East Tibetan populations of
<taxonomicName id="5A9B4D23FD01FFFEFDFEFD5758B8FD27" authorityName="Stitz" authorityYear="1930" box="[603,757,702,726]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="75" phylum="Arthropoda" rank="species" species="obscuratus">
<emphasis id="AFEFEAB2FD01FFFEFDFEFD5758B8FD27" box="[603,757,702,726]" italics="true" pageId="60" pageNumber="75">L. obscuratus</emphasis>
</taxonomicName>
. Coloration: all body parts dark brown, mandibles and tarsae slightly lighter with a yellowish tinge.
</paragraph>
</subSubSection>
<subSubSection id="D581652BFD01FFFEFF0BFCC858BCFC4D" pageId="60" pageNumber="75" type="biology_ecology">
<paragraph id="9D2436A0FD01FFFEFF0BFCC85BCEFCC9" blockId="60.[151,764,241,956]" box="[174,387,800,824]" pageId="60" pageNumber="75">
<emphasis id="AFEFEAB2FD01FFFEFF0BFCC85B4EFCC9" bold="true" box="[174,259,800,824]" pageId="60" pageNumber="75">Biology</emphasis>
. Unknown.
</paragraph>
<paragraph id="9D2436A0FD01FFFEFF0BFCA958BCFC4D" blockId="60.[151,764,241,956]" pageId="60" pageNumber="75">
Comments. 96% of
<specimenCount id="8B9DFD29FD01FFFEFE2AFCA95847FCA8" box="[399,522,833,857]" count="25" pageId="60" pageNumber="75" type="worker">25 workers</specimenCount>
of
<taxonomicName id="5A9B4D23FD01FFFEFD91FCAA58E4FCA8" authorityName="Seifert" authorityYear="1992" box="[564,681,833,857]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="75" phylum="Arthropoda" rank="species" species="lawarai">
<emphasis id="AFEFEAB2FD01FFFEFD91FCAA58E4FCA8" box="[564,681,833,857]" italics="true" pageId="60" pageNumber="75">L. lawarai</emphasis>
</taxonomicName>
and
<taxonomicName id="5A9B4D23FD01FFFEFD40FCAA5ABFFC8B" authorityName="Seifert" authorityYear="1992" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="75" phylum="Arthropoda" rank="species" species="wittmeri">
<emphasis id="AFEFEAB2FD01FFFEFD40FCAA5ABFFC8B" italics="true" pageId="60" pageNumber="75">L. wittmeri</emphasis>
</taxonomicName>
for which data of the full character set were available were correctly classified in a LOOCV-LDA using the first three components of a PCA as input data.
</paragraph>
</subSubSection>
</treatment>
</document>