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<document id="D3E178B67DE68D01CEA0CEEB4BFB948D" ID-CLB-Dataset="27519" ID-DOI="10.11646/zootaxa.4576.1.6" ID-GBIF-Dataset="7603d0a7-0f1c-42e7-be44-9de473c1ffc9" ID-ISSN="1175-5326" ID-Zenodo-Dep="2624732" ID-ZooBank="92C910D5-3966-4CC1-B3DC-876B978C53E5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554272946661" checkinUser="plazi" docAuthor="Hoskin, Conrad J., Bertola, Lorenzo V. &amp; Higgie, Megan" docDate="2019" docId="03FD7D72FFDEFFC9FF41FE54DC1AF861" docLanguage="en" docName="zootaxa.4576.1.6.pdf" docOrigin="Zootaxa 4576 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phyllurus pinnaclensis Hoskin &amp; Bertola &amp; Higgie 2019, sp. nov." docType="treatment" docUuid="2885CFC2-F807-40D3-A834-0F48DDBF9C43" docUuidSource="ZooBank" docVersion="9" lastPageNumber="132" masterDocId="FFC4050AFFDDFFCCFFD6FFE2D811FF94" masterDocTitle="A new species of Phyllurus leaf-tailed gecko (Lacertilia: Carphodactylidae) from The Pinnacles, north-east Australia" masterLastPageNumber="139" masterPageNumber="127" pageNumber="130" updateTime="1698716523061" updateUser="ExternalLinkService">
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<mods:title id="95034DA8CC55C701AE3EAA7AEDC9E211">A new species of Phyllurus leaf-tailed gecko (Lacertilia: Carphodactylidae) from The Pinnacles, north-east Australia</mods:title>
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<mods:namePart id="09C6D1A73EA4075A32CBFA41759F5827">Hoskin, Conrad J.</mods:namePart>
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<mods:namePart id="1202258C9867F484723FFB0CAEA4F28D">Bertola, Lorenzo V.</mods:namePart>
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<mods:namePart id="48240488D399838A2A4300094B981350">Higgie, Megan</mods:namePart>
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<treatment id="03FD7D72FFDEFFC9FF41FE54DC1AF861" ID-DOI="http://doi.org/10.5281/zenodo.5932410" ID-GBIF-Taxon="154532613" ID-Zenodo-Dep="5932410" LSID="urn:lsid:zoobank.org:act:2885CFC2-F807-40D3-A834-0F48DDBF9C43" httpUri="http://treatment.plazi.org/id/03FD7D72FFDEFFC9FF41FE54DC1AF861" lastPageId="5" lastPageNumber="132" pageId="3" pageNumber="130">
<subSubSection id="C34E9FEFFFDEFFCFFF41FE54D9C5FE66" pageId="3" pageNumber="130" type="nomenclature">
<paragraph id="8BEBCC64FFDEFFCFFF41FE54DA00FE44" blockId="3.[151,529,438,534]" box="[151,529,438,464]" pageId="3" pageNumber="130">
<heading id="D0A37B08FFDEFFCFFF41FE54DA00FE44" bold="true" box="[151,529,438,464]" fontSize="11" level="1" pageId="3" pageNumber="130" reason="1">
<emphasis id="B9201076FFDEFFCFFF41FE54DA00FE44" bold="true" box="[151,529,438,464]" pageId="3" pageNumber="130">
<taxonomicName id="4C54B7E7FFDEFFCFFF41FE54D9BDFE44" authority="Hoskin &amp; Bertola &amp; Higgie, 2019" authorityName="Hoskin &amp; Bertola &amp; Higgie" authorityYear="2019" box="[151,428,438,464]" class="Reptilia" family="Gekkonidae" genus="Phyllurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="130" phylum="Chordata" rank="species" species="pinnaclensis" status="sp. nov.">
<emphasis id="B9201076FFDEFFCFFF41FE54D9BDFE44" bold="true" box="[151,428,438,464]" italics="true" pageId="3" pageNumber="130">Phyllurus pinnaclensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A213AD0DFFDEFFCFFE62FE54DA00FE44" box="[436,529,438,464]" pageId="3" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEBCC64FFDEFFCFFF41FE3BD9C5FE66" blockId="3.[151,529,438,534]" box="[151,468,473,498]" pageId="3" pageNumber="130">Pinnacles Leaf-tailed Gecko</paragraph>
</subSubSection>
<subSubSection id="C34E9FEFFFDEFFCFFF41FE1FD97AFD82" box="[151,363,509,534]" pageId="3" pageNumber="130" type="description">
<paragraph id="8BEBCC64FFDEFFCFFF41FE1FD97AFD82" blockId="3.[151,529,438,534]" box="[151,363,509,534]" pageId="3" pageNumber="130">
(
<figureCitation id="136FD0E1FFDEFFCFFF49FE1FD8F6FD82" box="[159,231,509,534]" captionStart="FIGURE 2" captionStartId="5.[151,250,1532,1554]" captionTargetBox="[151,1436,301,1511]" captionTargetId="figure@5.[151,1436,301,1511]" captionTargetPageId="5" captionText="FIGURE 2. (A) Holotype of P. pinnaclensis sp. nov. in life (photo: Conrad Hoskin). (B) P. pinnaclensis sp. nov. with regenerated tail (photo: Lorenzo Bertola)." httpUri="https://zenodo.org/record/2624736/files/figure.png" pageId="3" pageNumber="130">Figs 2</figureCitation>
,
<figureCitation id="136FD0E1FFDEFFCFFF25FE1CD912FD82" box="[243,259,510,534]" captionStart="FIGURE 3" captionStartId="6.[151,250,1036,1058]" captionTargetBox="[151,1436,193,1024]" captionTargetId="figure@6.[151,1436,193,1024]" captionTargetPageId="6" captionText="FIGURE 3. Holotype of P. pinnaclensis sp. nov. in life (photo: Conrad Hoskin)." httpUri="https://zenodo.org/record/2624738/files/figure.png" pageId="3" pageNumber="130">3</figureCitation>
,
<figureCitation id="136FD0E1FFDEFFCFFED9FE1CD925FD82" box="[271,308,510,534]" captionStart="FIGURE 4" captionStartId="7.[151,250,1444,1466]" captionTargetBox="[151,1436,411,1409]" captionTargetId="figure@7.[151,1436,411,1409]" captionTargetPageId="7" captionText="FIGURE 4. Holotypes of (A) P. pinnaclensis sp. nov. (QMJ96418) and (B) P. gulbaru (QMJ75778) (photo: Conrad Hoskin)." httpUri="https://zenodo.org/record/2624740/files/figure.png" pageId="3" pageNumber="130">4A</figureCitation>
,
<figureCitation id="136FD0E1FFDEFFCFFE96FE1FD972FD82" box="[320,355,509,534]" captionStart="FIGURE 5" captionStartId="8.[151,250,1966,1988]" captionTargetBox="[194,1388,193,1949]" captionTargetId="figure@8.[194,1388,193,1949]" captionTargetPageId="8" captionText="FIGURE 5. Traced outline of internasal scales of holotypes of (A) P. pinnaclensis sp. nov. (QMJ96418) and P. gulbaru (QMJ75778) (photo: Conrad Hoskin)." httpUri="https://zenodo.org/record/2624742/files/figure.png" pageId="3" pageNumber="130">5A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34E9FEFFFDEFFCFFF41FDA7DA81FD15" pageId="3" pageNumber="130" type="materials_examined">
<paragraph id="8BEBCC64FFDEFFCFFF41FDA7DA81FD15" blockId="3.[151,1437,581,1946]" pageId="3" pageNumber="130">
<emphasis id="B9201076FFDEFFCFFF41FDA7D9EFFDCA" bold="true" box="[151,510,581,606]" pageId="3" pageNumber="130">
Material examined.
<typeStatus id="54EF72C6FFDEFFCFFE5CFDA7D9E8FDCA" box="[394,505,581,606]" pageId="3" pageNumber="130" type="holotype">Holotype</typeStatus>
.
</emphasis>
QMJ96418, adult male, The Pinnacles (
<geoCoordinate id="EE60AAA3FFDEFFCFFC16FDA4DC3DFDCA" box="[960,1068,582,606]" direction="south" orientation="latitude" pageId="3" pageNumber="130" precision="55" value="-19.41">19.410°S</geoCoordinate>
,
<geoCoordinate id="EE60AAA3FFDEFFCFFBEEFDA7DCA5FDCA" box="[1080,1204,581,606]" direction="east" orientation="longitude" pageId="3" pageNumber="130" precision="55" value="146.614">146.614°E</geoCoordinate>
),
<date id="FFEAEAA4FFDEFFCFFB1EFDA4DD89FDCA" box="[1224,1432,581,606]" pageId="3" pageNumber="130" value="2015-11-02">2 November 2015</date>
, C. J. Hoskin &amp; M. Higgie.
<emphasis id="B9201076FFDEFFCFFE10FD8BDA54FD16" bold="true" box="[454,581,617,642]" pageId="3" pageNumber="130">
<typeStatus id="54EF72C6FFDEFFCFFE10FD8BDA51FD16" box="[454,576,617,642]" pageId="3" pageNumber="130" type="paratype">Paratypes</typeStatus>
.
</emphasis>
None.
</paragraph>
</subSubSection>
<subSubSection id="C34E9FEFFFDEFFCFFF11FD6FDD65FD64" pageId="3" pageNumber="130" type="diagnosis">
<paragraph id="8BEBCC64FFDEFFCFFF11FD6FDD65FD64" blockId="3.[151,1437,581,1946]" pageId="3" pageNumber="130">
<emphasis id="B9201076FFDEFFCFFF11FD6FD950FD32" bold="true" box="[199,321,653,678]" pageId="3" pageNumber="130">Diagnosis.</emphasis>
Distinguished from congeners by the following characters: cylindrical, tapering tail; rostral scale partially divided by a single groove; high internasal count (710, typically 9 or 10); high subdigital lamellae count under the 4
<superScript id="7C21612CFFDEFFCFFEC3FD37D930FD77" attach="left" box="[277,289,725,739]" fontSize="6" pageId="3" pageNumber="130">th</superScript>
toe (1921); original tail long (&gt; 85% SVL) and largely devoid of tubercles after the basal quarter.
</paragraph>
</subSubSection>
<subSubSection id="C34E9FEFFFDEFFCFFF11FD1EDCFDFC81" box="[199,1260,764,789]" pageId="3" pageNumber="130" type="etymology">
<paragraph id="8BEBCC64FFDEFFCFFF11FD1EDCFDFC81" blockId="3.[151,1437,581,1946]" box="[199,1260,764,789]" pageId="3" pageNumber="130">
<emphasis id="B9201076FFDEFFCFFF11FD1ED95FFC81" bold="true" box="[199,334,764,789]" pageId="3" pageNumber="130">Etymology.</emphasis>
Refers to The Pinnacles, the locality to which this species appears to be restricted.
</paragraph>
</subSubSection>
<subSubSection id="C34E9FEFFFDEFFC9FF11FCC2DC66F9E4" lastPageId="5" lastPageNumber="132" pageId="3" pageNumber="130" type="description">
<paragraph id="8BEBCC64FFDEFFCFFF11FCC2D8F2F80E" blockId="3.[151,1437,581,1946]" pageId="3" pageNumber="130">
<emphasis id="B9201076FFDEFFCFFF11FCC2DB71FCA3" bold="true" box="[199,864,799,825]" pageId="3" pageNumber="130">
Description of
<typeStatus id="54EF72C6FFDEFFCFFEACFCC2D9F2FCAD" box="[378,483,800,825]" pageId="3" pageNumber="130" type="holotype">holotype</typeStatus>
.
<emphasis id="B9201076FFDEFFCFFE26FCC2DB71FCA3" bold="true" box="[496,864,799,824]" italics="true" pageId="3" pageNumber="130">Measurements and scale counts.</emphasis>
</emphasis>
SVL =
<quantity id="4CAC6181FFDEFFCFFC6AFCC2DC21FCAC" box="[956,1072,800,825]" metricMagnitude="-2" metricUnit="m" metricValue="8.522" pageId="3" pageNumber="130" unit="mm" value="85.22">85.22 mm</quantity>
, T (original) =
<quantity id="4CAC6181FFDEFFCFFB0CFCC2DD5FFCAC" box="[1242,1358,800,824]" metricMagnitude="-2" metricUnit="m" metricValue="7.428" pageId="3" pageNumber="130" unit="mm" value="74.28">74.28 mm</quantity>
, TW =
<quantity id="4CAC6181FFDEFFCFFF41FCA6D8EAFCC8" box="[151,251,836,861]" metricMagnitude="-3" metricUnit="m" metricValue="6.94" pageId="3" pageNumber="130" unit="mm" value="6.94">6.94 mm</quantity>
, TD =
<quantity id="4CAC6181FFDEFFCFFE9EFCA6D9BCFCC8" box="[328,429,836,860]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="3" pageNumber="130" unit="mm" value="4.9">4.90 mm</quantity>
, AG =
<quantity id="4CAC6181FFDEFFCFFE2BFCA6DA61FCC8" box="[509,624,836,861]" metricMagnitude="-2" metricUnit="m" metricValue="3.876" pageId="3" pageNumber="130" unit="mm" value="38.76">38.76 mm</quantity>
, NL =
<quantity id="4CAC6181FFDEFFCFFD6BFCA6DB3FFCC8" box="[701,814,836,860]" metricMagnitude="-2" metricUnit="m" metricValue="1.732" pageId="3" pageNumber="130" unit="mm" value="17.32">17.32 mm</quantity>
, HL =
<quantity id="4CAC6181FFDEFFCFFCAAFCA6DBFFFCC8" box="[892,1006,836,861]" metricMagnitude="-2" metricUnit="m" metricValue="2.269" pageId="3" pageNumber="130" unit="mm" value="22.69">22.69 mm</quantity>
, HW =
<quantity id="4CAC6181FFDEFFCFFB93FCA6DCA6FCC8" box="[1093,1207,836,860]" metricMagnitude="-2" metricUnit="m" metricValue="1.918" pageId="3" pageNumber="130" unit="mm" value="19.18">19.18 mm</quantity>
, HD =
<quantity id="4CAC6181FFDEFFCFFAD1FCA6DD7DFCC8" box="[1287,1388,836,860]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="3" pageNumber="130" unit="mm" value="8.2">8.20 mm</quantity>
, S =
<quantity id="4CAC6181FFDEFFCFFF41FC8AD918FC14" box="[151,265,872,896]" metricMagnitude="-2" metricUnit="m" metricValue="1.098" pageId="3" pageNumber="130" unit="mm" value="10.98">10.98 mm</quantity>
, L1 =
<quantity id="4CAC6181FFDEFFCFFE87FC8AD9D2FC14" box="[337,451,872,896]" metricMagnitude="-2" metricUnit="m" metricValue="3.79" pageId="3" pageNumber="130" unit="mm" value="37.9">37.90 mm</quantity>
, FLL = 16.00 mm, L2 =
<quantity id="4CAC6181FFDEFFCFFD01FC8ADB58FC14" box="[727,841,872,896]" metricMagnitude="-2" metricUnit="m" metricValue="4.91" pageId="3" pageNumber="130" unit="mm" value="49.1">49.10 mm</quantity>
, HLL =
<quantity id="4CAC6181FFDEFFCFFC7EFC8ADC0BFC14" box="[936,1050,872,896]" metricMagnitude="-2" metricUnit="m" metricValue="1.91" pageId="3" pageNumber="130" unit="mm" value="19.1">19.10 mm</quantity>
, internasals = 10, supralabials = 16 (left side) and 15 (right side), infralabials = 15, subdigital lamellae under fingers (listed 1
<superScript id="7C21612CFFDEFFCFFBA1FC69DC90FC0D" attach="none" box="[1143,1153,907,921]" fontSize="6" pageId="3" pageNumber="130">st</superScript>
5
<superScript id="7C21612CFFDEFFCFFB4AFC69DCB6FC0D" attach="none" box="[1180,1191,907,921]" fontSize="6" pageId="3" pageNumber="130">th</superScript>
) = 12, 14, 17, 17, 16, subdigital lamellae under toes (listed 1
<superScript id="7C21612CFFDEFFCFFD97FC50DA5AFC54" attach="none" box="[577,587,946,960]" fontSize="6" pageId="3" pageNumber="130">st</superScript>
5
<superScript id="7C21612CFFDEFFCFFDB0FC50DA60FC54" attach="none" box="[614,625,946,960]" fontSize="6" pageId="3" pageNumber="130">th</superScript>
) = 13, 17, 19, 20, 20.
<emphasis id="B9201076FFDEFFCFFCBEFC57DBBDFC5A" bold="true" box="[872,940,949,974]" pageId="3" pageNumber="130">
<emphasis id="B9201076FFDEFFCFFCBEFC57DBB4FC5A" bold="true" box="[872,933,949,974]" italics="true" pageId="3" pageNumber="130">Head</emphasis>
.
</emphasis>
Large, depressed, triangular; covered in very small granules with larger, pale, conical tubercles at back and sides of head (
<figureCitation id="136FD0E1FFDEFFCFFC0CFC3BDC27FC66" box="[986,1078,985,1010]" captionStart="FIGURE 2" captionStartId="5.[151,250,1532,1554]" captionTargetBox="[151,1436,301,1511]" captionTargetId="figure@5.[151,1436,301,1511]" captionTargetPageId="5" captionText="FIGURE 2. (A) Holotype of P. pinnaclensis sp. nov. in life (photo: Conrad Hoskin). (B) P. pinnaclensis sp. nov. with regenerated tail (photo: Lorenzo Bertola)." httpUri="https://zenodo.org/record/2624736/files/figure.png" pageId="3" pageNumber="130">Figs 2A</figureCitation>
,
<figureCitation id="136FD0E1FFDEFFCFFB97FC38DC41FC66" box="[1089,1104,986,1010]" captionStart="FIGURE 3" captionStartId="6.[151,250,1036,1058]" captionTargetBox="[151,1436,193,1024]" captionTargetId="figure@6.[151,1436,193,1024]" captionTargetPageId="6" captionText="FIGURE 3. Holotype of P. pinnaclensis sp. nov. in life (photo: Conrad Hoskin)." httpUri="https://zenodo.org/record/2624738/files/figure.png" pageId="3" pageNumber="130">3</figureCitation>
,
<figureCitation id="136FD0E1FFDEFFCFFB8BFC38DC93FC66" box="[1117,1154,986,1010]" captionStart="FIGURE 4" captionStartId="7.[151,250,1444,1466]" captionTargetBox="[151,1436,411,1409]" captionTargetId="figure@7.[151,1436,411,1409]" captionTargetPageId="7" captionText="FIGURE 4. Holotypes of (A) P. pinnaclensis sp. nov. (QMJ96418) and (B) P. gulbaru (QMJ75778) (photo: Conrad Hoskin)." httpUri="https://zenodo.org/record/2624740/files/figure.png" pageId="3" pageNumber="130">4A</figureCitation>
); a bony ridge with larger granules extends forward from upper anterior margin of each eye towards naris (
<figureCitation id="136FD0E1FFDEFFCFFBDEFC1FDC5BFB82" box="[1032,1098,1021,1046]" captionStart="FIGURE 3" captionStartId="6.[151,250,1036,1058]" captionTargetBox="[151,1436,193,1024]" captionTargetId="figure@6.[151,1436,193,1024]" captionTargetPageId="6" captionText="FIGURE 3. Holotype of P. pinnaclensis sp. nov. in life (photo: Conrad Hoskin)." httpUri="https://zenodo.org/record/2624738/files/figure.png" pageId="3" pageNumber="130">Fig. 3</figureCitation>
); skin of head co-ossified with skull; deep, vertical groove partially (25%) dividing rostral scale (
<figureCitation id="136FD0E1FFDEFFCFFCB6FBC3DBA6FBAE" box="[864,951,1057,1082]" captionStart="FIGURE 5" captionStartId="8.[151,250,1966,1988]" captionTargetBox="[194,1388,193,1949]" captionTargetId="figure@8.[194,1388,193,1949]" captionTargetPageId="8" captionText="FIGURE 5. Traced outline of internasal scales of holotypes of (A) P. pinnaclensis sp. nov. (QMJ96418) and P. gulbaru (QMJ75778) (photo: Conrad Hoskin)." httpUri="https://zenodo.org/record/2624742/files/figure.png" pageId="3" pageNumber="130">Fig. 5A</figureCitation>
); rostral not in contact with nostril; 10 scales along the dorsal margin of rostral shield (
<figureCitation id="136FD0E1FFDEFFCFFD83FBA7DABDFBCA" box="[597,684,1093,1118]" captionStart="FIGURE 5" captionStartId="8.[151,250,1966,1988]" captionTargetBox="[194,1388,193,1949]" captionTargetId="figure@8.[194,1388,193,1949]" captionTargetPageId="8" captionText="FIGURE 5. Traced outline of internasal scales of holotypes of (A) P. pinnaclensis sp. nov. (QMJ96418) and P. gulbaru (QMJ75778) (photo: Conrad Hoskin)." httpUri="https://zenodo.org/record/2624742/files/figure.png" pageId="3" pageNumber="130">Fig. 5A</figureCitation>
); first supralabial scale taller than wide, remaining supralabials broad and steadily decreasing in size; first infralabial scale taller than wide, remaining infralabials broad and steadily decreasing in size; granular scales of chin and throat largest along the edge of the mental and infralabials (particularly abutting infralabials 13) and diminishing in size to minute granules on the throat; ear opening inconspicuous, elliptical, vertical, much less than half as large as eye.
<emphasis id="B9201076FFDEFFCFFD32FB37DB35FB7A" bold="true" box="[740,804,1237,1262]" italics="true" pageId="3" pageNumber="130">Neck.</emphasis>
Broad; covered in small granules that are intermixed with larger pale conical tubercles, including prominent, sharp tubercles on the back of the head and anterior neck (
<figureCitation id="136FD0E1FFDEFFCFFAEBFB1BDD89FA86" box="[1341,1432,1273,1298]" captionStart="FIGURE 2" captionStartId="5.[151,250,1532,1554]" captionTargetBox="[151,1436,301,1511]" captionTargetId="figure@5.[151,1436,301,1511]" captionTargetPageId="5" captionText="FIGURE 2. (A) Holotype of P. pinnaclensis sp. nov. in life (photo: Conrad Hoskin). (B) P. pinnaclensis sp. nov. with regenerated tail (photo: Lorenzo Bertola)." httpUri="https://zenodo.org/record/2624736/files/figure.png" pageId="3" pageNumber="130">Figs 2A</figureCitation>
,
<figureCitation id="136FD0E1FFDEFFCFFF41FAFCD8B7FAA2" box="[151,166,1310,1334]" captionStart="FIGURE 3" captionStartId="6.[151,250,1036,1058]" captionTargetBox="[151,1436,193,1024]" captionTargetId="figure@6.[151,1436,193,1024]" captionTargetPageId="6" captionText="FIGURE 3. Holotype of P. pinnaclensis sp. nov. in life (photo: Conrad Hoskin)." httpUri="https://zenodo.org/record/2624738/files/figure.png" pageId="3" pageNumber="130">3</figureCitation>
,
<figureCitation id="136FD0E1FFDEFFCFFF64FAFCD8C6FAA2" box="[178,215,1310,1334]" captionStart="FIGURE 4" captionStartId="7.[151,250,1444,1466]" captionTargetBox="[151,1436,411,1409]" captionTargetId="figure@7.[151,1436,411,1409]" captionTargetPageId="7" captionText="FIGURE 4. Holotypes of (A) P. pinnaclensis sp. nov. (QMJ96418) and (B) P. gulbaru (QMJ75778) (photo: Conrad Hoskin)." httpUri="https://zenodo.org/record/2624740/files/figure.png" pageId="3" pageNumber="130">4A</figureCitation>
).
<emphasis id="B9201076FFDEFFCFFF3FFAFFD939FAA2" bold="true" box="[233,296,1309,1334]" italics="true" pageId="3" pageNumber="130">Body.</emphasis>
Depressed, covered in small granules; flank granules intermixed with larger pale, conical tubercles (
<figureCitation id="136FD0E1FFDEFFCFFABAFAFFD8AAFACE" captionStart="FIGURE 2" captionStartId="5.[151,250,1532,1554]" captionTargetBox="[151,1436,301,1511]" captionTargetId="figure@5.[151,1436,301,1511]" captionTargetPageId="5" captionText="FIGURE 2. (A) Holotype of P. pinnaclensis sp. nov. in life (photo: Conrad Hoskin). (B) P. pinnaclensis sp. nov. with regenerated tail (photo: Lorenzo Bertola)." httpUri="https://zenodo.org/record/2624736/files/figure.png" pageId="3" pageNumber="130">Figs 2A</figureCitation>
,
<figureCitation id="136FD0E1FFDEFFCFFF1EFAA0D8C6FACE" box="[200,215,1346,1370]" captionStart="FIGURE 3" captionStartId="6.[151,250,1036,1058]" captionTargetBox="[151,1436,193,1024]" captionTargetId="figure@6.[151,1436,193,1024]" captionTargetPageId="6" captionText="FIGURE 3. Holotype of P. pinnaclensis sp. nov. in life (photo: Conrad Hoskin)." httpUri="https://zenodo.org/record/2624738/files/figure.png" pageId="3" pageNumber="130">3</figureCitation>
,
<figureCitation id="136FD0E1FFDEFFCFFF30FAA0D91AFACE" box="[230,267,1346,1370]" captionStart="FIGURE 4" captionStartId="7.[151,250,1444,1466]" captionTargetBox="[151,1436,411,1409]" captionTargetId="figure@7.[151,1436,411,1409]" captionTargetPageId="7" captionText="FIGURE 4. Holotypes of (A) P. pinnaclensis sp. nov. (QMJ96418) and (B) P. gulbaru (QMJ75778) (photo: Conrad Hoskin)." httpUri="https://zenodo.org/record/2624740/files/figure.png" pageId="3" pageNumber="130">4A</figureCitation>
); tubercles on back very small; ventral surface covered in even, smooth, fine granules, with no enlarged tubercles; preanal pores absent; large testicular bulge; conspicuous cloacal spurs, consisting of a cluster of white triangular scales, on the anterior-lateral margin (
<figureCitation id="136FD0E1FFDEFFCFFD66FA6BDB1BFA36" box="[688,778,1417,1442]" captionStart="FIGURE 4" captionStartId="7.[151,250,1444,1466]" captionTargetBox="[151,1436,411,1409]" captionTargetId="figure@7.[151,1436,411,1409]" captionTargetPageId="7" captionText="FIGURE 4. Holotypes of (A) P. pinnaclensis sp. nov. (QMJ96418) and (B) P. gulbaru (QMJ75778) (photo: Conrad Hoskin)." httpUri="https://zenodo.org/record/2624740/files/figure.png" pageId="3" pageNumber="130">Fig. 4A</figureCitation>
); axilla deeply invaginated.
<emphasis id="B9201076FFDEFFCFFB93FA6BDC83FA36" bold="true" box="[1093,1170,1417,1442]" italics="true" pageId="3" pageNumber="130">Limbs.</emphasis>
Long and very slender, covered in small pointed tubercles dorsally (
<figureCitation id="136FD0E1FFDEFFCFFD58FA4FDAFCFA52" box="[654,749,1453,1478]" captionStart="FIGURE 2" captionStartId="5.[151,250,1532,1554]" captionTargetBox="[151,1436,301,1511]" captionTargetId="figure@5.[151,1436,301,1511]" captionTargetPageId="5" captionText="FIGURE 2. (A) Holotype of P. pinnaclensis sp. nov. in life (photo: Conrad Hoskin). (B) P. pinnaclensis sp. nov. with regenerated tail (photo: Lorenzo Bertola)." httpUri="https://zenodo.org/record/2624736/files/figure.png" pageId="3" pageNumber="130">Figs 2A</figureCitation>
,
<figureCitation id="136FD0E1FFDEFFCFFD2DFA4CDB1BFA52" box="[763,778,1454,1478]" captionStart="FIGURE 3" captionStartId="6.[151,250,1036,1058]" captionTargetBox="[151,1436,193,1024]" captionTargetId="figure@6.[151,1436,193,1024]" captionTargetPageId="6" captionText="FIGURE 3. Holotype of P. pinnaclensis sp. nov. in life (photo: Conrad Hoskin)." httpUri="https://zenodo.org/record/2624738/files/figure.png" pageId="3" pageNumber="130">3</figureCitation>
,
<figureCitation id="136FD0E1FFDEFFCFFCCCFA4CDB2EFA52" box="[794,831,1454,1478]" captionStart="FIGURE 4" captionStartId="7.[151,250,1444,1466]" captionTargetBox="[151,1436,411,1409]" captionTargetId="figure@7.[151,1436,411,1409]" captionTargetPageId="7" captionText="FIGURE 4. Holotypes of (A) P. pinnaclensis sp. nov. (QMJ96418) and (B) P. gulbaru (QMJ75778) (photo: Conrad Hoskin)." httpUri="https://zenodo.org/record/2624740/files/figure.png" pageId="3" pageNumber="130">4A</figureCitation>
); lacking enlarged tubercles on ventral surface; digits strongly compressed distally; dorsal surface of hands, feet and digits without enlarged conical tubercles (
<figureCitation id="136FD0E1FFDEFFCFFA91FA33DD9FFA7E" box="[1351,1422,1489,1514]" captionStart="FIGURE 3" captionStartId="6.[151,250,1036,1058]" captionTargetBox="[151,1436,193,1024]" captionTargetId="figure@6.[151,1436,193,1024]" captionTargetPageId="6" captionText="FIGURE 3. Holotype of P. pinnaclensis sp. nov. in life (photo: Conrad Hoskin)." httpUri="https://zenodo.org/record/2624738/files/figure.png" pageId="3" pageNumber="130">Fig. 3</figureCitation>
).
<emphasis id="B9201076FFDEFFCFFF41FA17D939F99A" bold="true" box="[151,296,1525,1550]" italics="true" pageId="3" pageNumber="130">Original tail.</emphasis>
Long, cylindrical, tapering (
<figureCitation id="136FD0E1FFDEFFCFFDB6FA17DAADF99A" box="[608,700,1525,1550]" captionStart="FIGURE 2" captionStartId="5.[151,250,1532,1554]" captionTargetBox="[151,1436,301,1511]" captionTargetId="figure@5.[151,1436,301,1511]" captionTargetPageId="5" captionText="FIGURE 2. (A) Holotype of P. pinnaclensis sp. nov. in life (photo: Conrad Hoskin). (B) P. pinnaclensis sp. nov. with regenerated tail (photo: Lorenzo Bertola)." httpUri="https://zenodo.org/record/2624736/files/figure.png" pageId="3" pageNumber="130">Figs 2A</figureCitation>
,
<figureCitation id="136FD0E1FFDEFFCFFD11FA14DAFDF99A" box="[711,748,1526,1550]" captionStart="FIGURE 4" captionStartId="7.[151,250,1444,1466]" captionTargetBox="[151,1436,411,1409]" captionTargetId="figure@7.[151,1436,411,1409]" captionTargetPageId="7" captionText="FIGURE 4. Holotypes of (A) P. pinnaclensis sp. nov. (QMJ96418) and (B) P. gulbaru (QMJ75778) (photo: Conrad Hoskin)." httpUri="https://zenodo.org/record/2624740/files/figure.png" pageId="3" pageNumber="130">4A</figureCitation>
); slightly depressed at base; distal quarter of tail a very narrow attenuated tip, terminating with a minute, rounded, white 'knob'; dorsal surface covered in minute granules and fine tubercles; prominent enlarged spinose tubercles restricted to the basal quarter; ventral surface smooth with a mosaic of fine round scales.
<emphasis id="B9201076FFDEFFCFFEB5F983DA7DF9EE" bold="true" box="[355,620,1633,1658]" italics="true" pageId="3" pageNumber="130">Colour pattern in spirit</emphasis>
(
<figureCitation id="136FD0E1FFDEFFCFFDABF983DAC6F9EE" box="[637,727,1633,1658]" captionStart="FIGURE 4" captionStartId="7.[151,250,1444,1466]" captionTargetBox="[151,1436,411,1409]" captionTargetId="figure@7.[151,1436,411,1409]" captionTargetPageId="7" captionText="FIGURE 4. Holotypes of (A) P. pinnaclensis sp. nov. (QMJ96418) and (B) P. gulbaru (QMJ75778) (photo: Conrad Hoskin)." httpUri="https://zenodo.org/record/2624740/files/figure.png" pageId="3" pageNumber="130">Fig. 4A</figureCitation>
). Dorsal base colour tan, with irregular dark brown blotches on head, body and limbs; blotches largest on body and neck, smallest on head; eyelids bluish grey; pale band from lower anterior margin of eye diagonally to midway along jaw, and similar pale band from lower posterior margin of eye diagonally to just behind jaw; pale band anterior to base of tail, and inconspicuous, irregular, pale transverse bars on back; top of hands and feet pale; most digits strongly banded; inner anterior digit almost completely white; original tail finely mottled light brown, with five prominent, regular, bright white transverse bands and fine white knob on tip; all ventral surfaces pale cream, with a few light brown smudges on throat, underside of limbs and testicular bulge.
<emphasis id="B9201076FFDEFFCFFA9BF8DBD926F8E2" bold="true" italics="true" pageId="3" pageNumber="130">Colour pattern in life.</emphasis>
As above but base colour of body much lighter (whitish grey) and brown stippling on tail darker (
<figureCitation id="136FD0E1FFDEFFCFFABAF8BFD8AAF80E" captionStart="FIGURE 2" captionStartId="5.[151,250,1532,1554]" captionTargetBox="[151,1436,301,1511]" captionTargetId="figure@5.[151,1436,301,1511]" captionTargetPageId="5" captionText="FIGURE 2. (A) Holotype of P. pinnaclensis sp. nov. in life (photo: Conrad Hoskin). (B) P. pinnaclensis sp. nov. with regenerated tail (photo: Lorenzo Bertola)." httpUri="https://zenodo.org/record/2624736/files/figure.png" pageId="3" pageNumber="130">Figs 2A</figureCitation>
,
<figureCitation id="136FD0E1FFDEFFCFFF13F860D8C5F80E" box="[197,212,1922,1946]" captionStart="FIGURE 3" captionStartId="6.[151,250,1036,1058]" captionTargetBox="[151,1436,193,1024]" captionTargetId="figure@6.[151,1436,193,1024]" captionTargetPageId="6" captionText="FIGURE 3. Holotype of P. pinnaclensis sp. nov. in life (photo: Conrad Hoskin)." httpUri="https://zenodo.org/record/2624738/files/figure.png" pageId="3" pageNumber="130">3</figureCitation>
).
</paragraph>
<caption id="DF2B9CECFFD9FFC8FF41FF7ADCA3FF5B" pageId="4" pageNumber="131">
<paragraph id="8BEBCC64FFD9FFC8FF41FF7ADCA3FF5B" blockId="4.[151,1436,151,207]" pageId="4" pageNumber="131">
<emphasis id="B9201076FFD9FFC8FF41FF7AD91BFF3B" bold="true" box="[151,266,152,175]" pageId="4" pageNumber="131">TABLE 1.</emphasis>
Field data for select morphometric and scale traits for
<taxonomicName id="4C54B7E7FFD9FFC8FC8FFF7ADBE9FF3B" authority="Hoskin &amp; Bertola &amp; Higgie, 2019" authorityName="Hoskin &amp; Bertola &amp; Higgie" authorityYear="2019" box="[857,1016,151,175]" class="Reptilia" family="Gekkonidae" genus="Phyllurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="131" phylum="Chordata" rank="species" species="pinnaclensis" status="sp. nov.">
<emphasis id="B9201076FFD9FFC8FC8FFF7ADBE9FF3B" box="[857,1016,151,175]" italics="true" pageId="4" pageNumber="131">P. pinnaclensis</emphasis>
</taxonomicName>
<emphasis id="B9201076FFD9FFC8FBD6FF7ADC47FF3B" bold="true" box="[1024,1110,152,175]" pageId="4" pageNumber="131">
<taxonomicNameLabel id="A213AD0DFFD9FFC8FBD6FF7ADC47FF3B" box="[1024,1110,152,175]" pageId="4" pageNumber="131" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C54B7E7FFD9FFC8FB5DFF7ADCE8FF3B" box="[1163,1273,151,175]" class="Reptilia" family="Gekkonidae" genus="Phyllurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="131" phylum="Chordata" rank="species" species="gulbaru">
<emphasis id="B9201076FFD9FFC8FB5DFF7ADCE8FF3B" box="[1163,1273,151,175]" italics="true" pageId="4" pageNumber="131">P. gulbaru</emphasis>
</taxonomicName>
. Traits marked in bold have obviously different means and/or no overlap in range values between the two species.
</paragraph>
</caption>
<paragraph id="8BEBCC64FFD9FFC8FE23FF09DD5EFC7C" pageId="4" pageNumber="131">
<table id="F9543EC4FFD90033FF49FF08DD41FC7C" box="[159,1360,234,1000]" gridcols="7" gridrows="19" pageId="4" pageNumber="131">
<tr id="3564CE26FFD90033FF49FF08DD41FE94" box="[159,1360,234,256]" gridrow="0" pageId="4" pageNumber="131" rowspan-0="1" rowspan-3="1" rowspan-5="1" rowspan-6="1">
<th id="76B5A75AFFD90033FE23FF08DAE5FE94" box="[501,756,234,256]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="4" pageNumber="131">
<taxonomicName id="4C54B7E7FFD9FFC8FE23FF09DA9CFE94" authority="Hoskin &amp; Bertola &amp; Higgie, 2019" authorityName="Hoskin &amp; Bertola &amp; Higgie" authorityYear="2019" box="[501,653,235,256]" class="Reptilia" family="Gekkonidae" genus="Phyllurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="131" phylum="Chordata" rank="species" species="pinnaclensis" status="sp. nov.">
<emphasis id="B9201076FFD9FFC8FE23FF09DA9CFE94" box="[501,653,235,256]" italics="true" pageId="4" pageNumber="131">P. pinnaclensis</emphasis>
</taxonomicName>
<emphasis id="B9201076FFD9FFC8FD45FF08DAF2FE94" bold="true" box="[659,739,234,256]" pageId="4" pageNumber="131">
<taxonomicNameLabel id="A213AD0DFFD9FFC8FD45FF08DAF2FE94" box="[659,739,234,256]" pageId="4" pageNumber="131" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<th id="76B5A75AFFD90033FC14FF08DC3BFE94" box="[962,1066,234,256]" gridcol="4" gridrow="0" pageId="4" pageNumber="131">
<taxonomicName id="4C54B7E7FFD9FFC8FC14FF09DC3BFE94" box="[962,1066,235,256]" class="Reptilia" family="Gekkonidae" genus="Phyllurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="131" phylum="Chordata" rank="species" species="gulbaru">
<emphasis id="B9201076FFD9FFC8FC14FF09DC3BFE94" box="[962,1066,235,256]" italics="true" pageId="4" pageNumber="131">P. gulbaru</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="3564CE26FFD90033FF49FEF6DD41FEBE" box="[159,1360,276,298]" gridrow="1" pageId="4" pageNumber="131">
<th id="76B5A75AFFD90033FF49FEF6D99FFEBE" box="[159,398,276,298]" gridcol="0" gridrow="1" pageId="4" pageNumber="131">
<emphasis id="B9201076FFD9FFC8FF49FEF6D8C4FEBE" bold="true" box="[159,213,276,298]" pageId="4" pageNumber="131">Trait</emphasis>
</th>
<td id="76B5A75AFFD90033FE23FEF6DA21FEBE" box="[501,560,276,298]" gridcol="1" gridrow="1" pageId="4" pageNumber="131">
<emphasis id="B9201076FFD9FFC8FE23FEF6DA21FEBE" bold="true" box="[501,560,276,298]" pageId="4" pageNumber="131">mean</emphasis>
</td>
<td id="76B5A75AFFD90033FD58FEF6DAE5FEBE" box="[654,756,276,298]" gridcol="2" gridrow="1" pageId="4" pageNumber="131">
<emphasis id="B9201076FFD9FFC8FD58FEF6DADDFEBE" bold="true" box="[654,716,276,298]" pageId="4" pageNumber="131">range</emphasis>
</td>
<td id="76B5A75AFFD90033FCE2FEF6DB5FFEBE" box="[820,846,276,298]" gridcol="3" gridrow="1" pageId="4" pageNumber="131">
<emphasis id="B9201076FFD9FFC8FCE2FEF6DB57FEBE" bold="true" box="[820,838,276,298]" pageId="4" pageNumber="131">N</emphasis>
</td>
<td id="76B5A75AFFD90033FC14FEF6DC3BFEBE" box="[962,1066,276,298]" gridcol="4" gridrow="1" pageId="4" pageNumber="131">
<emphasis id="B9201076FFD9FFC8FC14FEF6DBECFEBE" bold="true" box="[962,1021,276,298]" pageId="4" pageNumber="131">mean</emphasis>
</td>
<td id="76B5A75AFFD90033FBB0FEF6DCDDFEBE" box="[1126,1228,276,298]" gridcol="5" gridrow="1" pageId="4" pageNumber="131">
<emphasis id="B9201076FFD9FFC8FBB0FEF6DCB5FEBE" bold="true" box="[1126,1188,276,298]" pageId="4" pageNumber="131">range</emphasis>
</td>
<td id="76B5A75AFFD90033FAE0FEF6DD41FEBE" box="[1334,1360,276,298]" gridcol="6" gridrow="1" pageId="4" pageNumber="131">
<emphasis id="B9201076FFD9FFC8FAE0FEF6DD59FEBE" bold="true" box="[1334,1352,276,298]" pageId="4" pageNumber="131">N</emphasis>
</td>
</tr>
<tr id="3564CE26FFD90033FF49FEDFDD41FEC7" box="[159,1360,317,339]" gridrow="2" pageId="4" pageNumber="131">
<th id="76B5A75AFFD90033FF49FEDFD99FFEC7" box="[159,398,317,339]" gridcol="0" gridrow="2" pageId="4" pageNumber="131">SVL (mm)</th>
<td id="76B5A75AFFD90033FE23FEDFDA21FEC7" box="[501,560,317,339]" gridcol="1" gridrow="2" pageId="4" pageNumber="131">90.0</td>
<td id="76B5A75AFFD90033FD58FEDFDAE5FEC7" box="[654,756,317,339]" gridcol="2" gridrow="2" pageId="4" pageNumber="131">8793</td>
<td id="76B5A75AFFD90033FCE2FEDFDB5FFEC7" box="[820,846,317,339]" gridcol="3" gridrow="2" pageId="4" pageNumber="131">7</td>
<td id="76B5A75AFFD90033FC14FEDFDC3BFEC7" box="[962,1066,317,339]" gridcol="4" gridrow="2" pageId="4" pageNumber="131">90.1</td>
<td id="76B5A75AFFD90033FBB0FEDFDCDDFEC7" box="[1126,1228,317,339]" gridcol="5" gridrow="2" pageId="4" pageNumber="131">7798</td>
<td id="76B5A75AFFD90033FAE0FEDFDD41FEC7" box="[1334,1360,317,339]" gridcol="6" gridrow="2" pageId="4" pageNumber="131">22</td>
</tr>
<tr id="3564CE26FFD90033FF49FE84DD41FEE8" box="[159,1360,358,380]" gridrow="3" pageId="4" pageNumber="131">
<th id="76B5A75AFFD90033FF49FE84D99FFEE8" box="[159,398,358,380]" gridcol="0" gridrow="3" pageId="4" pageNumber="131">AG/SVL</th>
<td id="76B5A75AFFD90033FE23FE84DA21FEE8" box="[501,560,358,380]" gridcol="1" gridrow="3" pageId="4" pageNumber="131">0.46</td>
<td id="76B5A75AFFD90033FD58FE84DAE5FEE8" box="[654,756,358,380]" gridcol="2" gridrow="3" pageId="4" pageNumber="131">0.430.49</td>
<td id="76B5A75AFFD90033FCE2FE84DB5FFEE8" box="[820,846,358,380]" gridcol="3" gridrow="3" pageId="4" pageNumber="131">2</td>
<td id="76B5A75AFFD90033FC14FE84DC3BFEE8" box="[962,1066,358,380]" gridcol="4" gridrow="3" pageId="4" pageNumber="131">0.44</td>
<td id="76B5A75AFFD90033FBB0FE84DCDDFEE8" box="[1126,1228,358,380]" gridcol="5" gridrow="3" pageId="4" pageNumber="131">0.410.50</td>
<td id="76B5A75AFFD90033FAE0FE84DD41FEE8" box="[1334,1360,358,380]" gridcol="6" gridrow="3" pageId="4" pageNumber="131">18</td>
</tr>
<tr id="3564CE26FFD90033FF49FE72DD41FE32" box="[159,1360,400,422]" gridrow="4" pageId="4" pageNumber="131">
<th id="76B5A75AFFD90033FF49FE72D99FFE32" box="[159,398,400,422]" gridcol="0" gridrow="4" pageId="4" pageNumber="131">FL/SVL</th>
<td id="76B5A75AFFD90033FE23FE72DA21FE32" box="[501,560,400,422]" gridcol="1" gridrow="4" pageId="4" pageNumber="131">0.19</td>
<td id="76B5A75AFFD90033FD58FE72DAE5FE32" box="[654,756,400,422]" gridcol="2" gridrow="4" pageId="4" pageNumber="131">0.180.20</td>
<td id="76B5A75AFFD90033FCE2FE72DB5FFE32" box="[820,846,400,422]" gridcol="3" gridrow="4" pageId="4" pageNumber="131">8</td>
<td id="76B5A75AFFD90033FC14FE72DC3BFE32" box="[962,1066,400,422]" gridcol="4" gridrow="4" pageId="4" pageNumber="131">0.18</td>
<td id="76B5A75AFFD90033FBB0FE72DCDDFE32" box="[1126,1228,400,422]" gridcol="5" gridrow="4" pageId="4" pageNumber="131">0.160.19</td>
<td id="76B5A75AFFD90033FAE0FE72DD41FE32" box="[1334,1360,400,422]" gridcol="6" gridrow="4" pageId="4" pageNumber="131">18</td>
</tr>
<tr id="3564CE26FFD90033FF49FE5BDD41FE5B" box="[159,1360,441,463]" gridrow="5" pageId="4" pageNumber="131">
<th id="76B5A75AFFD90033FF49FE5BD99FFE5B" box="[159,398,441,463]" gridcol="0" gridrow="5" pageId="4" pageNumber="131">HLL/SVL</th>
<td id="76B5A75AFFD90033FE23FE5BDA21FE5B" box="[501,560,441,463]" gridcol="1" gridrow="5" pageId="4" pageNumber="131">0.21</td>
<td id="76B5A75AFFD90033FD58FE5BDAE5FE5B" box="[654,756,441,463]" gridcol="2" gridrow="5" pageId="4" pageNumber="131">0.200.22</td>
<td id="76B5A75AFFD90033FCE2FE5BDB5FFE5B" box="[820,846,441,463]" gridcol="3" gridrow="5" pageId="4" pageNumber="131">8</td>
<td id="76B5A75AFFD90033FC14FE5BDC3BFE5B" box="[962,1066,441,463]" gridcol="4" gridrow="5" pageId="4" pageNumber="131">0.21</td>
<td id="76B5A75AFFD90033FBB0FE5BDCDDFE5B" box="[1126,1228,441,463]" gridcol="5" gridrow="5" pageId="4" pageNumber="131">0.200.21</td>
<td id="76B5A75AFFD90033FAE0FE5BDD41FE5B" box="[1334,1360,441,463]" gridcol="6" gridrow="5" pageId="4" pageNumber="131">18</td>
</tr>
<tr id="3564CE26FFD90033FF49FE00DD41FE6C" box="[159,1360,482,504]" gridrow="6" pageId="4" pageNumber="131">
<th id="76B5A75AFFD90033FF49FE00D99FFE6C" box="[159,398,482,504]" gridcol="0" gridrow="6" pageId="4" pageNumber="131">
<emphasis id="B9201076FFD9FFC8FF49FE00D91CFE6C" bold="true" box="[159,269,482,504]" pageId="4" pageNumber="131">T(O)/SVL</emphasis>
</th>
<td id="76B5A75AFFD90033FE23FE00DA21FE6C" box="[501,560,482,504]" gridcol="1" gridrow="6" pageId="4" pageNumber="131">
<emphasis id="B9201076FFD9FFC8FE23FE00DA30FE6C" bold="true" box="[501,545,482,504]" pageId="4" pageNumber="131">0.90</emphasis>
</td>
<td id="76B5A75AFFD90033FD58FE00DAE5FE6C" box="[654,756,482,504]" gridcol="2" gridrow="6" pageId="4" pageNumber="131">
<emphasis id="B9201076FFD9FFC8FD58FE00DAE5FE6C" bold="true" box="[654,756,482,504]" pageId="4" pageNumber="131">0.880.91</emphasis>
</td>
<td id="76B5A75AFFD90033FCE2FE00DB5FFE6C" box="[820,846,482,504]" gridcol="3" gridrow="6" pageId="4" pageNumber="131">5</td>
<td id="76B5A75AFFD90033FC14FE00DC3BFE6C" box="[962,1066,482,504]" gridcol="4" gridrow="6" pageId="4" pageNumber="131">
<emphasis id="B9201076FFD9FFC8FC14FE00DBFFFE6C" bold="true" box="[962,1006,482,504]" pageId="4" pageNumber="131">0.80</emphasis>
</td>
<td id="76B5A75AFFD90033FBB0FE00DCDDFE6C" box="[1126,1228,482,504]" gridcol="5" gridrow="6" pageId="4" pageNumber="131">
<emphasis id="B9201076FFD9FFC8FBB0FE00DCDDFE6C" bold="true" box="[1126,1228,482,504]" pageId="4" pageNumber="131">0.710.84</emphasis>
</td>
<td id="76B5A75AFFD90033FAE0FE00DD41FE6C" box="[1334,1360,482,504]" gridcol="6" gridrow="6" pageId="4" pageNumber="131">11</td>
</tr>
<tr id="3564CE26FFD90033FF49FDEEDD41FDB6" box="[159,1360,524,546]" gridrow="7" pageId="4" pageNumber="131">
<th id="76B5A75AFFD90033FF49FDEED99FFDB6" box="[159,398,524,546]" gridcol="0" gridrow="7" pageId="4" pageNumber="131">
<emphasis id="B9201076FFD9FFC8FF49FDEED91DFDB6" bold="true" box="[159,268,524,546]" pageId="4" pageNumber="131">T(R)/SVL</emphasis>
</th>
<td id="76B5A75AFFD90033FE23FDEEDA21FDB6" box="[501,560,524,546]" gridcol="1" gridrow="7" pageId="4" pageNumber="131">
<emphasis id="B9201076FFD9FFC8FE23FDEEDA30FDB6" bold="true" box="[501,545,524,546]" pageId="4" pageNumber="131">0.73</emphasis>
</td>
<td id="76B5A75AFFD90033FD58FDEEDAE5FDB6" box="[654,756,524,546]" gridcol="2" gridrow="7" pageId="4" pageNumber="131">
<emphasis id="B9201076FFD9FFC8FD58FDEEDAE2FDB6" bold="true" box="[654,755,524,546]" pageId="4" pageNumber="131">0.730.73</emphasis>
</td>
<td id="76B5A75AFFD90033FCE2FDEEDB5FFDB6" box="[820,846,524,546]" gridcol="3" gridrow="7" pageId="4" pageNumber="131">2</td>
<td id="76B5A75AFFD90033FC14FDEEDC3BFDB6" box="[962,1066,524,546]" gridcol="4" gridrow="7" pageId="4" pageNumber="131">
<emphasis id="B9201076FFD9FFC8FC14FDEEDBFFFDB6" bold="true" box="[962,1006,524,546]" pageId="4" pageNumber="131">0.61</emphasis>
</td>
<td id="76B5A75AFFD90033FBB0FDEEDCDDFDB6" box="[1126,1228,524,546]" gridcol="5" gridrow="7" pageId="4" pageNumber="131">
<emphasis id="B9201076FFD9FFC8FBB0FDEEDCDDFDB6" bold="true" box="[1126,1228,524,546]" pageId="4" pageNumber="131">0.500.71</emphasis>
</td>
<td id="76B5A75AFFD90033FAE0FDEEDD41FDB6" box="[1334,1360,524,546]" gridcol="6" gridrow="7" pageId="4" pageNumber="131">13</td>
</tr>
<tr id="3564CE26FFD90033FF49FDD7DD41FDDF" box="[159,1360,565,587]" gridrow="8" pageId="4" pageNumber="131">
<th id="76B5A75AFFD90033FF49FDD7D99FFDDF" box="[159,398,565,587]" gridcol="0" gridrow="8" pageId="4" pageNumber="131">TW(O)/TD(O)</th>
<td id="76B5A75AFFD90033FE23FDD7DA21FDDF" box="[501,560,565,587]" gridcol="1" gridrow="8" pageId="4" pageNumber="131">1.80</td>
<td id="76B5A75AFFD90033FD58FDD7DAE5FDDF" box="[654,756,565,587]" gridcol="2" gridrow="8" pageId="4" pageNumber="131">1.532.06</td>
<td id="76B5A75AFFD90033FCE2FDD7DB5FFDDF" box="[820,846,565,587]" gridcol="3" gridrow="8" pageId="4" pageNumber="131">2</td>
<td id="76B5A75AFFD90033FC14FDD7DC3BFDDF" box="[962,1066,565,587]" gridcol="4" gridrow="8" pageId="4" pageNumber="131">1.60</td>
<td id="76B5A75AFFD90033FBB0FDD7DCDDFDDF" box="[1126,1228,565,587]" gridcol="5" gridrow="8" pageId="4" pageNumber="131">1.342.06</td>
<td id="76B5A75AFFD90033FAE0FDD7DD41FDDF" box="[1334,1360,565,587]" gridcol="6" gridrow="8" pageId="4" pageNumber="131">10</td>
</tr>
<tr id="3564CE26FFD90033FF49FDBCDD41FDE0" box="[159,1360,606,628]" gridrow="9" pageId="4" pageNumber="131">
<th id="76B5A75AFFD90033FF49FDBCD99FFDE0" box="[159,398,606,628]" gridcol="0" gridrow="9" pageId="4" pageNumber="131">HW/HL</th>
<td id="76B5A75AFFD90033FE23FDBCDA21FDE0" box="[501,560,606,628]" gridcol="1" gridrow="9" pageId="4" pageNumber="131">0.85</td>
<td id="76B5A75AFFD90033FD58FDBCDAE5FDE0" box="[654,756,606,628]" gridcol="2" gridrow="9" pageId="4" pageNumber="131">0.840.86</td>
<td id="76B5A75AFFD90033FCE2FDBCDB5FFDE0" box="[820,846,606,628]" gridcol="3" gridrow="9" pageId="4" pageNumber="131">2</td>
<td id="76B5A75AFFD90033FC14FDBCDC3BFDE0" box="[962,1066,606,628]" gridcol="4" gridrow="9" pageId="4" pageNumber="131">0.81</td>
<td id="76B5A75AFFD90033FBB0FDBCDCDDFDE0" box="[1126,1228,606,628]" gridcol="5" gridrow="9" pageId="4" pageNumber="131">0.760.88</td>
<td id="76B5A75AFFD90033FAE0FDBCDD41FDE0" box="[1334,1360,606,628]" gridcol="6" gridrow="9" pageId="4" pageNumber="131">18</td>
</tr>
<tr id="3564CE26FFD90033FF49FD6ADD41FD0A" box="[159,1360,648,670]" gridrow="10" pageId="4" pageNumber="131">
<th id="76B5A75AFFD90033FF49FD6AD99FFD0A" box="[159,398,648,670]" gridcol="0" gridrow="10" pageId="4" pageNumber="131">HD/HL</th>
<td id="76B5A75AFFD90033FE23FD6ADA21FD0A" box="[501,560,648,670]" gridcol="1" gridrow="10" pageId="4" pageNumber="131">0.39</td>
<td id="76B5A75AFFD90033FD58FD6ADAE5FD0A" box="[654,756,648,670]" gridcol="2" gridrow="10" pageId="4" pageNumber="131">0.390.39</td>
<td id="76B5A75AFFD90033FCE2FD6ADB5FFD0A" box="[820,846,648,670]" gridcol="3" gridrow="10" pageId="4" pageNumber="131">2</td>
<td id="76B5A75AFFD90033FC14FD6ADC3BFD0A" box="[962,1066,648,670]" gridcol="4" gridrow="10" pageId="4" pageNumber="131">0.36</td>
<td id="76B5A75AFFD90033FBB0FD6ADCDDFD0A" box="[1126,1228,648,670]" gridcol="5" gridrow="10" pageId="4" pageNumber="131">0.320.41</td>
<td id="76B5A75AFFD90033FAE0FD6ADD41FD0A" box="[1334,1360,648,670]" gridcol="6" gridrow="10" pageId="4" pageNumber="131">18</td>
</tr>
<tr id="3564CE26FFD90033FF49FD53DD41FD53" box="[159,1360,689,711]" gridrow="11" pageId="4" pageNumber="131">
<th id="76B5A75AFFD90033FF49FD53D99FFD53" box="[159,398,689,711]" gridcol="0" gridrow="11" pageId="4" pageNumber="131">S/HL</th>
<td id="76B5A75AFFD90033FE23FD53DA21FD53" box="[501,560,689,711]" gridcol="1" gridrow="11" pageId="4" pageNumber="131">0.13</td>
<td id="76B5A75AFFD90033FD58FD53DAE5FD53" box="[654,756,689,711]" gridcol="2" gridrow="11" pageId="4" pageNumber="131">0.130.13</td>
<td id="76B5A75AFFD90033FCE2FD53DB5FFD53" box="[820,846,689,711]" gridcol="3" gridrow="11" pageId="4" pageNumber="131">2</td>
<td id="76B5A75AFFD90033FC14FD53DC3BFD53" box="[962,1066,689,711]" gridcol="4" gridrow="11" pageId="4" pageNumber="131">0.13</td>
<td id="76B5A75AFFD90033FBB0FD53DCDDFD53" box="[1126,1228,689,711]" gridcol="5" gridrow="11" pageId="4" pageNumber="131">0.110.14</td>
<td id="76B5A75AFFD90033FAE0FD53DD41FD53" box="[1334,1360,689,711]" gridcol="6" gridrow="11" pageId="4" pageNumber="131">18</td>
</tr>
<tr id="3564CE26FFD90033FF49FD38DD41FD64" box="[159,1360,730,752]" gridrow="12" pageId="4" pageNumber="131">
<th id="76B5A75AFFD90033FF49FD38D99FFD64" box="[159,398,730,752]" gridcol="0" gridrow="12" pageId="4" pageNumber="131">
<emphasis id="B9201076FFD9FFC8FF49FD38D8EBFD64" bold="true" box="[159,250,730,752]" pageId="4" pageNumber="131">NL/SVL</emphasis>
</th>
<td id="76B5A75AFFD90033FE23FD38DA21FD64" box="[501,560,730,752]" gridcol="1" gridrow="12" pageId="4" pageNumber="131">
<emphasis id="B9201076FFD9FFC8FE23FD38DA30FD64" bold="true" box="[501,545,730,752]" pageId="4" pageNumber="131">0.17</emphasis>
</td>
<td id="76B5A75AFFD90033FD58FD38DAE5FD64" box="[654,756,730,752]" gridcol="2" gridrow="12" pageId="4" pageNumber="131">
<emphasis id="B9201076FFD9FFC8FD58FD38DAE2FD64" bold="true" box="[654,755,730,752]" pageId="4" pageNumber="131">0.170.18</emphasis>
</td>
<td id="76B5A75AFFD90033FCE2FD38DB5FFD64" box="[820,846,730,752]" gridcol="3" gridrow="12" pageId="4" pageNumber="131">2</td>
<td id="76B5A75AFFD90033FC14FD38DC3BFD64" box="[962,1066,730,752]" gridcol="4" gridrow="12" pageId="4" pageNumber="131">
<emphasis id="B9201076FFD9FFC8FC14FD38DBFFFD64" bold="true" box="[962,1006,730,752]" pageId="4" pageNumber="131">0.20</emphasis>
</td>
<td id="76B5A75AFFD90033FBB0FD38DCDDFD64" box="[1126,1228,730,752]" gridcol="5" gridrow="12" pageId="4" pageNumber="131">
<emphasis id="B9201076FFD9FFC8FBB0FD38DCDAFD64" bold="true" box="[1126,1227,730,752]" pageId="4" pageNumber="131">0.180.23</emphasis>
</td>
<td id="76B5A75AFFD90033FAE0FD38DD41FD64" box="[1334,1360,730,752]" gridcol="6" gridrow="12" pageId="4" pageNumber="131">18</td>
</tr>
<tr id="3564CE26FFD90033FF49FCE6DD41FC8E" box="[159,1360,772,794]" gridrow="13" pageId="4" pageNumber="131">
<th id="76B5A75AFFD90033FF49FCE6D99FFC8E" box="[159,398,772,794]" gridcol="0" gridrow="13" pageId="4" pageNumber="131">
<emphasis id="B9201076FFD9FFC8FF49FCE6D97AFC8E" bold="true" box="[159,363,772,794]" pageId="4" pageNumber="131">Internasals (count)</emphasis>
</th>
<td id="76B5A75AFFD90033FE23FCE6DA21FC8E" box="[501,560,772,794]" gridcol="1" gridrow="13" pageId="4" pageNumber="131">
<emphasis id="B9201076FFD9FFC8FE23FCE6DA04FC8E" bold="true" box="[501,533,772,794]" pageId="4" pageNumber="131">9.1</emphasis>
</td>
<td id="76B5A75AFFD90033FD58FCE6DAE5FC8E" box="[654,756,772,794]" gridcol="2" gridrow="13" pageId="4" pageNumber="131">
<emphasis id="B9201076FFD9FFC8FD58FCE6DAD0FC8E" bold="true" box="[654,705,772,794]" pageId="4" pageNumber="131">710</emphasis>
</td>
<td id="76B5A75AFFD90033FCE2FCE6DB5FFC8E" box="[820,846,772,794]" gridcol="3" gridrow="13" pageId="4" pageNumber="131">10</td>
<td id="76B5A75AFFD90033FC14FCE6DC3BFC8E" box="[962,1066,772,794]" gridcol="4" gridrow="13" pageId="4" pageNumber="131">
<emphasis id="B9201076FFD9FFC8FC14FCE6DBF3FC8E" bold="true" box="[962,994,772,794]" pageId="4" pageNumber="131">6.5</emphasis>
</td>
<td id="76B5A75AFFD90033FBB0FCE6DCDDFC8E" box="[1126,1228,772,794]" gridcol="5" gridrow="13" pageId="4" pageNumber="131">
<emphasis id="B9201076FFD9FFC8FBB0FCE6DC9DFC8E" bold="true" box="[1126,1164,772,794]" pageId="4" pageNumber="131">58</emphasis>
</td>
<td id="76B5A75AFFD90033FAE0FCE6DD41FC8E" box="[1334,1360,772,794]" gridcol="6" gridrow="13" pageId="4" pageNumber="131">36</td>
</tr>
<tr id="3564CE26FFD90033FF49FCCFDD41FCD7" box="[159,1360,813,835]" gridrow="14" pageId="4" pageNumber="131">
<th id="76B5A75AFFD90033FF49FCCFD99FFCD7" box="[159,398,813,835]" gridcol="0" gridrow="14" pageId="4" pageNumber="131">Rostral groove (%)</th>
<td id="76B5A75AFFD90033FE23FCCFDA21FCD7" box="[501,560,813,835]" gridcol="1" gridrow="14" pageId="4" pageNumber="131">0.45</td>
<td id="76B5A75AFFD90033FD58FCCFDAE5FCD7" box="[654,756,813,835]" gridcol="2" gridrow="14" pageId="4" pageNumber="131">0.250.60</td>
<td id="76B5A75AFFD90033FCE2FCCFDB5FFCD7" box="[820,846,813,835]" gridcol="3" gridrow="14" pageId="4" pageNumber="131">9</td>
<td id="76B5A75AFFD90033FC14FCCFDC3BFCD7" box="[962,1066,813,835]" gridcol="4" gridrow="14" pageId="4" pageNumber="131">0.44</td>
<td id="76B5A75AFFD90033FBB0FCCFDCDDFCD7" box="[1126,1228,813,835]" gridcol="5" gridrow="14" pageId="4" pageNumber="131">0.100.75</td>
<td id="76B5A75AFFD90033FAE0FCCFDD41FCD7" box="[1334,1360,813,835]" gridcol="6" gridrow="14" pageId="4" pageNumber="131">36</td>
</tr>
<tr id="3564CE26FFD90033FF49FCB4DD41FCF8" box="[159,1360,854,876]" gridrow="15" pageId="4" pageNumber="131">
<th id="76B5A75AFFD90033FF49FCB4D99FFCF8" box="[159,398,854,876]" gridcol="0" gridrow="15" pageId="4" pageNumber="131">Supralabials (count)</th>
<td id="76B5A75AFFD90033FE23FCB4DA21FCF8" box="[501,560,854,876]" gridcol="1" gridrow="15" pageId="4" pageNumber="131">15.3</td>
<td id="76B5A75AFFD90033FD58FCB4DAE5FCF8" box="[654,756,854,876]" gridcol="2" gridrow="15" pageId="4" pageNumber="131">1417</td>
<td id="76B5A75AFFD90033FCE2FCB4DB5FFCF8" box="[820,846,854,876]" gridcol="3" gridrow="15" pageId="4" pageNumber="131">9</td>
<td id="76B5A75AFFD90033FC14FCB4DC3BFCF8" box="[962,1066,854,876]" gridcol="4" gridrow="15" pageId="4" pageNumber="131">13.5</td>
<td id="76B5A75AFFD90033FBB0FCB4DCDDFCF8" box="[1126,1228,854,876]" gridcol="5" gridrow="15" pageId="4" pageNumber="131">1115</td>
<td id="76B5A75AFFD90033FAE0FCB4DD41FCF8" box="[1334,1360,854,876]" gridcol="6" gridrow="15" pageId="4" pageNumber="131">32</td>
</tr>
<tr id="3564CE26FFD90033FF49FC62DD41FC02" box="[159,1360,896,918]" gridrow="16" pageId="4" pageNumber="131">
<th id="76B5A75AFFD90033FF49FC62D99FFC02" box="[159,398,896,918]" gridcol="0" gridrow="16" pageId="4" pageNumber="131">Infralabials (count)</th>
<td id="76B5A75AFFD90033FE23FC62DA21FC02" box="[501,560,896,918]" gridcol="1" gridrow="16" pageId="4" pageNumber="131">15.9</td>
<td id="76B5A75AFFD90033FD58FC62DAE5FC02" box="[654,756,896,918]" gridcol="2" gridrow="16" pageId="4" pageNumber="131">1517</td>
<td id="76B5A75AFFD90033FCE2FC62DB5FFC02" box="[820,846,896,918]" gridcol="3" gridrow="16" pageId="4" pageNumber="131">9</td>
<td id="76B5A75AFFD90033FC14FC62DC3BFC02" box="[962,1066,896,918]" gridcol="4" gridrow="16" pageId="4" pageNumber="131">14.3</td>
<td id="76B5A75AFFD90033FBB0FC62DCDDFC02" box="[1126,1228,896,918]" gridcol="5" gridrow="16" pageId="4" pageNumber="131">1216</td>
<td id="76B5A75AFFD90033FAE0FC62DD41FC02" box="[1334,1360,896,918]" gridcol="6" gridrow="16" pageId="4" pageNumber="131">32</td>
</tr>
<tr id="3564CE26FFD90033FF49FC4BDD41FC2B" box="[159,1360,937,959]" gridrow="17" pageId="4" pageNumber="131">
<th id="76B5A75AFFD90033FF49FC4BD99FFC2B" box="[159,398,937,959]" gridcol="0" gridrow="17" pageId="4" pageNumber="131">
<emphasis id="B9201076FFD9FFC8FF49FC4BD99FFC2B" bold="true" box="[159,398,937,959]" pageId="4" pageNumber="131">Subdig 4th toe (count)</emphasis>
</th>
<td id="76B5A75AFFD90033FE23FC4BDA21FC2B" box="[501,560,937,959]" gridcol="1" gridrow="17" pageId="4" pageNumber="131">
<emphasis id="B9201076FFD9FFC8FE23FC4BDA1EFC2B" bold="true" box="[501,527,937,959]" pageId="4" pageNumber="131">20</emphasis>
</td>
<td id="76B5A75AFFD90033FD58FC4BDAE5FC2B" box="[654,756,937,959]" gridcol="2" gridrow="17" pageId="4" pageNumber="131">
<emphasis id="B9201076FFD9FFC8FD58FC4BDADCFC2B" bold="true" box="[654,717,937,959]" pageId="4" pageNumber="131">1921</emphasis>
</td>
<td id="76B5A75AFFD90033FCE2FC4BDB5FFC2B" box="[820,846,937,959]" gridcol="3" gridrow="17" pageId="4" pageNumber="131">8</td>
<td id="76B5A75AFFD90033FC14FC4BDC3BFC2B" box="[962,1066,937,959]" gridcol="4" gridrow="17" pageId="4" pageNumber="131">
<emphasis id="B9201076FFD9FFC8FC14FC4BDBFFFC2B" bold="true" box="[962,1006,937,959]" pageId="4" pageNumber="131">18.0</emphasis>
</td>
<td id="76B5A75AFFD90033FBB0FC4BDCDDFC2B" box="[1126,1228,937,959]" gridcol="5" gridrow="17" pageId="4" pageNumber="131">
<emphasis id="B9201076FFD9FFC8FBB0FC4BDCB7FC2B" bold="true" box="[1126,1190,937,959]" pageId="4" pageNumber="131">1619</emphasis>
</td>
<td id="76B5A75AFFD90033FAE0FC4BDD41FC2B" box="[1334,1360,937,959]" gridcol="6" gridrow="17" pageId="4" pageNumber="131">28</td>
</tr>
<tr id="3564CE26FFD90033FF49FC30DD41FC7C" box="[159,1360,978,1000]" gridrow="18" pageId="4" pageNumber="131">
<th id="76B5A75AFFD90033FF49FC30D99FFC7C" box="[159,398,978,1000]" gridcol="0" gridrow="18" pageId="4" pageNumber="131">Tail bands (count)</th>
<td id="76B5A75AFFD90033FE23FC30DA21FC7C" box="[501,560,978,1000]" gridcol="1" gridrow="18" pageId="4" pageNumber="131">5.4</td>
<td id="76B5A75AFFD90033FD58FC30DAE5FC7C" box="[654,756,978,1000]" gridcol="2" gridrow="18" pageId="4" pageNumber="131">57</td>
<td id="76B5A75AFFD90033FCE2FC30DB5FFC7C" box="[820,846,978,1000]" gridcol="3" gridrow="18" pageId="4" pageNumber="131">7</td>
<td id="76B5A75AFFD90033FC14FC30DC3BFC7C" box="[962,1066,978,1000]" gridcol="4" gridrow="18" pageId="4" pageNumber="131">6.9</td>
<td id="76B5A75AFFD90033FBB0FC30DCDDFC7C" box="[1126,1228,978,1000]" gridcol="5" gridrow="18" pageId="4" pageNumber="131">510</td>
<td id="76B5A75AFFD90033FAE0FC30DD41FC7C" box="[1334,1360,978,1000]" gridcol="6" gridrow="18" pageId="4" pageNumber="131">14</td>
</tr>
</table>
</paragraph>
<paragraph id="8BEBCC64FFD9FFC8FF41FBEBD900FB14" blockId="4.[151,1435,1033,1152]" pageId="4" pageNumber="131">Note: Abbreviations of measurements are: SVL, Snout to vent length; AG, axilla to groin length; FL, forearm length; HLL, lower hindlimb length; T(O), original tail length; T(R), regenerated tail length; TW(O), original tail width; TD(O), original tail depth; HW, head width; HL, head length; HD, head depth; S, snout length; NL, neck length. See Methods for definitions.</paragraph>
<paragraph id="8BEBCC64FFD9FFC8FF11FB33DD8DFAEE" blockId="4.[151,1436,1233,1402]" pageId="4" pageNumber="131">
<emphasis id="B9201076FFD9FFC8FF11FB33D957FB7E" bold="true" box="[199,326,1233,1258]" pageId="4" pageNumber="131">Field data.</emphasis>
Data was collected from an additional nine individuals in the field.
<emphasis id="B9201076FFD9FFC8FB97FB33DD8AFB7E" bold="true" box="[1089,1435,1233,1258]" italics="true" pageId="4" pageNumber="131">Morphometrics and scalation.</emphasis>
See
<tableCitation id="C6D6F9DFFFD9FFC8FF1CFB17D930FA9A" box="[202,289,1269,1294]" captionStart="TABLE 1" captionStartId="4.[151,239,152,175]" captionText="TABLE 1. Field data for select morphometric and scale traits for P. pinnaclensis sp. nov. and P. gulbaru. Traits marked in bold have obviously different means and/or no overlap in range values between the two species." pageId="4" pageNumber="131">Table 1</tableCitation>
.
<emphasis id="B9201076FFD9FFC8FEE7FB16D9D8FA99" bold="true" box="[305,457,1268,1293]" italics="true" pageId="4" pageNumber="131">Original tail.</emphasis>
Shape as for
<typeStatus id="54EF72C6FFD9FFC8FDB8FB17DAC1FA9A" box="[622,720,1269,1294]" pageId="4" pageNumber="131" type="holotype">holotype</typeStatus>
but slightly more depressed at base in some individuals. Tail colouration generally as for
<typeStatus id="54EF72C6FFD9FFC8FE0FFAFBDA2FFAA6" box="[473,574,1305,1330]" pageId="4" pageNumber="131" type="holotype">holotype</typeStatus>
, with all (N = 7) individuals having five prominent white bands (excluding white tip). Two of these individuals also had one or two less conspicuous thin, pale bands on tail. In most individuals, base colour (between the white bands) is darker than for
<typeStatus id="54EF72C6FFD9FFC8FBD5FA83DC79FAEE" box="[1027,1128,1377,1402]" pageId="4" pageNumber="131" type="holotype">holotype</typeStatus>
, generally almost black.
</paragraph>
<paragraph id="8BEBCC64FFD9FFC8FF41FA41D8F0F901" blockId="4.[151,1437,1443,2011]" pageId="4" pageNumber="131">
<emphasis id="B9201076FFD9FFC8FF41FA41D948FA28" bold="true" box="[151,345,1443,1468]" italics="true" pageId="4" pageNumber="131">Regenerated tail</emphasis>
(
<figureCitation id="136FD0E1FFD9FFC8FEBAFA46D9D8FA28" box="[364,457,1444,1469]" captionStart="FIGURE 2" captionStartId="5.[151,250,1532,1554]" captionTargetBox="[151,1436,301,1511]" captionTargetId="figure@5.[151,1436,301,1511]" captionTargetPageId="5" captionText="FIGURE 2. (A) Holotype of P. pinnaclensis sp. nov. in life (photo: Conrad Hoskin). (B) P. pinnaclensis sp. nov. with regenerated tail (photo: Lorenzo Bertola)." httpUri="https://zenodo.org/record/2624736/files/figure.png" pageId="4" pageNumber="131">Fig. 2B</figureCitation>
). Slightly depressed, cylindrical, bulbous at base and tapering to tip; covered with uniform granules aligned in fine concentric rings for the full length of the tail; no spinose tubercles. Regenerated tail heavily mottled dark brown on a tan base colour.
<emphasis id="B9201076FFD9FFC8FD2AFA09DBBEF997" bold="true" box="[764,943,1515,1540]" italics="true" pageId="4" pageNumber="131">Colour pattern.</emphasis>
As described for the
<typeStatus id="54EF72C6FFD9FFC8FB7EFA0EDD1BF991" box="[1192,1290,1516,1541]" pageId="4" pageNumber="131" type="holotype">holotype</typeStatus>
but with the following variation. Base colour of some individuals is brown, giving the whole animal a much darker appearance (e.g.,
<figureCitation id="136FD0E1FFD9FFC8FF0EF9D6D920F9D8" box="[216,305,1588,1613]" captionStart="FIGURE 2" captionStartId="5.[151,250,1532,1554]" captionTargetBox="[151,1436,301,1511]" captionTargetId="figure@5.[151,1436,301,1511]" captionTargetPageId="5" captionText="FIGURE 2. (A) Holotype of P. pinnaclensis sp. nov. in life (photo: Conrad Hoskin). (B) P. pinnaclensis sp. nov. with regenerated tail (photo: Lorenzo Bertola)." httpUri="https://zenodo.org/record/2624736/files/figure.png" pageId="4" pageNumber="131">Fig. 2B</figureCitation>
). Some individuals, have prominent pale spots or blotches on the dorsal surfaces of the body, head and limbs; in juveniles and subadults these are bright white. Variation in tail colour pattern described immediately above.
</paragraph>
<paragraph id="8BEBCC64FFD9FFC9FF11F97DD9FEFF6C" blockId="4.[151,1437,1443,2011]" lastBlockId="5.[151,1437,151,248]" lastPageId="5" lastPageNumber="132" pageId="4" pageNumber="131">
<emphasis id="B9201076FFD9FFC8FF11F97DD970F92C" bold="true" box="[199,353,1695,1720]" pageId="4" pageNumber="131">Comparison.</emphasis>
<taxonomicName id="4C54B7E7FFD9FFC8FEB8F942DA62F92C" authority="Hoskin &amp; Bertola &amp; Higgie, 2019" authorityName="Hoskin &amp; Bertola &amp; Higgie" authorityYear="2019" box="[366,627,1696,1720]" class="Reptilia" family="Gekkonidae" genus="Phyllurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="131" phylum="Chordata" rank="species" species="pinnaclensis" status="sp. nov.">
<emphasis id="B9201076FFD9FFC8FEB8F942DA62F92C" box="[366,627,1696,1720]" italics="true" pageId="4" pageNumber="131">Phyllurus pinnaclensis</emphasis>
</taxonomicName>
<emphasis id="B9201076FFD9FFC8FDA9F97DDACFF92C" bold="true" box="[639,734,1695,1720]" pageId="4" pageNumber="131">
<taxonomicNameLabel id="A213AD0DFFD9FFC8FDA9F97DDACFF92C" box="[639,734,1695,1720]" pageId="4" pageNumber="131" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
could only be confused with congeners with a cylindrical, tapering tail. Of these, it is readily distinguished from
<taxonomicName id="4C54B7E7FFD9FFC8FD25F927DC9DF949" authority="Covacevich, 1975" authorityName="Covacevich" authorityYear="1975" box="[755,1164,1732,1757]" class="Reptilia" family="Gekkonidae" genus="Phyllurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="131" phylum="Chordata" rank="species" species="caudiannulatus">
<emphasis id="B9201076FFD9FFC8FD25F927DBAAF948" box="[755,955,1732,1756]" italics="true" pageId="4" pageNumber="131">P. caudiannulatus</emphasis>
<bibRefCitation id="EFC5B195FFD9FFC8FC14F926DC9DF949" author="Covacevich, J." box="[962,1164,1732,1757]" pageId="4" pageNumber="131" pagination="293 - 303" refId="ref5607" refString="Covacevich, J. (1975) A review of the genus Phyllurus (Lacertilia: Gekkonidae). Memoirs of the Queensland Museum, 17, 293 - 303." type="journal article" year="1975">Covacevich, 1975</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C54B7E7FFD9FFC8FB17F927D9A2F894" authority="Couper, Hamley &amp; Hoskin, 2008" authorityName="Couper, Hamley &amp; Hoskin" authorityYear="2008" class="Reptilia" family="Gekkonidae" genus="Phyllurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="131" phylum="Chordata" rank="species" species="kabikabi">
<emphasis id="B9201076FFD9FFC8FB17F927DD2DF948" box="[1217,1340,1732,1756]" italics="true" pageId="4" pageNumber="131">P. kabikabi</emphasis>
<bibRefCitation id="EFC5B195FFD9FFC8FA92F926D9A2F894" author="Couper, P. J. &amp; Hamley, B. &amp; Hoskin C. J." pageId="4" pageNumber="131" pagination="139 - 147" refId="ref5391" refString="Couper, P. J., Hamley, B. &amp; Hoskin C. J. (2008) A new species of Phyllurus (Lacertilia: Gekkonidae) from the Kilkivan district of south-eastern Queensland. Memoirs of the Queensland Museum, 52, 139 - 147." type="journal article" year="2008">Couper, Hamley &amp; Hoskin, 2008</bibRefCitation>
</taxonomicName>
by a partially divided rostral scale (
<emphasis id="B9201076FFD9FFC8FC8DF90BDBB2F894" box="[859,931,1769,1792]" italics="true" pageId="4" pageNumber="131">versus</emphasis>
completely divided). From
<taxonomicName id="4C54B7E7FFD9FFC8FB3EF90BDD4EF894" box="[1256,1375,1768,1792]" class="Reptilia" family="Gekkonidae" genus="Phyllurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="131" phylum="Chordata" rank="species" species="gulbaru">
<emphasis id="B9201076FFD9FFC8FB3EF90BDD4EF894" box="[1256,1375,1768,1792]" italics="true" pageId="4" pageNumber="131">P. gulbaru</emphasis>
</taxonomicName>
, it is distinguished by a high internasal count (
<taxonomicName id="4C54B7E7FFD9FFC8FDB1F8EFDACDF8B0" box="[615,732,1804,1828]" class="Reptilia" family="Gekkonidae" genus="Phyllurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="131" phylum="Chordata" rank="species" species="gulbaru">
<emphasis id="B9201076FFD9FFC8FDB1F8EFDACDF8B0" box="[615,732,1804,1828]" italics="true" pageId="4" pageNumber="131">P. gulbaru</emphasis>
</taxonomicName>
mean 6.5, range 58;
<taxonomicName id="4C54B7E7FFD9FFC8FC01F8EFDC6EF8B0" authority="Hoskin &amp; Bertola &amp; Higgie, 2019" authorityName="Hoskin &amp; Bertola &amp; Higgie" authorityYear="2019" box="[983,1151,1804,1828]" class="Reptilia" family="Gekkonidae" genus="Phyllurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="131" phylum="Chordata" rank="species" species="pinnaclensis" status="sp. nov.">
<emphasis id="B9201076FFD9FFC8FC01F8EFDC6EF8B0" box="[983,1151,1804,1828]" italics="true" pageId="4" pageNumber="131">P. pinnaclensis</emphasis>
</taxonomicName>
<emphasis id="B9201076FFD9FFC8FB51F8EEDCF0F8B1" bold="true" box="[1159,1249,1804,1829]" pageId="4" pageNumber="131">
<taxonomicNameLabel id="A213AD0DFFD9FFC8FB51F8EEDCF0F8B1" box="[1159,1249,1804,1829]" pageId="4" pageNumber="131" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
mean 9.1, range 710); high subdigital lamellae count under the 4
<superScript id="7C21612CFFD9FFC8FD6DF8CDDAD6F8A9" attach="left" box="[699,711,1839,1853]" fontSize="6" pageId="4" pageNumber="131">th</superScript>
toe (
<taxonomicName id="4C54B7E7FFD9FFC8FCD6F8D1DB65F8DF" box="[768,884,1843,1867]" class="Reptilia" family="Gekkonidae" genus="Phyllurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="131" phylum="Chordata" rank="species" species="gulbaru">
<emphasis id="B9201076FFD9FFC8FCD6F8D1DB65F8DF" box="[768,884,1843,1867]" italics="true" pageId="4" pageNumber="131">P. gulbaru</emphasis>
</taxonomicName>
mean 17.9, range 1619;
<taxonomicName id="4C54B7E7FFD9FFC8FB43F8D1DD2DF8DF" authority="Hoskin &amp; Bertola &amp; Higgie, 2019" authorityName="Hoskin &amp; Bertola &amp; Higgie" authorityYear="2019" box="[1173,1340,1843,1867]" class="Reptilia" family="Gekkonidae" genus="Phyllurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="131" phylum="Chordata" rank="species" species="pinnaclensis" status="sp. nov.">
<emphasis id="B9201076FFD9FFC8FB43F8D1DD2DF8DF" box="[1173,1340,1843,1867]" italics="true" pageId="4" pageNumber="131">P. pinnaclensis</emphasis>
</taxonomicName>
<emphasis id="B9201076FFD9FFC8FA95F8D0DD8DF8DF" bold="true" box="[1347,1436,1842,1867]" pageId="4" pageNumber="131">
<taxonomicNameLabel id="A213AD0DFFD9FFC8FA95F8D0DD8DF8DF" box="[1347,1436,1842,1867]" pageId="4" pageNumber="131" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
mean 20.0, range 1921); longer original tail (
<taxonomicName id="4C54B7E7FFD9FFC8FD7EF8BADB0FF8FB" box="[680,798,1879,1903]" class="Reptilia" family="Gekkonidae" genus="Phyllurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="131" phylum="Chordata" rank="species" species="gulbaru">
<emphasis id="B9201076FFD9FFC8FD7EF8BADB0FF8FB" box="[680,798,1879,1903]" italics="true" pageId="4" pageNumber="131">P. gulbaru</emphasis>
</taxonomicName>
T/SVL mean 0.80, range 0.710.84;
<taxonomicName id="4C54B7E7FFD9FFC8FB1FF8BADD63F8FB" authority="Hoskin &amp; Bertola &amp; Higgie, 2019" authorityName="Hoskin &amp; Bertola &amp; Higgie" authorityYear="2019" box="[1225,1394,1879,1903]" class="Reptilia" family="Gekkonidae" genus="Phyllurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="131" phylum="Chordata" rank="species" species="pinnaclensis" status="sp. nov.">
<emphasis id="B9201076FFD9FFC8FB1FF8BADD63F8FB" box="[1225,1394,1879,1903]" italics="true" pageId="4" pageNumber="131">P. pinnaclensis</emphasis>
</taxonomicName>
<emphasis id="B9201076FFD9FFC8FAACF8B4D8D9F807" bold="true" pageId="4" pageNumber="131">
<taxonomicNameLabel id="A213AD0DFFD9FFC8FAACF8B4D8D9F807" pageId="4" pageNumber="131" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
T/SVL mean 0.90, range 0.880.91) and regenerated tail (
<taxonomicName id="4C54B7E7FFD9FFC8FCB0F899DBCDF807" box="[870,988,1915,1939]" class="Reptilia" family="Gekkonidae" genus="Phyllurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="131" phylum="Chordata" rank="species" species="gulbaru">
<emphasis id="B9201076FFD9FFC8FCB0F899DBCDF807" box="[870,988,1915,1939]" italics="true" pageId="4" pageNumber="131">P. gulbaru</emphasis>
</taxonomicName>
T/SVL mean 0.61, range 0.500.71;
<taxonomicName id="4C54B7E7FFD9FFC8FA51F899D932F823" authority="Hoskin &amp; Bertola &amp; Higgie, 2019" authorityName="Hoskin &amp; Bertola &amp; Higgie" authorityYear="2019" class="Reptilia" family="Gekkonidae" genus="Phyllurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="131" phylum="Chordata" rank="species" species="pinnaclensis" status="sp. nov.">
<emphasis id="B9201076FFD9FFC8FA51F899D932F823" italics="true" pageId="4" pageNumber="131">P. pinnaclensis</emphasis>
</taxonomicName>
<emphasis id="B9201076FFD9FFC8FEF9F87CD99CF823" bold="true" box="[303,397,1950,1975]" pageId="4" pageNumber="131">
<taxonomicNameLabel id="A213AD0DFFD9FFC8FEF9F87CD99CF823" box="[303,397,1950,1975]" pageId="4" pageNumber="131" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
T/SVL mean 0.73; range 0.73) (
<tableCitation id="C6D6F9DFFFD9FFC8FCC1F87CDB61F823" box="[791,880,1950,1975]" captionStart="TABLE 1" captionStartId="4.[151,239,152,175]" captionText="TABLE 1. Field data for select morphometric and scale traits for P. pinnaclensis sp. nov. and P. gulbaru. Traits marked in bold have obviously different means and/or no overlap in range values between the two species." pageId="4" pageNumber="131">Table 1</tableCitation>
); and original tail largely devoid of prominent tubercles past the basal quarter (
<emphasis id="B9201076FFD9FFC8FDC1F821DA4FF84E" box="[535,606,1987,2010]" italics="true" pageId="4" pageNumber="131">versus</emphasis>
prominent tubercles on at least basal half in
<taxonomicName id="4C54B7E7FFD9FFC8FBA8F821DCE2F84F" box="[1150,1267,1987,2011]" class="Reptilia" family="Gekkonidae" genus="Phyllurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="131" phylum="Chordata" rank="species" species="gulbaru">
<emphasis id="B9201076FFD9FFC8FBA8F821DCE2F84F" box="[1150,1267,1987,2011]" italics="true" pageId="4" pageNumber="131">P. gulbaru</emphasis>
</taxonomicName>
). A relatively straightforward and near-diagnostic trait to assess in the field is the number of internasals, with&gt; 90% of
<taxonomicName id="4C54B7E7FFD8FFC9FA51FF7BD8E1FF40" class="Reptilia" family="Gekkonidae" genus="Phyllurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="132" phylum="Chordata" rank="species" species="gulbaru">
<emphasis id="B9201076FFD8FFC9FA51FF7BD8E1FF40" italics="true" pageId="5" pageNumber="132">P. gulbaru</emphasis>
</taxonomicName>
individuals (N = 36) having &lt;8 (57) and 90% of
<taxonomicName id="4C54B7E7FFD8FFC9FCF9FF5FDBC6FF40" authority="Hoskin &amp; Bertola &amp; Higgie, 2019" authorityName="Hoskin &amp; Bertola &amp; Higgie" authorityYear="2019" box="[815,983,188,212]" class="Reptilia" family="Gekkonidae" genus="Phyllurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="132" phylum="Chordata" rank="species" species="pinnaclensis" status="sp. nov.">
<emphasis id="B9201076FFD8FFC9FCF9FF5FDBC6FF40" box="[815,983,188,212]" italics="true" pageId="5" pageNumber="132">P. pinnaclensis</emphasis>
</taxonomicName>
<emphasis id="B9201076FFD8FFC9FC08FF5EDC29FF41" bold="true" box="[990,1080,188,213]" pageId="5" pageNumber="132">
<taxonomicNameLabel id="A213AD0DFFD8FFC9FC08FF5EDC29FF41" box="[990,1080,188,213]" pageId="5" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(N = 10) having&gt; 8 (910; but one aberrant individual had 7).
</paragraph>
<caption id="DF2B9CECFFD8FFC9FF41FA1EDA2FF9A5" httpUri="https://zenodo.org/record/2624736/files/figure.png" pageId="5" pageNumber="132" startId="5.[151,250,1532,1554]" targetBox="[151,1436,301,1511]" targetPageId="5">
<paragraph id="8BEBCC64FFD8FFC9FF41FA1EDA2FF9A5" blockId="5.[151,1436,1532,1585]" pageId="5" pageNumber="132">
<emphasis id="B9201076FFD8FFC9FF41FA1ED900F986" bold="true" box="[151,273,1532,1554]" pageId="5" pageNumber="132">FIGURE 2</emphasis>
. (A) Holotype of
<taxonomicName id="4C54B7E7FFD8FFC9FE0CFA1FDA67F985" authority="Hoskin &amp; Bertola &amp; Higgie, 2019" authorityName="Hoskin &amp; Bertola &amp; Higgie" authorityYear="2019" box="[474,630,1532,1554]" class="Reptilia" family="Gekkonidae" genus="Phyllurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="132" phylum="Chordata" rank="species" species="pinnaclensis" status="sp. nov.">
<emphasis id="B9201076FFD8FFC9FE0CFA1FDA67F985" box="[474,630,1532,1554]" italics="true" pageId="5" pageNumber="132">P. pinnaclensis</emphasis>
</taxonomicName>
<emphasis id="B9201076FFD8FFC9FD57FA1EDAC7F986" bold="true" box="[641,726,1532,1554]" pageId="5" pageNumber="132">
<taxonomicNameLabel id="A213AD0DFFD8FFC9FD57FA1EDAC7F986" box="[641,726,1532,1554]" pageId="5" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in life (photo: Conrad Hoskin). (B)
<taxonomicName id="4C54B7E7FFD8FFC9FBB1FA1FDD12F985" authority="Hoskin &amp; Bertola &amp; Higgie, 2019" authorityName="Hoskin &amp; Bertola &amp; Higgie" authorityYear="2019" box="[1127,1283,1532,1554]" class="Reptilia" family="Gekkonidae" genus="Phyllurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="132" phylum="Chordata" rank="species" species="pinnaclensis" status="sp. nov.">
<emphasis id="B9201076FFD8FFC9FBB1FA1FDD12F985" box="[1127,1283,1532,1554]" italics="true" pageId="5" pageNumber="132">P. pinnaclensis</emphasis>
</taxonomicName>
<emphasis id="B9201076FFD8FFC9FAD8FA1EDD72F986" bold="true" box="[1294,1379,1532,1554]" pageId="5" pageNumber="132">
<taxonomicNameLabel id="A213AD0DFFD8FFC9FAD8FA1EDD72F986" box="[1294,1379,1532,1554]" pageId="5" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
with regenerated tail (photo: Lorenzo Bertola).
</paragraph>
</caption>
<paragraph id="8BEBCC64FFD8FFC9FF11F9BADC66F9E4" blockId="5.[151,1437,1624,2037]" box="[199,1143,1624,1649]" pageId="5" pageNumber="132">
<emphasis id="B9201076FFD8FFC9FF11F9BAD924F9E5" bold="true" box="[199,309,1624,1649]" pageId="5" pageNumber="132">Genetics.</emphasis>
Approximately 6.4% divergent from
<taxonomicName id="4C54B7E7FFD8FFC9FD0DF9BBDB5EF9E4" box="[731,847,1624,1648]" class="Reptilia" family="Gekkonidae" genus="Phyllurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="132" phylum="Chordata" rank="species" species="gulbaru">
<emphasis id="B9201076FFD8FFC9FD0DF9BBDB5EF9E4" box="[731,847,1624,1648]" italics="true" pageId="5" pageNumber="132">P. gulbaru</emphasis>
</taxonomicName>
for ND2 mtDNA (
<figureCitation id="136FD0E1FFD8FFC9FBF3F9BADC77F9E4" box="[1061,1126,1624,1649]" captionStart="FIGURE 1" captionStartId="2.[151,250,1806,1828]" captionTargetBox="[198,1380,195,1784]" captionTargetId="figure@2.[188,1380,195,1784]" captionTargetPageId="2" captionText="FIGURE 1. Left panel: A consensus tree of 1,041 base pairs of ND2 mtDNA produced using MrBayes. Nodes with Bayesian inference posterior probability of 1.0 are noted by an asterisk (*). Locality abbreviations are: HA, Mt. Halifax; BW, Bluewater Range; NP, North Patterson Gorge; SP, South Patterson Gorge; HR, Hervey Range; PI, The Pinnacles. The average uncorrected p-distance between The Pinnacles sequences and all sequences of P. gulbaru is 6.4%. Right panel: Map showing the known populations of the two species (symbols and abbreviations), topography (background shading), and the Species Distribution Model (SDM) (both species modelled together). Coloured pixels show predicted probability of presence as indicated by the legend. See Bertola et al. (2018) for further details of phylogenetics and SDM." httpUri="https://zenodo.org/record/2624734/files/figure.png" pageId="5" pageNumber="132">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34E9FEFFFD8FFC9FF11F99EDC1AF861" pageId="5" pageNumber="132" type="distribution">
<paragraph id="8BEBCC64FFD8FFC9FF11F99EDB16F8F9" blockId="5.[151,1437,1624,2037]" pageId="5" pageNumber="132">
<emphasis id="B9201076FFD8FFC9FF11F99ED971F901" bold="true" box="[199,352,1660,1685]" pageId="5" pageNumber="132">Distribution.</emphasis>
Appears to be restricted to The Pinnacles,
<quantity id="4CAC6181FFD8FFC9FCE8F99EDB94F901" box="[830,901,1660,1685]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="132" unit="km" value="25.0">25 km</quantity>
south-west of Townsville (
<figureCitation id="136FD0E1FFD8FFC9FB62F99EDCE4F901" box="[1204,1269,1660,1685]" captionStart="FIGURE 1" captionStartId="2.[151,250,1806,1828]" captionTargetBox="[198,1380,195,1784]" captionTargetId="figure@2.[188,1380,195,1784]" captionTargetPageId="2" captionText="FIGURE 1. Left panel: A consensus tree of 1,041 base pairs of ND2 mtDNA produced using MrBayes. Nodes with Bayesian inference posterior probability of 1.0 are noted by an asterisk (*). Locality abbreviations are: HA, Mt. Halifax; BW, Bluewater Range; NP, North Patterson Gorge; SP, South Patterson Gorge; HR, Hervey Range; PI, The Pinnacles. The average uncorrected p-distance between The Pinnacles sequences and all sequences of P. gulbaru is 6.4%. Right panel: Map showing the known populations of the two species (symbols and abbreviations), topography (background shading), and the Species Distribution Model (SDM) (both species modelled together). Coloured pixels show predicted probability of presence as indicated by the legend. See Bertola et al. (2018) for further details of phylogenetics and SDM." httpUri="https://zenodo.org/record/2624734/files/figure.png" pageId="5" pageNumber="132">Fig. 1</figureCitation>
). Known from three areas of habitat on the western and southern Pinnacles (
<figureCitation id="136FD0E1FFD8FFC9FCB2F942DBB8F92C" box="[868,937,1696,1721]" captionStart="FIGURE 1" captionStartId="2.[151,250,1806,1828]" captionTargetBox="[198,1380,195,1784]" captionTargetId="figure@2.[188,1380,195,1784]" captionTargetPageId="2" captionText="FIGURE 1. Left panel: A consensus tree of 1,041 base pairs of ND2 mtDNA produced using MrBayes. Nodes with Bayesian inference posterior probability of 1.0 are noted by an asterisk (*). Locality abbreviations are: HA, Mt. Halifax; BW, Bluewater Range; NP, North Patterson Gorge; SP, South Patterson Gorge; HR, Hervey Range; PI, The Pinnacles. The average uncorrected p-distance between The Pinnacles sequences and all sequences of P. gulbaru is 6.4%. Right panel: Map showing the known populations of the two species (symbols and abbreviations), topography (background shading), and the Species Distribution Model (SDM) (both species modelled together). Coloured pixels show predicted probability of presence as indicated by the legend. See Bertola et al. (2018) for further details of phylogenetics and SDM." httpUri="https://zenodo.org/record/2624734/files/figure.png" pageId="5" pageNumber="132">Fig. 1</figureCitation>
). Each of these small patches of habitat is isolated by unsuitable habitat that is too dry and/or not sufficiently rocky. Surveys of rainforest habitat in the northeastern Pinnacles (-19.397, 146.642) have failed to find
<taxonomicName id="4C54B7E7FFD8FFC9FCDDF90BDBA3F894" authority="Hoskin &amp; Bertola &amp; Higgie, 2019" authorityName="Hoskin &amp; Bertola &amp; Higgie" authorityYear="2019" box="[779,946,1768,1792]" class="Reptilia" family="Gekkonidae" genus="Phyllurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="132" phylum="Chordata" rank="species" species="pinnaclensis" status="sp. nov.">
<emphasis id="B9201076FFD8FFC9FCDDF90BDBA3F894" box="[779,946,1768,1792]" italics="true" pageId="5" pageNumber="132">P. pinnaclensis</emphasis>
</taxonomicName>
<emphasis id="B9201076FFD8FFC9FC6FF90ADC03F895" bold="true" box="[953,1042,1768,1793]" pageId="5" pageNumber="132">
<taxonomicNameLabel id="A213AD0DFFD8FFC9FC6FF90ADC03F895" box="[953,1042,1768,1793]" pageId="5" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
. This area has very high suitability in the SDM (dark green pixel in north-east Pinnacles in
<figureCitation id="136FD0E1FFD8FFC9FC96F8EEDBB3F8B1" box="[832,930,1804,1829]" captionStart="FIGURE 1" captionStartId="2.[151,250,1806,1828]" captionTargetBox="[198,1380,195,1784]" captionTargetId="figure@2.[188,1380,195,1784]" captionTargetPageId="2" captionText="FIGURE 1. Left panel: A consensus tree of 1,041 base pairs of ND2 mtDNA produced using MrBayes. Nodes with Bayesian inference posterior probability of 1.0 are noted by an asterisk (*). Locality abbreviations are: HA, Mt. Halifax; BW, Bluewater Range; NP, North Patterson Gorge; SP, South Patterson Gorge; HR, Hervey Range; PI, The Pinnacles. The average uncorrected p-distance between The Pinnacles sequences and all sequences of P. gulbaru is 6.4%. Right panel: Map showing the known populations of the two species (symbols and abbreviations), topography (background shading), and the Species Distribution Model (SDM) (both species modelled together). Coloured pixels show predicted probability of presence as indicated by the legend. See Bertola et al. (2018) for further details of phylogenetics and SDM." httpUri="https://zenodo.org/record/2624734/files/figure.png" pageId="5" pageNumber="132">Figure 1</figureCitation>
) but the rock in this area is probably not sufficiently layered to have provided long-term climatic buffering. Potentially suitable rocky rainforest gullies in the south-eastern Pinnacles have not yet been surveyed.
</paragraph>
<paragraph id="8BEBCC64FFD8FFC9FF11F89ADC1AF861" blockId="5.[151,1437,1624,2037]" pageId="5" pageNumber="132">
<emphasis id="B9201076FFD8FFC9FF11F89AD9A5F805" bold="true" box="[199,436,1912,1937]" pageId="5" pageNumber="132">Habitat and habits.</emphasis>
Layered rock with associated rainforest (vine-forest) vegetation. One site consists of piled, angular rock (
<figureCitation id="136FD0E1FFD8FFC9FEAFF87BD9C0F826" box="[377,465,1945,1970]" captionStart="FIGURE 6" captionStartId="9.[151,250,1924,1946]" captionTargetBox="[151,1436,218,1881]" captionTargetId="figure@9.[151,1436,218,1881]" captionTargetPageId="9" captionText="FIGURE 6. Photos of habitat of P. pinnaclensis sp. nov., showing (A) angular rock scree at the type locality (photo: Anders Zimny) and (B) piled granite boulder habitat at another site (photo: Lorenzo Bertola). The white arrow on B points to a gecko foraging in a typical position on a vertical rock surface at night." httpUri="https://zenodo.org/record/2624744/files/figure.png" pageId="5" pageNumber="132">Fig. 6A</figureCitation>
) associated with a gully, another site consists of piled, rounded, granite boulders along a gully (
<figureCitation id="136FD0E1FFD8FFC9FF22F859D95DF840" box="[244,332,1979,2004]" captionStart="FIGURE 6" captionStartId="9.[151,250,1924,1946]" captionTargetBox="[151,1436,218,1881]" captionTargetId="figure@9.[151,1436,218,1881]" captionTargetPageId="9" captionText="FIGURE 6. Photos of habitat of P. pinnaclensis sp. nov., showing (A) angular rock scree at the type locality (photo: Anders Zimny) and (B) piled granite boulder habitat at another site (photo: Lorenzo Bertola). The white arrow on B points to a gecko foraging in a typical position on a vertical rock surface at night." httpUri="https://zenodo.org/record/2624744/files/figure.png" pageId="5" pageNumber="132">Fig. 6B</figureCitation>
), and the third site consists of deeply piled, granite boulders along a gully and adjacent slopes. At all sites,
<taxonomicName id="4C54B7E7FFD8FFC9FF2CF83FD9B0F860" authority="Hoskin &amp; Bertola &amp; Higgie, 2019" authorityName="Hoskin &amp; Bertola &amp; Higgie" authorityYear="2019" box="[250,417,2012,2036]" class="Reptilia" family="Gekkonidae" genus="Phyllurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="132" phylum="Chordata" rank="species" species="pinnaclensis" status="sp. nov.">
<emphasis id="B9201076FFD8FFC9FF2CF83FD9B0F860" box="[250,417,2012,2036]" italics="true" pageId="5" pageNumber="132">P. pinnaclensis</emphasis>
</taxonomicName>
<emphasis id="B9201076FFD8FFC9FE7EF83EDA13F861" bold="true" box="[424,514,2012,2037]" pageId="5" pageNumber="132">
<taxonomicNameLabel id="A213AD0DFFD8FFC9FE7EF83EDA13F861" box="[424,514,2012,2037]" pageId="5" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was found on rock surfaces at night (
<figureCitation id="136FD0E1FFD8FFC9FC70F83EDBEFF861" box="[934,1022,2012,2037]" captionStart="FIGURE 6" captionStartId="9.[151,250,1924,1946]" captionTargetBox="[151,1436,218,1881]" captionTargetId="figure@9.[151,1436,218,1881]" captionTargetPageId="9" captionText="FIGURE 6. Photos of habitat of P. pinnaclensis sp. nov., showing (A) angular rock scree at the type locality (photo: Anders Zimny) and (B) piled granite boulder habitat at another site (photo: Lorenzo Bertola). The white arrow on B points to a gecko foraging in a typical position on a vertical rock surface at night." httpUri="https://zenodo.org/record/2624744/files/figure.png" pageId="5" pageNumber="132">Fig. 6B</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>