349 lines
38 KiB
XML
349 lines
38 KiB
XML
<document id="1187519407D67EC1EDBAECECB5ED806B" ID-CLB-Dataset="20325" ID-DOI="10.11646/zootaxa.5104.3.4" ID-GBIF-Dataset="17de5cf1-e498-4883-b0e7-8bcd5e976d47" ID-ISSN="1175-5326" ID-Zenodo-Dep="6332094" ID-ZooBank="9A9BA43F-3B32-4595-A95A-853B8F72FD90" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1646594675974" checkinUser="plazi" docAuthor="Kaneko, Naoki & Wada, Kaoru" docDate="2022" docId="03EBE556FFFCFFA0379590BDAFD9FEF7" docLanguage="en" docName="zootaxa.5104.3.4.pdf" docOrigin="Zootaxa 5104 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Malaia (M) simulatrix Heller 1892" docType="treatment" docVersion="8" lastPageNumber="431" masterDocId="FFD29D2EFFFFFFA53702931EAB7AFFB6" masterDocTitle="A new species of the genus Malaia Heller, 1892 (Scarabaeidae: Rutelinae: Anomalini) from Sulawesi Island, Indonesia" masterLastPageNumber="432" masterPageNumber="426" pageNumber="429" updateTime="1699316905168" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="52BCE7C590B5AAB1C3D020CAA3630270">A new species of the genus Malaia Heller, 1892 (Scarabaeidae: Rutelinae: Anomalini) from Sulawesi Island, Indonesia</mods:title>
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<mods:name id="2064848A49E9477F68C098C9AD9B1BDB" type="personal">
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<mods:namePart id="0133DD780A195D3DC25D7AD63EFDE7CA">Kaneko, Naoki</mods:namePart>
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<mods:namePart id="C723BE5C0D66D88EAB9A635FCD7E6CA8">Wada, Kaoru</mods:namePart>
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<mods:title id="988782116AC64FCF32255110DFE4E3B9">Zootaxa</mods:title>
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<mods:part id="F42AAA481D5B9F48800162BE8B717B8A">
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<mods:date id="362EBD6E643D1FDD3C15FEDF1F0DDDD9">2022</mods:date>
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<mods:number id="EA4796B28F4101DF816B01EB2656A754">2022-02-28</mods:number>
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<mods:number id="928853A4C373C69287DC0453DF1362C7">5104</mods:number>
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<treatment id="03EBE556FFFCFFA0379590BDAFD9FEF7" ID-DOI="http://doi.org/10.5281/zenodo.6343752" ID-GBIF-Taxon="193892593" ID-Zenodo-Dep="6343752" LSID="urn:lsid:plazi:treatment:03EBE556FFFCFFA0379590BDAFD9FEF7" httpUri="http://treatment.plazi.org/id/03EBE556FFFCFFA0379590BDAFD9FEF7" lastPageId="5" lastPageNumber="431" pageId="3" pageNumber="429">
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<subSubSection id="C35807CBFFFCFFA6379590BDA851FC08" box="[151,811,931,958]" pageId="3" pageNumber="429" type="nomenclature">
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<paragraph id="8BFD5440FFFCFFA6379590BDA851FC08" blockId="3.[151,811,931,958]" box="[151,811,931,958]" pageId="3" pageNumber="429">
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<heading id="D0B5E32CFFFCFFA6379590BDA851FC08" bold="true" box="[151,811,931,958]" fontSize="11" level="1" pageId="3" pageNumber="429" reason="1">
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<emphasis id="B9368852FFFCFFA6379590BDA851FC08" bold="true" box="[151,811,931,958]" pageId="3" pageNumber="429">
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<taxonomicName id="4C422FC3FFFCFFA6379590BDA930FC08" authority="Heller, 1892" authorityName="Heller" authorityYear="1892" box="[151,586,931,958]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="429" phylum="Arthropoda" rank="genus">
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<emphasis id="B9368852FFFCFFA6379590BDAB97FC08" bold="true" box="[151,237,931,958]" italics="true" pageId="3" pageNumber="429">Malaia</emphasis>
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(
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<emphasis id="B9368852FFFCFFA637FD90BAAA63FC08" bold="true" box="[255,281,932,958]" italics="true" pageId="3" pageNumber="429">M</emphasis>
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.)
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||
<emphasis id="B9368852FFFCFFA6363390BDAAD7FC08" bold="true" box="[305,429,931,958]" italics="true" pageId="3" pageNumber="429">simulatrix</emphasis>
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<bibRefCitation id="EFD329B1FFFCFFA636B690BAA930FC08" author="Heller, K. M." box="[436,586,932,958]" pageId="3" pageNumber="429" pagination="289 - 306" refId="ref4902" refString="Heller, K. M. (1892) Die mit der Ruteliden - Gattung Singhala Burm. verwandten Gattungen und Arten. Deutsche Entomologische Zeitschrift, 1891 (2), 289 - 306." type="journal article" year="1892">Heller, 1892</bibRefCitation>
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</taxonomicName>
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(
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<figureCitation id="137948C5FFFCFFA6355B90BAA9B0FC08" box="[601,714,932,958]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1678,1702]" captionTargetBox="[162,1427,189,1638]" captionTargetId="figure-17@2.[151,1436,179,1652]" captionTargetPageId="2" captionText="FIGURES 1–9. Habitus of Malaia (M.) medea Kaneko & Wada, new species (1–6) and M. (M.) simulatrix Heller, 1892 ♂ from Pulau Peleng, Sulawesi (7–9). 1, holotype, dorsal view; 2, holotype, ventral view; 3, holotype, lateral view; 4, paratype, dorsal view (color variability); 5, allotype, dorsal view; 6, allotype, ventral view; 7, specimen from RIEB, dorsal view; 8, the same, ventral view; 9, the same, lateral view. Scale: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6332096" httpUri="https://zenodo.org/record/6332096/files/figure.png" pageId="3" pageNumber="429">Figs. 7–9</figureCitation>
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,
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<figureCitation id="137948C5FFFCFFA635DA90BAA858FC08" box="[728,802,932,958]" captionStart="FIGURES 10–13" captionStartId="4.[151,264,1911,1935]" captionTargetBox="[234,1358,191,1880]" captionTargetId="figure-17@4.[215,1371,181,1887]" captionTargetPageId="4" captionText="FIGURES 10–13. Aedeagus of Malaia (M.) medea Kaneko & Wada, new species (10–11) and M. (M.) simulatrix Heller, 1892 ♂ from Pulau Peleng, Sulawesi (12–13). 10, holotype, dorsal view; 11, holotype, lateral view; 12, specimen from RIEB, dorsal view; 13, the same, lateral view. Scale: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6332099" httpUri="https://zenodo.org/record/6332099/files/figure.png" pageId="3" pageNumber="429">12–13</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C35807CBFFFCFFA6379590F6A9F7FBB6" box="[151,653,1000,1025]" pageId="3" pageNumber="429" type="reference_group">
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<paragraph id="8BFD5440FFFCFFA6379590F6A9F7FBB6" blockId="3.[151,653,1000,1025]" box="[151,653,1000,1025]" pageId="3" pageNumber="429">
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<treatmentCitationGroup id="AB52736EFFFCFFA6379590F6A9F7FBB6" box="[151,653,1000,1025]" pageId="3" pageNumber="429">
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<treatmentCitation id="0AE37251FFFCFFA6379590F6A940FBB6" author="Heller, K. M." box="[151,570,1000,1025]" page="305" pageId="3" pageNumber="429" year="1892">
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<taxonomicName id="4C422FC3FFFCFFA6379590F6A940FBB6" authority="Heller, 1892: 303 - 305" authorityName="Heller" authorityPageNumber="303 - 305" authorityYear="1892" box="[151,570,1000,1025]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="429" phylum="Arthropoda" rank="species" species="simulatrix">
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||
<emphasis id="B9368852FFFCFFA6379590F6AA34FBB6" box="[151,334,1000,1024]" italics="true" pageId="3" pageNumber="429">Malaia simulatrix</emphasis>
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||
<bibRefCitation id="EFD329B1FFFCFFA6365690F6A940FBB6" author="Heller, K. M." box="[340,570,1000,1025]" pageId="3" pageNumber="429" pagination="289 - 306" refId="ref4902" refString="Heller, K. M. (1892) Die mit der Ruteliden - Gattung Singhala Burm. verwandten Gattungen und Arten. Deutsche Entomologische Zeitschrift, 1891 (2), 289 - 306." type="journal article" year="1892">Heller, 1892: 303–305</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
, fig. 16.
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||
</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35807CBFFFCFFA637959731AF76F91B" pageId="3" pageNumber="429" type="materials_examined">
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<paragraph id="8BFD5440FFFCFFA637959731A8CBFBFF" blockId="3.[151,1437,1070,2033]" box="[151,945,1070,1097]" pageId="3" pageNumber="429">
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<emphasis id="B9368852FFFCFFA637959731AA4CFBFE" bold="true" box="[151,310,1070,1097]" pageId="3" pageNumber="429">
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||
<typeStatus id="54F9EAE2FFFCFFA637959731ABA8FBFF" box="[151,210,1071,1097]" pageId="3" pageNumber="429">Type</typeStatus>
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locality.
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</emphasis>
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||
“Insula Bangkai [= Pulau Banggai (
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<taxonomicName id="4C422FC3FFFCFFA635C99731A8EFFBFF" authority="Tengah" authorityName="Tengah" box="[715,917,1070,1097]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="429" phylum="Arthropoda" rank="species" species="sulawesi">Sulawesi Tengah</taxonomicName>
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)]”.
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</paragraph>
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||
<paragraph id="8BFD5440FFFCFFA637C5974DA940FBDB" blockId="3.[151,1437,1070,2033]" box="[199,570,1106,1133]" pageId="3" pageNumber="429">
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<emphasis id="B9368852FFFCFFA637C5974DAA95FBDA" bold="true" box="[199,495,1106,1133]" pageId="3" pageNumber="429">
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||
<typeStatus id="54F9EAE2FFFCFFA637C5974DAA78FBDB" box="[199,258,1107,1133]" pageId="3" pageNumber="429">Type</typeStatus>
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material examined.
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</emphasis>
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None.
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</paragraph>
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<paragraph id="8BFD5440FFFCFFA637C59768AAFBFA97" blockId="3.[151,1437,1070,2033]" pageId="3" pageNumber="429">
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||
<emphasis id="B9368852FFFCFFA637C59768A94EFB26" bold="true" box="[199,564,1142,1168]" pageId="3" pageNumber="429">Additional material examined.</emphasis>
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||
<materialsCitation id="3B2A5E1DFFFCFFA635399769AF99FB27" ID-GBIF-Occurrence="3698900307" box="[571,1251,1143,1169]" location="Palopo" municipality="Puncak" pageId="3" pageNumber="429" specimenCount="3" specimenCount-female="3" stateProvince="Sulawesi Utara" typeStatus="holotype">
|
||
<specimenCount id="9D449FC9FFFCFFA635399769A906FB27" box="[571,636,1143,1169]" pageId="3" pageNumber="429" type="female">3 ♀♀</specimenCount>
|
||
: “
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||
<collectingMunicipality id="6B99CE3AFFFCFFA635909769A99DFB27" box="[658,743,1143,1169]" pageId="3" pageNumber="429">Puncak</collectingMunicipality>
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||
/
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||
<location id="8E9D029BFFFCFFA635FC9769A836FB27" LSID="urn:lsid:plazi:treatment:03EBE556FFFCFFA0379590BDAFD9FEF7:8E9D029BFFFCFFA635FC9769A836FB27" box="[766,844,1143,1169]" municipality="Puncak" name="Palopo" pageId="3" pageNumber="429" stateProvince="Sulawesi Utara">Palopo</location>
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||
/ IX-X. 1990 /
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||
<collectingRegion id="49869AA2FFFCFFA634FA9769AF0EFB27" box="[1016,1140,1143,1169]" country="Indonesia" name="Sulawesi Utara" pageId="3" pageNumber="429">CELEBES</collectingRegion>
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||
” (PSMF)
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="3B2A5E1DFFFCFFA633EC9769A99FFB03" ID-GBIF-Occurrence="3698900301" collectingDate="1993-07" country="Indonesia" location="Palu" pageId="3" pageNumber="429" specimenCount="1" specimenCount-female="1" stateProvince="Sulawesi Utara" typeStatus="holotype">
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<specimenCount id="9D449FC9FFFCFFA633EC9769AE60FB27" box="[1262,1306,1143,1169]" pageId="3" pageNumber="429" type="female">1 ♀</specimenCount>
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: “near
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||
<location id="8E9D029BFFFCFFA632689769AEE6FB27" LSID="urn:lsid:plazi:treatment:03EBE556FFFCFFA0379590BDAFD9FEF7:8E9D029BFFFCFFA632689769AEE6FB27" box="[1386,1436,1143,1169]" country="Indonesia" name="Palu" pageId="3" pageNumber="429" stateProvince="Sulawesi Utara">Palu</location>
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||
/
|
||
<collectingRegion id="49869AA2FFFCFFA637A49785AA4BFB03" box="[166,305,1179,1205]" country="Indonesia" name="Sulawesi Utara" pageId="3" pageNumber="429">SULAWESI</collectingRegion>
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/
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<collectingCountry id="F35514D0FFFCFFA636459785AAA4FB03" box="[327,478,1179,1205]" name="Indonesia" pageId="3" pageNumber="429">INDONESIA</collectingCountry>
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/
|
||
<date id="FFFC7280FFFCFFA636F19785A914FB03" box="[499,622,1179,1205]" pageId="3" pageNumber="429" value="1993-07">
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||
<collectingDate id="EFB88B68FFFCFFA636F19785A914FB03" box="[499,622,1179,1205]" pageId="3" pageNumber="429" value="1993-07">JUL. 1993</collectingDate>
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</date>
|
||
.” (PCMF)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B2A5E1DFFFCFFA635ED9785AAF7FB6F" ID-GBIF-Occurrence="3698900312" collectingDate="2001-01" country="Indonesia" location="Palolo Palu" pageId="3" pageNumber="429" specimenCount="7" specimenCount-female="5" specimenCount-male="2" stateProvince="Sulawesi Utara" typeStatus="holotype">
|
||
<specimenCount id="9D449FC9FFFCFFA635ED9785A84AFB03" box="[751,816,1179,1205]" pageId="3" pageNumber="429" type="male">2 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D449FC9FFFCFFA6343E9784A807FB03" box="[828,893,1178,1205]" pageId="3" pageNumber="429" type="female">5 ♀♀</specimenCount>
|
||
: “
|
||
<collectingCountry id="F35514D0FFFCFFA634919785AF49FB03" box="[915,1075,1179,1205]" name="Indonesia" pageId="3" pageNumber="429">INDONESIA</collectingCountry>
|
||
: / C.
|
||
<collectingRegion id="49869AA2FFFCFFA6336E9785AF8DFB03" box="[1132,1271,1179,1205]" country="Indonesia" name="Sulawesi Utara" pageId="3" pageNumber="429">SULAWESI</collectingRegion>
|
||
/
|
||
<location id="8E9D029BFFFCFFA6320F9785AEF4FB03" LSID="urn:lsid:plazi:treatment:03EBE556FFFCFFA0379590BDAFD9FEF7:8E9D029BFFFCFFA6320F9785AEF4FB03" box="[1293,1422,1179,1205]" country="Indonesia" name="Palolo Palu" pageId="3" pageNumber="429" stateProvince="Sulawesi Utara">Palolo Palu</location>
|
||
/
|
||
<date id="FFFC7280FFFCFFA6379597A1AA6CFB6F" box="[151,278,1215,1241]" pageId="3" pageNumber="429" value="2001-01">
|
||
<collectingDate id="EFB88B68FFFCFFA6379597A1AA6CFB6F" box="[151,278,1215,1241]" pageId="3" pageNumber="429" value="2001-01">JAN. 2001</collectingDate>
|
||
</date>
|
||
.” (PCMF)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B2A5E1DFFFCFFA6369B97A1AE33FB6F" ID-GBIF-Occurrence="3698900305" box="[409,1353,1215,1241]" collectingDate="2009-01" country="Indonesia" location="Palolo Palu" pageId="3" pageNumber="429" specimenCount="10" specimenCount-female="6" specimenCount-male="4" stateProvince="Sulawesi Utara" typeStatus="holotype">
|
||
<specimenCount id="9D449FC9FFFCFFA6369B97A1AAA0FB6F" box="[409,474,1215,1241]" pageId="3" pageNumber="429" type="male">4 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D449FC9FFFCFFA636E597A1A952FB6F" box="[487,552,1215,1241]" pageId="3" pageNumber="429" type="female">6 ♀♀</specimenCount>
|
||
: “
|
||
<collectingCountry id="F35514D0FFFCFFA6353D97A1A9A5FB6F" box="[575,735,1215,1241]" name="Indonesia" pageId="3" pageNumber="429">INDONESIA</collectingCountry>
|
||
: / C.
|
||
<collectingRegion id="49869AA2FFFCFFA6341897A1A8DFFB6F" box="[794,933,1215,1241]" country="Indonesia" name="Sulawesi Utara" pageId="3" pageNumber="429">SULAWESI</collectingRegion>
|
||
/
|
||
<location id="8E9D029BFFFCFFA634BE97A1AF47FB6F" LSID="urn:lsid:plazi:treatment:03EBE556FFFCFFA0379590BDAFD9FEF7:8E9D029BFFFCFFA634BE97A1AF47FB6F" box="[956,1085,1215,1241]" country="Indonesia" name="Palolo Palu" pageId="3" pageNumber="429" stateProvince="Sulawesi Utara">Palolo Palu</location>
|
||
/
|
||
<date id="FFFC7280FFFCFFA6335197A1AFA9FB6F" box="[1107,1235,1215,1241]" pageId="3" pageNumber="429" value="2009-01">
|
||
<collectingDate id="EFB88B68FFFCFFA6335197A1AFA9FB6F" box="[1107,1235,1215,1241]" pageId="3" pageNumber="429" value="2009-01">JAN. 2009</collectingDate>
|
||
</date>
|
||
.” (PCMF)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B2A5E1DFFFCFFA6325797A1AF95FB4B" ID-GBIF-Occurrence="3698900310" collectingDate="2017-10" collectorName="Native Collector" country="Indonesia" location="Puncak Palopo" pageId="3" pageNumber="429" specimenCount="2" specimenCount-female="2" stateProvince="Sulawesi Tengah" typeStatus="holotype">
|
||
<specimenCount id="9D449FC9FFFCFFA6325797A1AEECFB6F" box="[1365,1430,1215,1241]" pageId="3" pageNumber="429" type="female">2 ♀♀</specimenCount>
|
||
: “[
|
||
<collectingCountry id="F35514D0FFFCFFA637B697FDAA5BFB4B" box="[180,289,1251,1277]" name="Indonesia" pageId="3" pageNumber="429">Indonesia</collectingCountry>
|
||
] /
|
||
<location id="8E9D029BFFFCFFA6364B97FDAA8AFB4B" LSID="urn:lsid:plazi:treatment:03EBE556FFFCFFA0379590BDAFD9FEF7:8E9D029BFFFCFFA6364B97FDAA8AFB4B" box="[329,496,1251,1277]" country="Indonesia" name="Puncak Palopo" pageId="3" pageNumber="429" stateProvince="Sulawesi Tengah">Puncak Palopo</location>
|
||
pass, /
|
||
<collectingRegion id="49869AA2FFFCFFA6354797FDA872FB4B" box="[581,776,1251,1277]" country="Indonesia" name="Sulawesi Tengah" pageId="3" pageNumber="429">Central Sulawesi</collectingRegion>
|
||
. /
|
||
<date id="FFFC7280FFFCFFA6342097FDA8FBFB4B" box="[802,897,1251,1277]" pageId="3" pageNumber="429" value="2017-10">
|
||
<collectingDate id="EFB88B68FFFCFFA6342097FDA8FBFB4B" box="[802,897,1251,1277]" pageId="3" pageNumber="429" value="2017-10">X. 2017</collectingDate>
|
||
</date>
|
||
. /
|
||
<collectorName id="26B73196FFFCFFA6349897FDAF2EFB4B" box="[922,1108,1251,1277]" pageId="3" pageNumber="429">Native Collector</collectorName>
|
||
leg.” (RIEB)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B2A5E1DFFFCFFA633F997FDAA07FA97" ID-GBIF-Occurrence="3698900308" country="Indonesia" location="Peleng" pageId="3" pageNumber="429" specimenCount="1" specimenCount-male="1" stateProvince="Sulawesi Utara" typeStatus="holotype">
|
||
<specimenCount id="9D449FC9FFFCFFA633F997FDAE52FB4B" box="[1275,1320,1251,1277]" pageId="3" pageNumber="429" type="male">1 ♂</specimenCount>
|
||
: “
|
||
<location id="8E9D029BFFFCFFA6323C97FDAEF7FB4B" LSID="urn:lsid:plazi:treatment:03EBE556FFFCFFA0379590BDAFD9FEF7:8E9D029BFFFCFFA6323C97FDAEF7FB4B" box="[1342,1421,1251,1277]" country="Indonesia" name="Peleng" pageId="3" pageNumber="429" stateProvince="Sulawesi Utara">Peleng</location>
|
||
/ nr.
|
||
<collectingRegion id="49869AA2FFFCFFA637B99619AA6CFA97" box="[187,278,1287,1313]" country="Indonesia" name="Sulawesi Utara" pageId="3" pageNumber="429">Celebes</collectingRegion>
|
||
” (RIEB)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BFD5440FFFCFFA637C59634AF76F91B" blockId="3.[151,1437,1070,2033]" pageId="3" pageNumber="429">
|
||
<emphasis id="B9368852FFFCFFA637C59634AAAEFAF3" bold="true" box="[199,468,1322,1349]" pageId="3" pageNumber="429">Diagnostic characters.</emphasis>
|
||
Body shape ovoid, slightly bent laterally (
|
||
<figureCitation id="137948C5FFFCFFA634CA9635AF43FAF3" box="[968,1081,1323,1349]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1678,1702]" captionTargetBox="[162,1427,189,1638]" captionTargetId="figure-17@2.[151,1436,179,1652]" captionTargetPageId="2" captionText="FIGURES 1–9. Habitus of Malaia (M.) medea Kaneko & Wada, new species (1–6) and M. (M.) simulatrix Heller, 1892 ♂ from Pulau Peleng, Sulawesi (7–9). 1, holotype, dorsal view; 2, holotype, ventral view; 3, holotype, lateral view; 4, paratype, dorsal view (color variability); 5, allotype, dorsal view; 6, allotype, ventral view; 7, specimen from RIEB, dorsal view; 8, the same, ventral view; 9, the same, lateral view. Scale: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6332096" httpUri="https://zenodo.org/record/6332096/files/figure.png" pageId="3" pageNumber="429">Figs. 7–9</figureCitation>
|
||
). Color almost blackish-brown with a dull vitreous green gloss; legs black with green vitreous luster; elytra black with two or four reddish-brown maculae, or dark blue. Head distinctly convex; clypeus trapezoidal with rounded angles, weakly reflexed anterior margin, weakly concave at the center; frons shallowly depressed and without a median groove extending to the vertex. Pronotum with a pair of shallow oblique depressions in the middle to the anterior part of the lateral sides; anterior angles acute and protruding weakly; lateral sides of pronotum with dense short appressed yellowish-white scaly hairs along anterior to posterior angles. Pygidium convex and apex broadly rounded. Mesoventrites covered with dense appressed yellowish-white hairs. Abdominal ventrites covered with dense appressed yellowish-white hairs. Protibia with bidentate, apical, and proximal tooth developed. Meso- and metatibia short and fusiform. Aedeagus curved in lateral view, shape simple as shown in
|
||
<figureCitation id="137948C5FFFCFFA634379571A8C4F93F" box="[821,958,1647,1673]" captionStart="FIGURES 10–13" captionStartId="4.[151,264,1911,1935]" captionTargetBox="[234,1358,191,1880]" captionTargetId="figure-17@4.[215,1371,181,1887]" captionTargetPageId="4" captionText="FIGURES 10–13. Aedeagus of Malaia (M.) medea Kaneko & Wada, new species (10–11) and M. (M.) simulatrix Heller, 1892 ♂ from Pulau Peleng, Sulawesi (12–13). 10, holotype, dorsal view; 11, holotype, lateral view; 12, specimen from RIEB, dorsal view; 13, the same, lateral view. Scale: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6332099" httpUri="https://zenodo.org/record/6332099/files/figure.png" pageId="3" pageNumber="429">Figs. 12–13</figureCitation>
|
||
. Parameres almost symmetrical, with each lobe protruding at the apex, and multiple sparse short reddish setae at the apex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35807CBFFFCFFA637C595A8AFD0F964" box="[199,1194,1718,1746]" pageId="3" pageNumber="429" type="distribution">
|
||
<paragraph id="8BFD5440FFFCFFA637C595A8AFD0F964" blockId="3.[151,1437,1070,2033]" box="[199,1194,1718,1746]" pageId="3" pageNumber="429">
|
||
<emphasis id="B9368852FFFCFFA637C595A8AA1BF966" bold="true" box="[199,353,1718,1744]" pageId="3" pageNumber="429">Distribution.</emphasis>
|
||
<collectingCountry id="F35514D0FFFCFFA6366595A9AAA2F967" box="[359,472,1719,1745]" name="Indonesia" pageId="3" pageNumber="429">Indonesia</collectingCountry>
|
||
:
|
||
<collectingRegion id="49869AA2FFFCFFA636E195A9A932F967" box="[483,584,1719,1745]" country="Indonesia" name="Sulawesi Utara" pageId="3" pageNumber="429">Sulawesi</collectingRegion>
|
||
(
|
||
<taxonomicName id="4C422FC3FFFCFFA6355595A9AFE7F964" authority="Tengah, Pulau Banggai, and Pulau Peleng" authorityName="Tengah, Pulau Banggai, and Pulau Peleng" box="[599,1181,1719,1746]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="429" phylum="Arthropoda" rank="species" species="sulawesi">Sulawesi Tengah, Pulau Banggai, and Pulau Peleng</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35807CBFFFCFFA037C595C5AFD9FEF7" lastPageId="5" lastPageNumber="431" pageId="3" pageNumber="429" type="discussion">
|
||
<paragraph id="8BFD5440FFFCFFA637C595C5A9D5F847" blockId="3.[151,1437,1070,2033]" pageId="3" pageNumber="429">
|
||
<emphasis id="B9368852FFFCFFA637C595C5AA41F943" bold="true" box="[199,315,1755,1781]" pageId="3" pageNumber="429">Remarks.</emphasis>
|
||
<taxonomicName id="4C422FC3FFFCFFA6364195C5A974F943" authorityName="Heller" authorityYear="1892" box="[323,526,1755,1781]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="429" phylum="Arthropoda" rank="species" species="simulatrix">
|
||
<emphasis id="B9368852FFFCFFA6364195C5A974F943" box="[323,526,1755,1781]" italics="true" pageId="3" pageNumber="429">Malaia simulatrix</emphasis>
|
||
</taxonomicName>
|
||
was described by
|
||
<bibRefCitation id="EFD329B1FFFCFFA635E295C5A806F943" author="Heller, K. M." box="[736,892,1755,1781]" pageId="3" pageNumber="429" pagination="289 - 306" refId="ref4902" refString="Heller, K. M. (1892) Die mit der Ruteliden - Gattung Singhala Burm. verwandten Gattungen und Arten. Deutsche Entomologische Zeitschrift, 1891 (2), 289 - 306." type="journal article" year="1892">Heller (1892)</bibRefCitation>
|
||
. Later,
|
||
<bibRefCitation id="EFD329B1FFFCFFA634D395C5AFCDF943" author="Machatschke, J. W." box="[977,1207,1754,1781]" pageId="3" pageNumber="429" pagination="1 - 219" refId="ref4965" refString="Machatschke, J. W. (1957) Coleoptera Lamellicornia Fam. Scarabaeidae. Subfam. Rutelinae. Tribus Anomalini. In: Wytsman, P. A. G. (Ed.), Genera Insectorum. Fasc. 199 (B). Desmet-Verteneuil, Bruxelles, pp. 1 - 219, vi pls." type="book chapter" year="1957">Machatschke (1957)</bibRefCitation>
|
||
classified the genus
|
||
<taxonomicName id="4C422FC3FFFCFFA6379595E1AB92F8AF" authorityName="Heller" authorityYear="1892" box="[151,232,1791,1817]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="429" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9368852FFFCFFA6379595E1AB92F8AF" box="[151,232,1791,1817]" italics="true" pageId="3" pageNumber="429">Malaia</emphasis>
|
||
</taxonomicName>
|
||
into three subgenera, and assigned
|
||
<taxonomicName id="4C422FC3FFFCFFA6357C95E1A86DF8AF" authorityName="Heller" authorityYear="1892" box="[638,791,1791,1817]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="429" phylum="Arthropoda" rank="species" species="simulatrix">
|
||
<emphasis id="B9368852FFFCFFA6357C95E1A9EFF8AE" box="[638,661,1791,1816]" italics="true" pageId="3" pageNumber="429">M</emphasis>
|
||
.
|
||
<emphasis id="B9368852FFFCFFA635A795E1A86DF8AF" box="[677,791,1791,1817]" italics="true" pageId="3" pageNumber="429">simulatrix</emphasis>
|
||
</taxonomicName>
|
||
to the subgenus
|
||
<taxonomicName id="4C422FC3FFFCFFA634D595E1AF52F8AF" authorityName="Heller" authorityYear="1892" box="[983,1064,1791,1817]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="429" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9368852FFFCFFA634D595E1AF52F8AF" box="[983,1064,1791,1817]" italics="true" pageId="3" pageNumber="429">Malaia</emphasis>
|
||
</taxonomicName>
|
||
. Additionally,
|
||
<taxonomicName id="4C422FC3FFFCFFA633D295E1AE13F8AF" authorityName="Heller" authorityYear="1892" box="[1232,1385,1791,1817]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="429" phylum="Arthropoda" rank="species" species="simulatrix">
|
||
<emphasis id="B9368852FFFCFFA633D295E1AF9DF8AE" box="[1232,1255,1791,1816]" italics="true" pageId="3" pageNumber="429">M</emphasis>
|
||
.
|
||
<emphasis id="B9368852FFFCFFA633F595E1AE13F8AF" box="[1271,1385,1791,1817]" italics="true" pageId="3" pageNumber="429">simulatrix</emphasis>
|
||
</taxonomicName>
|
||
was assigned to the simulatrix group, one of the species groups included in the subgenus
|
||
<taxonomicName id="4C422FC3FFFCFFA6333B943DAFF0F88B" authorityName="Heller" authorityYear="1892" box="[1081,1162,1827,1853]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="429" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9368852FFFCFFA6333B943DAFF0F88B" box="[1081,1162,1827,1853]" italics="true" pageId="3" pageNumber="429">Malaia</emphasis>
|
||
</taxonomicName>
|
||
. However,
|
||
<bibRefCitation id="EFD329B1FFFCFFA6320B943DAB98F8D7" author="Machatschke, J. W." pageId="3" pageNumber="429" pagination="1 - 219" refId="ref4965" refString="Machatschke, J. W. (1957) Coleoptera Lamellicornia Fam. Scarabaeidae. Subfam. Rutelinae. Tribus Anomalini. In: Wytsman, P. A. G. (Ed.), Genera Insectorum. Fasc. 199 (B). Desmet-Verteneuil, Bruxelles, pp. 1 - 219, vi pls." type="book chapter" year="1957">Machatschke (1957)</bibRefCitation>
|
||
established five species groups in the subgenus
|
||
<taxonomicName id="4C422FC3FFFCFFA634019459A82EF8D7" authorityName="Heller" authorityYear="1892" box="[771,852,1863,1889]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="429" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9368852FFFCFFA634019459A82EF8D7" box="[771,852,1863,1889]" italics="true" pageId="3" pageNumber="429">Malaia</emphasis>
|
||
</taxonomicName>
|
||
without clearly defining the characteristics of those species groups, so the features that characterized each group remain unknown. Therefore, it will be necessary to review all species in the genus
|
||
<taxonomicName id="4C422FC3FFFCFFA636E99491A946F81F" authorityName="Heller" authorityYear="1892" box="[491,572,1935,1961]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="429" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9368852FFFCFFA636E99491A946F81F" box="[491,572,1935,1961]" italics="true" pageId="3" pageNumber="429">Malaia</emphasis>
|
||
</taxonomicName>
|
||
in the future to establish a definition of these species groups. Our examination showed some important characters for differentiating
|
||
<taxonomicName id="4C422FC3FFFCFFA6341194ADA894F87B" authorityName="Heller" authorityYear="1892" box="[787,1006,1971,1997]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="429" phylum="Arthropoda" rank="species" species="simulatrix" subGenus="M.">
|
||
<emphasis id="B9368852FFFCFFA6341194ADA850F87A" box="[787,810,1971,1996]" italics="true" pageId="3" pageNumber="429">M</emphasis>
|
||
. (
|
||
<emphasis id="B9368852FFFCFFA6344A94ADA825F87A" box="[840,863,1971,1996]" italics="true" pageId="3" pageNumber="429">M</emphasis>
|
||
.)
|
||
<emphasis id="B9368852FFFCFFA6347E94ADA894F87B" box="[892,1006,1971,1997]" italics="true" pageId="3" pageNumber="429">simulatrix</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<emphasis id="B9368852FFFCFFA6333C94ADAF2FF87A" box="[1086,1109,1971,1996]" italics="true" pageId="3" pageNumber="429">M</emphasis>
|
||
.
|
||
<taxonomicName id="4C422FC3FFFCFFA6336B94ADAF95F87B" authorityName="Kaneko & Wada" authorityYear="2022" box="[1129,1263,1971,1997]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="429" phylum="Arthropoda" rank="species" species="medea" subGenus="M">
|
||
(
|
||
<emphasis id="B9368852FFFCFFA6337094ADAFF3F87A" box="[1138,1161,1971,1996]" italics="true" pageId="3" pageNumber="429">M</emphasis>
|
||
.)
|
||
<emphasis id="B9368852FFFCFFA633A494ADAF95F87B" box="[1190,1263,1971,1997]" italics="true" pageId="3" pageNumber="429">medea</emphasis>
|
||
</taxonomicName>
|
||
. For a clearer differentiation, see the identification key below.
|
||
</paragraph>
|
||
<caption id="DF3D04C8FFFBFFA137959469A907F860" ID-DOI="http://doi.org/10.5281/zenodo.6332099" ID-Zenodo-Dep="6332099" httpUri="https://zenodo.org/record/6332099/files/figure.png" pageId="4" pageNumber="430" startId="4.[151,264,1911,1935]" targetBox="[234,1358,191,1880]" targetPageId="4">
|
||
<paragraph id="8BFD5440FFFBFFA137959469A907F860" blockId="4.[151,1437,1911,2007]" pageId="4" pageNumber="430">
|
||
<emphasis id="B9368852FFFBFFA137959469AA2EF839" bold="true" box="[151,340,1911,1935]" pageId="4" pageNumber="430">FIGURES 10–13.</emphasis>
|
||
Aedeagus of
|
||
<taxonomicName id="4C422FC3FFFBFFA136DC9469A95DF839" authorityName="Heller" authorityYear="1892" box="[478,551,1911,1935]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="430" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9368852FFFBFFA136DC9469A95DF839" box="[478,551,1911,1935]" italics="true" pageId="4" pageNumber="430">Malaia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicName id="4C422FC3FFFBFFA1352C9469A829F839" authority="Kaneko & Wada" authorityName="Kaneko & Wada" authorityYear="2022" box="[558,851,1911,1935]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="430" phylum="Arthropoda" rank="species" species="medea" status="sp. nov." subGenus="M">
|
||
(
|
||
<emphasis id="B9368852FFFBFFA135349466A931F839" box="[566,587,1912,1935]" italics="true" pageId="4" pageNumber="430">M</emphasis>
|
||
.)
|
||
<emphasis id="B9368852FFFBFFA135629469A9D8F839" box="[608,674,1911,1935]" italics="true" pageId="4" pageNumber="430">medea</emphasis>
|
||
Kaneko & Wada
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2053529FFFBFFA1345F9469A8ACF839" box="[861,982,1911,1935]" pageId="4" pageNumber="430" rank="species">new species</taxonomicNameLabel>
|
||
(10–11) and
|
||
<emphasis id="B9368852FFFBFFA1335F9466AF00F839" box="[1117,1146,1912,1935]" italics="true" pageId="4" pageNumber="430">M.</emphasis>
|
||
<taxonomicName id="4C422FC3FFFBFFA1337D9469AEE7F839" authority="Heller, 1892" authorityName="Heller" authorityYear="1892" box="[1151,1437,1911,1935]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="430" phylum="Arthropoda" rank="species" species="simulatrix" subGenus="M">
|
||
(
|
||
<emphasis id="B9368852FFFBFFA133859466AFE6F839" box="[1159,1180,1912,1935]" italics="true" pageId="4" pageNumber="430">M</emphasis>
|
||
.)
|
||
<emphasis id="B9368852FFFBFFA133B09469AE63F839" box="[1202,1305,1911,1935]" italics="true" pageId="4" pageNumber="430">simulatrix</emphasis>
|
||
<bibRefCitation id="EFD329B1FFFBFFA1321D9469AEE7F839" author="Heller, K. M." box="[1311,1437,1911,1935]" pageId="4" pageNumber="430" pagination="289 - 306" refId="ref4902" refString="Heller, K. M. (1892) Die mit der Ruteliden - Gattung Singhala Burm. verwandten Gattungen und Arten. Deutsche Entomologische Zeitschrift, 1891 (2), 289 - 306." type="journal article" year="1892">Heller, 1892</bibRefCitation>
|
||
</taxonomicName>
|
||
♂ from Pulau Peleng, Sulawesi (12–13). 10, holotype, dorsal view; 11, holotype, lateral view; 12, specimen from RIEB, dorsal view; 13, the same, lateral view. Scale: 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFD5440FFFAFFA037C59389AFD9FEF7" blockId="5.[151,1436,151,321]" pageId="5" pageNumber="431">
|
||
Unfortunately, we were not able to examine the
|
||
<typeStatus id="54F9EAE2FFFAFFA035FB9389A821FF07" box="[761,859,151,177]" pageId="5" pageNumber="431" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C422FC3FFFAFFA034849389AF26FF07" authorityName="Heller" authorityYear="1892" box="[902,1116,151,177]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="429" phylum="Arthropoda" rank="species" species="simulatrix" subGenus="M.">
|
||
<emphasis id="B9368852FFFAFFA034849389A8E7FF06" box="[902,925,151,176]" italics="true" pageId="5" pageNumber="431">M</emphasis>
|
||
. (
|
||
<emphasis id="B9368852FFFAFFA034BA9389A8B5FF06" box="[952,975,151,176]" italics="true" pageId="5" pageNumber="431">M</emphasis>
|
||
.)
|
||
<emphasis id="B9368852FFFAFFA034E89389AF26FF07" box="[1002,1116,151,177]" italics="true" pageId="5" pageNumber="431">simulatrix</emphasis>
|
||
</taxonomicName>
|
||
in this study. We contacted Senckenberg Natural History Collections Dresden (
|
||
<collectingCountry id="F35514D0FFFAFFA035DB93A5A83FFF63" box="[729,837,187,213]" name="Germany" pageId="5" pageNumber="431">Germany</collectingCountry>
|
||
) and Dr. Guido Sabatinelli (Natural History Museum of
|
||
<collectingRegion id="49869AA2FFFAFFA037B793C1AA77FF4F" box="[181,269,223,249]" country="Switzerland" name="Geneve" pageId="5" pageNumber="431">Geneva</collectingRegion>
|
||
,
|
||
<collectingCountry id="F35514D0FFFAFFA0361A93C1AAD8FF4C" box="[280,418,223,250]" name="Switzerland" pageId="5" pageNumber="431">Switzerland</collectingCountry>
|
||
), who had knowledge of whereabouts of Heller’s type material, but the
|
||
<typeStatus id="54F9EAE2FFFAFFA033C293C1AE58FF4F" box="[1216,1314,223,249]" pageId="5" pageNumber="431" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C422FC3FFFAFFA0324493C1AA73FEAB" authorityName="Heller" authorityYear="1892" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="429" phylum="Arthropoda" rank="species" species="simulatrix" subGenus="M.">
|
||
<emphasis id="B9368852FFFAFFA0324493C1AE27FF4E" box="[1350,1373,223,248]" italics="true" pageId="5" pageNumber="431">M</emphasis>
|
||
. (
|
||
<emphasis id="B9368852FFFAFFA0327793C1AEF6FF4E" box="[1397,1420,223,248]" italics="true" pageId="5" pageNumber="431">M</emphasis>
|
||
.)
|
||
<emphasis id="B9368852FFFAFFA03795921DAA73FEAB" box="[151,265,259,285]" italics="true" pageId="5" pageNumber="431">simulatrix</emphasis>
|
||
</taxonomicName>
|
||
had not been found and seemed to be missing. Therefore, we identified
|
||
<taxonomicName id="4C422FC3FFFAFFA0333A921DAE72FEAB" authorityName="Heller" authorityYear="1892" box="[1080,1288,259,285]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="429" phylum="Arthropoda" rank="species" species="simulatrix" subGenus="M.">
|
||
<emphasis id="B9368852FFFAFFA0333A921DAF35FEAA" box="[1080,1103,259,284]" italics="true" pageId="5" pageNumber="431">M</emphasis>
|
||
. (
|
||
<emphasis id="B9368852FFFAFFA03365921DAF04FEAA" box="[1127,1150,259,284]" italics="true" pageId="5" pageNumber="431">M</emphasis>
|
||
.)
|
||
<emphasis id="B9368852FFFAFFA03394921DAE72FEAB" box="[1174,1288,259,285]" italics="true" pageId="5" pageNumber="431">simulatrix</emphasis>
|
||
</taxonomicName>
|
||
based on the original description and the photograph of
|
||
<emphasis id="B9368852FFFAFFA035749239A9F7FEF6" box="[630,653,295,320]" italics="true" pageId="5" pageNumber="431">M</emphasis>
|
||
.
|
||
<taxonomicName id="4C422FC3FFFAFFA035999239AFD9FEF7" authority="Heller, 1892 sensu Sabatinelli." authorityName="Heller" authorityYear="1892" box="[667,1187,295,321]" class="Insecta" family="Rutelidae" genus="Malaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="431" phylum="Arthropoda" rank="species" species="simulatrix" subGenus="M">
|
||
(
|
||
<emphasis id="B9368852FFFAFFA035A79239A9C6FEF6" box="[677,700,295,320]" italics="true" pageId="5" pageNumber="431">M</emphasis>
|
||
.)
|
||
<emphasis id="B9368852FFFAFFA035D19239A83FFEF7" box="[723,837,295,321]" italics="true" pageId="5" pageNumber="431">simulatrix</emphasis>
|
||
<bibRefCitation id="EFD329B1FFFAFFA0344E9239A8ADFEF7" author="Heller, K. M." box="[844,983,295,321]" pageId="5" pageNumber="431" pagination="289 - 306" refId="ref4902" refString="Heller, K. M. (1892) Die mit der Ruteliden - Gattung Singhala Burm. verwandten Gattungen und Arten. Deutsche Entomologische Zeitschrift, 1891 (2), 289 - 306." type="journal article" year="1892">Heller, 1892</bibRefCitation>
|
||
sensu Sabatinelli.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |