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<document ID-DOI="10.5733/afin.054.0213" ID-ISSN="2305-2562" ID-Zenodo-Dep="7918661" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1683583268648" checkinUser="felipe" docAuthor="Niekerk, P. van &amp; Dippenaar-Schoeman, A. S." docDate="2013" docId="03D8D254FFE2A9242251FAAAFE3FF96A" docLanguage="en" docName="AfrInvertebr.54.2.447-476.pdf" docOrigin="African Invertebrates 54 (2)" docSource="http://www.bioone.org/doi/abs/10.5733/afin.054.0213" docStyle="DocumentStyle:B78C63EC8CEBBB75364A82B35BB1F83D.4:AfrInvertebr.2008-2015.journal_article.0cover" docStyleId="B78C63EC8CEBBB75364A82B35BB1F83D" docStyleName="AfrInvertebr.2008-2015.journal_article.0cover" docStyleVersion="4" docTitle="Heriaeus copricola Niekerk &amp; Dippenaar-Schoeman 2013, sp. n." docType="treatment" docVersion="4" lastPageNumber="458" masterDocId="FFE1AA2CFFEBA92F2055FF99FFE4FFE6" masterDocTitle="A revision of the crab spider genus Heriaeus Simon, 1875 (Araneae: Thomisidae) in the Afrotropical Region" masterLastPageNumber="447" masterPageNumber="447" pageNumber="456" updateTime="1683701492856" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the crab spider genus Heriaeus Simon, 1875 (Araneae: Thomisidae) in the Afrotropical Region</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Niekerk, P. van</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dippenaar-Schoeman, A. S.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>African Invertebrates</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="pubDate">
<mods:number>2013-11-14</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>54</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>447</mods:start>
<mods:end>447</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://www.bioone.org/doi/abs/10.5733/afin.054.0213</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5733/afin.054.0213</mods:identifier>
<mods:identifier type="ISSN">2305-2562</mods:identifier>
<mods:identifier type="Zenodo-Dep">7918661</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7917977" ID-Zenodo-Dep="7917977" LSID="urn:lsid:plazi:treatment:03D8D254FFE2A9242251FAAAFE3FF96A" httpUri="http://treatment.plazi.org/id/03D8D254FFE2A9242251FAAAFE3FF96A" lastPageId="11" lastPageNumber="458" pageId="9" pageNumber="456">
<subSubSection box="[516,807,1331,1358]" pageId="9" pageNumber="456" type="nomenclature">
<paragraph blockId="9.[516,807,1331,1398]" box="[516,807,1331,1358]" pageId="9" pageNumber="456">
<heading box="[516,807,1331,1358]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="456" reason="2">
<taxonomicName ID-CoL="6M458" authority="Niekerk &amp; Dippenaar-Schoeman, 2013" authorityName="Niekerk &amp; Dippenaar-Schoeman" authorityYear="2013" box="[516,740,1331,1357]" class="Arachnida" family="Thomisidae" genus="Heriaeus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="456" phylum="Arthropoda" rank="species" species="copricola" status="sp. n.">
<emphasis bold="true" box="[516,740,1331,1357]" pageId="9" pageNumber="456">Heriaeus copricola</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[747,807,1332,1358]" pageId="9" pageNumber="456" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="11" lastPageNumber="458" pageId="9" pageNumber="456" type="description">
<paragraph blockId="9.[516,807,1331,1398]" box="[562,761,1371,1398]" pageId="9" pageNumber="456">
<figureCitation box="[562,633,1372,1398]" captionStart="Figs 1–9" captionStartId="3.[181,220,1090,1112]" captionTargetBox="[187,1132,204,1065]" captionTargetId="figure-241@3.[187,1132,204,1065]" captionTargetPageId="3" captionText="Figs 1–9. Heriaeus spp., habitus:(1) Heriaeus sp. long spiniform abdominal setae; (2) H.allenjonesi antoni sp. n. female; (3) H.antoni sp. n. female; (4) H.copricola sp. n. male; (5) H. crassispinus Lawrence, 1942 male; (6) H. foordi sp. n. male; (7) H. latifrons Lessert, 1919 female; (8) H.madagascar sp. n. male; (9) H. muizenberg sp. n. female." figureDoi="http://doi.org/10.5281/zenodo.7918663" httpUri="https://zenodo.org/record/7918663/files/figure.png" pageId="9" pageNumber="456">Figs 4</figureCitation>
,
<figureCitation box="[645,721,1371,1398]" captionStart="Figs 22–29" captionStartId="10.[144,183,1291,1313]" captionTargetBox="[178,1086,207,1264]" captionTargetId="figure-133@10.[103,1161,204,1272]" captionTargetPageId="10" captionText="Figs 22–29. (22–25) Heriaeus copricola sp. n.: (22) male palp, ventral view; (23) male palp, lateral view; (24) epigyne, ventral view; (25) epigyne, dorsal view; (26–29) Heriaeus crassispinus Lawrence, 1942: (26) male palp, ventral view; (27) male palp, lateral view; (28) epigyne, ventral view; (29) epigyne, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7918669" httpUri="https://zenodo.org/record/7918669/files/figure.png" pageId="9" pageNumber="456">22–25</figureCitation>
,
<figureCitation box="[733,761,1371,1398]" captionStart="Figs 57–59" captionStartId="27.[347,386,1654,1676]" captionTargetBox="[181,1141,609,1633]" captionTargetId="figure-176@27.[132,1190,602,1636]" captionTargetPageId="27" captionText="Figs 57–59. Distribution of Heriaeus spp. in the Afrotropical Region." figureDoi="http://doi.org/10.5281/zenodo.7918679" httpUri="https://zenodo.org/record/7918679/files/figure.png" pageId="9" pageNumber="456">59</figureCitation>
</paragraph>
<paragraph blockId="9.[181,1142,1414,1505]" pageId="9" pageNumber="456">
Etymology: From Latin
<emphasis box="[454,529,1415,1440]" italics="true" pageId="9" pageNumber="456">copros</emphasis>
(dung) and
<emphasis box="[665,734,1414,1440]" italics="true" pageId="9" pageNumber="456">colere</emphasis>
(to dwell in). The species name was suggested by Dr R.F. Lawrence (
<emphasis box="[558,620,1446,1472]" italics="true" pageId="9" pageNumber="456">in litt</emphasis>
.), who marked the
<typeStatus box="[844,892,1447,1473]" pageId="9" pageNumber="456">type</typeStatus>
specimens as being a new species but did not publish its description.
</paragraph>
<paragraph blockId="9.[181,1142,1521,1675]" lastBlockId="10.[144,1104,1431,1522]" lastPageId="10" lastPageNumber="457" pageId="9" pageNumber="456">
Diagnosis: The species is recognized by the combination of long, slender, blunt abdominal setae and the epigyne having a shallow hood over a small round opening (
<figureCitation captionStart="Figs 22–29" captionStartId="10.[144,183,1291,1313]" captionTargetBox="[178,1086,207,1264]" captionTargetId="figure-133@10.[103,1161,204,1272]" captionTargetPageId="10" captionText="Figs 22–29. (22–25) Heriaeus copricola sp. n.: (22) male palp, ventral view; (23) male palp, lateral view; (24) epigyne, ventral view; (25) epigyne, dorsal view; (26–29) Heriaeus crassispinus Lawrence, 1942: (26) male palp, ventral view; (27) male palp, lateral view; (28) epigyne, ventral view; (29) epigyne, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7918669" httpUri="https://zenodo.org/record/7918669/files/figure.png" pageId="9" pageNumber="456">Fig. 24</figureCitation>
), with the internal organs visible as two light brown kidney-shaped structures below the hood (
<figureCitation box="[290,372,1617,1643]" captionStart="Figs 22–29" captionStartId="10.[144,183,1291,1313]" captionTargetBox="[178,1086,207,1264]" captionTargetId="figure-133@10.[103,1161,204,1272]" captionTargetPageId="10" captionText="Figs 22–29. (22–25) Heriaeus copricola sp. n.: (22) male palp, ventral view; (23) male palp, lateral view; (24) epigyne, ventral view; (25) epigyne, dorsal view; (26–29) Heriaeus crassispinus Lawrence, 1942: (26) male palp, ventral view; (27) male palp, lateral view; (28) epigyne, ventral view; (29) epigyne, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7918669" httpUri="https://zenodo.org/record/7918669/files/figure.png" pageId="9" pageNumber="456">Fig. 24</figureCitation>
). The embolus of the male palp is short, winding less than once around tegulum, with the tip slender and straight; VTA broad (
<figureCitation box="[798,881,1649,1675]" captionStart="Figs 22–29" captionStartId="10.[144,183,1291,1313]" captionTargetBox="[178,1086,207,1264]" captionTargetId="figure-133@10.[103,1161,204,1272]" captionTargetPageId="10" captionText="Figs 22–29. (22–25) Heriaeus copricola sp. n.: (22) male palp, ventral view; (23) male palp, lateral view; (24) epigyne, ventral view; (25) epigyne, dorsal view; (26–29) Heriaeus crassispinus Lawrence, 1942: (26) male palp, ventral view; (27) male palp, lateral view; (28) epigyne, ventral view; (29) epigyne, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7918669" httpUri="https://zenodo.org/record/7918669/files/figure.png" pageId="9" pageNumber="456">Fig. 23</figureCitation>
); RTA with short distal tooth when viewed laterally (
<figureCitation box="[473,558,1431,1457]" captionStart="Figs 22–29" captionStartId="10.[144,183,1291,1313]" captionTargetBox="[178,1086,207,1264]" captionTargetId="figure-133@10.[103,1161,204,1272]" captionTargetPageId="10" captionText="Figs 22–29. (22–25) Heriaeus copricola sp. n.: (22) male palp, ventral view; (23) male palp, lateral view; (24) epigyne, ventral view; (25) epigyne, dorsal view; (26–29) Heriaeus crassispinus Lawrence, 1942: (26) male palp, ventral view; (27) male palp, lateral view; (28) epigyne, ventral view; (29) epigyne, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7918669" httpUri="https://zenodo.org/record/7918669/files/figure.png" pageId="10" pageNumber="457">Fig. 22</figureCitation>
). The male palp resembles that of
<taxonomicName authority="Niekerk &amp; Dippenaar-Schoeman, 2013" authorityName="Niekerk &amp; Dippenaar-Schoeman" authorityYear="2013" box="[946,1035,1432,1457]" class="Arachnida" family="Thomisidae" genus="Heriaeus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="457" phylum="Arthropoda" rank="species" species="zanii" status="sp. n.">
<emphasis box="[946,1035,1432,1457]" italics="true" pageId="10" pageNumber="457">H. zanii</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1043,1104,1432,1458]" pageId="10" pageNumber="457" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation box="[152,235,1463,1490]" captionStart="Figs 53–56" captionStartId="26.[144,183,962,984]" captionTargetBox="[160,1088,205,942]" captionTargetId="figure-284@26.[95,1153,202,945]" captionTargetPageId="26" captionText="Figs 53–56.Heriaeus zanii sp.n.: (53) male palp, ventral view; (54) male lateral view; (55) epigyne, ventral view; (56) epigyne, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7918677" httpUri="https://zenodo.org/record/7918677/files/figure.png" pageId="10" pageNumber="457">Fig. 53</figureCitation>
) but differs in the shape of the RTA, having a single tooth and the uniquely shaped abdominal setae (
<figureCitation box="[425,494,1495,1521]" captionStart="Figs 1–9" captionStartId="3.[181,220,1090,1112]" captionTargetBox="[187,1132,204,1065]" captionTargetId="figure-241@3.[187,1132,204,1065]" captionTargetPageId="3" captionText="Figs 1–9. Heriaeus spp., habitus:(1) Heriaeus sp. long spiniform abdominal setae; (2) H.allenjonesi antoni sp. n. female; (3) H.antoni sp. n. female; (4) H.copricola sp. n. male; (5) H. crassispinus Lawrence, 1942 male; (6) H. foordi sp. n. male; (7) H. latifrons Lessert, 1919 female; (8) H.madagascar sp. n. male; (9) H. muizenberg sp. n. female." figureDoi="http://doi.org/10.5281/zenodo.7918663" httpUri="https://zenodo.org/record/7918663/files/figure.png" pageId="10" pageNumber="457">Fig. 4</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7918669" ID-Zenodo-Dep="7918669" httpUri="https://zenodo.org/record/7918669/files/figure.png" pageId="10" pageNumber="457" startId="10.[144,183,1291,1313]" targetBox="[178,1086,207,1264]" targetPageId="10" targetType="figure">
<paragraph blockId="10.[144,1104,1291,1385]" pageId="10" pageNumber="457">
Figs 22–29. (22–25)
<taxonomicName authority="Niekerk &amp; Dippenaar-Schoeman, 2013" authorityName="Niekerk &amp; Dippenaar-Schoeman" authorityYear="2013" box="[339,515,1291,1313]" class="Arachnida" family="Thomisidae" genus="Heriaeus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="457" phylum="Arthropoda" rank="species" species="copricola" status="sp. n.">
<emphasis box="[339,515,1291,1313]" italics="true" pageId="10" pageNumber="457">Heriaeus copricola</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[520,571,1292,1313]" pageId="10" pageNumber="457" rank="species">sp. n.</taxonomicNameLabel>
: (22) male palp, ventral view; (23) male palp, lateral view; (24) epigyne, ventral view; (25) epigyne, dorsal view; (26–29)
<taxonomicName authority="Lawrence, 1942" authorityName=": Lawrence" authorityYear="1942" class="Arachnida" family="Thomisidae" genus="Heriaeus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="457" phylum="Arthropoda" rank="species" species="crassispinus">
<emphasis box="[799,1001,1316,1337]" italics="true" pageId="10" pageNumber="457">Heriaeus crassispinus</emphasis>
<bibRefCitation author="LAWRENCE, R. F." pageId="10" pageNumber="457" pagination="141 - 190" refId="ref19954" refString="LAWRENCE, R. F. 1942. A contribution to the araneid fauna of Natal and Zululand. Annals of the Natal Museum 10: 141 - 190." type="journal article" year="1942">Lawrence, 1942</bibRefCitation>
</taxonomicName>
: (26) male palp, ventral view; (27) male palp, lateral view; (28) epigyne, ventral view; (29) epigyne, dorsal view.
</paragraph>
</caption>
<paragraph blockId="10.[144,1103,1538,1676]" box="[144,283,1538,1564]" pageId="10" pageNumber="457">Description:</paragraph>
<paragraph blockId="10.[144,1103,1538,1676]" box="[144,351,1578,1604]" pageId="10" pageNumber="457">
<emphasis box="[144,228,1578,1604]" italics="true" pageId="10" pageNumber="457">Female</emphasis>
(
<typeStatus box="[243,339,1578,1604]" pageId="10" pageNumber="457">allotype</typeStatus>
).
</paragraph>
<paragraph blockId="10.[144,1103,1538,1676]" lastBlockId="11.[181,1142,204,678]" lastPageId="11" lastPageNumber="458" pageId="10" pageNumber="457">
<emphasis box="[144,189,1618,1644]" italics="true" pageId="10" pageNumber="457">Size</emphasis>
: TL 4.88, CL 2.31, CW 2.04.
<emphasis box="[515,594,1618,1644]" italics="true" pageId="10" pageNumber="457">Colour</emphasis>
: Carapace with two longitudinal brown bands extending to PEs; setae brown; clypeus
<emphasis box="[590,609,1651,1676]" italics="true" pageId="10" pageNumber="457">w</emphasis>
ith brown marks; sternum white along edge; chelicerae yellow-brown; eye region white. Abdomen beige, mottled with white, with faint transverse lines posteriorly as well as a number of dark brown markings; venter white with fine hairs, with some brown spots laterally. Legs beige with brown spot on proximal end of tibiae and distal ends of metatarsi III and IV; some white mottling on coxae.
<emphasis box="[259,369,332,358]" italics="true" pageId="11" pageNumber="458">Carapace</emphasis>
:
<emphasis box="[383,589,332,357]" italics="true" pageId="11" pageNumber="458">Eye measurements</emphasis>
: AME–AME 0.14, ALE–AME 0.30, PME–PME 0.22, PLE–PME 0.31, AME–PME 0.24, MOQL AME–PME 0.31, MOQAW AME– AME 0.36, MOQPW PME–PME 0.28.
<emphasis box="[626,833,396,421]" italics="true" pageId="11" pageNumber="458">Leg measurements</emphasis>
: leg I Fe 1.89, Pat 1.04, Tib 1.76, Mt 1.80,Ta 0.51, total 7.0; leg II Fe 1,50, Pat 0.82, Tib 1.23, Mt 1.23, Ta 0.36, total 5.14; leg III Fe 0.92, Pat 0.59, Tib 0.74, Mt 0.74, Ta 0.55, total 3.54; leg IV Fe 1.37, Pat 0.41, Tib 0.94, Mt 0.76, Ta 0.48, total 3.96.
<emphasis box="[724,830,492,518]" italics="true" pageId="11" pageNumber="458">Abdomen</emphasis>
: Oval, truncated anteriorly, with long slender either transparent or pale brown setae with blunt tips; darker brown setae along lateral edge of dorsum.
<emphasis box="[578,670,556,581]" italics="true" pageId="11" pageNumber="458">Epigyne</emphasis>
: Hood shallow with small round opening, internal organs visible as two pale brown kidney-shaped structures, touching anteriorly and narrower posteriorly (
<figureCitation box="[464,544,620,646]" captionStart="Figs 22–29" captionStartId="10.[144,183,1291,1313]" captionTargetBox="[178,1086,207,1264]" captionTargetId="figure-133@10.[103,1161,204,1272]" captionTargetPageId="10" captionText="Figs 22–29. (22–25) Heriaeus copricola sp. n.: (22) male palp, ventral view; (23) male palp, lateral view; (24) epigyne, ventral view; (25) epigyne, dorsal view; (26–29) Heriaeus crassispinus Lawrence, 1942: (26) male palp, ventral view; (27) male palp, lateral view; (28) epigyne, ventral view; (29) epigyne, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7918669" httpUri="https://zenodo.org/record/7918669/files/figure.png" pageId="11" pageNumber="458">Fig. 24</figureCitation>
); copulatory ducts large and bulbous anteriorly, tubular posteriorly (
<figureCitation box="[318,402,651,678]" captionStart="Figs 22–29" captionStartId="10.[144,183,1291,1313]" captionTargetBox="[178,1086,207,1264]" captionTargetId="figure-133@10.[103,1161,204,1272]" captionTargetPageId="10" captionText="Figs 22–29. (22–25) Heriaeus copricola sp. n.: (22) male palp, ventral view; (23) male palp, lateral view; (24) epigyne, ventral view; (25) epigyne, dorsal view; (26–29) Heriaeus crassispinus Lawrence, 1942: (26) male palp, ventral view; (27) male palp, lateral view; (28) epigyne, ventral view; (29) epigyne, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7918669" httpUri="https://zenodo.org/record/7918669/files/figure.png" pageId="11" pageNumber="458">Fig. 25</figureCitation>
).
</paragraph>
<paragraph blockId="11.[181,370,697,723]" box="[181,370,697,723]" pageId="11" pageNumber="458">
<emphasis box="[181,239,697,723]" italics="true" pageId="11" pageNumber="458">Male</emphasis>
(
<typeStatus box="[254,359,697,723]" pageId="11" pageNumber="458">holotype</typeStatus>
).
</paragraph>
<paragraph blockId="11.[181,1142,740,1214]" pageId="11" pageNumber="458">
<emphasis box="[181,226,740,766]" italics="true" pageId="11" pageNumber="458">Size</emphasis>
: TL 3.88, CL 1.89, CW 1.70.
<emphasis box="[553,632,740,766]" italics="true" pageId="11" pageNumber="458">Colour</emphasis>
: Carapace with two longitudinal brown bands extending to PEs and into eye region; setae brown (
<figureCitation box="[764,834,772,798]" captionStart="Figs 1–9" captionStartId="3.[181,220,1090,1112]" captionTargetBox="[187,1132,204,1065]" captionTargetId="figure-241@3.[187,1132,204,1065]" captionTargetPageId="3" captionText="Figs 1–9. Heriaeus spp., habitus:(1) Heriaeus sp. long spiniform abdominal setae; (2) H.allenjonesi antoni sp. n. female; (3) H.antoni sp. n. female; (4) H.copricola sp. n. male; (5) H. crassispinus Lawrence, 1942 male; (6) H. foordi sp. n. male; (7) H. latifrons Lessert, 1919 female; (8) H.madagascar sp. n. male; (9) H. muizenberg sp. n. female." figureDoi="http://doi.org/10.5281/zenodo.7918663" httpUri="https://zenodo.org/record/7918663/files/figure.png" pageId="11" pageNumber="458">Fig. 4</figureCitation>
); chelicerae yellow-brown; clypeus with brown marks; eye region white. Abdomen brown, mottled with white; laterally with some brown marks; setae on abdomen dark brown with dark spots at their bases laterally; venter white, covered with fine hairs. Legs yellow­brown, with white spot on each coxa.
<emphasis box="[458,568,900,926]" italics="true" pageId="11" pageNumber="458">Carapace</emphasis>
:
<emphasis box="[581,788,900,925]" italics="true" pageId="11" pageNumber="458">Eye measurements</emphasis>
: AME–AME 0.23, ALE–AME 0.53, PME–PME 0.23, PLE–PME 0.30, AME–PME 0.21, MOQL AME–PME 0.29, MOQAW AME–AME 0.28, MOQPW PME–PME 0.29.
<emphasis box="[829,1038,964,989]" italics="true" pageId="11" pageNumber="458">Leg measurements</emphasis>
: leg I Fe 2.69, Pat 0.76, Tib 2.23, Mt 2.17, Ta 1.06, total 8.91; leg II Fe 2.26, Pat 0.64, Tib 1.76, Mt 1.56, Ta 0.99, total 7.21; leg III Fe 1.31, Pat 0.44, Tib 1.02, Mt 0.71, Ta 0.58, total 4.06; leg IV Fe 1.62, Pat 0.37, Tib 0.97, Mt 0.87, Ta 0.54, total 4.37.
<emphasis box="[959,1065,1060,1086]" italics="true" pageId="11" pageNumber="458">Abdomen</emphasis>
: Oval, truncated anteriorly, but narrower than that of female.
<emphasis box="[760,870,1092,1118]" italics="true" pageId="11" pageNumber="458">Male palp</emphasis>
: Embolus short, winding less than once around tegulum; embolus tip slender and straight (
<figureCitation box="[913,997,1124,1150]" captionStart="Figs 22–29" captionStartId="10.[144,183,1291,1313]" captionTargetBox="[178,1086,207,1264]" captionTargetId="figure-133@10.[103,1161,204,1272]" captionTargetPageId="10" captionText="Figs 22–29. (22–25) Heriaeus copricola sp. n.: (22) male palp, ventral view; (23) male palp, lateral view; (24) epigyne, ventral view; (25) epigyne, dorsal view; (26–29) Heriaeus crassispinus Lawrence, 1942: (26) male palp, ventral view; (27) male palp, lateral view; (28) epigyne, ventral view; (29) epigyne, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7918669" httpUri="https://zenodo.org/record/7918669/files/figure.png" pageId="11" pageNumber="458">Fig. 22</figureCitation>
); VTA short, somewhat flattened but variable in shape; RTA broad anteriorly, ending in short distal tooth viewed laterally (
<figureCitation box="[441,525,1188,1214]" captionStart="Figs 22–29" captionStartId="10.[144,183,1291,1313]" captionTargetBox="[178,1086,207,1264]" captionTargetId="figure-133@10.[103,1161,204,1272]" captionTargetPageId="10" captionText="Figs 22–29. (22–25) Heriaeus copricola sp. n.: (22) male palp, ventral view; (23) male palp, lateral view; (24) epigyne, ventral view; (25) epigyne, dorsal view; (26–29) Heriaeus crassispinus Lawrence, 1942: (26) male palp, ventral view; (27) male palp, lateral view; (28) epigyne, ventral view; (29) epigyne, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7918669" httpUri="https://zenodo.org/record/7918669/files/figure.png" pageId="11" pageNumber="458">Fig. 23</figureCitation>
).
</paragraph>
<paragraph blockId="11.[181,1141,1233,1320]" box="[181,706,1233,1260]" pageId="11" pageNumber="458">
<emphasis box="[181,293,1233,1259]" italics="true" pageId="11" pageNumber="458">Juveniles.</emphasis>
Recognized by long and blunt setae.
</paragraph>
<paragraph blockId="11.[181,1141,1233,1320]" pageId="11" pageNumber="458">
<materialsCitation collectingDate="1942-11" collectionCode="NMSA" collectorName="R. F. Lawrence" country="South Africa" latitude="-29.6" location="Pietermaritzburg" longLatPrecision="736" longitude="30.38" pageId="11" pageNumber="458" specimenCode="NMSA 3844" specimenCount="1" specimenCount-male="1" stateProvince="KwaZulu-Natal" typeStatus="holotype">
<typeStatus box="[181,267,1275,1297]" pageId="11" pageNumber="458">Holotype</typeStatus>
:
<specimenCount box="[274,291,1275,1296]" pageId="11" pageNumber="458" type="male">♂</specimenCount>
<collectingCountry box="[295,457,1275,1296]" name="South Africa" pageId="11" pageNumber="458">SOUTH AFRICA</collectingCountry>
:
<emphasis box="[464,602,1275,1296]" italics="true" pageId="11" pageNumber="458">
<collectingRegion box="[464,602,1275,1296]" country="South Africa" name="KwaZulu-Natal" pageId="11" pageNumber="458">KwaZulu­Natal</collectingRegion>
</emphasis>
:
<location LSID="urn:lsid:plazi:treatment:03D8D254FFE2A9242251FAAAFE3FF96A:8EAE3599FFE0A9242231FB62FD1CFAF7" box="[612,760,1275,1297]" country="South Africa" latitude="-29.6" longLatPrecision="736" longitude="30.38" name="Pietermaritzburg" pageId="11" pageNumber="458" stateProvince="KwaZulu-Natal">Pietermaritzburg</location>
(
<geoCoordinate box="[771,843,1274,1296]" degrees="29.60" direction="south" orientation="latitude" pageId="11" pageNumber="458" precision="555" value="-29.6">29.60°S</geoCoordinate>
<geoCoordinate box="[847,923,1275,1296]" degrees="30.38" direction="east" orientation="longitude" pageId="11" pageNumber="458" precision="555" value="30.38">30.38°E</geoCoordinate>
),
<date box="[936,1006,1275,1297]" pageId="11" pageNumber="458" value="1942-11">
<collectingDate box="[936,1006,1275,1297]" pageId="11" pageNumber="458" value="1942-11">xi.1942</collectingDate>
</date>
,
<collectorName box="[1013,1141,1275,1296]" pageId="11" pageNumber="458">R.F. Lawrence</collectorName>
(
<specimenCode box="[187,308,1299,1320]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="11" pageNumber="458" type="Museum">NMSA 3844</specimenCode>
a).
</materialsCitation>
</paragraph>
<paragraph blockId="11.[181,660,1337,1359]" box="[181,660,1337,1359]" pageId="11" pageNumber="458">
<materialsCitation box="[181,660,1337,1359]" collectingDate="1942-11" collectionCode="NMSA" collectorName="R. F. Lawrence" country="South Africa" latitude="-29.6" location="Pietermaritzburg" longLatPrecision="736" longitude="30.38" pageId="11" pageNumber="458" specimenCode="NMSA 3844" specimenCount="1" specimenCount-female="1" stateProvince="KwaZulu-Natal" typeStatus="allotype">
<typeStatus box="[181,264,1337,1359]" pageId="11" pageNumber="458">Allotype</typeStatus>
:
<specimenCount box="[273,301,1337,1358]" pageId="11" pageNumber="458" type="female">1♀</specimenCount>
same data as holotype (
<specimenCode box="[518,638,1337,1359]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="11" pageNumber="458" type="Museum">NMSA 3844</specimenCode>
b).
</materialsCitation>
</paragraph>
<paragraph blockId="11.[181,1142,1375,1520]" pageId="11" pageNumber="458">
<materialsCitation collectingDate="2003-12-02" collectionCode="NCA" collectorName="H. Roux" country="South Africa" latitude="-26.09" location="Krugersdorp" longLatPrecision="746" longitude="27.78" pageId="11" pageNumber="458" specimenCode="NCA 2009" specimenCount="1" specimenCount-male="1" stateProvince="Gauteng" typeStatus="paratype">
<typeStatus box="[181,274,1376,1397]" pageId="11" pageNumber="458">Paratypes</typeStatus>
:
<collectingCountry box="[285,457,1375,1396]" name="South Africa" pageId="11" pageNumber="458">SOUTH AFRICA</collectingCountry>
:
<emphasis box="[467,546,1376,1397]" italics="true" pageId="11" pageNumber="458">
<collectingRegion box="[467,546,1376,1397]" country="South Africa" name="Gauteng" pageId="11" pageNumber="458">Gauteng</collectingRegion>
</emphasis>
:
<specimenCount box="[561,590,1375,1396]" pageId="11" pageNumber="458" type="male">1♂</specimenCount>
<location LSID="urn:lsid:plazi:treatment:03D8D254FFE2A9242251FAAAFE3FF96A:8EAE3599FFE0A9242203FAC6FD2EFA93" box="[598,714,1375,1397]" country="South Africa" latitude="-26.09" longLatPrecision="746" longitude="27.78" name="Krugersdorp" pageId="11" pageNumber="458" stateProvince="Gauteng">Krugersdorp</location>
(
<geoCoordinate box="[729,804,1375,1397]" degrees="26.09" direction="south" orientation="latitude" pageId="11" pageNumber="458" precision="555" value="-26.09">26.09°S</geoCoordinate>
<geoCoordinate box="[811,890,1375,1396]" degrees="27.78" direction="east" orientation="longitude" pageId="11" pageNumber="458" precision="555" value="27.78">27.78°E</geoCoordinate>
),
<date box="[908,1004,1375,1397]" pageId="11" pageNumber="458" value="2003-12-02">
<collectingDate box="[908,1004,1375,1397]" pageId="11" pageNumber="458" value="2003-12-02">2.xii.2003</collectingDate>
</date>
,
<collectorName box="[1015,1098,1376,1397]" pageId="11" pageNumber="458">H. Roux</collectorName>
,
<collectingMethod pageId="11" pageNumber="458">pitfall trap</collectingMethod>
(
<specimenCode box="[271,377,1400,1422]" collectionCode="NCA" pageId="11" pageNumber="458">NCA 2009</specimenCode>
/1552)
</materialsCitation>
;
<materialsCitation collectingDate="2003-12-02" collectionCode="NCA" collectorName="H. Roux" country="South Africa" county="Krugersdorp" latitude="-25.88" location="Farm Nooitgedacht" longLatPrecision="746" longitude="27.52" municipality="Magaliesberg" pageId="11" pageNumber="458" specimenCode="NCA 2010" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Gauteng" typeStatus="paratype">
<specimenCount box="[450,478,1400,1421]" pageId="11" pageNumber="458" type="female">1♀</specimenCount>
<specimenCount box="[487,516,1400,1421]" pageId="11" pageNumber="458" type="male">1♂</specimenCount>
<collectingCounty box="[524,642,1400,1422]" pageId="11" pageNumber="458">Krugersdorp</collectingCounty>
,
<collectingMunicipality box="[655,782,1400,1422]" pageId="11" pageNumber="458">Magaliesberg</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03D8D254FFE2A9242251FAAAFE3FF96A:8EAE3599FFE0A924234FFAE1FC35FA68" box="[794,977,1400,1422]" country="South Africa" county="Krugersdorp" latitude="-25.88" longLatPrecision="746" longitude="27.52" municipality="Magaliesberg" name="Farm Nooitgedacht" pageId="11" pageNumber="458" stateProvince="Gauteng">Farm Nooitgedacht</location>
, 4171Q (
<geoCoordinate box="[1066,1141,1400,1422]" degrees="25.88" direction="south" orientation="latitude" pageId="11" pageNumber="458" precision="555" value="-25.88">25.88°S</geoCoordinate>
<geoCoordinate box="[181,260,1424,1446]" degrees="27.52" direction="east" orientation="longitude" pageId="11" pageNumber="458" precision="555" value="27.52">27.52°E</geoCoordinate>
),
<date box="[274,369,1424,1446]" pageId="11" pageNumber="458" value="2003-12-02">
<collectingDate box="[274,369,1424,1446]" pageId="11" pageNumber="458" value="2003-12-02">2.xii.2003</collectingDate>
</date>
,
<collectorName box="[377,456,1425,1446]" pageId="11" pageNumber="458">H. Roux</collectorName>
,
<collectingMethod box="[464,561,1424,1446]" pageId="11" pageNumber="458">pitfall trap</collectingMethod>
, grassland (
<specimenCode box="[666,767,1425,1446]" collectionCode="NCA" pageId="11" pageNumber="458">NCA 2010</specimenCode>
/2877)
</materialsCitation>
.
<materialsCitation collectingDate="1943-10" collectionCode="NMSA" collectorName="R. F. Lawrence" country="South Africa" latitude="-29.6" location="Scottsville" longLatPrecision="736" longitude="30.4" municipality="Pietermaritzburg" pageId="11" pageNumber="458" specimenCode="NMSA 3906" specimenCount="3" specimenCount-female="3" stateProvince="KwaZulu-Natal" typeStatus="paratype">
<emphasis box="[835,978,1424,1445]" italics="true" pageId="11" pageNumber="458">
<collectingRegion box="[835,978,1424,1445]" country="South Africa" name="KwaZulu-Natal" pageId="11" pageNumber="458">KwaZulu­Natal</collectingRegion>
</emphasis>
:
<specimenCount box="[989,1017,1425,1446]" pageId="11" pageNumber="458" type="female">3♀</specimenCount>
<collectingMunicipality pageId="11" pageNumber="458">Pietermaritzburg</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03D8D254FFE2A9242251FAAAFE3FF96A:8EAE3599FFE0A92420BEFA30FEA8FA59" box="[235,332,1449,1471]" country="South Africa" latitude="-29.6" longLatPrecision="736" longitude="30.4" municipality="Pietermaritzburg" name="Scottsville" pageId="11" pageNumber="458" stateProvince="KwaZulu-Natal">Scottsville</location>
(
<geoCoordinate box="[344,418,1449,1471]" degrees="29.60" direction="south" orientation="latitude" pageId="11" pageNumber="458" precision="555" value="-29.6">29.60°S</geoCoordinate>
<geoCoordinate box="[425,503,1449,1470]" degrees="30.40" direction="east" orientation="longitude" pageId="11" pageNumber="458" precision="555" value="30.4">30.40°E</geoCoordinate>
),
<date box="[519,584,1449,1470]" pageId="11" pageNumber="458" value="1943-10">
<collectingDate box="[519,584,1449,1470]" pageId="11" pageNumber="458" value="1943-10">x.1943</collectingDate>
</date>
,
<collectorName box="[594,727,1450,1471]" pageId="11" pageNumber="458">R.F. Lawrence</collectorName>
(
<specimenCode box="[740,862,1449,1471]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="11" pageNumber="458" type="Museum">NMSA 3906</specimenCode>
)
</materialsCitation>
.
<materialsCitation box="[877,961,1450,1471]" country="South Africa" location="Limpop" pageId="11" pageNumber="458" specimenCount="1" stateProvince="KwaZulu-Natal" typeStatus="paratype">
<emphasis box="[877,947,1450,1471]" italics="true" pageId="11" pageNumber="458">
<location LSID="urn:lsid:plazi:treatment:03D8D254FFE2A9242251FAAAFE3FF96A:8EAE3599FFE0A9242338FA33FC57FA59" box="[877,947,1450,1471]" country="South Africa" name="Limpop" pageId="11" pageNumber="458" stateProvince="KwaZulu-Natal">Limpop</location>
</emphasis>
o
</materialsCitation>
:
<materialsCitation collectingDate="2005-12-08" collectionCode="NCA" collectorName="T. Khoza &amp; M. Madiba" country="South Africa" latitude="-23.9" location="Polokwane Nat. Res." longLatPrecision="5578" longitude="29.47" pageId="11" pageNumber="458" specimenCode="NCA 2008" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="KwaZulu-Natal" typeStatus="paratype">
<specimenCount box="[971,999,1449,1470]" pageId="11" pageNumber="458" type="female">1♀</specimenCount>
<specimenCount box="[1005,1034,1449,1470]" pageId="11" pageNumber="458" type="male">1♂</specimenCount>
<location LSID="urn:lsid:plazi:treatment:03D8D254FFE2A9242251FAAAFE3FF96A:8EAE3599FFE0A9242445FA30FEE9FA31" country="South Africa" latitude="-23.9" longLatPrecision="5578" longitude="29.47" name="Polokwane Nat. Res." pageId="11" pageNumber="458" stateProvince="KwaZulu-Natal">Polokwane Nat. Res.</location>
(
<geoCoordinate box="[283,346,1474,1495]" degrees="23.9" direction="south" orientation="latitude" pageId="11" pageNumber="458" precision="5555" value="-23.9">23.9°S</geoCoordinate>
<geoCoordinate box="[353,431,1474,1495]" degrees="29.47" direction="east" orientation="longitude" pageId="11" pageNumber="458" precision="555" value="29.47">29.47°E</geoCoordinate>
),
<date box="[449,545,1474,1496]" pageId="11" pageNumber="458" value="2005-12-08">
<collectingDate box="[449,545,1474,1496]" pageId="11" pageNumber="458" value="2005-12-08">8.xii.2005</collectingDate>
</date>
,
<collectorName box="[556,641,1474,1496]" pageId="11" pageNumber="458">T. Khoza</collectorName>
&amp;
<collectorName box="[674,780,1474,1496]" pageId="11" pageNumber="458">M. Madiba</collectorName>
, hand sampling, false grassland (
<specimenCode collectionCode="NCA" pageId="11" pageNumber="458">NCA 2008</specimenCode>
/1792)
</materialsCitation>
.
</paragraph>
<paragraph blockId="11.[182,1054,1540,1567]" box="[182,1054,1540,1567]" pageId="11" pageNumber="458">
Distribution:
<collectingCountry box="[332,475,1540,1566]" name="South Africa" pageId="11" pageNumber="458">South Africa</collectingCountry>
(
<collectingRegion box="[489,588,1540,1566]" country="South Africa" name="Gauteng" pageId="11" pageNumber="458">Gauteng</collectingRegion>
,
<collectingRegion box="[599,775,1540,1566]" country="South Africa" name="KwaZulu-Natal" pageId="11" pageNumber="458">KwaZulu­Natal</collectingRegion>
and
<collectingRegion box="[829,934,1540,1566]" country="South Africa" name="Limpopo" pageId="11" pageNumber="458">Limpopo</collectingRegion>
) (
<figureCitation box="[957,1041,1540,1567]" captionStart="Figs 57–59" captionStartId="27.[347,386,1654,1676]" captionTargetBox="[181,1141,609,1633]" captionTargetId="figure-176@27.[132,1190,602,1636]" captionTargetPageId="27" captionText="Figs 57–59. Distribution of Heriaeus spp. in the Afrotropical Region." figureDoi="http://doi.org/10.5281/zenodo.7918679" httpUri="https://zenodo.org/record/7918679/files/figure.png" pageId="11" pageNumber="458">Fig. 59</figureCitation>
).
</paragraph>
<paragraph blockId="11.[181,1142,1586,1676]" pageId="11" pageNumber="458">Natural history: Most specimens were collected with pitfall traps in grassland and savanna. Adult females were collected in October and December, and males during November and December.</paragraph>
</subSubSection>
</treatment>
</document>
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