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<document id="FCBCE0684721F99574E133C101FA1466" ID-DOI="10.1080/0022293021000055441" ID-ISSN="1464-5262" ID-Zenodo-Dep="4653568" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617217246595" checkinUser="carolina" docAuthor="Peart, R. A." docDate="2004" docId="03CD6212D56EFFCFFD8FFA5BEE02F975" docLanguage="en" docName="JNATHIST.38.301-336.pdf" docOrigin="Journal of Natural History 38 (3)" docSource="http://www.tandfonline.com/doi/abs/10.1080/0022293021000055441" docStyle="DocumentStyle:443144A73DAE3E092C51A378E4CD0138.2:JNatHist.2003-2004.journal_article.1cover" docStyleId="443144A73DAE3E092C51A378E4CD0138" docStyleName="JNatHist.2003-2004.journal_article.1cover" docStyleVersion="2" docTitle="Cymadusa tattersalli Peart 2004, n. sp." docType="treatment" docVersion="4" lastPageNumber="319" masterDocId="FFF41A6AD560FFDCFFF0FFFBEA3AFFBD" masterDocTitle="A revision of the Cymadusa filosa complex (Crustacea: Amphipoda: Corophioidea: Ampithoidae)" masterLastPageNumber="336" masterPageNumber="301" pageNumber="314" updateTime="1698898288008" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="269E6D7EF1D62A42B81F8462B42B1691" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DBDE73AEE938591314DB90B181D36B57">
<mods:title id="F56412D2C25B1CDC2BFFA13348E38EC9">A revision of the Cymadusa filosa complex (Crustacea: Amphipoda: Corophioidea: Ampithoidae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="8DBF9DF41535F30D496AE0D7EDAB3499">Peart, R. A.</mods:namePart>
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<mods:title id="37277B3DFD35436FFA269A428FF96140">Journal of Natural History</mods:title>
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<mods:part id="C444E5803C0FF210B2429C943181CB2B">
<mods:date id="00D9D4F935D7E620B0D08B5534E36AB0">2004</mods:date>
<mods:detail id="EB5E5242AAE0FA33E19941AD86F1ABC3" type="pubDate">
<mods:number id="19B027A2C83B167BAB3DFB6852AED212">2004-02-10</mods:number>
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<mods:detail id="08893D0774850ECE083E6767CFDD2027" type="volume">
<mods:number id="D1567903E5A2E9E5A4E208443DBC2A19">38</mods:number>
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<mods:number id="22962764CB18E4809295D85AE4A10EF4">3</mods:number>
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<mods:classification id="48CE9CF6D09F74C4900E1C146D92B993">journal article</mods:classification>
<mods:identifier id="3718AC12DCFEBF468979A0132519F295" type="DOI">10.1080/0022293021000055441</mods:identifier>
<mods:identifier id="E103F0864D7628C0E2744343A24CAAF7" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="FCE45A353E83B41F0C14346BBB922961" type="Zenodo-Dep">4653568</mods:identifier>
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<treatment id="03CD6212D56EFFCFFD8FFA5BEE02F975" ID-DOI="http://doi.org/10.5281/zenodo.4653850" ID-Zenodo-Dep="4653850" LSID="urn:lsid:plazi:treatment:03CD6212D56EFFCFFD8FFA5BEE02F975" httpUri="http://treatment.plazi.org/id/03CD6212D56EFFCFFD8FFA5BEE02F975" lastPageId="19" lastPageNumber="319" pageId="14" pageNumber="314">
<subSubSection id="C37E808FD56EFFD2FD8FFA5BE95CFA64" pageId="14" pageNumber="314" type="nomenclature">
<paragraph id="8BDBD304D56EFFD2FD8FFA5BE994FA05" blockId="14.[313,999,1440,1560]" box="[639,942,1440,1464]" pageId="14" pageNumber="314">
<heading id="D0936468D56EFFD2FD8FFA5BE994FA05" box="[639,942,1440,1464]" centered="true" fontSize="10" level="2" pageId="14" pageNumber="314" reason="3">
<taxonomicName id="4C64A887D56EFFD2FD8FFA5BE95DFA05" ID-CoL="335QZ" authority="Peart, 2004" authorityName="Peart" authorityYear="2004" box="[639,871,1440,1464]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="14" pageNumber="314" phylum="Arthropoda" rank="species" species="tattersalli" status="sp. nov.">
<emphasis id="B9100F16D56EFFD2FD8FFA5BE95DFA05" bold="true" box="[639,871,1440,1464]" italics="true" pageId="14" pageNumber="314">Cymadusa tattersalli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A223B26DD56EFFD2FC9FFA5AE994FA05" box="[879,942,1441,1464]" pageId="14" pageNumber="314" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BDBD304D56EFFD2FD37FA3BE95CFA64" blockId="14.[313,999,1440,1560]" box="[711,870,1472,1497]" pageId="14" pageNumber="314">(figures 8–11)</paragraph>
</subSubSection>
<subSubSection id="C37E808FD56EFFD2FEC9FA19E9DDF9A5" pageId="14" pageNumber="314" type="reference_group">
<paragraph id="8BDBD304D56EFFD2FEC9FA19E895FA4A" blockId="14.[313,999,1440,1560]" box="[313,687,1506,1528]" pageId="14" pageNumber="314">
<treatmentCitationGroup id="AB74F42AD56EFFD2FEC9FA19E895FA4A" box="[313,687,1506,1528]" pageId="14" pageNumber="314">
<treatmentCitation id="0AC5F515D56EFFD2FEC9FA19E891FA4A" author="TATTERSALL, W. M." box="[313,683,1506,1528]" page="19" pageId="14" pageNumber="314" year="1922">
<taxonomicName id="4C64A887D56EFFD2FEC9FA19E891FA4A" ID-CoL="3HF77" authority="Tattersall, 1922: 1 - 19" authorityName="Tattersall" authorityPageNumber="1 - 19" authorityYear="1922" box="[313,683,1506,1528]" class="Malacostraca" family="Ampithoidae" genus="Grubia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="14" pageNumber="314" phylum="Arthropoda" rank="species" species="setosa">
<emphasis id="B9100F16D56EFFD2FEC9FA19EBF9FA45" box="[313,451,1506,1528]" italics="true" pageId="14" pageNumber="314">Grubia setosa</emphasis>
<bibRefCitation id="EFF5AEF5D56EFFD2FE3AFA19E891FA4A" author="TATTERSALL, W. M." box="[458,683,1506,1528]" pageId="14" pageNumber="314" pagination="1 - 19" refId="ref12429" refString="TATTERSALL, W. M., 1922, Amphipoda and Isopoda. The Percy Sladen Trust Expedition to the Abrolhos Islands (Indian Ocean), Journal of the Linnean Society of London, Zoology, 35, 1 - 19." type="journal article" year="1922">Tattersall, 1922: 1–19</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDBD304D56EFFD2FEC9F9F9E9DDF9A5" blockId="14.[313,999,1440,1560]" box="[313,999,1537,1560]" pageId="14" pageNumber="314">
<treatmentCitationGroup id="AB74F42AD56EFFD2FEC9F9F9E9DDF9A5" box="[313,999,1537,1560]" pageId="14" pageNumber="314">
<treatmentCitation id="0AC5F515D56EFFD2FEC9F9F9E8DCF9A5" author="LEDOYER, M." box="[313,742,1537,1560]" page="135" pageId="14" pageNumber="314" year="1982">
<taxonomicName id="4C64A887D56EFFD2FEC9F9F9E8DCF9A5" ID-CoL="335PS" authority=": Ledoyer, 1982: 130 - 135" authorityName=": Ledoyer" authorityPageNumber="130 - 135" authorityYear="1982" box="[313,742,1537,1560]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="14" pageNumber="314" phylum="Arthropoda" rank="species" species="filosa">
<emphasis id="B9100F16D56EFFD2FEC9F9F9EBE4F9AA" box="[313,478,1538,1559]" italics="true" pageId="14" pageNumber="314">Cymadusa filosa</emphasis>
:
<bibRefCitation id="EFF5AEF5D56EFFD2FE1DF9F9E8DCF9A5" author="LEDOYER, M." box="[493,742,1537,1560]" pageId="14" pageNumber="314" pagination="1 - 598" refId="ref11601" refString="LEDOYER, M., 1982, Crustaces amphipodes gammariens familles des Acanthonotozomatidae a Gammaridae, Faune de Madagascar, 59 (1), 1 - 598." type="journal article" year="1982">Ledoyer, 1982: 130–135</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figures 44–46 (form A).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C37E808FD56EFFCFFE91F9CAEE02F975" lastPageId="19" lastPageNumber="319" pageId="14" pageNumber="314" type="materials_examined">
<paragraph id="8BDBD304D56EFFD2FE91F9CAEEA1F975" blockId="14.[313,1268,1584,1736]" pageId="14" pageNumber="314">
<emphasis id="B9100F16D56EFFD2FE91F9CAE83EF9F5" box="[353,516,1584,1608]" italics="true" pageId="14" pageNumber="314">Type material.</emphasis>
<materialsCitation id="3B0CD959D56EFFD2FDFBF9CAE89DF935" collectingDate="1973-07-14" collectionCode="T" collectorName="M. Walker &amp; D. D. Bray" country="Australia" location="Woodman Point" municipality="Cockburn Sound" pageId="14" pageNumber="314" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<typeStatus id="54DF6DA6D56EFFD2FDFBF9CAE8BAF9F5" box="[523,640,1584,1608]" pageId="14" pageNumber="314" type="holotype">HOLOTYPE</typeStatus>
: male,
<quantity id="4C9C7EE1D56EFFD2FD23F9CBE924F9F5" box="[723,798,1584,1608]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="14" pageNumber="314" unit="mm" value="17.0">17 mm</quantity>
length, TMAG G3932,
<location id="8EBB85DFD56EFFD2FBC2F9CBEED5F9F5" LSID="urn:lsid:plazi:treatment:03CD6212D56EFFCFFD8FFA5BEE02F975:8EBB85DFD56EFFD2FBC2F9CBEED5F9F5" box="[1074,1263,1584,1608]" country="Australia" municipality="Cockburn Sound" name="Woodman Point" pageId="14" pageNumber="314" stateProvince="Western Australia">Woodman Point</location>
,
<collectingMunicipality id="6BBF497ED56EFFD2FEC9F9ABE83EF9D5" box="[313,516,1616,1640]" pageId="14" pageNumber="314">Cockburn Sound</collectingMunicipality>
,
<collectingRegion id="49A01DE6D56EFFD2FDE4F9ABE8D3F9D5" box="[532,745,1616,1640]" country="Australia" name="Western Australia" pageId="14" pageNumber="314">Western Australia</collectingRegion>
,
<collectingCountry id="F3739394D56EFFD2FD0AF9ABE95CF9D5" box="[762,870,1616,1640]" name="Australia" pageId="14" pageNumber="314">Australia</collectingCountry>
, 32°08∞S, 115°45∞E,
<date id="FFDAF5C4D56EFFD2FBA6F9ABEECAF9D5" box="[1110,1264,1616,1640]" pageId="14" pageNumber="314" value="1973-07-14">
<collectingDate id="EF9E0C2CD56EFFD2FBA6F9ABEECAF9D5" box="[1110,1264,1616,1640]" pageId="14" pageNumber="314" value="1973-07-14">14 July 1973</collectingDate>
</date>
,
<collectionCode id="ED754BC1D56EFFD2FEC9F98BEB74F93A" box="[313,334,1648,1671]" name="Tavera, Department of Geology and Geophysics" pageId="14" pageNumber="314">T</collectionCode>
.
<collectorName id="2691B6D2D56EFFD2FEADF98BEBE1F935" box="[349,475,1648,1672]" pageId="14" pageNumber="314">M. Walker</collectorName>
and
<collectorName id="2691B6D2D56EFFD2FDECF98BE89DF935" box="[540,679,1648,1672]" pageId="14" pageNumber="314">D. D. Bray</collectorName>
</materialsCitation>
.
<materialsCitation id="3B0CD959D56EFFD2FD46F98BEEADF975" accessionNumber="P51282" country="Australia" pageId="14" pageNumber="314" specimenCount="2" stateProvince="Western Australia" typeStatus="paratype">
<typeStatus id="54DF6DA6D56EFFD2FD46F98BE914F935" box="[694,814,1648,1672]" pageId="14" pageNumber="314" type="paratype">PARATYPES</typeStatus>
: TMAG G3931 (female), AM P51282 (female and male); TMAG G3933 (male); TMAG G3934-G3938 (females and males—five specimens), collection data is the same as that of the
<typeStatus id="54DF6DA6D56EFFD2FBC0F94BEEADF975" box="[1072,1175,1712,1736]" pageId="14" pageNumber="314" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
<caption id="DF1B838CD56FFFD3FEC9FA94EEDCFA1D" ID-DOI="http://doi.org/10.5281/zenodo.4653588" ID-Zenodo-Dep="4653588" httpUri="https://zenodo.org/record/4653588/files/figure.png" pageId="15" pageNumber="315" startId="15.[313,329,1391,1412]" targetBox="[316,1265,182,1359]" targetPageId="15">
<paragraph id="8BDBD304D56FFFD3FEC9FA94EEDCFA1D" blockId="15.[313,1269,1391,1440]" pageId="15" pageNumber="315">
FIG. 8.
<taxonomicName id="4C64A887D56FFFD3FE65FA94E85EFA39" authority="Peart, 2004" authorityName="Peart" authorityYear="2004" box="[405,612,1391,1412]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="15" pageNumber="315" phylum="Arthropoda" rank="species" species="tattersalli" status="sp. nov.">
<emphasis id="B9100F16D56FFFD3FE65FA94E85EFA39" box="[405,612,1391,1412]" italics="true" pageId="15" pageNumber="315">Cymadusa tattersalli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A223B26DD56FFFD3FD9CFA94E89EFA39" box="[620,676,1391,1412]" pageId="15" pageNumber="315" rank="species">n. sp.</taxonomicNameLabel>
Holotype male, 17 mm, TMAG G3932. Paratype female, 18 mm, TMAG G3931, Woodman Point, Cockburn Sound, Western Australia, Australia.
</paragraph>
</caption>
<paragraph id="8BDBD304D56FFFD3FE91FA0AEE1DF9B5" blockId="15.[313,1269,1520,1736]" box="[353,1063,1520,1544]" pageId="15" pageNumber="315">
<materialsCitation id="3B0CD959D56FFFD3FE91FA0AEE19F9B5" box="[353,1059,1520,1544]" country="Australia" location="Cockburn Sound" pageId="15" pageNumber="315" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<emphasis id="B9100F16D56FFFD3FE91FA0AEBC1F9B5" box="[353,507,1520,1544]" italics="true" pageId="15" pageNumber="315">
<typeStatus id="54DF6DA6D56FFFD3FE91FA0AEBA3F9B5" box="[353,409,1521,1544]" pageId="15" pageNumber="315">Type</typeStatus>
locality.
</emphasis>
<location id="8EBB85DFD56FFFD3FDF3FA0BE8F0F9B5" LSID="urn:lsid:plazi:treatment:03CD6212D56EFFCFFD8FFA5BEE02F975:8EBB85DFD56FFFD3FDF3FA0BE8F0F9B5" box="[515,714,1520,1544]" country="Australia" name="Cockburn Sound" pageId="15" pageNumber="315" stateProvince="Western Australia">Cockburn Sound</location>
,
<collectingRegion id="49A01DE6D56FFFD3FD28FA0AE992F9B5" box="[728,936,1520,1544]" country="Australia" name="Western Australia" pageId="15" pageNumber="315">Western Australia</collectingRegion>
,
<collectingCountry id="F3739394D56FFFD3FC46FA0BEE19F9B5" box="[950,1059,1520,1544]" name="Australia" pageId="15" pageNumber="315">Australia</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8BDBD304D56FFFD3FE91F9EAE96BF975" blockId="15.[313,1269,1520,1736]" pageId="15" pageNumber="315">
<emphasis id="B9100F16D56FFFD3FE91F9EAEBEFF995" box="[353,469,1553,1576]" italics="true" pageId="15" pageNumber="315">Diagnosis.</emphasis>
Antenna 1 with a two-articulate accessory flagellum. Antenna 2 with densely plumose setose margins. Gnathopod 1 with densely plumose setose margins; merus slightly produced to form a small, rounded, anteroventral lobe; carpus robust, broad and subtriangular. Gnathopod 2 with densely plumose setose margins; propodal palm with defining posterodistal, small and subacute tooth, with subquadrate mid-palmar tooth; dactylus shorter than palm.
</paragraph>
<caption id="DF1B838CD570FFCCFEC9FA8CE8D8FA7F" ID-DOI="http://doi.org/10.5281/zenodo.4653590" ID-Zenodo-Dep="4653590" httpUri="https://zenodo.org/record/4653590/files/figure.png" pageId="16" pageNumber="316" startId="16.[313,329,1399,1420]" targetBox="[316,1266,185,1369]" targetPageId="16">
<paragraph id="8BDBD304D570FFCCFEC9FA8CE8D8FA7F" blockId="16.[313,1269,1399,1474]" pageId="16" pageNumber="316">
FIG. 9.
<taxonomicName id="4C64A887D570FFCCFE64FA8CE85EFA31" authority="Peart, 2004" authorityName="Peart" authorityYear="2004" box="[404,612,1399,1420]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="16" pageNumber="316" phylum="Arthropoda" rank="species" species="tattersalli" status="sp. nov.">
<emphasis id="B9100F16D570FFCCFE64FA8CE85EFA31" box="[404,612,1399,1420]" italics="true" pageId="16" pageNumber="316">Cymadusa tattersalli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A223B26DD570FFCCFD9BFA8CE89EFA31" box="[619,676,1399,1420]" pageId="16" pageNumber="316" rank="species">n. sp.</taxonomicNameLabel>
Holotype male, 17 mm, TMAG G3932. Paratype female, 18 mm, TMAG G3931, Woodman Point, Cockburn Sound, Western Australia, Australia. Scales represent 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BDBD304D570FFCCFEC9FA0AEB82F9B5" blockId="16.[313,1268,1521,1736]" box="[313,440,1521,1544]" pageId="16" pageNumber="316">
<emphasis id="B9100F16D570FFCCFEC9FA0AEB82F9B5" box="[313,440,1521,1544]" italics="true" pageId="16" pageNumber="316">Description</emphasis>
</paragraph>
<paragraph id="8BDBD304D570FFCCFE91F9EBE8E2F995" blockId="16.[313,1268,1521,1736]" box="[353,728,1552,1576]" pageId="16" pageNumber="316">
Based on
<typeStatus id="54DF6DA6D570FFCCFE23F9EBE80CF995" box="[467,566,1552,1576]" pageId="16" pageNumber="316" type="holotype">holotype</typeStatus>
male,
<quantity id="4C9C7EE1D570FFCCFD76F9EBE8EEF995" box="[646,724,1552,1576]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="16" pageNumber="316" unit="mm" value="17.0">17 mm</quantity>
.
</paragraph>
<paragraph id="8BDBD304D570FFCDFE91F9CBEEF0F9DA" blockId="16.[313,1268,1521,1736]" lastBlockId="17.[313,1269,1520,1736]" lastPageId="17" lastPageNumber="317" pageId="16" pageNumber="316">
<emphasis id="B9100F16D570FFCCFE91F9CBE825F9F5" box="[353,543,1584,1608]" italics="true" pageId="16" pageNumber="316">Head. Antenna 1</emphasis>
longer than antenna 2; peduncular article 1 subequal to article 2, article 2 longer than article 3, article 3 shorter than article 1; primary flagellum with 46 articles; accessory flagellum with two articles.
<emphasis id="B9100F16D570FFCCFC69F98BEE33F935" box="[921,1033,1648,1672]" italics="true" pageId="16" pageNumber="316">Antenna 2</emphasis>
with dense, plumose setae on ventral margin; peduncular article 4 subequal to article 5.
<emphasis id="B9100F16D570FFCCFBCBF96AEE9FF915" box="[1083,1189,1680,1704]" italics="true" pageId="16" pageNumber="316">Upper lip</emphasis>
entire; distal setae present.
<emphasis id="B9100F16D570FFCCFDE9F94BE8B8F975" box="[537,642,1712,1736]" italics="true" pageId="16" pageNumber="316">Lower lip</emphasis>
outer plate, outer lobe longer than inner lobe.
<emphasis id="B9100F16D570FFCCFB7BF94BEECEF975" box="[1163,1268,1712,1736]" italics="true" pageId="16" pageNumber="316">Mandible</emphasis>
with nine robust setae in setal row; palp article 1 shorter than article 2, article 2 subequal to article 3, article 3 longer than article 1, setae on posterior margin and apex.
<emphasis id="B9100F16D571FFCDFE8FF9CBEBD5F9F5" box="[383,495,1584,1608]" italics="true" pageId="17" pageNumber="317">Maxilla 1</emphasis>
palp apical margin with robust setae; inner plate with five slender setae.
<emphasis id="B9100F16D571FFCDFE71F9ABEBCAF9D5" box="[385,496,1616,1640]" italics="true" pageId="17" pageNumber="317">Maxilla 2</emphasis>
outer lobe broader than inner lobe.
<emphasis id="B9100F16D571FFCDFC68F9ABEE2BF9D5" box="[920,1041,1616,1640]" italics="true" pageId="17" pageNumber="317">Maxilliped</emphasis>
strongly setose.
</paragraph>
<caption id="DF1B838CD571FFCDFEC9FA91E8D8FA09" ID-DOI="http://doi.org/10.5281/zenodo.4653592" ID-Zenodo-Dep="4653592" httpUri="https://zenodo.org/record/4653592/files/figure.png" pageId="17" pageNumber="317" startId="17.[313,329,1386,1407]" targetBox="[316,1266,182,1355]" targetPageId="17">
<paragraph id="8BDBD304D571FFCDFEC9FA91E8D8FA09" blockId="17.[313,1269,1386,1461]" pageId="17" pageNumber="317">
FIG. 10.
<taxonomicName id="4C64A887D571FFCDFE51FA91E855FAC2" authority="Peart, 2004" authorityName="Peart" authorityYear="2004" box="[417,623,1386,1407]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="17" pageNumber="317" phylum="Arthropoda" rank="species" species="tattersalli" status="sp. nov.">
<emphasis id="B9100F16D571FFCDFE51FA91E855FAC2" box="[417,623,1386,1407]" italics="true" pageId="17" pageNumber="317">Cymadusa tattersalli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A223B26DD571FFCDFD85FA91E897FAC2" box="[629,685,1386,1407]" pageId="17" pageNumber="317" rank="species">n. sp.</taxonomicNameLabel>
Holotype male, 17 mm, TMAG G3932. Paratype female, 18 mm, TMAG G3931, Woodman Point, Cockburn Sound, Western Australia, Australia. Scales represent 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BDBD304D571FFCFFE91F98AE9ACFEB3" blockId="17.[313,1269,1520,1736]" lastBlockId="19.[313,1269,181,1050]" lastPageId="19" lastPageNumber="319" pageId="17" pageNumber="317">
<emphasis id="B9100F16D571FFCDFE91F98AE86EF935" box="[353,596,1648,1672]" italics="true" pageId="17" pageNumber="317">Pereon. Gnathopod 1</emphasis>
smaller than gnathopod 2; with long plumose setae on margins; coxa produced forward, anterior margin convex, ventral margin with long, plumose setal fringe; basis shorter than the coxa, with a long plumose setal fringe, distal lobe absent; merus produced to form a small, rounded anteroventral lobe; carpus shorter than merus, longer than propodus, lateral margins divergent at distal end; carpal lobe truncated; propodus ovoid; palm entire, straight, with posterodistal tooth absent, and mid-palmar tooth absent; dactylus subequal to palm.
<emphasis id="B9100F16D572FFCEFB8BF9B5EB7CF93A" italics="true" pageId="18" pageNumber="318">Gnathopod 2</emphasis>
margins with long, plumose setae; coxa with long, plumose setal fringe on ventral margin only; basis longer than coxa, and with long plumose setae in a fringe, distal lobe absent; merus produced to form a small, rounded, anteroventral lobe; carpus longer than merus, shorter than propodus; propodus subquadrate; palm entire, straight, with small, subacute posterodistal tooth, subquadrate mid-palmar tooth present; dactylus shorter than palm, tapering evenly.
</paragraph>
<caption id="DF1B838CD572FFCEFEC9FAB1EEB5FAC7" ID-DOI="http://doi.org/10.5281/zenodo.4653594" ID-Zenodo-Dep="4653594" httpUri="https://zenodo.org/record/4653594/files/figure.png" pageId="18" pageNumber="318" startId="18.[313,329,1354,1375]" targetBox="[318,1266,182,1323]" targetPageId="18">
<paragraph id="8BDBD304D572FFCEFEC9FAB1EEB5FAC7" blockId="18.[313,1268,1354,1402]" pageId="18" pageNumber="318">
FIG. 11.
<taxonomicName id="4C64A887D572FFCEFE51FAB1E855FAE2" authority="Peart, 2004" authorityName="Peart" authorityYear="2004" box="[417,623,1354,1375]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="18" pageNumber="318" phylum="Arthropoda" rank="species" species="tattersalli" status="sp. nov.">
<emphasis id="B9100F16D572FFCEFE51FAB1E855FAE2" box="[417,623,1354,1375]" italics="true" pageId="18" pageNumber="318">Cymadusa tattersalli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A223B26DD572FFCEFD85FAB1E891FAE2" box="[629,683,1354,1375]" pageId="18" pageNumber="318" rank="species">n. sp.</taxonomicNameLabel>
Holotype male, 17 mm, TMAG G3932, Woodman Point, Cockburn Sound, Western Australia, Australia. Scales represent 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BDBD304D573FFCFFE91FEEDEBFAFE12" blockId="19.[313,1269,181,1050]" pageId="19" pageNumber="319">
<emphasis id="B9100F16D573FFCFFE91FEEDEBDDFE93" box="[353,487,278,302]" italics="true" pageId="19" pageNumber="319">Pereopods 3</emphasis>
and
<emphasis id="B9100F16D573FFCFFDD4FEEDE80BFE93" box="[548,561,278,302]" italics="true" pageId="19" pageNumber="319">4</emphasis>
similar in shape; coxae with a simple setal fringe.
<emphasis id="B9100F16D573FFCFFB88FEEDEECEFE93" box="[1144,1268,278,302]" italics="true" pageId="19" pageNumber="319">Pereopod 5</emphasis>
coxae with setal fringe on two-thirds of the ventral margin and with setae on the medial surface.
<emphasis id="B9100F16D573FFCFFE18FEACE858FED2" box="[488,610,343,367]" italics="true" pageId="19" pageNumber="319">Pereopod 6</emphasis>
basis with sparse medial setae.
<emphasis id="B9100F16D573FFCFFC30FEACEE00FED2" box="[960,1082,343,367]" italics="true" pageId="19" pageNumber="319">Pereopod 7</emphasis>
basis subrectangular.
<emphasis id="B9100F16D573FFCFFE78FE8CE82EFE32" box="[392,532,375,399]" italics="true" pageId="19" pageNumber="319">Pereopods 6</emphasis>
and
<emphasis id="B9100F16D573FFCFFDAAFE83E85DFE32" box="[602,615,376,399]" italics="true" pageId="19" pageNumber="319">7</emphasis>
much longer than pereopods 3–5; distal articles with dense setae.
</paragraph>
<paragraph id="8BDBD304D573FFCFFE91FE43EECFFDCE" blockId="19.[313,1269,181,1050]" pageId="19" pageNumber="319">
<emphasis id="B9100F16D573FFCFFE91FE43E800FE6C" box="[353,570,440,465]" italics="true" pageId="19" pageNumber="319">Pleon. Epimeron 3</emphasis>
with a small subacute tooth.
<emphasis id="B9100F16D573FFCFFC5DFE43EE21FE6C" box="[941,1051,440,465]" italics="true" pageId="19" pageNumber="319">Uropod 1</emphasis>
peduncle with 10 marginal robust setae; outer ramus shorter than inner ramus.
<emphasis id="B9100F16D573FFCFFC0AFE22EE58FE4C" box="[1018,1122,473,497]" italics="true" pageId="19" pageNumber="319">Uropod 2</emphasis>
outer ramus subequal to inner ramus.
<emphasis id="B9100F16D573FFCFFDAFFE02E8F3FDAC" box="[607,713,505,529]" italics="true" pageId="19" pageNumber="319">Uropod 3</emphasis>
with seven distal peduncular robust setae; outer ramus subequal to inner ramus, hooks both strongly recurved, setal fringe present on lateral margin; inner ramus with both robust and slender setae.
<taxonomicName id="4C64A887D573FFCFFBCDFDC1EEBFFDEF" authorityName="Pearse" authorityYear="1952" box="[1085,1157,570,594]" class="Maxillopoda" family="Telsidae" genus="Telson" kingdom="Animalia" order="Poecilostomatoida" pageId="19" pageNumber="319" phylum="Arthropoda" rank="genus">
<emphasis id="B9100F16D573FFCFFBCDFDC1EEBFFDEF" box="[1085,1157,570,594]" italics="true" pageId="19" pageNumber="319">Telson</emphasis>
</taxonomicName>
rounded, with both lateral and apical slender setae (not in oblique rows); apical cusps present.
</paragraph>
<paragraph id="8BDBD304D573FFCFFE91FD87EEF3FD29" blockId="19.[313,1269,181,1050]" box="[353,1225,636,660]" pageId="19" pageNumber="319">
<emphasis id="B9100F16D573FFCFFE91FD87EB82FD29" box="[353,440,636,660]" italics="true" pageId="19" pageNumber="319">Female.</emphasis>
Based on
<typeStatus id="54DF6DA6D573FFCFFDC3FD87E8ADFD29" box="[563,663,636,660]" pageId="19" pageNumber="319" type="paratype">paratype</typeStatus>
<quantity id="4C9C7EE1D573FFCFFD6FFD87E8D0FD2E" box="[671,746,636,660]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="19" pageNumber="319" unit="mm" value="18.0">18 mm</quantity>
in length. Sexually dimorphic characters.
</paragraph>
<paragraph id="8BDBD304D573FFCFFE91FD66EECFFBA7" blockId="19.[313,1269,181,1050]" pageId="19" pageNumber="319">
<emphasis id="B9100F16D573FFCFFE91FD66E868FD09" box="[353,594,668,692]" italics="true" pageId="19" pageNumber="319">Pereon. Gnathopod 1</emphasis>
subequal to gnathopod 2, poorly setose; coxa produced forward, anterior margin straight, posterior two-thirds of ventral margin with long, plumose setal fringe; basis shorter than the coxa, with a long plumose setal fringe; merus produced to form a small, subacute anteroventral lobe; carpus longer than merus, subequal to propodus, lateral margins divergent at the distal end; carpal lobe truncated with rounded corners; propodus ovoid; palm entire, straight, with defining posterodistal tooth absent, and mid-palmar tooth absent; dactylus subequal to palm.
<emphasis id="B9100F16D573FFCFFEC9FC7BEBF3FC25" box="[313,457,896,920]" italics="true" pageId="19" pageNumber="319">Gnathopod 2</emphasis>
weakly setose; coxa with long plumose setal fringe on posterior twothirds of ventral margin; basis shorter than coxa, and weakly setose; merus produced to form a small, rounded, anteroventral lobe; carpus subequal to merus, shorter than propodus; propodus ovoid; palm entire, sinuous, with small, subacute posterodistal tooth, mid-palmar tooth absent; dactylus longer than palm and tapered evenly.
</paragraph>
<paragraph id="8BDBD304D573FFCFFEC9FB9DEBA6FBC3" blockId="19.[313,1269,1126,1736]" box="[313,412,1126,1150]" pageId="19" pageNumber="319">
<emphasis id="B9100F16D573FFCFFEC9FB9DEBA6FBC3" box="[313,412,1126,1150]" italics="true" pageId="19" pageNumber="319">Remarks</emphasis>
</paragraph>
<paragraph id="8BDBD304D573FFCFFE91FB7DE9F7F998" blockId="19.[313,1269,1126,1736]" pageId="19" pageNumber="319">
<bibRefCitation id="EFF5AEF5D573FFCFFE91FB7DE81AFB23" author="TATTERSALL, W. M." box="[353,544,1158,1182]" pageId="19" pageNumber="319" pagination="1 - 19" refId="ref12429" refString="TATTERSALL, W. M., 1922, Amphipoda and Isopoda. The Percy Sladen Trust Expedition to the Abrolhos Islands (Indian Ocean), Journal of the Linnean Society of London, Zoology, 35, 1 - 19." type="journal article" year="1922">Tattersall (1922)</bibRefCitation>
in his description says that even though Haswell’s description was really short and was not illustrated, his specimens actually fit the description and therefore named his specimens
<taxonomicName id="4C64A887D573FFCFFD21FB3CE9ABFB62" baseAuthorityName="Haswell" baseAuthorityYear="1879" box="[721,913,1223,1247]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="19" pageNumber="319" phylum="Arthropoda" rank="species" species="setosa">
<emphasis id="B9100F16D573FFCFFD21FB3CE9ABFB62" box="[721,913,1223,1247]" italics="true" pageId="19" pageNumber="319">Cymadusa setosa</emphasis>
</taxonomicName>
. However, after looking at the specimens from the east coast of
<collectingCountry id="F3739394D573FFCFFD5AFB13E92CFABD" box="[682,790,1256,1280]" name="Australia" pageId="19" pageNumber="319">Australia</collectingCountry>
, there are a number of differences between Haswell’s and Tattersall’s species.
<taxonomicName id="4C64A887D573FFCFFD3DFAF3E982FA9D" authority="Peart, 2004" authorityName="Peart" authorityYear="2004" box="[717,952,1288,1312]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="19" pageNumber="319" phylum="Arthropoda" rank="species" species="tattersalli" status="sp. nov.">
<emphasis id="B9100F16D573FFCFFD3DFAF3E982FA9D" box="[717,952,1288,1312]" italics="true" pageId="19" pageNumber="319">Cymadusa tattersalli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A223B26DD573FFCFFC33FAF2EE3EFA9D" box="[963,1028,1289,1312]" pageId="19" pageNumber="319" rank="species">n. sp.</taxonomicNameLabel>
differs by maxilla 1 inner plate which has five slender setae (whereas
<taxonomicName id="4C64A887D573FFCFFC73FAD2E9D6FAFC" baseAuthorityName="Haswell" baseAuthorityYear="1879" box="[899,1004,1321,1345]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="19" pageNumber="319" phylum="Arthropoda" rank="species" species="setosa">
<emphasis id="B9100F16D573FFCFFC73FAD2E9D6FAFC" box="[899,1004,1321,1345]" italics="true" pageId="19" pageNumber="319">C. setosa</emphasis>
</taxonomicName>
has 11 slender setae); gnathopod 1 basis is shorter than the coxa (
<taxonomicName id="4C64A887D573FFCFFCBCFAB1E98FFADC" baseAuthorityName="Haswell" baseAuthorityYear="1879" box="[844,949,1354,1377]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="19" pageNumber="319" phylum="Arthropoda" rank="species" species="setosa">
<emphasis id="B9100F16D573FFCFFCBCFAB1E98FFADC" box="[844,949,1354,1377]" italics="true" pageId="19" pageNumber="319">C. setosa</emphasis>
</taxonomicName>
basis is much longer than coxa), and has a long, dense, plumose setal fringe (
<taxonomicName id="4C64A887D573FFCFFC51FA90EE30FA3F" baseAuthorityName="Haswell" baseAuthorityYear="1879" box="[929,1034,1387,1410]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="19" pageNumber="319" phylum="Arthropoda" rank="species" species="setosa">
<emphasis id="B9100F16D573FFCFFC51FA90EE30FA3F" box="[929,1034,1387,1410]" italics="true" pageId="19" pageNumber="319">C. setosa</emphasis>
</taxonomicName>
has a short stubby fringe on the margin), the merus is produced to form a small subrounded anteroventral lobe (
<taxonomicName id="4C64A887D573FFCFFE55FA57E830FA7E" baseAuthorityName="Haswell" baseAuthorityYear="1879" box="[421,522,1452,1475]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="19" pageNumber="319" phylum="Arthropoda" rank="species" species="setosa">
<emphasis id="B9100F16D573FFCFFE55FA57E830FA7E" box="[421,522,1452,1475]" italics="true" pageId="19" pageNumber="319">C. setosa</emphasis>
</taxonomicName>
merus is produced to form a long, subacute lobe), carpus is short and broad (
<taxonomicName id="4C64A887D573FFCFFE30FA37E81DFA59" baseAuthorityName="Haswell" baseAuthorityYear="1879" box="[448,551,1484,1508]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="19" pageNumber="319" phylum="Arthropoda" rank="species" species="setosa">
<emphasis id="B9100F16D573FFCFFE30FA37E81DFA59" box="[448,551,1484,1508]" italics="true" pageId="19" pageNumber="319">C. setosa</emphasis>
</taxonomicName>
carpus is elongated and narrow), the propodal palm is entire and convex (
<taxonomicName id="4C64A887D573FFCFFE3FFA16E802F9B9" baseAuthorityName="Haswell" baseAuthorityYear="1879" box="[463,568,1517,1540]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="19" pageNumber="319" phylum="Arthropoda" rank="species" species="setosa">
<emphasis id="B9100F16D573FFCFFE3FFA16E802F9B9" box="[463,568,1517,1540]" italics="true" pageId="19" pageNumber="319">C. setosa</emphasis>
</taxonomicName>
propodal palm is entire and straight); uropod 3 has fewer robust setae on the peduncle and rami than on
<emphasis id="B9100F16D573FFCFFCAEF9F6E9F7F998" box="[862,973,1549,1573]" italics="true" pageId="19" pageNumber="319">
<taxonomicName id="4C64A887D573FFCFFCAEF9F6E9F3F998" baseAuthorityName="Haswell" baseAuthorityYear="1879" box="[862,969,1549,1573]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="19" pageNumber="319" phylum="Arthropoda" rank="species" species="setosa">C. setosa</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8BDBD304D573FFCFFE91F9D6E93DF9D8" blockId="19.[313,1269,1126,1736]" pageId="19" pageNumber="319">
<emphasis id="B9100F16D573FFCFFE91F9D6EBDBF9F8" box="[353,481,1581,1605]" italics="true" pageId="19" pageNumber="319">Etymology.</emphasis>
Named for W. Tattersall who documented this animal for the first time from the Western Australian coast.
</paragraph>
<paragraph id="8BDBD304D573FFCFFE91F995E9FEF93B" blockId="19.[313,1269,1126,1736]" box="[353,964,1646,1670]" pageId="19" pageNumber="319">
<emphasis id="B9100F16D573FFCFFE91F995E895F93B" box="[353,687,1646,1670]" italics="true" pageId="19" pageNumber="319">
Habitat.
<taxonomicName id="4C64A887D573FFCFFE37F995E895F93B" authorityName="Peart" authorityYear="2004" box="[455,687,1646,1670]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="19" pageNumber="319" phylum="Arthropoda" rank="species" species="tattersalli">Cymadusa tattersalli</taxonomicName>
</emphasis>
occurs on brown algae.
</paragraph>
<paragraph id="8BDBD304D573FFCFFE91F974EE02F975" blockId="19.[313,1269,1126,1736]" pageId="19" pageNumber="319">
<emphasis id="B9100F16D573FFCFFE91F974EBD4F91A" box="[353,494,1679,1703]" italics="true" pageId="19" pageNumber="319">Distribution.</emphasis>
Cockburn Sound,
<collectingRegion id="49A01DE6D573FFCFFD3EF96BE9A5F91A" box="[718,927,1679,1703]" country="Australia" name="Western Australia" pageId="19" pageNumber="319">Western Australia</collectingRegion>
,
<collectingCountry id="F3739394D573FFCFFC5EF974EE20F91A" box="[942,1050,1679,1703]" name="Australia" pageId="19" pageNumber="319">Australia</collectingCountry>
; Abrohlos Islands,
<collectingRegion id="49A01DE6D573FFCFFEC9F94BE830F975" box="[313,522,1712,1736]" country="Australia" name="Western Australia" pageId="19" pageNumber="319">Western Australia</collectingRegion>
,
<collectingCountry id="F3739394D573FFCFFDE7F94BE8B9F975" box="[535,643,1712,1736]" name="Australia" pageId="19" pageNumber="319">Australia</collectingCountry>
;
<collectingCountry id="F3739394D573FFCFFD60F94BE924F975" box="[656,798,1712,1736]" name="Madagascar" pageId="19" pageNumber="319">Madagascar</collectingCountry>
, Indian Ocean (
<figureCitation id="135FCF81D573FFCFFC28F94BEE1FF975" box="[984,1061,1712,1736]" captionStart="MAP" captionStartId="11.[313,337,861,882]" captionTargetBox="[345,1236,179,831]" captionTargetId="figure-8@11.[343,1239,177,834]" captionTargetPageId="11" captionText="MAP 1. Global distribution of Cymadusa filosa Savigny, 1816 (*), Cymadusa ledoyeri n. sp. (•), Cymadusa tattersalli n. sp. (*), Cymadusa imbroglio Rabindranath, 1972 (Δ), Cymadusa setosa (Haswell, 1879) (v), Cymadusa vadosa Imbach, 1967 (&amp;)." figureDoi="http://doi.org/10.5281/zenodo.4653578" httpUri="https://zenodo.org/record/4653578/files/figure.png" pageId="19" pageNumber="319">Map 1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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