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<document id="FCEF84C93E2ED88518AC398B33A7B5FC" ID-DOI="10.11646/zootaxa.5336.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="8281432" ID-ZooBank="B0315010-41C9-4F5A-91D5-7A1D333E0D36" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1692950238990" checkinUser="plazi" docAuthor="Neto, Pedro Bonfá, Vilarino, Albane &amp; Salles, Frederico F." docDate="2023" docId="03AAE777C4254807FF46FD771154F8FD" docLanguage="en" docName="zootaxa.5336.3.1.pdf" docOrigin="Zootaxa 5336 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5336.3.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phylloicus guarani Neto &amp; Vilarino &amp; Salles 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="310" masterDocId="FF939F0FC42D480EFFD1FFF81338FFB1" masterDocTitle="Brevitentoria Weaver 1984 (Trichoptera: Integripalpia) of Espírito Santo State, Brazil: New records and new species" masterLastPageNumber="327" masterPageNumber="301" pageNumber="309" updateTime="1692950958001" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="7B439112CE7B41691E05E37F491975FB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EC4D79532C83D551245C1341E280827A">
<mods:title id="2D8D7D3784C41E21B34128310D986F31">Brevitentoria Weaver 1984 (Trichoptera: Integripalpia) of Espírito Santo State, Brazil: New records and new species</mods:title>
</mods:titleInfo>
<mods:name id="FA11FCD3FCD5742F7612AA84B1ACAE25" type="personal">
<mods:role id="EF49C08D1B51CC88AB459269F0228E35">
<mods:roleTerm id="7216CA96909CD4FF102E39D9D9DFEB05">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8839D072F681A2083D8D452B3C88C900">Neto, Pedro Bonfá</mods:namePart>
<mods:nameIdentifier id="5C8A104F28D3E9B3EF6977C4DAFC034E" type="ORCID">0000-0002-9328-1352</mods:nameIdentifier>
<mods:affiliation id="CE85331A2D822645C0FDFB86BC9244E9">Museu de Entomologia, Departamento de Entomologia, Universidade Federal de Viçosa, Av. P. H. Rolfs, s. n, Campus Universitário, CEP 36570 - 900, Viçosa, Minas Gerais, Brazil &amp; bonfa. pn @ gmail. com; https: // orcid. org / 0000 - 0002 - 9328 - 1352</mods:affiliation>
<mods:nameIdentifier id="202C519B3ED7024C7BA91AD05EF5C142" type="email">bonfa.pn@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="8F45FFFF6ECF1F1A3639CE2C7EA38417" type="personal">
<mods:role id="6968789C86BE9E152802C801DA72BB72">
<mods:roleTerm id="F4759BE03AB1D448418ED9E9CE9E11E5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="575C80A5843F370D52DFA886E43C0A86">Vilarino, Albane</mods:namePart>
<mods:nameIdentifier id="894BBD61FCC1DFDCC58FA3E37A1A533E" type="ORCID">0000-0003-3912-8928</mods:nameIdentifier>
<mods:affiliation id="2FBD67FD5B14FC80B74623CD7C6C2D6A">Museu de Entomologia, Departamento de Entomologia, Universidade Federal de Viçosa, Av. P. H. Rolfs, s. n, Campus Universitário, CEP 36570 - 900, Viçosa, Minas Gerais, Brazil &amp; albanevilarino @ gmail. com; https: // orcid. org / 0000 - 0003 - 3912 - 8928</mods:affiliation>
<mods:nameIdentifier id="3DEC0730CD6E20458F4C58B3B8CB5FE7" type="email">albanevilarino@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="1AA655B6C3E2B1E000427AE6BC5EA0FA" type="personal">
<mods:role id="0B8DC4C8E48DA124C52F4357523C09C2">
<mods:roleTerm id="A74120964EAA26C84F794234A864F018">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="19F296C205E48FC9098018EBD9C57705">Salles, Frederico F.</mods:namePart>
<mods:nameIdentifier id="1A61A4115BB9FD35C7DCF6801C464142" type="ORCID">0000-0001-8331-5929</mods:nameIdentifier>
<mods:affiliation id="87541BC270C19067818BF64519C7F461">Museu de Entomologia, Departamento de Entomologia, Universidade Federal de Viçosa, Av. P. H. Rolfs, s. n, Campus Universitário, CEP 36570 - 900, Viçosa, Minas Gerais, Brazil &amp; frederico. salles @ ufv. br; https: // orcid. org / 0000 - 0001 - 8331 - 5929</mods:affiliation>
<mods:nameIdentifier id="C7F39061741FB2F96D802A520B7CA690" type="email">frederico.salles@ufv.br</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="33A7E2ED1B8E090132692C59072EE41E">text</mods:typeOfResource>
<mods:relatedItem id="F85DA2A9EAA62359EB9A08A68E33B52C" type="host">
<mods:titleInfo id="304F34FC44703700646F3877D635EF31">
<mods:title id="1ABC722DEE51F5E2119E29F174132941">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B52780E19FE52487FE3E855D0971DD3D">
<mods:date id="45053FA4775EAF7B1886F87F55CA9B56">2023</mods:date>
<mods:detail id="BBFC78F9AE69D278087ED21D9417B88E" type="pubDate">
<mods:number id="43107396B7CA5A9B0137211F5E588C23">2023-08-23</mods:number>
</mods:detail>
<mods:detail id="A3662DE002192C2ACB325BD068952306" type="volume">
<mods:number id="B57FB8D3F85CFBBF9D0FCD2745264BD8">5336</mods:number>
</mods:detail>
<mods:detail id="7A04B3CB7D6599D635B9272A0C5F0100" type="issue">
<mods:number id="0191DFFD0B3347101195A2CB31D981DA">3</mods:number>
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<mods:start id="12D2F927BCD64C3FE84AC8C882EE59F1">301</mods:start>
<mods:end id="45A0E7CFD53D6FE246FE3BAD5AB5B826">327</mods:end>
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<mods:url id="F69BF8CE4E7550C7622BCBBD1F8BFFA0">http://dx.doi.org/10.11646/zootaxa.5336.3.1</mods:url>
</mods:location>
<mods:classification id="62033B3236AD371CD23078B1C16132AE">journal article</mods:classification>
<mods:identifier id="CFF90568851782757887ED2944C916A4" type="DOI">10.11646/zootaxa.5336.3.1</mods:identifier>
<mods:identifier id="3455E10B54D16B4D38C1A6591417E7D4" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="44BCE4AE9AB9E284F08B21D6E6D48F70" type="Zenodo-Dep">8281432</mods:identifier>
<mods:identifier id="C53B709CA55FD9822CE6DA73666F5896" type="ZooBank">B0315010-41C9-4F5A-91D5-7A1D333E0D36</mods:identifier>
</mods:mods>
<treatment id="03AAE777C4254807FF46FD771154F8FD" ID-DOI="http://doi.org/10.5281/zenodo.8281444" ID-Zenodo-Dep="8281444" LSID="urn:lsid:plazi:treatment:03AAE777C4254807FF46FD771154F8FD" httpUri="http://treatment.plazi.org/id/03AAE777C4254807FF46FD771154F8FD" lastPageId="9" lastPageNumber="310" pageId="8" pageNumber="309">
<subSubSection id="C31905EAC4254806FF46FD7712DBFD18" box="[151,483,655,682]" pageId="8" pageNumber="309" type="nomenclature">
<paragraph id="8BBC5661C4254806FF46FD7712DBFD18" blockId="8.[151,967,655,753]" box="[151,483,655,682]" pageId="8" pageNumber="309">
<heading id="D0F4E10DC4254806FF46FD7712DBFD18" bold="true" box="[151,483,655,682]" fontSize="11" level="1" pageId="8" pageNumber="309" reason="1">
<emphasis id="B9778A73C4254806FF46FD7712DBFD18" bold="true" box="[151,483,655,682]" pageId="8" pageNumber="309">
<taxonomicName id="4C032DE2C4254806FF46FD771246FD18" authority="Neto &amp; Vilarino &amp; Salles, 2023" authorityName="Neto &amp; Vilarino &amp; Salles" authorityYear="2023" box="[151,382,655,682]" class="Insecta" family="Calamoceratidae" genus="Phylloicus" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="guarani" status="sp. nov.">
<emphasis id="B9778A73C4254806FF46FD771246FD18" bold="true" box="[151,382,655,682]" italics="true" pageId="8" pageNumber="309">Phylloicus guarani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2443708C4254806FE54FD7712DBFD18" box="[389,483,655,681]" pageId="8" pageNumber="309" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31905EAC4254806FF46FD4B10FFFD40" pageId="8" pageNumber="309" type="description">
<paragraph id="8BBC5661C4254806FF46FD4B12B6FD7F" blockId="8.[151,967,655,753]" box="[151,398,691,718]" pageId="8" pageNumber="309">
(
<figureCitation id="13384AE4C4254806FF4EFD4B1214FD7C" box="[159,300,691,717]" captionStart="FIGURE 4" captionStartId="9.[151,250,1519,1543]" captionTargetBox="[151,1436,181,1495]" captionTargetId="figure-18@9.[151,1436,181,1495]" captionTargetPageId="9" captionText="FIGURE 4. Phylloicus guarani sp. nov., adult male, dorsal. 4A, habitus; 4B, head and thorax; 4C, head. Abbreviations: a.w. = anterior setal wart; am.w. = anteromesal setal wart; l.pr.w. = lateral pronotal setal wart; m.pr.w. = mesal pronotal setal wart; p.w. = posterior setal wart; pl.w. = posterolateral setal wart. Scale bar: A = 1 mm; B = 0.5 mm; C = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8281448" httpUri="https://zenodo.org/record/8281448/files/figure.png" pageId="8" pageNumber="309">Figs 4A–4C</figureCitation>
,
<figureCitation id="13384AE4C4254806FEE6FD4B12BEFD7F" box="[311,390,691,718]" captionStart="FIGURE 5" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[199,1388,191,1910]" captionTargetId="figure-28@10.[189,1399,181,1920]" captionTargetPageId="10" captionText="FIGURE 5. Phylloicus guarani sp. nov., adult male. 5A, right fore- and hind wing venation, dorsal. 5B–5F, genitalia: 5B, left lateral; 5C, dorsal; 5D, ventral; 5E, phallus, left lateral, everted; 5F, phallus, ventral, everted (above) and not everted (below)." figureDoi="http://doi.org/10.5281/zenodo.8281452" httpUri="https://zenodo.org/record/8281452/files/figure.png" pageId="8" pageNumber="309">5A–5F</figureCitation>
)
</paragraph>
<paragraph id="8BBC5661C4254806FF46FD2F10FFFD40" blockId="8.[151,967,655,753]" box="[151,967,727,753]" pageId="8" pageNumber="309">
<uri id="FF925A63C4254806FF46FD2F10FFFD40" box="[151,967,727,753]" pageId="8" pageNumber="309">
urn:lsid:zoobank.org:act:
<uuid id="FFA56CB4C4254806FE78FD2F10FFFD40" box="[425,967,727,753]" pageId="8" pageNumber="309">D324A06E-263F-482D-9E27-281ECAEC12F1</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C31905EAC4254806FF46FCE1126CFC58" pageId="8" pageNumber="309" type="materials_examined">
<paragraph id="8BBC5661C4254806FF46FCE1163EFCE6" blockId="8.[151,1437,792,2043]" pageId="8" pageNumber="309">
<materialsCitation id="3B6B5C3CC4254806FF46FCE1163EFCE6" collectingDate="2017-06-21" collectingDateMax="2017-07-26" collectingDateMin="2017-06-21" collectionCode="UFVB" country="Brazil" county="Santa Teresa" elevation="830" latitude="-19.867369" location="Bragacho" longLatPrecision="1" longitude="-40.559517" municipality="Reserva Biologica Augusto Ruschi" pageId="8" pageNumber="309" specimenCode="TR00082" specimenCount="1" stateProvince="Espirito Santo" typeStatus="holotype">
<emphasis id="B9778A73C4254806FF46FCE11275FC82" bold="true" box="[151,333,793,819]" pageId="8" pageNumber="309">
<typeStatus id="54B8E8C3C4254806FF46FCE1123CFC82" box="[151,260,793,819]" pageId="8" pageNumber="309">Holotype</typeStatus>
male.
</emphasis>
<collectingCountry id="F31416F1C4254806FE84FCE012A4FC83" box="[341,412,792,818]" name="Brazil" pageId="8" pageNumber="309">Brazil</collectingCountry>
,
<collectingRegion id="49C79883C4254806FE78FCE01175FC82" box="[425,589,792,819]" country="Brazil" name="Espirito Santo" pageId="8" pageNumber="309">Espírito Santo</collectingRegion>
,
<collectingCounty id="62DD2EEDC4254806FD88FCE111D3FC82" box="[601,747,793,819]" pageId="8" pageNumber="309">Santa Teresa</collectingCounty>
,
<collectingMunicipality id="6BD8CC1BC4254806FD27FCE117BDFC83" box="[758,1157,792,819]" pageId="8" pageNumber="309">Reserva Biológica Augusto Ruschi</collectingMunicipality>
,
<location id="8EDC00BAC4254806FB40FCE017C4FC83" LSID="urn:lsid:plazi:treatment:03AAE777C4254807FF46FD771154F8FD:8EDC00BAC4254806FB40FCE017C4FC83" box="[1169,1276,792,818]" country="Brazil" county="Santa Teresa" latitude="-19.867369" longLatPrecision="1" longitude="-40.559517" municipality="Reserva Biologica Augusto Ruschi" name="Bragacho" pageId="8" pageNumber="309" stateProvince="Espirito Santo">Bragacho</location>
stream (P13),
<geoCoordinate id="EE3730A6C4254806FF46FCC41209FCE6" box="[151,305,828,855]" degrees="19" direction="south" minutes="52" orientation="latitude" pageId="8" pageNumber="309" precision="1" seconds="2.53" value="-19.867369">19°52’2.53”S</geoCoordinate>
<geoCoordinate id="EE3730A6C4254806FEE9FCC512D2FCE6" box="[312,490,829,855]" degrees="40" direction="west" minutes="33" orientation="longitude" pageId="8" pageNumber="309" precision="1" seconds="34.27" value="-40.559517">40°33’34.27”W</geoCoordinate>
,
<quantity id="4CFBFB84C4254806FE27FCC51107FCE6" box="[502,575,829,855]" metricMagnitude="2" metricUnit="m" metricValue="8.3" pageId="8" pageNumber="309" unit="m" value="830.0">
<elevation id="002EB152C4254806FE27FCC51107FCE6" box="[502,575,829,855]" metricMagnitude="2" metricUnit="m" metricValue="8.3" pageId="8" pageNumber="309" unit="m" value="830.0">830 m</elevation>
</quantity>
,
<date id="FFBD70A1C4254806FD9BFCC4102CFCE7" box="[586,788,828,854]" pageId="8" pageNumber="309" value="2017-06-21" valueMax="2017-07-26" valueMin="2017-06-21">
<collectingDate id="EFF98949C4254806FD9BFCC4102CFCE7" box="[586,788,828,854]" pageId="8" pageNumber="309" value="2017-06-21" valueMax="2017-07-26" valueMin="2017-06-21">21.vi–26.vii.2017</collectingDate>
</date>
, Malaise, CEUNES col. (
<collectionCode id="ED12CEA4C4254806FBE0FCC517B9FCE6" box="[1073,1153,829,855]" country="Brazil" httpUri="http://grbio.org/cool/3i20-rx1w" name="Vicosa, Universidade Federal de Vicosa, Museum of Entomology" pageId="8" pageNumber="309">UFVB</collectionCode>
:
<specimenCode id="DBA5FE1AC4254806FB5DFCC517C3FCE6" box="[1164,1275,829,855]" collectionCode="UFVB" country="Brazil" httpUri="http://grbio.org/cool/3i20-rx1w" name="Vicosa, Universidade Federal de Vicosa, Museum of Entomology" pageId="8" pageNumber="309">TR00082</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BBC5661C4254806FF16FC991101FC2E" blockId="8.[151,1437,792,2043]" pageId="8" pageNumber="309">
<materialsCitation id="3B6B5C3CC4254806FF16FC99110DFC2E" collectingDate="2017-07-26" collectingDateMax="2017-11-18" collectingDateMin="2017-07-26" collectionCode="UFVB" country="Brazil" county="Santa Teresa" elevation="830" latitude="-19.867369" location="Bragacho" longLatPrecision="1" longitude="-40.559517" municipality="Reserva Biologica Augusto Ruschi" pageId="8" pageNumber="309" specimenCode="TR00083, TR00084" specimenCount="1" stateProvince="Espirito Santo" typeStatus="paratype">
<emphasis id="B9778A73C4254806FF16FC99127EFCCA" bold="true" box="[199,326,865,891]" pageId="8" pageNumber="309">
<typeStatus id="54B8E8C3C4254806FF16FC991279FCCA" box="[199,321,865,891]" pageId="8" pageNumber="309">Paratypes</typeStatus>
.
</emphasis>
Same data as holotype, except
<date id="FFBD70A1C4254806FD6CFC9810ABFCCB" box="[701,915,864,890]" pageId="8" pageNumber="309" value="2017-07-26" valueMax="2017-08-23" valueMin="2017-07-26">
<collectingDate id="EFF98949C4254806FD6CFC9810ABFCCB" box="[701,915,864,890]" pageId="8" pageNumber="309" value="2017-07-26" valueMax="2017-08-23" valueMin="2017-07-26">26.vii–23.viii.2017</collectingDate>
</date>
(1♁,
<collectionCode id="ED12CEA4C4254806FC0DFC991713FCCA" box="[988,1067,865,891]" country="Brazil" httpUri="http://grbio.org/cool/3i20-rx1w" name="Vicosa, Universidade Federal de Vicosa, Museum of Entomology" pageId="8" pageNumber="309">UFVB</collectionCode>
:
<specimenCode id="DBA5FE1AC4254806FBE9FC99179FFCCA" box="[1080,1191,865,891]" collectionCode="UFVB" country="Brazil" httpUri="http://grbio.org/cool/3i20-rx1w" name="Vicosa, Universidade Federal de Vicosa, Museum of Entomology" pageId="8" pageNumber="309">TR00083</specimenCode>
); same, except
<date id="FFBD70A1C4254806FA8CFC991237FC2F" pageId="8" pageNumber="309" value="2017-10-21" valueMax="2017-11-18" valueMin="2017-10-21">
<collectingDate id="EFF98949C4254806FA8CFC991237FC2F" pageId="8" pageNumber="309" value="2017-10-21" valueMax="2017-11-18" valueMin="2017-10-21">21.x– 18.xi.2017</collectingDate>
</date>
(2♁♁,
<collectionCode id="ED12CEA4C4254806FEB4FC7D128CFC2E" box="[357,436,901,927]" country="Brazil" httpUri="http://grbio.org/cool/3i20-rx1w" name="Vicosa, Universidade Federal de Vicosa, Museum of Entomology" pageId="8" pageNumber="309">UFVB</collectionCode>
:
<specimenCode id="DBA5FE1AC4254806FE6FFC7D1116FC2E" box="[446,558,901,927]" collectionCode="UFVB" country="Brazil" httpUri="http://grbio.org/cool/3i20-rx1w" name="Vicosa, Universidade Federal de Vicosa, Museum of Entomology" pageId="8" pageNumber="309">TR00084</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BBC5661C4254806FF16FC50126CFC58" blockId="8.[151,1437,792,2043]" pageId="8" pageNumber="309">
<emphasis id="B9778A73C4254806FF16FC5011BBFC73" bold="true" box="[199,643,936,962]" pageId="8" pageNumber="309">Additional material examined. P13:</emphasis>
<date id="FFBD70A1C4254806FD5EFC501008FC73" box="[655,816,936,962]" pageId="8" pageNumber="309" value="2017-06-21" valueMax="2017-06-22" valueMin="2017-06-21">21–22.vi.2017</date>
(1♁),
<date id="FFBD70A1C4254806FC57FC50173DFC73" box="[902,1029,936,962]" pageId="8" pageNumber="309" value="2017-12-28">28.xii.2017</date>
(1♁),
<date id="FFBD70A1C4254806FB8DFC50163EFC73" box="[1116,1286,936,962]" pageId="8" pageNumber="309" value="2018-01-17" valueMax="2018-02-20" valueMin="2018-01-17">17.i–20.ii.2018</date>
(1♁),
<date id="FFBD70A1C4254806FA8DFC501228FC57" pageId="8" pageNumber="309" value="2018-02-20" valueMax="2018-03-21" valueMin="2018-02-20">20.ii– 21.iii.2018</date>
(1♁).
</paragraph>
</subSubSection>
<subSubSection id="C31905EAC4254806FF16FC0810D2FA9A" pageId="8" pageNumber="309" type="diagnosis">
<paragraph id="8BBC5661C4254806FF16FC0810D2FA9A" blockId="8.[151,1437,792,2043]" pageId="8" pageNumber="309">
<emphasis id="B9778A73C4254806FF16FC08127AFBBB" bold="true" box="[199,322,1008,1034]" pageId="8" pageNumber="309">Diagnosis.</emphasis>
The new species is very similar to
<taxonomicName id="4C032DE2C4254806FD39FC08164EFBBA" authority="Barcelos-Silva, Desiderio &amp; Pes 2017" authorityName="Barcelos-Silva, Desiderio &amp; Pes" authorityYear="2017" box="[744,1398,1008,1035]" class="Insecta" family="Calamoceratidae" genus="Phylloicus" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="sallesi">
<emphasis id="B9778A73C4254806FD39FC081089FBBB" box="[744,945,1008,1034]" italics="true" pageId="8" pageNumber="309">Phylloicus sallesi</emphasis>
<bibRefCitation id="EF922B90C4254806FC6AFC08164EFBBA" author="Barcelos-Silva, P. &amp; Desiderio, G. R. &amp; Pes, A. M." box="[955,1398,1008,1035]" pageId="8" pageNumber="309" pagination="287 - 292" refId="ref15883" refString="Barcelos-Silva, P., Desiderio, G. R. &amp; Pes, A. M. (2017) A new species of Phylloicus M ¸ ller 1880 (Trichoptera: Calamoceratidae) from Espirito Santo State, southeastern Brazil. Zootaxa, 4238 (2), 287 - 292. https: // doi. org / 10.11646 / zootaxa. 4238.2.11" type="journal article" year="2017">Barcelos-Silva, Desidério &amp; Pes 2017</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C032DE2C4254806FA55FC091295FB9E" authority="Prather 2003" authorityName="Prather" authorityYear="2003" class="Insecta" family="Calamoceratidae" genus="Phylloicus" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="bidigitatus">
<emphasis id="B9778A73C4254806FA55FC0916ADFBBB" box="[1412,1429,1009,1034]" italics="true" pageId="8" pageNumber="309">P</emphasis>
.
<emphasis id="B9778A73C4254806FF46FBEC1237FB9F" box="[151,271,1044,1070]" italics="true" pageId="8" pageNumber="309">bidigitatus</emphasis>
<bibRefCitation id="EF922B90C4254806FEC9FBEC1295FB9E" author="Prather, A. L." box="[280,429,1044,1071]" pageId="8" pageNumber="309" pagination="1 - 214" refId="ref18694" refString="Prather, A. L. (2003) Revision of the Neotropical caddisfly genus Phylloicus (Trichoptera: Calamoceratidae). Zootaxa, 275 (1), 1 - 214. https: // doi. org / 10.11646 / zootaxa. 275.1.1" type="journal article" year="2003">Prather 2003</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C032DE2C4254806FE68FBED11F1FB9E" authority="Navas 1931" authorityName="Navas" authorityYear="1931" box="[441,713,1044,1071]" class="Insecta" family="Calamoceratidae" genus="Phylloicus" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="obliquus">
<emphasis id="B9778A73C4254806FE68FBED110FFB9F" box="[441,567,1044,1070]" italics="true" pageId="8" pageNumber="309">P. obliquus</emphasis>
<bibRefCitation id="EF922B90C4254806FD91FBED11F1FB9E" author="Navas, L." box="[576,713,1045,1071]" pageId="8" pageNumber="309" pagination="455 - 458" refId="ref18498" refString="Navas, L. (1931) Insectos del Brasil, 4 a serie. Revista do Museu Paulista, 17 (1), 455 - 458." type="journal article" year="1931">Navás 1931</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C032DE2C4254806FCD9FBED171BFB9E" authority="Flint 1983" authorityName="Flint" authorityYear="1983" box="[776,1059,1044,1071]" class="Insecta" family="Calamoceratidae" genus="Phylloicus" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="plaumanni">
<emphasis id="B9778A73C4254806FCD9FBED1021FB9F" box="[776,793,1045,1070]" italics="true" pageId="8" pageNumber="309">P</emphasis>
.
<emphasis id="B9778A73C4254806FCF8FBEC1099FB9F" box="[809,929,1044,1070]" italics="true" pageId="8" pageNumber="309">plaumanni</emphasis>
<bibRefCitation id="EF922B90C4254806FC7BFBEC171BFB9E" author="Flint, O. S. Jr." box="[938,1059,1044,1071]" pageId="8" pageNumber="309" pagination="1 - 100" refId="ref16759" refString="Flint, O. S. Jr. (1983) Studies of Neotropical caddisflies, XXXIII: New species from austral South America (Trichoptera). Smithsonian Contributions to Zoology, 377, 1 - 100. https: // doi. org / 10.5479 / si. 00810282.377" type="journal article" year="1983">Flint 1983</bibRefCitation>
</taxonomicName>
. All these species have a median notch at the apex of segment X; the apical joint of the inferior appendage is elongate, triangular, and with three small robust bristles at the tip.
<taxonomicName id="4C032DE2C4254806FE60FBA411B1FBC7" authority="Neto &amp; Vilarino &amp; Salles, 2023" authorityName="Neto &amp; Vilarino &amp; Salles" authorityYear="2023" box="[433,649,1116,1142]" class="Insecta" family="Calamoceratidae" genus="Phylloicus" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="guarani" status="sp. nov.">
<emphasis id="B9778A73C4254806FE60FBA411B1FBC7" box="[433,649,1116,1142]" italics="true" pageId="8" pageNumber="309">Phylloicus guarani</emphasis>
</taxonomicName>
<emphasis id="B9778A73C4254806FD41FBA511D3FBC6" bold="true" box="[656,747,1117,1143]" pageId="8" pageNumber="309">
<taxonomicNameLabel id="A2443708C4254806FD41FBA511D3FBC6" box="[656,747,1117,1143]" pageId="8" pageNumber="309" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has the arms formed by the cavity of the apex of segment X round like those of
<taxonomicName id="4C032DE2C4254806FEBDFB7912C7FB2B" authorityName="Prather" authorityYear="2003" box="[364,511,1152,1178]" class="Insecta" family="Calamoceratidae" genus="Phylloicus" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="bidigitatus">
<emphasis id="B9778A73C4254806FEBDFB7912C7FB2B" box="[364,511,1152,1178]" italics="true" pageId="8" pageNumber="309">P. bidigitatus</emphasis>
</taxonomicName>
and not pointed like those of
<taxonomicName id="4C032DE2C4254806FC95FB79109EFB2B" authorityName="Barcelos-Silva, Desiderio &amp; Pes" authorityYear="2017" box="[836,934,1152,1178]" class="Insecta" family="Calamoceratidae" genus="Phylloicus" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="sallesi">
<emphasis id="B9778A73C4254806FC95FB79109EFB2B" box="[836,934,1152,1178]" italics="true" pageId="8" pageNumber="309">P. sallesi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C032DE2C4254806FC62FB791715FB2B" authorityName="Navas" authorityYear="1931" box="[947,1069,1152,1178]" class="Insecta" family="Calamoceratidae" genus="Phylloicus" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="obliquus">
<emphasis id="B9778A73C4254806FC62FB791715FB2B" box="[947,1069,1152,1178]" italics="true" pageId="8" pageNumber="309">P. obliquus</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C032DE2C4254806FBB8FB7917C3FB2B" authorityName="Flint" authorityYear="1983" box="[1129,1275,1152,1178]" class="Insecta" family="Calamoceratidae" genus="Phylloicus" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="plaumanni">
<emphasis id="B9778A73C4254806FBB8FB7917C3FB2B" box="[1129,1275,1152,1178]" italics="true" pageId="8" pageNumber="309">P. plaumanni</emphasis>
</taxonomicName>
. However, the basal part of segment X in dorsal view in the new species has a convex prominence, as in
<taxonomicName id="4C032DE2C4254806FB50FB5D17DCFB0F" authorityName="Barcelos-Silva, Desiderio &amp; Pes" authorityYear="2017" box="[1153,1252,1188,1214]" class="Insecta" family="Calamoceratidae" genus="Phylloicus" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="sallesi">
<emphasis id="B9778A73C4254806FB50FB5D17DCFB0F" box="[1153,1252,1188,1214]" italics="true" pageId="8" pageNumber="309">P. sallesi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C032DE2C4254806FACBFB5D16ADFB0F" authorityName="Navas" authorityYear="1931" box="[1306,1429,1188,1214]" class="Insecta" family="Calamoceratidae" genus="Phylloicus" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="obliquus">
<emphasis id="B9778A73C4254806FACBFB5D16ADFB0F" box="[1306,1429,1188,1214]" italics="true" pageId="8" pageNumber="309">P. obliquus</emphasis>
</taxonomicName>
. The inferior appendages of
<taxonomicName id="4C032DE2C4254806FE04FB311174FB53" authority="Neto &amp; Vilarino &amp; Salles, 2023" authorityName="Neto &amp; Vilarino &amp; Salles" authorityYear="2023" box="[469,588,1225,1250]" class="Insecta" family="Calamoceratidae" genus="Phylloicus" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="guarani" status="sp. nov.">
<emphasis id="B9778A73C4254806FE04FB311174FB53" box="[469,588,1225,1250]" italics="true" pageId="8" pageNumber="309">P. guarani</emphasis>
</taxonomicName>
<emphasis id="B9778A73C4254806FD85FB311188FB52" bold="true" box="[596,688,1225,1251]" pageId="8" pageNumber="309">
<taxonomicNameLabel id="A2443708C4254806FD85FB311188FB52" box="[596,688,1225,1251]" pageId="8" pageNumber="309" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are relatively smaller than those of
<taxonomicName id="4C032DE2C4254806FB80FB31178EFB53" authorityName="Barcelos-Silva, Desiderio &amp; Pes" authorityYear="2017" box="[1105,1206,1224,1250]" class="Insecta" family="Calamoceratidae" genus="Phylloicus" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="sallesi">
<emphasis id="B9778A73C4254806FB80FB31178EFB53" box="[1105,1206,1224,1250]" italics="true" pageId="8" pageNumber="309">P. sallesi</emphasis>
</taxonomicName>
, approximately half the length of segment X. The new species also can be differentiated by not having the small curvature or globose expansion of the apicoventral margin of the phallobase, present in
<taxonomicName id="4C032DE2C4254806FC51FAE910DCFA9B" authorityName="Barcelos-Silva, Desiderio &amp; Pes" authorityYear="2017" box="[896,996,1296,1322]" class="Insecta" family="Calamoceratidae" genus="Phylloicus" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="sallesi">
<emphasis id="B9778A73C4254806FC51FAE910DCFA9B" box="[896,996,1296,1322]" italics="true" pageId="8" pageNumber="309">P. sallesi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C31905EAC4254806FF16FACC1003F84A" pageId="8" pageNumber="309" type="description">
<paragraph id="8BBC5661C4254806FF16FACC10F7F9FA" blockId="8.[151,1437,792,2043]" pageId="8" pageNumber="309">
<emphasis id="B9778A73C4254806FF16FACC1261FAFF" bold="true" box="[199,345,1332,1358]" pageId="8" pageNumber="309">Description.</emphasis>
<emphasis id="B9778A73C4254806FEB2FACC12D0FAFF" box="[355,488,1332,1358]" italics="true" pageId="8" pageNumber="309">Adult male.</emphasis>
Forewing length 9.0 mm (
<typeStatus id="54B8E8C3C4254806FCF5FACC10B5FAFF" box="[804,909,1332,1358]" pageId="8" pageNumber="309">holotype</typeStatus>
). Body color overall light brown, wings dark brown (in alcohol) (
<figureCitation id="13384AE4C4254806FEAEFAA012E5FAC2" box="[383,477,1368,1395]" captionStart="FIGURE 4" captionStartId="9.[151,250,1519,1543]" captionTargetBox="[151,1436,181,1495]" captionTargetId="figure-18@9.[151,1436,181,1495]" captionTargetPageId="9" captionText="FIGURE 4. Phylloicus guarani sp. nov., adult male, dorsal. 4A, habitus; 4B, head and thorax; 4C, head. Abbreviations: a.w. = anterior setal wart; am.w. = anteromesal setal wart; l.pr.w. = lateral pronotal setal wart; m.pr.w. = mesal pronotal setal wart; p.w. = posterior setal wart; pl.w. = posterolateral setal wart. Scale bar: A = 1 mm; B = 0.5 mm; C = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8281448" httpUri="https://zenodo.org/record/8281448/files/figure.png" pageId="8" pageNumber="309">Fig. 4A</figureCitation>
).
<emphasis id="B9778A73C4254806FE23FAA01100FAC3" box="[498,568,1368,1394]" italics="true" pageId="8" pageNumber="309">Head:</emphasis>
Central setal area dark brown; anteromesal setal wart single, heart-shaped; anterior setal wart small, round; posterior setal wart narrow, elongate. Maxillary palp setose. Antenna long, scape wide (
<figureCitation id="13384AE4C4254806FF31FA581203FA0A" box="[224,315,1440,1467]" captionStart="FIGURE 4" captionStartId="9.[151,250,1519,1543]" captionTargetBox="[151,1436,181,1495]" captionTargetId="figure-18@9.[151,1436,181,1495]" captionTargetPageId="9" captionText="FIGURE 4. Phylloicus guarani sp. nov., adult male, dorsal. 4A, habitus; 4B, head and thorax; 4C, head. Abbreviations: a.w. = anterior setal wart; am.w. = anteromesal setal wart; l.pr.w. = lateral pronotal setal wart; m.pr.w. = mesal pronotal setal wart; p.w. = posterior setal wart; pl.w. = posterolateral setal wart. Scale bar: A = 1 mm; B = 0.5 mm; C = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8281448" httpUri="https://zenodo.org/record/8281448/files/figure.png" pageId="8" pageNumber="309">Fig. 4B</figureCitation>
).
<emphasis id="B9778A73C4254806FE83FA581294FA0B" box="[338,428,1440,1466]" italics="true" pageId="8" pageNumber="309">Thorax:</emphasis>
Pronotum narrow with small round mesal and lateral pronotal setal warts. Mesonotum wide, with mesoscutal setal wart diffuse, reaching mesoscutellum; mesoscutellum heart-shaped (
<figureCitation id="13384AE4C4254806FB22FA3C1677FA6E" box="[1267,1359,1476,1503]" captionStart="FIGURE 4" captionStartId="9.[151,250,1519,1543]" captionTargetBox="[151,1436,181,1495]" captionTargetId="figure-18@9.[151,1436,181,1495]" captionTargetPageId="9" captionText="FIGURE 4. Phylloicus guarani sp. nov., adult male, dorsal. 4A, habitus; 4B, head and thorax; 4C, head. Abbreviations: a.w. = anterior setal wart; am.w. = anteromesal setal wart; l.pr.w. = lateral pronotal setal wart; m.pr.w. = mesal pronotal setal wart; p.w. = posterior setal wart; pl.w. = posterolateral setal wart. Scale bar: A = 1 mm; B = 0.5 mm; C = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8281448" httpUri="https://zenodo.org/record/8281448/files/figure.png" pageId="8" pageNumber="309">Fig. 4B</figureCitation>
). Legs light brown. Tibial spur formula 2-4-4. Forewing forks I–V present; hind wing forks II, III and V present (
<figureCitation id="13384AE4C4254806FAE7FA1016A8F9B2" box="[1334,1424,1512,1539]" captionStart="FIGURE 5" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[199,1388,191,1910]" captionTargetId="figure-28@10.[189,1399,181,1920]" captionTargetPageId="10" captionText="FIGURE 5. Phylloicus guarani sp. nov., adult male. 5A, right fore- and hind wing venation, dorsal. 5B–5F, genitalia: 5B, left lateral; 5C, dorsal; 5D, ventral; 5E, phallus, left lateral, everted; 5F, phallus, ventral, everted (above) and not everted (below)." figureDoi="http://doi.org/10.5281/zenodo.8281452" httpUri="https://zenodo.org/record/8281452/files/figure.png" pageId="8" pageNumber="309">Fig. 5A</figureCitation>
).
<emphasis id="B9778A73C4254806FF46F9F41239F997" box="[151,257,1548,1574]" italics="true" pageId="8" pageNumber="309">Abdomen</emphasis>
: Preterminalic abdominal terga I–VIII each without anteromesal notch. Corematic structures absent, terga III–VIII unmodified, without membranous lobes or sclerotized processes.
</paragraph>
<paragraph id="8BBC5661C4254806FF16F9AC1003F84A" blockId="8.[151,1437,792,2043]" pageId="8" pageNumber="309">
<emphasis id="B9778A73C4254806FF16F9AC1255F9DF" box="[199,365,1620,1646]" italics="true" pageId="8" pageNumber="309">Male genitalia</emphasis>
(
<figureCitation id="13384AE4C4254806FEACF9AC1131F9DE" box="[381,521,1620,1647]" captionStart="FIGURE 5" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[199,1388,191,1910]" captionTargetId="figure-28@10.[189,1399,181,1920]" captionTargetPageId="10" captionText="FIGURE 5. Phylloicus guarani sp. nov., adult male. 5A, right fore- and hind wing venation, dorsal. 5B–5F, genitalia: 5B, left lateral; 5C, dorsal; 5D, ventral; 5E, phallus, left lateral, everted; 5F, phallus, ventral, everted (above) and not everted (below)." figureDoi="http://doi.org/10.5281/zenodo.8281452" httpUri="https://zenodo.org/record/8281452/files/figure.png" pageId="8" pageNumber="309">Figs 5B–5F</figureCitation>
): Tergum IX with dorsomesal suture subtle and short; posterior margin roof-like (
<figureCitation id="13384AE4C4254806FF4EF98013C0F922" box="[159,248,1656,1683]" captionStart="FIGURE 5" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[199,1388,191,1910]" captionTargetId="figure-28@10.[189,1399,181,1920]" captionTargetPageId="10" captionText="FIGURE 5. Phylloicus guarani sp. nov., adult male. 5A, right fore- and hind wing venation, dorsal. 5B–5F, genitalia: 5B, left lateral; 5C, dorsal; 5D, ventral; 5E, phallus, left lateral, everted; 5F, phallus, ventral, everted (above) and not everted (below)." figureDoi="http://doi.org/10.5281/zenodo.8281452" httpUri="https://zenodo.org/record/8281452/files/figure.png" pageId="8" pageNumber="309">Fig. 5C</figureCitation>
); lateral suture present; dorsopleural setae present, ventropleural setae absent (
<figureCitation id="13384AE4C4254806FBA2F98017F4F922" box="[1139,1228,1656,1683]" captionStart="FIGURE 5" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[199,1388,191,1910]" captionTargetId="figure-28@10.[189,1399,181,1920]" captionTargetPageId="10" captionText="FIGURE 5. Phylloicus guarani sp. nov., adult male. 5A, right fore- and hind wing venation, dorsal. 5B–5F, genitalia: 5B, left lateral; 5C, dorsal; 5D, ventral; 5E, phallus, left lateral, everted; 5F, phallus, ventral, everted (above) and not everted (below)." figureDoi="http://doi.org/10.5281/zenodo.8281452" httpUri="https://zenodo.org/record/8281452/files/figure.png" pageId="8" pageNumber="309">Fig. 5B</figureCitation>
); sternum IX with mesolateral suture (
<figureCitation id="13384AE4C4254806FEA1F96412F0F906" box="[368,456,1692,1719]" captionStart="FIGURE 5" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[199,1388,191,1910]" captionTargetId="figure-28@10.[189,1399,181,1920]" captionTargetPageId="10" captionText="FIGURE 5. Phylloicus guarani sp. nov., adult male. 5A, right fore- and hind wing venation, dorsal. 5B–5F, genitalia: 5B, left lateral; 5C, dorsal; 5D, ventral; 5E, phallus, left lateral, everted; 5F, phallus, ventral, everted (above) and not everted (below)." figureDoi="http://doi.org/10.5281/zenodo.8281452" httpUri="https://zenodo.org/record/8281452/files/figure.png" pageId="8" pageNumber="309">Fig. 5D</figureCitation>
). Preanal appendage, in lateral view, not surpassing segment X, about as long as segment X, of uniform diameter throughout length, with setae long, but not filamentous or longer than appendage (
<figureCitation id="13384AE4C4254806FAE6F93816B7F96A" box="[1335,1423,1728,1755]" captionStart="FIGURE 5" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[199,1388,191,1910]" captionTargetId="figure-28@10.[189,1399,181,1920]" captionTargetPageId="10" captionText="FIGURE 5. Phylloicus guarani sp. nov., adult male. 5A, right fore- and hind wing venation, dorsal. 5B–5F, genitalia: 5B, left lateral; 5C, dorsal; 5D, ventral; 5E, phallus, left lateral, everted; 5F, phallus, ventral, everted (above) and not everted (below)." figureDoi="http://doi.org/10.5281/zenodo.8281452" httpUri="https://zenodo.org/record/8281452/files/figure.png" pageId="8" pageNumber="309">Fig. 5B</figureCitation>
). Tergum X without basal lobes; basodorsal and basolateral processes absent; in lateral view, subtriangular, apex narrow, round (
<figureCitation id="13384AE4C4254806FE95F8F012A5F892" box="[324,413,1800,1827]" captionStart="FIGURE 5" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[199,1388,191,1910]" captionTargetId="figure-28@10.[189,1399,181,1920]" captionTargetPageId="10" captionText="FIGURE 5. Phylloicus guarani sp. nov., adult male. 5A, right fore- and hind wing venation, dorsal. 5B–5F, genitalia: 5B, left lateral; 5C, dorsal; 5D, ventral; 5E, phallus, left lateral, everted; 5F, phallus, ventral, everted (above) and not everted (below)." figureDoi="http://doi.org/10.5281/zenodo.8281452" httpUri="https://zenodo.org/record/8281452/files/figure.png" pageId="8" pageNumber="309">Fig. 5B</figureCitation>
); in dorsal view, wide at base, narrow near midlength, apex notched, notch elliptical, deep, extending through 1/3 of segment length, forming digitate posterior lobes; tergum covered with short setae dorsally (
<figureCitation id="13384AE4C4254806FF4EF8A813C1F8DA" box="[159,249,1872,1899]" captionStart="FIGURE 5" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[199,1388,191,1910]" captionTargetId="figure-28@10.[189,1399,181,1920]" captionTargetPageId="10" captionText="FIGURE 5. Phylloicus guarani sp. nov., adult male. 5A, right fore- and hind wing venation, dorsal. 5B–5F, genitalia: 5B, left lateral; 5C, dorsal; 5D, ventral; 5E, phallus, left lateral, everted; 5F, phallus, ventral, everted (above) and not everted (below)." figureDoi="http://doi.org/10.5281/zenodo.8281452" httpUri="https://zenodo.org/record/8281452/files/figure.png" pageId="8" pageNumber="309">Fig. 5C</figureCitation>
). Inferior appendage bi-articulated, short; coxopodite short, round apicodorsally, covered with long setae; harpago in ventral view almost as long as coxopodite, sharply tapered with few apical spine-like setae, in lateral view short, hooked dorsad (
<figureCitation id="13384AE4C4254806FE04F860115EF802" box="[469,614,1944,1971]" captionStart="FIGURE 5" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[199,1388,191,1910]" captionTargetId="figure-28@10.[189,1399,181,1920]" captionTargetPageId="10" captionText="FIGURE 5. Phylloicus guarani sp. nov., adult male. 5A, right fore- and hind wing venation, dorsal. 5B–5F, genitalia: 5B, left lateral; 5C, dorsal; 5D, ventral; 5E, phallus, left lateral, everted; 5F, phallus, ventral, everted (above) and not everted (below)." figureDoi="http://doi.org/10.5281/zenodo.8281452" httpUri="https://zenodo.org/record/8281452/files/figure.png" pageId="8" pageNumber="309">Figs 5B, 5D</figureCitation>
). Phallus in lateral view slightly enlarged basally, curved caudoventrad; endotheca membranous, with paired apicolateral lobes, evident when everted; phallotremal sclerite hook-like in lateral view (
<figureCitation id="13384AE4C4254806FEFBF81812B8F84A" box="[298,384,2016,2043]" captionStart="FIGURE 5" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[199,1388,191,1910]" captionTargetId="figure-28@10.[189,1399,181,1920]" captionTargetPageId="10" captionText="FIGURE 5. Phylloicus guarani sp. nov., adult male. 5A, right fore- and hind wing venation, dorsal. 5B–5F, genitalia: 5B, left lateral; 5C, dorsal; 5D, ventral; 5E, phallus, left lateral, everted; 5F, phallus, ventral, everted (above) and not everted (below)." figureDoi="http://doi.org/10.5281/zenodo.8281452" httpUri="https://zenodo.org/record/8281452/files/figure.png" pageId="8" pageNumber="309">Fig. 5E</figureCitation>
), forceps-like in ventral view (
<figureCitation id="13384AE4C4254806FD08F8181016F84A" box="[729,814,2016,2043]" captionStart="FIGURE 5" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[199,1388,191,1910]" captionTargetId="figure-28@10.[189,1399,181,1920]" captionTargetPageId="10" captionText="FIGURE 5. Phylloicus guarani sp. nov., adult male. 5A, right fore- and hind wing venation, dorsal. 5B–5F, genitalia: 5B, left lateral; 5C, dorsal; 5D, ventral; 5E, phallus, left lateral, everted; 5F, phallus, ventral, everted (above) and not everted (below)." figureDoi="http://doi.org/10.5281/zenodo.8281452" httpUri="https://zenodo.org/record/8281452/files/figure.png" pageId="8" pageNumber="309">Fig. 5F</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF7C06E9C4244807FF46FA171792F9FE" ID-DOI="http://doi.org/10.5281/zenodo.8281448" ID-Zenodo-Dep="8281448" httpUri="https://zenodo.org/record/8281448/files/figure.png" pageId="9" pageNumber="310" startId="9.[151,250,1519,1543]" targetBox="[151,1436,181,1495]" targetPageId="9" targetType="figure">
<paragraph id="8BBC5661C4244807FF46FA171792F9FE" blockId="9.[151,1437,1519,1616]" pageId="9" pageNumber="310">
<emphasis id="B9778A73C4244807FF46FA17122BF9B6" bold="true" box="[151,275,1519,1543]" pageId="9" pageNumber="310">FIGURE 4.</emphasis>
<taxonomicName id="4C032DE2C4244807FEC8FA1712E3F9B6" authority="Neto &amp; Vilarino &amp; Salles, 2023" authorityName="Neto &amp; Vilarino &amp; Salles" authorityYear="2023" box="[281,475,1519,1543]" class="Insecta" family="Calamoceratidae" genus="Phylloicus" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="guarani" status="sp. nov.">
<emphasis id="B9778A73C4244807FEC8FA1712E3F9B6" box="[281,475,1519,1543]" italics="true" pageId="9" pageNumber="310">Phylloicus guarani</emphasis>
</taxonomicName>
<emphasis id="B9778A73C4244807FE30FA08110AF9B9" bold="true" box="[481,562,1520,1544]" pageId="9" pageNumber="310">
<taxonomicNameLabel id="A2443708C4244807FE30FA08110AF9B9" box="[481,562,1520,1544]" pageId="9" pageNumber="310" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult male, dorsal. 4A, habitus; 4B, head and thorax; 4C, head. Abbreviations: a.w. = anterior setal wart; am.w. = anteromesal setal wart; l.pr.w. = lateral pronotal setal wart; m.pr.w. = mesal pronotal setal wart; p.w. = posterior setal wart; pl.w. = posterolateral setal wart. Scale bar: A = 1 mm; B = 0.5 mm; C = 0.2 mm.
</paragraph>
</caption>
<subSubSection id="C31905EAC4244807FF16F95A112FF899" pageId="9" pageNumber="310" type="etymology">
<paragraph id="8BBC5661C4244807FF16F95A112FF899" blockId="9.[151,1436,1698,1868]" pageId="9" pageNumber="310">
<emphasis id="B9778A73C4244807FF16F95A1276F90D" bold="true" box="[199,334,1698,1724]" pageId="9" pageNumber="310">Etymology.</emphasis>
The specific epithet “guarani” is the name of the indigenous people who constitute one of the indigenous populations with the greatest territorial presence on the South American continent. In
<collectingRegion id="49C79883C4244807FB2BF93E13EAF8B5" country="Brazil" name="Espirito Santo" pageId="9" pageNumber="310">Espírito Santo State</collectingRegion>
, the Guarani people live on the coast of the municipality of Aracruz. The name is used here as a noun in apposition with unvarying gender.
</paragraph>
</subSubSection>
<subSubSection id="C31905EAC4244807FF16F8C91154F8FD" box="[199,620,1841,1868]" pageId="9" pageNumber="310" type="distribution">
<paragraph id="8BBC5661C4244807FF16F8C91154F8FD" blockId="9.[151,1436,1698,1868]" box="[199,620,1841,1868]" pageId="9" pageNumber="310">
<emphasis id="B9778A73C4244807FF16F8C91259F8FA" bold="true" box="[199,353,1841,1867]" pageId="9" pageNumber="310">Distribution.</emphasis>
<collectingCountry id="F31416F1C4244807FEB6F8CA1293F8FD" box="[359,427,1842,1868]" name="Brazil" pageId="9" pageNumber="310">Brazil</collectingCountry>
(
<collectingRegion id="49C79883C4244807FE6BF8CA1159F8FD" box="[442,609,1842,1868]" country="Brazil" name="Espirito Santo" pageId="9" pageNumber="310">Espírito Santo</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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