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<document ID-DOI="http://doi.org/10.5281/zenodo.3792500" ID-GBIF-Dataset="5b0d19ac-86a1-4e0f-adb3-b7c6230947b8" ID-GBIF-Taxon="163963689" ID-ISSN="13132970" ID-Zenodo-Dep="3792500" ID-ZooBank="93D507B3-AD8F-46D9-9E8C-F209C65A8D2A" LSID="urn:lsid:zoobank.org:act:7DFC5962-B47E-4848-9608-C387CE58049B" approvalRequired="1" approvalRequired_for_matCits="1" checkinTime="1587986346726" checkinUser="plazi" docAuthor="Pollock, Darren" docDate="2009" docId="03A74B2EF2160F526DE18E08FB85FB9C" docLanguage="en" docName="ZK_article_1968.pdf" docOrigin="ZooKeys 6 (6)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@60a754c6" docStyleName="zookeys.2008.journal_article" docTitle="Ischyomius escalonai Pollock 2009, sp. n." docType="treatment" docUuid="7DFC5962-B47E-4848-9608-C387CE58049B" docUuidSource="ZooBank" docVersion="4" lastPageId="4" lastPageNumber="73" masterDocId="FF9E3356F2170F566D6D8C21FFD6FFC8" masterDocTitle="A new species of Ischyomius from Venezuela (Coleoptera, Tenebrionoidea, Pythidae) with a revised key to world species" masterLastPageNumber="74" masterPageNumber="69" pageId="1" pageNumber="70" updateTime="1643567766553" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new species of Ischyomius from Venezuela (Coleoptera, Tenebrionoidea, Pythidae) with a revised key to world species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pollock, Darren</mods:namePart>
<mods:affiliation>Eastern New Mexico University, Portales, United States of America</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-03-11</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>6</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>6</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>69</mods:start>
<mods:end>74</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.6.81</mods:identifier>
<mods:identifier type="GBIF-Dataset">5b0d19ac-86a1-4e0f-adb3-b7c6230947b8</mods:identifier>
<mods:identifier type="ISSN">13132970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576434</mods:identifier>
<mods:identifier type="ZooBank">93D507B3-AD8F-46D9-9E8C-F209C65A8D2A</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3792500" ID-GBIF-Taxon="163963689" ID-Zenodo-Dep="3792500" LSID="urn:lsid:zoobank.org:act:7DFC5962-B47E-4848-9608-C387CE58049B" httpUri="http://treatment.plazi.org/id/03A74B2EF2160F526DE18E08FB85FB9C" lastPageId="4" lastPageNumber="73" pageId="1" pageNumber="70">
<subSubSection box="[140,562,553,580]" pageId="1" pageNumber="70" type="nomenclature">
<paragraph blockId="1.[140,966,553,650]" box="[140,562,553,580]" pageId="1" pageNumber="70">
<heading box="[140,562,553,580]" fontSize="11" level="2" pageId="1" pageNumber="70" reason="3">
<emphasis bold="true" box="[140,562,553,580]" pageId="1" pageNumber="70">
<taxonomicName authority="Pollock" authorityName="Pollock" authorityYear="2009" box="[140,484,553,580]" class="Insecta" family="Pythidae" genus="Ischyomius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="70" phylum="Arthropoda" rank="species" species="escalonai" status="sp. n.">
<emphasis bold="true" box="[140,385,553,580]" italics="true" pageId="1" pageNumber="70">
<emphasis box="[140,269,553,580]" inLineHeading="true" pageId="1" pageNumber="70" reason="1">Ischyomius</emphasis>
escalonai
</emphasis>
Pollock
</taxonomicName>
,
<taxonomicNameLabel box="[497,562,554,580]" pageId="1" pageNumber="70" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="70" type="description">
<paragraph blockId="1.[140,966,553,650]" box="[140,966,589,616]" pageId="1" pageNumber="70">
<uri box="[140,966,589,616]" pageId="1" pageNumber="70">
urn:lsid:zoobank.org:act:
<uuid box="[419,966,589,616]" pageId="1" pageNumber="70">7DFC5962-B47E-4848-9608-C387CE58049B</uuid>
</uri>
</paragraph>
<paragraph blockId="1.[140,966,553,650]" box="[140,257,624,650]" pageId="1" pageNumber="70">
(
<figureCitation box="[148,247,624,650]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="2.[140,174,1558,1580]" captionStartId-1="2.[140,174,1590,1612]" captionStartId-2="2.[140,174,1622,1644]" captionStartId-3="2.[140,174,1654,1676]" captionTargetBox-0="[140,617,165,1003]" captionTargetBox-1="[140,617,1033,1533]" captionTargetBox-2="[648,1108,165,533]" captionTargetBox-3="[648,1107,582,1534]" captionTargetId-0="figure@2.[140,618,165,1003]" captionTargetId-1="figure@2.[140,617,1033,1534]" captionTargetId-2="figure@2.[647,1108,165,533]" captionTargetId-3="figure@2.[647,1107,563,1535]" captionTargetPageId-0="2" captionTargetPageId-1="2" captionTargetPageId-2="2" captionTargetPageId-3="2" captionText-0="Fig. 1. Dorsal habitus of Ischyomius escalonai sp. n., male paratype. TL = 11 mm." captionText-1="Fig. 2. Ischyomius escalonai sp. n., male, detail of forebody. Scale bar = 1 mm." captionText-2="Fig. 3. Ischyomius escalonai sp. n., male, detail of elytral apices. Scale bar = 1 mm." captionText-3="Fig. 4. Ischyomius escalonai sp. n., aedeagus, dorsal view. Scale bar = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3787150" figureDoi-1="http://doi.org/10.5281/zenodo.3787152" figureDoi-2="http://doi.org/10.5281/zenodo.3787154" figureDoi-3="http://doi.org/10.5281/zenodo.3787156" httpUri-0="https://zenodo.org/record/3787150/files/figure.png" httpUri-1="https://zenodo.org/record/3787152/files/figure.png" httpUri-2="https://zenodo.org/record/3787154/files/figure.png" httpUri-3="https://zenodo.org/record/3787156/files/figure.png" pageId="1" pageNumber="70">Figs. 1-4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="70" type="materials_examined">
<paragraph blockId="1.[140,1108,694,1671]" pageId="1" pageNumber="70">
<emphasis bold="true" box="[140,273,694,721]" pageId="1" pageNumber="70">
<emphasis box="[140,199,695,721]" inLineHeading="true" pageId="1" pageNumber="70" reason="1">Type</emphasis>
series
</emphasis>
.
<materialsCitation ID-GBIF-Occurrence="2610415301" collectingDate="2005-06-26" collectorName="P. N. Yacambu &amp; Hermes Escalona" country="Venezuela" elevation="1400" location="Lara" pageId="1" pageNumber="70" specimenCount="2" stateProvince="Lara" typeStatus="holotype">
<typeStatus box="[288,398,694,720]" pageId="1" pageNumber="70">Holotype</typeStatus>
male, labeled: “
<collectingCountry box="[580,759,695,721]" name="Venezuela" pageId="1" pageNumber="70">VENEZUELA</collectingCountry>
,
<collectingRegion box="[771,825,695,721]" country="Venezuela" name="Lara" pageId="1" pageNumber="70">Lara</collectingRegion>
,
<collectorName box="[836,1002,694,721]" pageId="1" pageNumber="70">P.N. Yacambú</collectorName>
, El Blanquito,
<quantity box="[217,308,730,756]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="70" unit="m" value="1400.0">
<elevation box="[217,308,730,756]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="70" unit="m" value="1400.0">1400 m</elevation>
</quantity>
<date box="[317,455,729,756]" pageId="1" pageNumber="70" value="2005-06-26">
<collectingDate box="[317,455,729,756]" pageId="1" pageNumber="70" value="2005-06-26">26.VI.2005</collectingDate>
</date>
;
<collectorName box="[468,671,730,756]" pageId="1" pageNumber="70">Hermes Escalona</collectorName>
, collector” / “coll. on bird of paradise (
<taxonomicName authorityName="Banks" authorityYear="1788" box="[150,251,765,791]" class="Liliopsida" family="Strelitziaceae" genus="Strelitzia" kingdom="Plantae" order="Zingiberales" pageId="1" pageNumber="70" phylum="Tracheophyta" rank="genus">
<emphasis box="[150,251,765,791]" italics="true" pageId="1" pageNumber="70">Strelitzia</emphasis>
</taxonomicName>
) dry leaves” / “
<typeStatus box="[439,598,765,791]" pageId="1" pageNumber="70">HOLOTYPE</typeStatus>
<taxonomicName authorityName="Chevrolat" authorityYear="1878" box="[609,740,765,791]" class="Insecta" family="Pythidae" genus="Ischyomius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="70" phylum="Arthropoda" rank="genus">Ischyomius</taxonomicName>
♁ escalonai,
<taxonomicNameLabel box="[906,971,765,791]" pageId="1" pageNumber="70" rank="species">sp. n.</taxonomicNameLabel>
D. A.
<bibRefCitation author="Pollock DA" pageId="1" pageNumber="70" pagination="277 - 280" refId="ref2182" refString="Pollock DA (2007) A new species of Ischyomius Chevrolat (Coleoptera: Pythidae) from Costa Rica. Coleopterists Bulletin 61 (2): 277 - 280." type="journal article" year="2007">Pollock 2007</bibRefCitation>
</materialsCitation>
.
<typeStatus box="[289,387,800,826]" pageId="1" pageNumber="70">Allotype</typeStatus>
female, same labels as
<typeStatus box="[665,767,800,826]" pageId="1" pageNumber="70">holotype</typeStatus>
(with “
<typeStatus box="[858,1006,800,826]" pageId="1" pageNumber="70">ALLOTYPE</typeStatus>
” instead of “
<typeStatus box="[181,343,835,861]" pageId="1" pageNumber="70">HOLOTYPE</typeStatus>
” on third label). Two
<typeStatus box="[602,714,836,862]" pageId="1" pageNumber="70">paratypes</typeStatus>
, one male, one female, with same labels as
<typeStatus box="[246,356,870,896]" pageId="1" pageNumber="70">Holotype</typeStatus>
(with “
<typeStatus box="[446,592,871,897]" pageId="1" pageNumber="70">PARATYPE</typeStatus>
” instead of “
<typeStatus box="[747,909,871,897]" pageId="1" pageNumber="70">HOLOTYPE</typeStatus>
” on third label).
<typeStatus box="[140,253,906,932]" pageId="1" pageNumber="70">Holotype</typeStatus>
,
<typeStatus box="[268,359,906,932]" pageId="1" pageNumber="70">allotype</typeStatus>
and female
<typeStatus box="[512,611,906,932]" pageId="1" pageNumber="70">paratype</typeStatus>
deposited in Museo del Instituto de Zoología Agrícola “Francisco Fernández Yepez”, UCV, Maracay, Venezuela (Luis J. Joly, curator). Male
<typeStatus box="[314,413,977,1003]" pageId="1" pageNumber="70">paratype</typeStatus>
deposited in collection of author.
</paragraph>
<paragraph blockId="1.[140,1108,694,1671]" pageId="1" pageNumber="70">
<emphasis bold="true" box="[185,537,1011,1037]" pageId="1" pageNumber="70">
<emphasis box="[185,315,1011,1037]" inLineHeading="true" pageId="1" pageNumber="70" reason="1">Derivation</emphasis>
of specific epithet
</emphasis>
. I am very pleased to name this new species after friend and colleague Hermes Escalona, a fellow specialist on the “salpingid group” of tenebrionoid families, and who also collected all known specimens of this species.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="70" type="diagnosis">
<paragraph blockId="1.[140,1108,694,1671]" pageId="1" pageNumber="70">
<emphasis bold="true" box="[185,304,1117,1143]" inLineHeading="true" pageId="1" pageNumber="70" reason="1">Diagnosis</emphasis>
. This new species of
<taxonomicName authorityName="Chevrolat" authorityYear="1878" box="[551,666,1117,1143]" class="Insecta" family="Pythidae" genus="Ischyomius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis box="[551,666,1117,1143]" italics="true" pageId="1" pageNumber="70">Ischyomius</emphasis>
</taxonomicName>
may be separated from the rest of the genus by the following combination of characters: posterior pronotal bead indistinct; lateral pronotal bead not attaining anterior bead; color uniformly light to medium brown; abdominal ventrites with short but obvious setation; elytral apices rounded, without spine (
<figureCitation box="[314,382,1258,1284]" captionStart="Fig" captionStartId="2.[140,174,1622,1644]" captionTargetBox="[648,1108,165,533]" captionTargetId="figure@2.[647,1108,165,533]" captionTargetPageId="2" captionText="Fig. 3. Ischyomius escalonai sp. n., male, detail of elytral apices. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3787154" httpUri="https://zenodo.org/record/3787154/files/figure.png" pageId="1" pageNumber="70">Fig. 3</figureCitation>
) aedeagus relatively short and stout, accessory lobes very short, inserted midlength on the tegmen (
<figureCitation box="[540,609,1293,1319]" captionStart="Fig" captionStartId="2.[140,174,1654,1676]" captionTargetBox="[648,1107,582,1534]" captionTargetId="figure@2.[647,1107,563,1535]" captionTargetPageId="2" captionText="Fig. 4. Ischyomius escalonai sp. n., aedeagus, dorsal view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3787156" httpUri="https://zenodo.org/record/3787156/files/figure.png" pageId="1" pageNumber="70">Fig. 4</figureCitation>
); known only from
<collectingCountry box="[835,952,1293,1319]" name="Venezuela" pageId="1" pageNumber="70">Venezuela</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="72" pageId="1" pageNumber="70" type="description">
<paragraph blockId="1.[140,1108,694,1671]" lastBlockId="3.[140,1108,166,1214]" lastPageId="3" lastPageNumber="72" pageId="1" pageNumber="70">
<emphasis bold="true" box="[185,332,1328,1354]" inLineHeading="true" pageId="1" pageNumber="70" reason="1">Description</emphasis>
. With generic characteristics of
<taxonomicName authorityName="Chevrolat" authorityYear="1878" box="[707,827,1328,1354]" class="Insecta" family="Pythidae" genus="Ischyomius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis box="[707,827,1328,1354]" italics="true" pageId="1" pageNumber="70">Ischyomius</emphasis>
</taxonomicName>
(see
<bibRefCitation author="Pollock DA" pageId="1" pageNumber="70" pagination="245 - 264" refId="ref2141" refString="Pollock DA (1998) Revision of the Neotropical genus Ischyomius with a discussion on its systematic position (Insecta: Coleoptera: Tenebrionoidea: Pythidae). Annals of Carnegie Museum 67 (3): 245 - 264." type="journal article" year="1998">Pollock 1998: 246- 250</bibRefCitation>
), plus the following: total length (TL)
<quantity box="[675,838,1363,1390]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" metricValueMax="1.15" metricValueMin="0.99" pageId="1" pageNumber="70" unit="mm" value="10.7" valueMax="11.5" valueMin="9.9">9.9-11.5 mm</quantity>
; greatest elytral width (GEW)
<quantity box="[235,378,1399,1425]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.2" metricValueMin="2.8" pageId="1" pageNumber="70" unit="mm" value="3.0" valueMax="3.2" valueMin="2.8">2.8-3.2 mm</quantity>
; body (
<figureCitation box="[464,530,1398,1425]" captionStart="Fig" captionStartId="2.[140,174,1558,1580]" captionTargetBox="[140,617,165,1003]" captionTargetId="figure@2.[140,618,165,1003]" captionTargetPageId="2" captionText="Fig. 1. Dorsal habitus of Ischyomius escalonai sp. n., male paratype. TL = 11 mm." figureDoi="http://doi.org/10.5281/zenodo.3787150" httpUri="https://zenodo.org/record/3787150/files/figure.png" pageId="1" pageNumber="70">Fig. 1</figureCitation>
) elongate (TL / GEW = 3.5-3.8) color uniformly light to medium brown dorsally, without obvious color contrasts, ventrally, thorax and abdominal ventrites slightly darker than legs and elytral epipleuron; eyes moderately protuberant; antennae relatively long, antennomeres filiform, slightly widened from antennomere 5; male lacking pit on mentum; entire dorsal surface of body with shallow, sparse punctation, without any obvious microsculpture; pronotum wider than long (GPW / PL = 1.19-1.21); posterior pronotal bead indistinct; lateral pronotal carinae distinct, smooth, without tubercles, prolonged anteriorly to, and in contact with, anterior pronotal bead; lateral pronotal margins parallelsided basally, then relatively abruptly widened, greatest width anterior of midlength (
<figureCitation box="[148,218,237,263]" captionStart="Fig" captionStartId="2.[140,174,1590,1612]" captionTargetBox="[140,617,1033,1533]" captionTargetId="figure@2.[140,617,1033,1534]" captionTargetPageId="2" captionText="Fig. 2. Ischyomius escalonai sp. n., male, detail of forebody. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3787152" httpUri="https://zenodo.org/record/3787152/files/figure.png" pageId="3" pageNumber="72">Fig. 2</figureCitation>
); anterolateral angles of pronotum rounded, not angulate; disc of pronotum moderately convex with indistinct posterolateral depressions; elytra convex, slightly flattened dorsally, without distinct setation; outer apical elytral spine absent (
<figureCitation captionStart="Fig" captionStartId="2.[140,174,1622,1644]" captionTargetBox="[648,1108,165,533]" captionTargetId="figure@2.[647,1108,165,533]" captionTargetPageId="2" captionText="Fig. 3. Ischyomius escalonai sp. n., male, detail of elytral apices. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3787154" httpUri="https://zenodo.org/record/3787154/files/figure.png" pageId="3" pageNumber="72">Fig. 3</figureCitation>
); thoracic sterna and abdominal ventrites with distinct, almost isodiametric microsculpture; abdominal ventrites relatively coarsely punctate, with short but evident setation; aedeagus (
<figureCitation box="[433,505,413,439]" captionStart="Fig" captionStartId="2.[140,174,1654,1676]" captionTargetBox="[648,1107,582,1534]" captionTargetId="figure@2.[647,1107,563,1535]" captionTargetPageId="2" captionText="Fig. 4. Ischyomius escalonai sp. n., aedeagus, dorsal view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3787156" httpUri="https://zenodo.org/record/3787156/files/figure.png" pageId="3" pageNumber="72">Fig. 4</figureCitation>
) stout, apicale slightly shorter than basale; apicale broad, tapered distally; accessory lobes narrow, very short, not extended to end of apicale, inserted about midlength of apicale.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3787150" ID-Zenodo-Dep="3787150" box="[140,891,1558,1580]" httpUri="https://zenodo.org/record/3787150/files/figure.png" pageId="2" pageNumber="71" startId="2.[140,174,1558,1580]" targetBox="[140,617,165,1003]" targetPageId="2">
<paragraph blockId="2.[140,891,1558,1580]" box="[140,891,1558,1580]" pageId="2" pageNumber="71">
<emphasis bold="true" box="[140,204,1558,1580]" pageId="2" pageNumber="71">Fig. 1.</emphasis>
Dorsal habitus of
<taxonomicName authority="Pollock, 2009" authorityName="Pollock" authorityYear="2009" box="[376,554,1558,1579]" class="Insecta" family="Pythidae" genus="Ischyomius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="71" phylum="Arthropoda" rank="species" species="escalonai" status="sp. n.">
<emphasis box="[376,554,1558,1579]" italics="true" pageId="2" pageNumber="71">Ischyomius escalonai</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[561,612,1559,1580]" pageId="2" pageNumber="71" rank="species">sp. n.</taxonomicNameLabel>
, male
<typeStatus box="[673,755,1559,1580]" pageId="2" pageNumber="71">paratype</typeStatus>
. TL =
<quantity box="[816,887,1558,1580]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="71" unit="mm" value="11.0">11 mm</quantity>
.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3787152" ID-Zenodo-Dep="3787152" box="[140,856,1590,1612]" httpUri="https://zenodo.org/record/3787152/files/figure.png" pageId="2" pageNumber="71" startId="2.[140,174,1590,1612]" targetBox="[140,617,1033,1533]" targetPageId="2">
<paragraph blockId="2.[140,856,1590,1612]" box="[140,856,1590,1612]" pageId="2" pageNumber="71">
<emphasis bold="true" box="[140,205,1590,1612]" pageId="2" pageNumber="71">Fig. 2.</emphasis>
<taxonomicName authority="Pollock, 2009" authorityName="Pollock" authorityYear="2009" box="[210,389,1590,1611]" class="Insecta" family="Pythidae" genus="Ischyomius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="71" phylum="Arthropoda" rank="species" species="escalonai" status="sp. n.">
<emphasis box="[210,389,1590,1611]" italics="true" pageId="2" pageNumber="71">Ischyomius escalonai</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[395,447,1591,1612]" pageId="2" pageNumber="71" rank="species">sp. n.</taxonomicNameLabel>
, male, detail of forebody. Scale bar =
<quantity box="[793,852,1590,1612]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="71" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3787154" ID-Zenodo-Dep="3787154" box="[140,893,1622,1644]" httpUri="https://zenodo.org/record/3787154/files/figure.png" pageId="2" pageNumber="71" startId="2.[140,174,1622,1644]" targetBox="[648,1108,165,533]" targetPageId="2">
<paragraph blockId="2.[140,893,1622,1644]" box="[140,893,1622,1644]" pageId="2" pageNumber="71">
<emphasis bold="true" box="[140,205,1622,1644]" pageId="2" pageNumber="71">Fig. 3.</emphasis>
<taxonomicName authority="Pollock, 2009" authorityName="Pollock" authorityYear="2009" box="[210,389,1622,1643]" class="Insecta" family="Pythidae" genus="Ischyomius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="71" phylum="Arthropoda" rank="species" species="escalonai" status="sp. n.">
<emphasis box="[210,389,1622,1643]" italics="true" pageId="2" pageNumber="71">Ischyomius escalonai</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[395,447,1623,1644]" pageId="2" pageNumber="71" rank="species">sp. n.</taxonomicNameLabel>
, male, detail of elytral apices. Scale bar =
<quantity box="[831,890,1622,1644]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="71" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3787156" ID-Zenodo-Dep="3787156" box="[140,851,1654,1676]" httpUri="https://zenodo.org/record/3787156/files/figure.png" pageId="2" pageNumber="71" startId="2.[140,174,1654,1676]" targetBox="[648,1107,582,1534]" targetPageId="2">
<paragraph blockId="2.[140,851,1654,1676]" box="[140,851,1654,1676]" pageId="2" pageNumber="71">
<emphasis bold="true" box="[140,205,1654,1676]" pageId="2" pageNumber="71">Fig. 4.</emphasis>
<taxonomicName authority="Pollock, 2009" authorityName="Pollock" authorityYear="2009" box="[210,389,1654,1675]" class="Insecta" family="Pythidae" genus="Ischyomius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="71" phylum="Arthropoda" rank="species" species="escalonai" status="sp. n.">
<emphasis box="[210,389,1654,1675]" italics="true" pageId="2" pageNumber="71">Ischyomius escalonai</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[395,447,1655,1676]" pageId="2" pageNumber="71" rank="species">sp. n.</taxonomicNameLabel>
, aedeagus, dorsal view. Scale bar =
<quantity box="[770,847,1654,1676]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="71" unit="mm" value="0.5">0.5 mm</quantity>
.
</paragraph>
</caption>
<subSubSection pageId="3" pageNumber="72" type="distribution">
<paragraph blockId="3.[140,1108,166,1214]" pageId="3" pageNumber="72">
<emphasis bold="true" box="[185,336,518,544]" inLineHeading="true" pageId="3" pageNumber="72" reason="1">Distribution</emphasis>
. The four
<typeStatus box="[466,514,519,545]" pageId="3" pageNumber="72">type</typeStatus>
specimens were collected from a single locality, in Yacambú National Park,
<collectingRegion box="[422,537,554,580]" country="Venezuela" name="Lara" pageId="3" pageNumber="72">Lara State</collectingRegion>
, in easternmost
<collectingCountry box="[720,837,554,580]" name="Venezuela" pageId="3" pageNumber="72">Venezuela</collectingCountry>
.
</paragraph>
<paragraph blockId="3.[140,1108,166,1214]" pageId="3" pageNumber="72">
<emphasis bold="true" box="[185,372,589,615]" pageId="3" pageNumber="72">
<emphasis box="[185,279,589,615]" inLineHeading="true" pageId="3" pageNumber="72" reason="1">Natural</emphasis>
history
</emphasis>
. All four specimens were collected by beating withered leaves of “Bird of Paradise” plants (
<taxonomicName authorityName="Banks" authorityYear="1788" box="[442,543,624,650]" class="Liliopsida" family="Strelitziaceae" genus="Strelitzia" kingdom="Plantae" order="Zingiberales" pageId="3" pageNumber="72" phylum="Tracheophyta" rank="genus">
<emphasis box="[442,543,624,650]" italics="true" pageId="3" pageNumber="72">Strelitzia</emphasis>
</taxonomicName>
sp.) (Hermes Escalona, pers. comm.). Very little is known about any other species of
<taxonomicName authorityName="Chevrolat" authorityYear="1878" box="[561,679,659,685]" class="Insecta" family="Pythidae" genus="Ischyomius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis box="[561,679,659,685]" italics="true" pageId="3" pageNumber="72">Ischyomius</emphasis>
</taxonomicName>
, although they seem to be associated with dead or dying vegetation of
<taxonomicName authorityName="Durande" authorityYear="1782" box="[546,657,695,721]" class="Liliopsida" family="Musaceae" kingdom="Plantae" order="Zingiberales" pageId="3" pageNumber="72" phylum="Tracheophyta" rank="family">Musaceae</taxonomicName>
or Palmae (
<bibRefCitation author="Champion GC" box="[806,1007,694,720]" pageId="3" pageNumber="72" pagination="75 - 83" refId="ref2054" refString="Champion GC (1916) Notes on Melandryidae (3). Entomologist's Monthly Magazine (Ser. 3) 2: 75 - 83." type="journal article" year="1916">Champion 1916</bibRefCitation>
;
<bibRefCitation author="Pollock DA" pageId="3" pageNumber="72" pagination="245 - 264" refId="ref2141" refString="Pollock DA (1998) Revision of the Neotropical genus Ischyomius with a discussion on its systematic position (Insecta: Coleoptera: Tenebrionoidea: Pythidae). Annals of Carnegie Museum 67 (3): 245 - 264." type="journal article" year="1998">Pollock 1998</bibRefCitation>
). Although the genus
<taxonomicName authorityName="Banks" authorityYear="1788" box="[473,574,730,756]" class="Liliopsida" family="Strelitziaceae" genus="Strelitzia" kingdom="Plantae" order="Zingiberales" pageId="3" pageNumber="72" phylum="Tracheophyta" rank="genus">
<emphasis box="[473,574,730,756]" italics="true" pageId="3" pageNumber="72">Strelitzia</emphasis>
</taxonomicName>
is native to
<collectingCountry box="[730,883,730,756]" name="South Africa" pageId="3" pageNumber="72">South Africa</collectingCountry>
, and therefore was introduced to the Venezuelan
<typeStatus box="[487,537,765,791]" pageId="3" pageNumber="72">type</typeStatus>
locality, the family
<taxonomicName authorityName="Hutchinson" authorityYear="1934" baseAuthorityName="K.Schum." box="[765,914,765,791]" class="Liliopsida" family="Strelitziaceae" kingdom="Plantae" order="Zingiberales" pageId="3" pageNumber="72" phylum="Tracheophyta" rank="family">Strelitziaceae</taxonomicName>
is closely related to
<taxonomicName authorityName="Durande" authorityYear="1782" box="[170,281,801,827]" class="Liliopsida" family="Musaceae" kingdom="Plantae" order="Zingiberales" pageId="3" pageNumber="72" phylum="Tracheophyta" rank="family">Musaceae</taxonomicName>
(which are also introduced), in the “banana and ginger” group of plants (
<bibRefCitation author="Kress WJ &amp; Hahn WJ" box="[147,408,835,861]" pageId="3" pageNumber="72" refId="ref2098" refString="Kress WJ, Hahn WJ (1997) Zingiberanae. The bananas, gingers, and relatives. Version 01 January 1997. http: // tolweb. org / Zingiberanae / 21339 / 1997.01.01 (accessed 19 December, 2008)." type="url" year="1997">Kress and Hahn 1997</bibRefCitation>
). It seems probable that
<taxonomicName authorityName="Pollock" authorityYear="2009" box="[694,817,835,861]" class="Insecta" family="Pythidae" genus="Ischyomius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="72" phylum="Arthropoda" rank="species" species="escalonai">
<emphasis box="[694,817,835,861]" italics="true" pageId="3" pageNumber="72">I. escalonai</emphasis>
</taxonomicName>
will be found in association with
<taxonomicName authorityName="Durande" authorityYear="1782" box="[255,370,871,897]" class="Liliopsida" family="Musaceae" kingdom="Plantae" order="Zingiberales" pageId="3" pageNumber="72" phylum="Tracheophyta" rank="family">Musaceae</taxonomicName>
, also.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="72" type="discussion">
<paragraph blockId="3.[140,1108,166,1214]" pageId="3" pageNumber="72">
<emphasis bold="true" box="[185,317,905,931]" inLineHeading="true" pageId="3" pageNumber="72" reason="1">Discussion</emphasis>
. The description of
<taxonomicName authorityName="Pollock" authorityYear="2009" box="[555,679,906,932]" class="Insecta" family="Pythidae" genus="Ischyomius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="72" phylum="Arthropoda" rank="species" species="escalonai">
<emphasis box="[555,565,906,931]" italics="true" pageId="3" pageNumber="72">I</emphasis>
.
<emphasis box="[582,679,906,932]" italics="true" pageId="3" pageNumber="72">escalonai</emphasis>
</taxonomicName>
brings the total known species in
<taxonomicName authorityName="Chevrolat" authorityYear="1878" class="Insecta" family="Pythidae" genus="Ischyomius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="3" pageNumber="72">Ischyomius</emphasis>
</taxonomicName>
to eight. The first couplet of
<bibRefCitation author="Pollock DA" box="[590,775,941,967]" pageId="3" pageNumber="72" pagination="245 - 264" refId="ref2141" refString="Pollock DA (1998) Revision of the Neotropical genus Ischyomius with a discussion on its systematic position (Insecta: Coleoptera: Tenebrionoidea: Pythidae). Annals of Carnegie Museum 67 (3): 245 - 264." type="journal article" year="1998">Pollocks (1998)</bibRefCitation>
key divided the species into two groups: those that have a distinct, posterior pronotal bead and distinct vestiture on abdominal ventrites and those without posterior bead and lacking ventral abdominal vestiture. Each of the four
<typeStatus box="[484,533,1047,1073]" pageId="3" pageNumber="72">type</typeStatus>
specimens of
<taxonomicName authorityName="Pollock" authorityYear="2009" box="[693,815,1046,1073]" class="Insecta" family="Pythidae" genus="Ischyomius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="72" phylum="Arthropoda" rank="species" species="escalonai">
<emphasis box="[693,703,1047,1072]" italics="true" pageId="3" pageNumber="72">I</emphasis>
.
<emphasis box="[718,815,1046,1072]" italics="true" pageId="3" pageNumber="72">escalonai</emphasis>
</taxonomicName>
are intermediate between these two groups,
<emphasis box="[349,357,1082,1107]" italics="true" pageId="3" pageNumber="72">i</emphasis>
.
<emphasis box="[365,375,1082,1107]" italics="true" pageId="3" pageNumber="72">e</emphasis>
. without the pronotal bead, but with short, but distinct ventral abdominal vestiture. Therefore, the original key of
<bibRefCitation author="Pollock DA" box="[718,891,1117,1143]" pageId="3" pageNumber="72" pagination="245 - 264" refId="ref2141" refString="Pollock DA (1998) Revision of the Neotropical genus Ischyomius with a discussion on its systematic position (Insecta: Coleoptera: Tenebrionoidea: Pythidae). Annals of Carnegie Museum 67 (3): 245 - 264." type="journal article" year="1998">Pollock (1998)</bibRefCitation>
, revised by
<bibRefCitation author="Pollock DA" pageId="3" pageNumber="72" pagination="277 - 280" refId="ref2182" refString="Pollock DA (2007) A new species of Ischyomius Chevrolat (Coleoptera: Pythidae) from Costa Rica. Coleopterists Bulletin 61 (2): 277 - 280." type="journal article" year="2007">Pollock (2007)</bibRefCitation>
for inclusion of
<taxonomicName box="[410,513,1152,1179]" class="Insecta" family="Pythidae" genus="Ischyomius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="72" phylum="Arthropoda" rank="species" species="hovorei">
<emphasis box="[410,420,1153,1178]" italics="true" pageId="3" pageNumber="72">I</emphasis>
.
<emphasis box="[435,513,1152,1178]" italics="true" pageId="3" pageNumber="72">hovorei</emphasis>
</taxonomicName>
, must be amended, as follows. References to figures for species other than
<taxonomicName authorityName="Pollock" authorityYear="2009" box="[392,515,1187,1214]" class="Insecta" family="Pythidae" genus="Ischyomius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="72" phylum="Arthropoda" rank="species" species="escalonai">
<emphasis box="[392,402,1188,1213]" italics="true" pageId="3" pageNumber="72">I</emphasis>
.
<emphasis box="[418,515,1187,1213]" italics="true" pageId="3" pageNumber="72">escalonai</emphasis>
</taxonomicName>
refer to figures in
<bibRefCitation author="Pollock DA" box="[725,899,1187,1213]" pageId="3" pageNumber="72" pagination="245 - 264" refId="ref2141" refString="Pollock DA (1998) Revision of the Neotropical genus Ischyomius with a discussion on its systematic position (Insecta: Coleoptera: Tenebrionoidea: Pythidae). Annals of Carnegie Museum 67 (3): 245 - 264." type="journal article" year="1998">Pollock (1998)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="73" pageId="3" pageNumber="72" type="key">
<key lastPageId="4" lastPageNumber="73" pageId="3" pageNumber="72">
<keyStep pageId="3" pageNumber="72">
<paragraph blockId="3.[140,1107,1258,1355]" pageId="3" pageNumber="72">
<keyLead pageId="3" pageNumber="72">
1 Pronotum with distinct bead along entire posterior margin (in most speci-mens)..........................................................................................................
<emphasis bold="true" box="[1092,1107,1293,1319]" pageId="3" pageNumber="72">2</emphasis>
</keyLead>
</paragraph>
<paragraph blockId="3.[140,1107,1258,1355]" box="[140,1107,1328,1355]" pageId="3" pageNumber="72">
<keyLead box="[140,1107,1328,1355]" pageId="3" pageNumber="72">
1' Pronotum without posterior bead, or present laterally only.........................
<emphasis bold="true" box="[1092,1107,1328,1354]" pageId="3" pageNumber="72">4</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="3" pageNumber="72">
<paragraph blockId="3.[140,1108,1398,1531]" pageId="3" pageNumber="72">
<keyLead pageId="3" pageNumber="72">
2(1) Antennomeres filiform; elytral apex with conspicuous, outer spine (
<figureCitation box="[986,1045,1398,1425]" captionStart="Fig" captionStartId="2.[140,174,1558,1580]" captionTargetBox="[140,617,165,1003]" captionTargetId="figure@2.[140,618,165,1003]" captionTargetPageId="2" captionText="Fig. 1. Dorsal habitus of Ischyomius escalonai sp. n., male paratype. TL = 11 mm." figureDoi="http://doi.org/10.5281/zenodo.3787150" httpUri="https://zenodo.org/record/3787150/files/figure.png" pageId="3" pageNumber="72">Fig. 1</figureCitation>
A-D, F)................................................................................................................
<emphasis bold="true" box="[1092,1107,1433,1459]" pageId="3" pageNumber="72">3</emphasis>
</keyLead>
</paragraph>
<paragraph blockId="3.[140,1108,1398,1531]" pageId="3" pageNumber="72">
<keyLead pageId="3" pageNumber="72">
2' Antennomeres short, subserrate; elytral apex without conspicuous, outer spine (
<figureCitation box="[320,402,1504,1530]" captionStart="Fig" captionStartId="2.[140,174,1558,1580]" captionTargetBox="[140,617,165,1003]" captionTargetId="figure@2.[140,618,165,1003]" captionTargetPageId="2" captionText="Fig. 1. Dorsal habitus of Ischyomius escalonai sp. n., male paratype. TL = 11 mm." figureDoi="http://doi.org/10.5281/zenodo.3787150" httpUri="https://zenodo.org/record/3787150/files/figure.png" pageId="3" pageNumber="72">Fig. 1E</figureCitation>
) .............................................................
<taxonomicName authority="Champion" authorityName="Champion" box="[865,1107,1503,1530]" class="Insecta" family="Pythidae" genus="Ischyomius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="72" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis bold="true" box="[865,1107,1503,1530]" pageId="3" pageNumber="72">
<emphasis bold="true" box="[865,972,1503,1530]" italics="true" pageId="3" pageNumber="72">I. bicolor</emphasis>
Champion
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep lastPageId="4" lastPageNumber="73" pageId="3" pageNumber="72">
<paragraph blockId="3.[140,1108,1574,1636]" lastBlockId="4.[140,1108,166,369]" lastPageId="4" lastPageNumber="73" pageId="3" pageNumber="72">
<keyLead lastPageId="4" lastPageNumber="73" pageId="3" pageNumber="72">
3(2) Antennomeres 2-10 piceous to black, contrasting in color to antennomeres 1 and 11; lateral margins of pronotum with several long setae; body testa-ceous, with median, longitudinal dark vitta extended from frons to elytral apex (
<figureCitation box="[312,392,202,228]" captionStart="Fig" captionStartId="2.[140,174,1558,1580]" captionTargetBox="[140,617,165,1003]" captionTargetId="figure@2.[140,618,165,1003]" captionTargetPageId="2" captionText="Fig. 1. Dorsal habitus of Ischyomius escalonai sp. n., male paratype. TL = 11 mm." figureDoi="http://doi.org/10.5281/zenodo.3787150" httpUri="https://zenodo.org/record/3787150/files/figure.png" pageId="4" pageNumber="73">Fig. 1D</figureCitation>
).............................................................
<taxonomicName authority="Pollock" authorityName="Pollock" box="[859,1107,201,228]" class="Insecta" family="Pythidae" genus="Ischyomius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="championi">
<emphasis bold="true" box="[859,1107,201,228]" pageId="4" pageNumber="73">
<emphasis bold="true" box="[859,1010,201,228]" italics="true" pageId="4" pageNumber="73">I. championi</emphasis>
Pollock
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="4.[140,1108,166,369]" pageId="4" pageNumber="73">
<keyLead pageId="4" pageNumber="73">
3' Antennomeres 1-11 concolorous, rufous; lateral margins of pronotum without long setae; body testaceous to rufous with elytral infuscation, around scutellum and rectangular to diamond-shaped, transverse dark area slightly posterior of elytral midlength (
<figureCitation box="[592,674,342,369]" captionStart="Fig" captionStartId="2.[140,174,1558,1580]" captionTargetBox="[140,617,165,1003]" captionTargetId="figure@2.[140,618,165,1003]" captionTargetPageId="2" captionText="Fig. 1. Dorsal habitus of Ischyomius escalonai sp. n., male paratype. TL = 11 mm." figureDoi="http://doi.org/10.5281/zenodo.3787150" httpUri="https://zenodo.org/record/3787150/files/figure.png" pageId="4" pageNumber="73">Fig. 1F</figureCitation>
) .....................
<taxonomicName authority="Pollock" authorityName="Pollock" box="[842,1107,342,369]" class="Insecta" family="Pythidae" genus="Ischyomius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="nevermanni">
<emphasis bold="true" box="[842,1107,342,369]" pageId="4" pageNumber="73">
<emphasis bold="true" box="[842,1010,342,369]" italics="true" pageId="4" pageNumber="73">I. nevermanni</emphasis>
Pollock
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="4" pageNumber="73">
<paragraph blockId="4.[140,1107,413,510]" pageId="4" pageNumber="73">
<keyLead pageId="4" pageNumber="73">
4(1') Anterolateral angles of pronotum produced, square to slightly acute (
<figureCitation box="[254,339,448,474]" captionStart="Fig" captionStartId="2.[140,174,1558,1580]" captionTargetBox="[140,617,165,1003]" captionTargetId="figure@2.[140,618,165,1003]" captionTargetPageId="2" captionText="Fig. 1. Dorsal habitus of Ischyomius escalonai sp. n., male paratype. TL = 11 mm." figureDoi="http://doi.org/10.5281/zenodo.3787150" httpUri="https://zenodo.org/record/3787150/files/figure.png" pageId="4" pageNumber="73">Fig. 1C</figureCitation>
)................................................................
<taxonomicName authority="Champion" authorityName="Champion" box="[826,1107,447,474]" class="Insecta" family="Pythidae" genus="Ischyomius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="denticollis">
<emphasis bold="true" box="[826,1107,447,474]" pageId="4" pageNumber="73">
<emphasis bold="true" box="[826,972,447,474]" italics="true" pageId="4" pageNumber="73">I. denticollis</emphasis>
Champion
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="4.[140,1107,413,510]" box="[140,1107,483,510]" pageId="4" pageNumber="73">
<keyLead box="[140,1107,483,510]" pageId="4" pageNumber="73">
4' Anterolateral angles of pronotum not produced, more or less rounded .......
<emphasis bold="true" box="[1092,1107,483,509]" pageId="4" pageNumber="73">5</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="4" pageNumber="73">
<paragraph blockId="4.[140,1107,553,651]" pageId="4" pageNumber="73">
<keyLead pageId="4" pageNumber="73">
5(4') Lateral pronotal bead attaining anterior pronotal margin, continuous with anterior bead (
<figureCitation box="[410,495,589,615]" captionStart="Fig" captionStartId="2.[140,174,1654,1676]" captionTargetBox="[648,1107,582,1534]" captionTargetId="figure@2.[647,1107,563,1535]" captionTargetPageId="2" captionText="Fig. 4. Ischyomius escalonai sp. n., aedeagus, dorsal view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3787156" httpUri="https://zenodo.org/record/3787156/files/figure.png" pageId="4" pageNumber="73">Fig. 4B</figureCitation>
)................................................................................
<emphasis bold="true" box="[1092,1107,589,615]" pageId="4" pageNumber="73">6</emphasis>
</keyLead>
</paragraph>
<paragraph blockId="4.[140,1107,553,651]" box="[140,1107,624,651]" pageId="4" pageNumber="73">
<keyLead box="[140,1107,624,651]" pageId="4" pageNumber="73">
5' Lateral pronotal bead not attaining anterior pronotal margin (
<figureCitation box="[943,1033,624,650]" captionStart="Fig" captionStartId="2.[140,174,1654,1676]" captionTargetBox="[648,1107,582,1534]" captionTargetId="figure@2.[647,1107,563,1535]" captionTargetPageId="2" captionText="Fig. 4. Ischyomius escalonai sp. n., aedeagus, dorsal view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3787156" httpUri="https://zenodo.org/record/3787156/files/figure.png" pageId="4" pageNumber="73">Fig. 4D</figureCitation>
) ......
<emphasis bold="true" box="[1092,1107,624,650]" pageId="4" pageNumber="73">7</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="4" pageNumber="73">
<paragraph blockId="4.[140,1108,694,967]" pageId="4" pageNumber="73">
<keyLead pageId="4" pageNumber="73">
6(5) Apex of elytron with distinct outer spine (
<figureCitation box="[712,797,694,721]" captionStart="Fig" captionStartId="2.[140,174,1558,1580]" captionTargetBox="[140,617,165,1003]" captionTargetId="figure@2.[140,618,165,1003]" captionTargetPageId="2" captionText="Fig. 1. Dorsal habitus of Ischyomius escalonai sp. n., male paratype. TL = 11 mm." figureDoi="http://doi.org/10.5281/zenodo.3787150" httpUri="https://zenodo.org/record/3787150/files/figure.png" pageId="4" pageNumber="73">Fig. 1A</figureCitation>
); abdominal ventrites without obvious setation (setae, if present, hidden within punctation); accessory lobes of aedeagus long, greater than half length of apicale, extended to apex of apicale (Fig. 7D)...................................................
<taxonomicName authority="Chevrolat" authorityName="Chevrolat" box="[841,1107,799,826]" class="Insecta" family="Pythidae" genus="Ischyomius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="singularis">
<emphasis bold="true" box="[841,1107,799,826]" pageId="4" pageNumber="73">
<emphasis bold="true" box="[841,982,799,826]" italics="true" pageId="4" pageNumber="73">I. singularis</emphasis>
Chevrolat
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="4.[140,1108,694,967]" pageId="4" pageNumber="73">
<keyLead pageId="4" pageNumber="73">
6' Apex of elytron smooth, without outer spine (
<figureCitation box="[785,855,835,861]" captionStart="Fig" captionStartId="2.[140,174,1622,1644]" captionTargetBox="[648,1108,165,533]" captionTargetId="figure@2.[647,1108,165,533]" captionTargetPageId="2" captionText="Fig. 3. Ischyomius escalonai sp. n., male, detail of elytral apices. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3787154" httpUri="https://zenodo.org/record/3787154/files/figure.png" pageId="4" pageNumber="73">Fig. 3</figureCitation>
); abdominal ventrites with short but obvious setation; accessory lobes of aedeagus short, distinctly less than half length of apicale, not extended to apex of apicale (
<figureCitation box="[964,1030,906,932]" captionStart="Fig" captionStartId="2.[140,174,1654,1676]" captionTargetBox="[648,1107,582,1534]" captionTargetId="figure@2.[647,1107,563,1535]" captionTargetPageId="2" captionText="Fig. 4. Ischyomius escalonai sp. n., aedeagus, dorsal view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3787156" httpUri="https://zenodo.org/record/3787156/files/figure.png" pageId="4" pageNumber="73">Fig. 4</figureCitation>
) ........ ...........................................................................
<emphasis bold="true" box="[797,1107,940,967]" pageId="4" pageNumber="73">
<taxonomicName authority="Pollock" authorityName="Pollock" authorityYear="2009" box="[797,1030,940,967]" class="Insecta" family="Pythidae" genus="Ischyomius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="escalonai" status="sp. n.">
<emphasis bold="true" box="[797,931,940,967]" italics="true" pageId="4" pageNumber="73">I. escalonai</emphasis>
Pollock
</taxonomicName>
,
<taxonomicNameLabel box="[1043,1107,941,967]" pageId="4" pageNumber="73" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="4" pageNumber="73">
<paragraph blockId="4.[140,1107,1011,1108]" box="[140,1107,1011,1038]" pageId="4" pageNumber="73">
<keyLead box="[140,1107,1011,1038]" pageId="4" pageNumber="73">
7(5') Body color uniform, from testaceous to rufous .......
<taxonomicName authority="Champion" authorityName="Champion" box="[829,1107,1011,1038]" class="Insecta" family="Pythidae" genus="Ischyomius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="chevrolati">
<emphasis bold="true" box="[829,1107,1011,1038]" pageId="4" pageNumber="73">
<emphasis bold="true" box="[829,973,1011,1038]" italics="true" pageId="4" pageNumber="73">I. chevrolati</emphasis>
Champion
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="4.[140,1107,1011,1108]" pageId="4" pageNumber="73">
<keyLead pageId="4" pageNumber="73">
7' Body color orange-red except for elytral apices, femora and tibiae piceous to nearly black......................................................................
<taxonomicName authority="Pollock" authorityName="Pollock" box="[898,1107,1081,1108]" class="Insecta" family="Pythidae" genus="Ischyomius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="hovorei">
<emphasis bold="true" box="[898,1107,1081,1108]" pageId="4" pageNumber="73">
<emphasis bold="true" box="[898,1010,1081,1108]" italics="true" pageId="4" pageNumber="73">I. hovorei</emphasis>
Pollock
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>