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<document id="B9537F1C094DD6D3ACB1C9077D98438D" ID-CLB-Dataset="5285" ID-DOI="10.11646/zootaxa.3835.4.3" ID-GBIF-Dataset="dbf8de20-9c7a-4b66-9130-615b6b42e84e" ID-ISSN="1175-5326" ID-Zenodo-Dep="4921623" ID-ZooBank="AD845CF3-CB19-4924-891F-330AAE283D07" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623283883149" checkinUser="felipe" docAuthor="Yoshizawa, Kazunori, Lienhard, Charles &amp; Abd, Idris" docDate="2014" docId="039F5A5FFFE8FFE2FF20FAABFDDC3DD6" docLanguage="en" docName="zt03835p500.pdf" docOrigin="Zootaxa 3835 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trichadenotecnum anomalum Yoshizawa &amp; Lienhard &amp; Abd 2014, sp.n." docType="treatment" docVersion="7" lastPageNumber="493" masterDocId="FFA62227FFFFFFFAFFB7FF9FFF893E1E" masterDocTitle="Trichadenotecnum species from Peninsular Malaysia and Singapore (Insecta: Psocodea: ' Psocoptera': Psocidae)" masterLastPageNumber="500" masterPageNumber="469" pageNumber="492" updateTime="1699009618324" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="1DE9A7D70721050622FB5A9366D4BCAF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D9815448FCA1C644F8C8DBF6B9FEFE25">
<mods:title id="DD11F5B732DD65A927E32BF48DB431CD">Trichadenotecnum species from Peninsular Malaysia and Singapore (Insecta: Psocodea: ' Psocoptera': Psocidae)</mods:title>
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<mods:name id="812B387C018B7C244425E75183ADC936" type="personal">
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<mods:roleTerm id="322A5B604468FFEDD88FC7E42D3FC7E1">Author</mods:roleTerm>
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<mods:namePart id="80334BC7BA7B79BE1C36AC0F8CB1CBA6">Yoshizawa, Kazunori</mods:namePart>
<mods:affiliation id="079AE9BC467FE4F9A3DEB0F055A0D268">Systematic Entomology, School of Agriculture, Hokkaido University, Sapporo 060 - 8589, Japan.</mods:affiliation>
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<mods:namePart id="A70C729FAD149C6D7F9524F47085D251">Lienhard, Charles</mods:namePart>
<mods:affiliation id="B7C15990680648CCB80EF13469A68A25">Muséum d'histoire naturelle, C. P. 6434, CH- 1211 Genève 6, Switzerland</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="7B5392DEB95A5C60DE4407215C324190">Abd, Idris</mods:namePart>
<mods:affiliation id="D7ABBE041F4320BE47B7931403CEBC08">Center for Insect Systematics, Faculty of Sciences and Technology, Universiti Kebangsaan Malaysia, 43600 Bangi, Malaysia</mods:affiliation>
</mods:name>
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<mods:titleInfo id="24AFE26B93C234E08A14530704587CD5">
<mods:title id="D663AEABF0F295C25C6EF7E4D7BE5C81">Zootaxa</mods:title>
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<mods:part id="4A50FFBE423A11ECD9AF0D15C02A61F7">
<mods:date id="B14C20A82BFB4B64F7A4D92A3851BBC6">2014</mods:date>
<mods:detail id="1EF7A0CD2EE6044F8F970F5B4E14DB2B" type="pubDate">
<mods:number id="0C89DFD3CDB08AC93C2837FC126A20F3">2014-07-14</mods:number>
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<mods:detail id="D17E6AE65A3ECAFDCEA995D069483F34" type="volume">
<mods:number id="944920E0BEAFCDBF78F113D4398CE95C">3835</mods:number>
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<mods:number id="985A7EAACFF2321945493CF886B8EBF4">4</mods:number>
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<mods:identifier id="F8518450C853AD8E0A0D885CBC51CDCD" type="CLB-Dataset">5285</mods:identifier>
<mods:identifier id="DD0023BD9D571410EC9E31AB1431C425" type="DOI">10.11646/zootaxa.3835.4.3</mods:identifier>
<mods:identifier id="02A1C4B99864B6D581C9CBCC77775FFC" type="GBIF-Dataset">dbf8de20-9c7a-4b66-9130-615b6b42e84e</mods:identifier>
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<treatment id="039F5A5FFFE8FFE2FF20FAABFDDC3DD6" ID-DOI="http://doi.org/10.5281/zenodo.4921967" ID-GBIF-Taxon="183731228" ID-Zenodo-Dep="4921967" LSID="urn:lsid:plazi:treatment:039F5A5FFFE8FFE2FF20FAABFDDC3DD6" httpUri="http://treatment.plazi.org/id/039F5A5FFFE8FFE2FF20FAABFDDC3DD6" lastPageId="24" lastPageNumber="493" pageId="23" pageNumber="492">
<subSubSection id="C32CB8C2FFE8FFEDFF20FAABFCFF3B50" box="[151,886,1332,1358]" pageId="23" pageNumber="492" type="nomenclature">
<paragraph id="8B89EB49FFE8FFEDFF20FAABFCFF3B50" blockId="23.[151,886,1332,1392]" box="[151,886,1332,1358]" pageId="23" pageNumber="492">
<heading id="D0C15C25FFE8FFEDFF20FAABFCFF3B50" bold="true" box="[151,886,1332,1358]" fontSize="11" level="1" pageId="23" pageNumber="492" reason="1">
<emphasis id="B942375BFFE8FFEDFF20FAABFCFF3B50" bold="true" box="[151,886,1332,1358]" pageId="23" pageNumber="492">
<taxonomicName id="4C3690CAFFE8FFEDFF20FAABFCA73B50" ID-CoL="587BJ" authority="Yoshizawa &amp; Lienhard" authorityName="Yoshizawa &amp; Lienhard &amp; Abd" authorityYear="2014" box="[151,814,1332,1358]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="23" pageNumber="492" phylum="Arthropoda" rank="species" species="anomalum" status="sp. nov.">
<emphasis id="B942375BFFE8FFEDFF20FAABFD8F3B50" bold="true" box="[151,518,1332,1358]" italics="true" pageId="23" pageNumber="492">Trichadenotecnum anomalum</emphasis>
Yoshizawa &amp; Lienhard
</taxonomicName>
,
<taxonomicNameLabel id="A2718A20FFE8FFEDFC8CFAABFCFF3B50" box="[827,886,1332,1358]" pageId="23" pageNumber="492" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32CB8C2FFE8FFEDFF20FAC8FEA23B6E" box="[151,299,1367,1392]" pageId="23" pageNumber="492" type="description">
<paragraph id="8B89EB49FFE8FFEDFF20FAC8FEA23B6E" blockId="23.[151,886,1332,1392]" box="[151,299,1367,1392]" pageId="23" pageNumber="492">
(
<figureCitation id="130DF7CCFFE8FFEDFF28FAC8FF733B6E" box="[159,250,1367,1392]" captionStart="FIGURE 1.1" captionStartId="2.[151,250,1786,1808]" captionTargetBox="[151,1435,196,1746]" captionTargetId="figure-94@2.[151,1436,193,1765]" captionTargetPageId="2" captionText="FIGURE 1.1. Forewings of Trichadenotecnum spp.: A: T. circularoides, female; B: T. cinnamonum, male; C: ditto, female; D: T. imrum, male; E: T.pardus, female; F: T. yatai, holotype male; G: T. kojimai, holotype male; H: T. adika, male; I: T. endangae, holotype male; J: T. hammani, female; K: T. sp.My19, female; L: T. gombakense, female; M: T. malayense, male." figureDoi="http://doi.org/10.5281/zenodo.4921625" httpUri="https://zenodo.org/record/4921625/files/figure.png" pageId="23" pageNumber="492">Figs 1R</figureCitation>
,
<figureCitation id="130DF7CCFFE8FFEDFEB1FAC8FEAA3B6E" box="[262,291,1367,1392]" captionStart="FIGURE 25" captionStartId="26.[151,250,1193,1215]" captionTargetBox="[152,1426,258,1160]" captionTargetId="figure-88@26.[151,1436,251,1171]" captionTargetPageId="26" captionText="FIGURE 25. Male terminalia of Trichadenotecnum anomalum. A: terminalia, lateral view; B: epiproct, dorsal view; C: hypandrium, ventral view; D: phallosome, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4921678" httpUri="https://zenodo.org/record/4921678/files/figure.png" pageId="23" pageNumber="492">25</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32CB8C2FFE8FFEDFF20FA00FE8E3BC2" pageId="23" pageNumber="492" type="materials_examined">
<paragraph id="8B89EB49FFE8FFEDFF20FA00FE8E3BC2" blockId="23.[151,1436,1439,2004]" pageId="23" pageNumber="492">
<materialsCitation id="3B5EE114FFE8FFEDFF20FA00FE8E3BC2" ID-GBIF-Occurrence="3326724305" collectingDate="1969-10-26" collectionCode="MHNG" collectorName="D. H. Murphy" country="Singapore" location="Male" municipality="Bukit Timah forest" pageId="23" pageNumber="492" specimenCount="1" typeStatus="holotype">
<emphasis id="B942375BFFE8FFEDFF20FA00FE823BA6" bold="true" box="[151,267,1439,1464]" inLineHeading="true" pageId="23" pageNumber="492" reason="1">
<typeStatus id="548D55EBFFE8FFEDFF20FA00FE8F3BA6" box="[151,262,1439,1464]" pageId="23" pageNumber="492" type="holotype">Holotype</typeStatus>
.
</emphasis>
<location id="8EE9BD92FFE8FFEDFEA1FA00FEDB3BA6" LSID="urn:lsid:plazi:treatment:039F5A5FFFE8FFE2FF20FAABFDDC3DD6:8EE9BD92FFE8FFEDFEA1FA00FEDB3BA6" box="[278,338,1439,1464]" country="Singapore" municipality="Bukit Timah forest" name="Male" pageId="23" pageNumber="492">Male</location>
,
<collectingCountry id="F321ABD9FFE8FFEDFED4FA00FE513BA6" box="[355,472,1439,1464]" name="Singapore" pageId="23" pageNumber="492">Singapore</collectingCountry>
,
<collectingMunicipality id="6BED7133FFE8FFEDFE50FA00FD403BA6" box="[487,713,1439,1464]" pageId="23" pageNumber="492">Bukit Timah forest</collectingMunicipality>
, 24 hr survey,
<collectingMethod id="5277935EFFE8FFEDFCC8FA00FB8F3BA9" box="[895,1030,1439,1464]" pageId="23" pageNumber="492">section trap</collectingMethod>
59 am,
<date id="FF88CD89FFE8FFEDFBCAFA00FB663BA6" box="[1149,1263,1439,1464]" pageId="23" pageNumber="492" value="1969-10-26">
<collectingDate id="EFCC3461FFE8FFEDFBCAFA00FB663BA6" box="[1149,1263,1439,1464]" pageId="23" pageNumber="492" value="1969-10-26">26.x.1969</collectingDate>
</date>
,
<collectorName id="26C38E9FFFE8FFEDFAB7FA3FFA153BA6" box="[1280,1436,1439,1464]" pageId="23" pageNumber="492">D.H. Murphy</collectorName>
(
<collectionCode id="ED27738CFFE8FFEDFF28FA5CFF723BC2" box="[159,251,1475,1500]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="23" pageNumber="492" type="Museum">MHNG</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32CB8C2FFE8FFE2FF70FA78FE2B3FEB" lastPageId="24" lastPageNumber="493" pageId="23" pageNumber="492" type="description">
<paragraph id="8B89EB49FFE8FFEDFF70FA78FA1F3872" blockId="23.[151,1436,1439,2004]" pageId="23" pageNumber="492">
<emphasis id="B942375BFFE8FFEDFF70FA78FCFA381E" box="[199,883,1511,1536]" inLineHeading="true" pageId="23" pageNumber="492" reason="1">
<emphasis id="B942375BFFE8FFEDFF70FA78FED8381E" bold="true" box="[199,337,1511,1536]" pageId="23" pageNumber="492">Description</emphasis>
(note that the condition of the specimen is bad).
</emphasis>
<emphasis id="B942375BFFE8FFEDFCCDFA78FC3E381E" bold="true" box="[890,951,1511,1536]" pageId="23" pageNumber="492">Male</emphasis>
.
<emphasis id="B942375BFFE8FFEDFC72FA78FB85381E" bold="true" box="[965,1036,1511,1536]" pageId="23" pageNumber="492">Head.</emphasis>
Yellowish in ground color; vertical markings and orbital markings obscure; coronal suture black; epicranial suture faintly bordered with brown band dorsally; frons with faint.markings; eye black, IO/D= 1.1; ocelli white, ocellar field brown; gena mostly yellowish; postclypeus with seven rows of spots dorsomedially; anteclypeus brown. Antenna pale brown. Mouthparts brown.
</paragraph>
<paragraph id="8B89EB49FFE8FFEDFF70F9E8FB2D38C6" blockId="23.[151,1436,1439,2004]" pageId="23" pageNumber="492">
<emphasis id="B942375BFFE8FFEDFF70F9E8FEAF3891" box="[199,294,1655,1680]" inLineHeading="true" pageId="23" pageNumber="492" reason="1">
<emphasis id="B942375BFFE8FFEDFF70F9E8FEA9388E" bold="true" box="[199,288,1655,1680]" pageId="23" pageNumber="492">Thorax</emphasis>
.
</emphasis>
Prothorax brown. Mesonotum yellowish, lateral lobe of scutum with faint marking, anterolateral corners of scutellum brown. Metanotum yellowish, lateral lobe of scutum with faint marking, lateral corner of scutellum brown. Meso- and metapleuron pale brown except for white membranous regions.
</paragraph>
<paragraph id="8B89EB49FFE8FFEDFF70F97CFCF738E2" blockId="23.[151,1436,1439,2004]" box="[199,894,1763,1788]" pageId="23" pageNumber="492">
<emphasis id="B942375BFFE8FFEDFF70F97CFF7638E2" bold="true" box="[199,255,1763,1788]" pageId="23" pageNumber="492">Legs</emphasis>
. Mostly pale brown, foretibia and tarsi of all legs darker.
</paragraph>
<paragraph id="8B89EB49FFE8FFEDFF70F898FE073979" blockId="23.[151,1436,1439,2004]" pageId="23" pageNumber="492">
<emphasis id="B942375BFFE8FFEDFF70F898FE30393E" box="[199,441,1799,1824]" inLineHeading="true" pageId="23" pageNumber="492" reason="1">
<emphasis id="B942375BFFE8FFEDFF70F898FEB1393E" bold="true" box="[199,312,1799,1824]" pageId="23" pageNumber="492">Forewing</emphasis>
(
<figureCitation id="130DF7CCFFE8FFEDFEFAF898FE22393E" box="[333,427,1799,1824]" captionStart="FIGURE 1.1" captionStartId="2.[151,250,1786,1808]" captionTargetBox="[151,1435,196,1746]" captionTargetId="figure-94@2.[151,1436,193,1765]" captionTargetPageId="2" captionText="FIGURE 1.1. Forewings of Trichadenotecnum spp.: A: T. circularoides, female; B: T. cinnamonum, male; C: ditto, female; D: T. imrum, male; E: T.pardus, female; F: T. yatai, holotype male; G: T. kojimai, holotype male; H: T. adika, male; I: T. endangae, holotype male; J: T. hammani, female; K: T. sp.My19, female; L: T. gombakense, female; M: T. malayense, male." figureDoi="http://doi.org/10.5281/zenodo.4921625" httpUri="https://zenodo.org/record/4921625/files/figure.png" pageId="23" pageNumber="492">Fig. 1R</figureCitation>
).
</emphasis>
Hyaline, lacking almost all marking elements characteristics for
<taxonomicName id="4C3690CAFFE8FFEDFB7BF898FA15393E" authorityName="Enderlein" authorityYear="1909" box="[1228,1436,1799,1824]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="23" pageNumber="492" phylum="Arthropoda" rank="genus">
<emphasis id="B942375BFFE8FFEDFB7BF898FA15393E" box="[1228,1436,1799,1824]" italics="true" pageId="23" pageNumber="492">Trichadenotecnum</emphasis>
</taxonomicName>
except following: opposing spots in cell r distinct, fused to single marking; basal band only distinct around Rs-M fusion and in cell cua.
</paragraph>
<paragraph id="8B89EB49FFE8FFE2FF70F8ECFD103FB5" blockId="23.[151,1436,1439,2004]" lastBlockId="24.[151,1437,151,968]" lastPageId="24" lastPageNumber="493" pageId="23" pageNumber="492">
<emphasis id="B942375BFFE8FFEDFF70F8ECFED83995" box="[199,337,1907,1932]" inLineHeading="true" pageId="23" pageNumber="492" reason="1">
<emphasis id="B942375BFFE8FFEDFF70F8ECFEC33992" bold="true" box="[199,330,1907,1932]" pageId="23" pageNumber="492">Terminalia</emphasis>
.
</emphasis>
8th sternum with pair of sclerotized portions (
<figureCitation id="130DF7CCFFE8FFEDFCE9F8ECFC393992" box="[862,944,1907,1932]" captionStart="FIGURE 25" captionStartId="26.[151,250,1193,1215]" captionTargetBox="[152,1426,258,1160]" captionTargetId="figure-88@26.[151,1436,251,1171]" captionTargetPageId="26" captionText="FIGURE 25. Male terminalia of Trichadenotecnum anomalum. A: terminalia, lateral view; B: epiproct, dorsal view; C: hypandrium, ventral view; D: phallosome, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4921678" httpUri="https://zenodo.org/record/4921678/files/figure.png" pageId="23" pageNumber="492">Fig. 25</figureCitation>
AC). Clunium (
<figureCitation id="130DF7CCFFE8FFEDFBE8F8ECFB4F3992" box="[1119,1222,1907,1932]" captionStart="FIGURE 25" captionStartId="26.[151,250,1193,1215]" captionTargetBox="[152,1426,258,1160]" captionTargetId="figure-88@26.[151,1436,251,1171]" captionTargetPageId="26" captionText="FIGURE 25. Male terminalia of Trichadenotecnum anomalum. A: terminalia, lateral view; B: epiproct, dorsal view; C: hypandrium, ventral view; D: phallosome, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4921678" httpUri="https://zenodo.org/record/4921678/files/figure.png" pageId="23" pageNumber="492">Fig. 25A</figureCitation>
) with broad lateral arms, about parallel sided and with strong dorsal expansion apically. Epiproct (
<figureCitation id="130DF7CCFFE8FFEDFB8DF808FB0639AE" box="[1082,1167,1943,1968]" captionStart="FIGURE 25" captionStartId="26.[151,250,1193,1215]" captionTargetBox="[152,1426,258,1160]" captionTargetId="figure-88@26.[151,1436,251,1171]" captionTargetPageId="26" captionText="FIGURE 25. Male terminalia of Trichadenotecnum anomalum. A: terminalia, lateral view; B: epiproct, dorsal view; C: hypandrium, ventral view; D: phallosome, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4921678" httpUri="https://zenodo.org/record/4921678/files/figure.png" pageId="23" pageNumber="492">Fig. 25</figureCitation>
AB) strongly expanded dorsally and anteriorly over clunium, posteriorly with small conical projection medially; dorsal margin of epiproct lobe broad and posterior surface convex. Paraproct (
<figureCitation id="130DF7CCFFE7FFE2FD44FF08FCD53EAE" box="[755,860,151,176]" captionStart="FIGURE 25" captionStartId="26.[151,250,1193,1215]" captionTargetBox="[152,1426,258,1160]" captionTargetId="figure-88@26.[151,1436,251,1171]" captionTargetPageId="26" captionText="FIGURE 25. Male terminalia of Trichadenotecnum anomalum. A: terminalia, lateral view; B: epiproct, dorsal view; C: hypandrium, ventral view; D: phallosome, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4921678" httpUri="https://zenodo.org/record/4921678/files/figure.png" pageId="24" pageNumber="493">Fig. 25A</figureCitation>
) with small distal lobe and with short and almost straight distal process directed posteriorly. Hypandrium (
<figureCitation id="130DF7CCFFE7FFE2FCAEFF23FCF73ECB" box="[793,894,188,213]" captionStart="FIGURE 25" captionStartId="26.[151,250,1193,1215]" captionTargetBox="[152,1426,258,1160]" captionTargetId="figure-88@26.[151,1436,251,1171]" captionTargetPageId="26" captionText="FIGURE 25. Male terminalia of Trichadenotecnum anomalum. A: terminalia, lateral view; B: epiproct, dorsal view; C: hypandrium, ventral view; D: phallosome, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4921678" httpUri="https://zenodo.org/record/4921678/files/figure.png" pageId="24" pageNumber="493">Fig. 25C</figureCitation>
) asymmetrical, without median tongue but with tongue-shaped projection medially near anterior margin, anterolaterally with pair of areas densely covered with tiny denticles, distally with two processes, one arises medially (= right process), acute conical in shape, the other arises left to this process (= left process), broad with diagonally truncated distal end; right posterolateral corner with denticulated longitudinal keel and with broadly expanded plate with serrated distal and internal margins. Phallosome (
<figureCitation id="130DF7CCFFE7FFE2FE9DFEF0FE1B3F96" box="[298,402,367,392]" captionStart="FIGURE 25" captionStartId="26.[151,250,1193,1215]" captionTargetBox="[152,1426,258,1160]" captionTargetId="figure-88@26.[151,1436,251,1171]" captionTargetPageId="26" captionText="FIGURE 25. Male terminalia of Trichadenotecnum anomalum. A: terminalia, lateral view; B: epiproct, dorsal view; C: hypandrium, ventral view; D: phallosome, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4921678" httpUri="https://zenodo.org/record/4921678/files/figure.png" pageId="24" pageNumber="493">Fig. 25D</figureCitation>
) with broad distal margin slightly concave medially, about parallel sided and narrowing to pointed anterior end, lacking anterior process.
</paragraph>
<paragraph id="8B89EB49FFE7FFE2FF70FE28FD0D3FCE" blockId="24.[151,1437,151,968]" box="[199,644,439,464]" pageId="24" pageNumber="493">
<emphasis id="B942375BFFE7FFE2FF70FE28FEF43FD1" box="[199,381,439,464]" inLineHeading="true" pageId="24" pageNumber="493" reason="1">
<emphasis id="B942375BFFE7FFE2FF70FE28FEFF3FCE" bold="true" box="[199,374,439,464]" pageId="24" pageNumber="493">Measurements</emphasis>
.
</emphasis>
B 1.5, Fw 2.0, Hw 1.5.
</paragraph>
<paragraph id="8B89EB49FFE7FFE2FF70FE43FE2B3FEB" blockId="24.[151,1437,151,968]" box="[199,418,476,501]" pageId="24" pageNumber="493">
<emphasis id="B942375BFFE7FFE2FF70FE43FEAC3FED" box="[199,293,476,501]" inLineHeading="true" pageId="24" pageNumber="493" reason="1">
<emphasis id="B942375BFFE7FFE2FF70FE43FE973FEB" bold="true" box="[199,286,476,501]" pageId="24" pageNumber="493">Female</emphasis>
.
</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C32CB8C2FFE7FFE2FF70FE60FDDC3DD6" pageId="24" pageNumber="493" type="discussion">
<paragraph id="8B89EB49FFE7FFE2FF70FE60FDDC3DD6" blockId="24.[151,1437,151,968]" pageId="24" pageNumber="493">
<emphasis id="B942375BFFE7FFE2FF70FE60FEB23C09" box="[199,315,511,536]" inLineHeading="true" pageId="24" pageNumber="493" reason="1">
<emphasis id="B942375BFFE7FFE2FF70FE60FEBD3C06" bold="true" box="[199,308,511,536]" pageId="24" pageNumber="493">Remarks</emphasis>
.
</emphasis>
This species is very unusual in its forewing markings (
<figureCitation id="130DF7CCFFE7FFE2FC00FE60FB983C06" box="[951,1041,511,536]" captionStart="FIGURE 1.1" captionStartId="2.[151,250,1786,1808]" captionTargetBox="[151,1435,196,1746]" captionTargetId="figure-94@2.[151,1436,193,1765]" captionTargetPageId="2" captionText="FIGURE 1.1. Forewings of Trichadenotecnum spp.: A: T. circularoides, female; B: T. cinnamonum, male; C: ditto, female; D: T. imrum, male; E: T.pardus, female; F: T. yatai, holotype male; G: T. kojimai, holotype male; H: T. adika, male; I: T. endangae, holotype male; J: T. hammani, female; K: T. sp.My19, female; L: T. gombakense, female; M: T. malayense, male." figureDoi="http://doi.org/10.5281/zenodo.4921625" httpUri="https://zenodo.org/record/4921625/files/figure.png" pageId="24" pageNumber="493">Fig. 1R</figureCitation>
). Although density is variable, the forewings of all known
<taxonomicName id="4C3690CAFFE7FFE2FE16FDBBFD433C23" box="[417,714,548,573]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="24" pageNumber="493" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B942375BFFE7FFE2FE16FDBBFDF83C23" box="[417,625,548,573]" italics="true" pageId="24" pageNumber="493">Trichadenotecnum</emphasis>
species
</taxonomicName>
are broadly spotted (
<figureCitation id="130DF7CCFFE7FFE2FC01FDBBFC7A3C23" box="[950,1011,548,573]" captionStart="FIGURE 1.1" captionStartId="2.[151,250,1786,1808]" captionTargetBox="[151,1435,196,1746]" captionTargetId="figure-94@2.[151,1436,193,1765]" captionTargetPageId="2" captionText="FIGURE 1.1. Forewings of Trichadenotecnum spp.: A: T. circularoides, female; B: T. cinnamonum, male; C: ditto, female; D: T. imrum, male; E: T.pardus, female; F: T. yatai, holotype male; G: T. kojimai, holotype male; H: T. adika, male; I: T. endangae, holotype male; J: T. hammani, female; K: T. sp.My19, female; L: T. gombakense, female; M: T. malayense, male." figureDoi="http://doi.org/10.5281/zenodo.4921625" httpUri="https://zenodo.org/record/4921625/files/figure.png" pageId="24" pageNumber="493">Fig. 1</figureCitation>
A-Q, S-U). Especially, in less heavily spotted species, six characteristic submarginal spots are always clearly visible, and this is considered to be an autapomorphy of the genus. In contrast, the forewings of
<taxonomicName id="4C3690CAFFE7FFE2FC8EFDF2FC473C9B" authorityName="Yoshizawa &amp; Lienhard &amp; Abd" authorityYear="2014" box="[825,974,620,645]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="24" pageNumber="493" phylum="Arthropoda" rank="species" species="anomalum">
<emphasis id="B942375BFFE7FFE2FC8EFDF2FC473C9B" box="[825,974,620,645]" italics="true" pageId="24" pageNumber="493">T. anomalum</emphasis>
</taxonomicName>
are mostly transparent except for some markings in the basal half (
<figureCitation id="130DF7CCFFE7FFE2FE65FD10FDA23CB6" box="[466,555,655,680]" captionStart="FIGURE 1.1" captionStartId="2.[151,250,1786,1808]" captionTargetBox="[151,1435,196,1746]" captionTargetId="figure-94@2.[151,1436,193,1765]" captionTargetPageId="2" captionText="FIGURE 1.1. Forewings of Trichadenotecnum spp.: A: T. circularoides, female; B: T. cinnamonum, male; C: ditto, female; D: T. imrum, male; E: T.pardus, female; F: T. yatai, holotype male; G: T. kojimai, holotype male; H: T. adika, male; I: T. endangae, holotype male; J: T. hammani, female; K: T. sp.My19, female; L: T. gombakense, female; M: T. malayense, male." figureDoi="http://doi.org/10.5281/zenodo.4921625" httpUri="https://zenodo.org/record/4921625/files/figure.png" pageId="24" pageNumber="493">Fig. 1R</figureCitation>
). The condition of the
<typeStatus id="548D55EBFFE7FFE2FC82FD10FC1E3CB6" box="[821,919,655,680]" pageId="24" pageNumber="493" type="holotype">holotype</typeStatus>
specimen is poor, with many body markings faded. However, the basal band of the forewing can be seen clearly in the
<typeStatus id="548D55EBFFE7FFE2FC6DFD2BFBB53CD3" box="[986,1084,692,717]" pageId="24" pageNumber="493" type="holotype">holotype</typeStatus>
so that absence of the marking characteristics of
<taxonomicName id="4C3690CAFFE7FFE2FEDFFD48FDB13CEE" authorityName="Enderlein" authorityYear="1909" box="[360,568,727,752]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="24" pageNumber="493" phylum="Arthropoda" rank="genus">
<emphasis id="B942375BFFE7FFE2FEDFFD48FDB13CEE" box="[360,568,727,752]" italics="true" pageId="24" pageNumber="493">Trichadenotecnum</emphasis>
</taxonomicName>
, such as the submarginal row of spots, is probably not due to the poor condition of the specimen. The 9th tergite of
<taxonomicName id="4C3690CAFFE7FFE2FD14FD62FCBE3D0B" authorityName="Yoshizawa &amp; Lienhard &amp; Abd" authorityYear="2014" box="[675,823,764,789]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="24" pageNumber="493" phylum="Arthropoda" rank="species" species="anomalum">
<emphasis id="B942375BFFE7FFE2FD14FD62FCBE3D0B" box="[675,823,764,789]" italics="true" pageId="24" pageNumber="493">T. anomalum</emphasis>
</taxonomicName>
possesses posterolateral arms (
<figureCitation id="130DF7CCFFE7FFE2FB28FD63FA803D0B" box="[1183,1289,764,789]" captionStart="FIGURE 25" captionStartId="26.[151,250,1193,1215]" captionTargetBox="[152,1426,258,1160]" captionTargetId="figure-88@26.[151,1436,251,1171]" captionTargetPageId="26" captionText="FIGURE 25. Male terminalia of Trichadenotecnum anomalum. A: terminalia, lateral view; B: epiproct, dorsal view; C: hypandrium, ventral view; D: phallosome, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4921678" httpUri="https://zenodo.org/record/4921678/files/figure.png" pageId="24" pageNumber="493">Fig. 25A</figureCitation>
), a character which is uniquely observed in
<taxonomicName id="4C3690CAFFE7FFE2FE44FC80FD4A3D26" authorityName="Enderlein" authorityYear="1909" box="[499,707,799,824]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="24" pageNumber="493" phylum="Arthropoda" rank="genus">
<emphasis id="B942375BFFE7FFE2FE44FC80FD4A3D26" box="[499,707,799,824]" italics="true" pageId="24" pageNumber="493">Trichadenotecnum</emphasis>
</taxonomicName>
among
<taxonomicName id="4C3690CAFFE7FFE2FC97FC80FC0E3D26" authorityName="Yoshizawa &amp; Lienhard &amp; Abd" authorityYear="2014" box="[800,903,799,824]" class="Insecta" family="Psocidae" kingdom="Animalia" order="Psocodea" pageId="24" pageNumber="493" phylum="Arthropoda" rank="family">Psocidae</taxonomicName>
. Other diagnostic characters of the genus, such as the triangular areola postica, can also be seen (
<figureCitation id="130DF7CCFFE7FFE2FD0CFCDBFC9B3D43" box="[699,786,836,861]" captionStart="FIGURE 1.1" captionStartId="2.[151,250,1786,1808]" captionTargetBox="[151,1435,196,1746]" captionTargetId="figure-94@2.[151,1436,193,1765]" captionTargetPageId="2" captionText="FIGURE 1.1. Forewings of Trichadenotecnum spp.: A: T. circularoides, female; B: T. cinnamonum, male; C: ditto, female; D: T. imrum, male; E: T.pardus, female; F: T. yatai, holotype male; G: T. kojimai, holotype male; H: T. adika, male; I: T. endangae, holotype male; J: T. hammani, female; K: T. sp.My19, female; L: T. gombakense, female; M: T. malayense, male." figureDoi="http://doi.org/10.5281/zenodo.4921625" httpUri="https://zenodo.org/record/4921625/files/figure.png" pageId="24" pageNumber="493">Fig. 1R</figureCitation>
). Therefore, the species is here considered as a member of
<taxonomicName id="4C3690CAFFE7FFE2FF20FCF8FEEE3D9E" authorityName="Enderlein" authorityYear="1909" box="[151,359,871,896]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="24" pageNumber="493" phylum="Arthropoda" rank="genus">
<emphasis id="B942375BFFE7FFE2FF20FCF8FEEE3D9E" box="[151,359,871,896]" italics="true" pageId="24" pageNumber="493">Trichadenotecnum</emphasis>
</taxonomicName>
. The species seems to be closely related to
<taxonomicName id="4C3690CAFFE7FFE2FCFCFCF6FC4A3D9E" authorityName="Thornton" authorityYear="1961" box="[843,963,873,896]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="24" pageNumber="493" phylum="Arthropoda" rank="species" species="apertum">
<emphasis id="B942375BFFE7FFE2FCFCFCF6FC4A3D9E" box="[843,963,873,896]" italics="true" pageId="24" pageNumber="493">T. apertum</emphasis>
</taxonomicName>
and its relatives because of the absence of the hypandrial median tongue and other resemblances of the terminal structures. The unique forewing markings are probably autapomorphic to this species.
</paragraph>
</subSubSection>
</treatment>
</document>