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<document id="2D3E2DDD666B40715203BE8B9CB1389B" ID-CLB-Dataset="26389" ID-DOI="10.5852/ejt.2019.533" ID-GBIF-Dataset="e1990f2e-89e4-476e-8407-8d736c02f47a" ID-Zenodo-Dep="3258383" ID-ZooBank="E98D3A33-49A7-45A8-94BC-F2AC96A7622B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1561645484001" checkinUser="plazi" docAuthor="Gnohossou, Pierre, Piscart, Christophe &amp; Umr, Ecobio -" docDate="2019" docId="03958E15CC70FFB68361FDF6F0B6F868" docLanguage="en" docName="ejt-533_gnohossou_piscart.pdf.imf" docOrigin="European Journal of Taxonomy 533" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Quadrivisio laleyei Gnohossou &amp; Piscart &amp; Umr 2019, sp. nov." docType="treatment" docUuid="AE890355-182D-4473-ABF2-A90DFE1E1208" docUuidSource="ZooBank" docVersion="8" lastPageNumber="10" masterDocId="FFACF66DCC73FFBF8116FF83F217FFC6" masterDocTitle="A new species of Quadrivisio (Amphipoda, Maeridae) from coastal tropical lagoons (Benin, West Africa)" masterLastPageNumber="14" masterPageNumber="1" pageNumber="4" updateTime="1698736886963" updateUser="ExternalLinkService">
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<mods:title id="65E62264E95A9FD443031F0F8CD10CBA">A new species of Quadrivisio (Amphipoda, Maeridae) from coastal tropical lagoons (Benin, West Africa)</mods:title>
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<mods:namePart id="A67EFE8872863537F44505BD20773863">Gnohossou, Pierre</mods:namePart>
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<mods:namePart id="A4373915F2B66112E59D346FD7388D6D">Piscart, Christophe</mods:namePart>
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<mods:roleTerm id="DCF2AD677E1457178642CE930C10768C">Author</mods:roleTerm>
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<mods:namePart id="5CBCBE155744555632DCC40970E7578D">Umr, Ecobio -</mods:namePart>
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<mods:title id="3E8400CE40E89F6FCDA0A8DAC5E69368">European Journal of Taxonomy</mods:title>
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<mods:date id="92F615AD547B3F3B45F7500B865E2580">2019</mods:date>
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<mods:number id="927BB526FE770576979B601E73842CEB">2019-06-26</mods:number>
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<subSubSection id="C3266C88CC70FFBC8361FDF6F6D3FD75" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="8B833F03CC70FFBC8361FDF6F1AAFD56" blockId="3.[367,1220,629,726]" box="[631,957,629,656]" pageId="3" pageNumber="4">
<heading id="D0CB886FCC70FFBC8361FDF6F1AAFD56" box="[631,957,629,656]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="4" reason="2">
<taxonomicName id="4C3C4480CC70FFBC8361FDF6F14DFD56" authority="Gnohossou &amp; Piscart &amp; Umr, 2019" authorityName="Gnohossou &amp; Piscart &amp; Umr" authorityYear="2019" box="[631,858,629,656]" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="laleyei" status="sp. nov.">
<emphasis id="B948E311CC70FFBC8361FDF6F14DFD56" bold="true" box="[631,858,629,656]" italics="true" pageId="3" pageNumber="4">Quadrivisio laleyei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B5E6ACC70FFBC8274FDF5F1AAFD56" box="[866,957,630,656]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
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<subSubSection id="C3266C88CC70FFBC83B6FD3FF183FD10" box="[672,916,699,726]" pageId="3" pageNumber="4" type="description">
<paragraph id="8B833F03CC70FFBC83B6FD3FF183FD10" blockId="3.[367,1220,629,726]" box="[672,916,699,726]" pageId="3" pageNumber="4">
<figureCitation id="13072386CC70FFBC83B6FD3FF11FFD10" box="[672,776,699,726]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="4.[189,232,1930,1956]" captionStartId-1="5.[189,232,1966,1992]" captionStartId-2="6.[189,232,1930,1956]" captionStartId-3="7.[189,232,1930,1956]" captionTargetBox-0="[213,1374,317,1884]" captionTargetBox-1="[221,1389,593,1908]" captionTargetBox-2="[323,1276,583,1890]" captionTargetBox-3="[212,1354,579,1875]" captionTargetId-0="figure@4.[213,1374,317,1890]" captionTargetId-1="figure@5.[189,1398,583,1927]" captionTargetId-2="figure@6.[309,1279,573,1893]" captionTargetId-3="figure@7.[189,1398,575,1888]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="Fig. 2. Quadrivisio laleyei sp. nov., ♂, holotype, 7.5 mm, MNHN IU-2017-209. A. Habitus (scale 1). B. Lower lip (scale 3). C. First antenna (scale 2). D. Second antenna (scale 2). E. Left mandible (scale 4). F. Maxilliped palp (scale 3)." captionText-1="Fig. 3. Quadrivisio laleyei sp. nov. A., C. Holotype, ♂, 7.5 mm, MNHN IU-2017-209, gnathopod. B., D. Paratype, ♀, 7.0 mm, MNHN IU-2017-211, gnathopod." captionText-2="Fig. 4. Quadrivisio laleyi sp. nov. AB., DE. Holotype, ♂, 7.5 mm, MNHN IU-2017-209. C. Paratype, ♂, 7.6 mm, MNHN IU-2017-210. A. Pereopod 3. B. Pereopod 4. C. Pereopod 5. D. Pereopod 6. E. Pereopod 7." captionText-3="Fig. 5. Quadrivisio laleyi sp. nov. AD., F. Holotype, ♂, 7.5 mm, MNHN IU-2017-209. E. Paratype, ♀, 7.0 mm, MNHN IU-2017-211. A. Male uropod 1 (scale 1). B. Male uropod 2 (scale 1). C. Male urosome (scale 1). D. Male uropod 3 (scale 1). E. Female uropod 3 (scale 1). F. Male telson (scale 2)." httpUri-0="https://zenodo.org/record/3258387/files/figure.png" httpUri-1="https://zenodo.org/record/3258389/files/figure.png" httpUri-2="https://zenodo.org/record/3258391/files/figure.png" httpUri-3="https://zenodo.org/record/3258393/files/figure.png" pageId="3" pageNumber="4">Figs 25</figureCitation>
,
<tableCitation id="C6BE0AB8CC70FFBC8202FD3FF183FD10" box="[788,916,700,726]" captionStart-0="Table 1" captionStart-1="Table 2" captionStartId-0="10.[189,258,268,294]" captionStartId-1="11.[189,258,268,294]" captionTargetBox-0="[204,1380,406,1934]" captionTargetBox-1="[313,1251,441,796]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="Table 1. Relationship between abiotic environmental variables and densities of the amphipod Quadrivisio laleyei sp. nov. sampled in 20022003 at five stations of the Nokoué lagoon (basket sampling method; mean of three replicates at each station)." captionText-1="Table 2. Mean abundances of the amphipod Quadrivisio laleyei sp. nov., sampled with four different methods at five stations of the Nokoué lagoon. Bundle (ind.bundle-1) and basket (ind.m-2) methods: 7 monthly samplings in 20022003, n = 3; Macrophytes (ind.macrophyte-1) and Surber (ind.m-2) methods: 2 sampling periods (February and September 2006), (n = 5)." httpUri-0="http://table.plazi.org/id/DF436F8BCC79FFB581ABFE8FF080FEAB" httpUri-1="http://table.plazi.org/id/DF436F8BCC78FFB481ABFE8FF1E4FE56" pageId="3" pageNumber="4" tableUuid-0="DF436F8BCC79FFB581ABFE8FF080FEAB" tableUuid-1="DF436F8BCC78FFB481ABFE8FF1E4FE56">Tables 12</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3266C88CC70FFBC81ABFD7EF15AFBFA" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="8B833F03CC70FFBC81ABFD7EF321FCD1" blockId="3.[189,310,765,791]" box="[189,310,765,791]" pageId="3" pageNumber="4">
<heading id="D0CB886FCC70FFBC81ABFD7EF321FCD1" bold="true" box="[189,310,765,791]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="B948E311CC70FFBC81ABFD7EF321FCD1" bold="true" box="[189,310,765,791]" pageId="3" pageNumber="4">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B833F03CC70FFBC81ABFCAFF15AFBFA" blockId="3.[189,1399,812,1085]" pageId="3" pageNumber="4">
The new species belongs to the genus
<taxonomicName id="4C3C4480CC70FFBC8382FCAFF137FC80" authorityName="Stebbing" authorityYear="1907" box="[660,800,812,838]" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B948E311CC70FFBC8382FCAFF137FC80" box="[660,800,812,838]" italics="true" pageId="3" pageNumber="4">Quadrivisio</emphasis>
</taxonomicName>
, recognized by the shape of their eyes for which ommatidia are almost split into two pairs on each side of head, antenna 1 with a very long flagellum, propodus of male gnathopod 2 very large, uropod 3 with rami foliaceous. Accessory flagellum of antenna 1 with 46 articles. Article 3 of mandibular palp slightly shorter than article 2 and armed with two setae. Propodus of male gnathopod 1 almost rectangular. Gnathopod 2 sexually dimorphic. Epimeral plates 23 with a small tooth at posterodistal angle, but lacking supernumerary teeth on posterior margin. Pleonite and urosomites smooth. Urosomite 2 with posterodorsal robust setae (one on each side). Telson without spines or setae along inner margin of each lobe.
</paragraph>
</subSubSection>
<subSubSection id="C3266C88CC70FFBC81ABFBE7F62DFB16" pageId="3" pageNumber="4" type="etymology">
<paragraph id="8B833F03CC70FFBC81ABFBE7F353FBB8" blockId="3.[189,324,1124,1150]" box="[189,324,1124,1150]" pageId="3" pageNumber="4">
<heading id="D0CB886FCC70FFBC81ABFBE7F353FBB8" bold="true" box="[189,324,1124,1150]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="B948E311CC70FFBC81ABFBE7F353FBB8" bold="true" box="[189,324,1124,1150]" pageId="3" pageNumber="4">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B833F03CC70FFBC81ABFB11F62DFB16" blockId="3.[189,1398,1170,1232]" pageId="3" pageNumber="4">
The epithet
<taxonomicName id="4C3C4480CC70FFBC805AFB11F38DFB6A" authorityName="Gnohossou &amp; Piscart &amp; Umr" authorityYear="2019" box="[332,410,1170,1196]" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="laleyei">
<emphasis id="B948E311CC70FFBC805AFB11F38DFB6A" box="[332,410,1170,1196]" italics="true" pageId="3" pageNumber="4">laleyei</emphasis>
</taxonomicName>
refers to the name of the Professor Philippe A. Lalèyè who initiated the studies of macrofauna from lagoons in
<collectingCountry id="F32B7F93CC70FFBC8306FB36F04DFB09" box="[528,602,1205,1231]" name="Benin" pageId="3" pageNumber="4">Benin</collectingCountry>
. The name is a noun in the genitive case.
</paragraph>
</subSubSection>
<subSubSection id="C3266C88CC70FFBC81ABFB75F72EF8CD" pageId="3" pageNumber="4" type="materials_examined">
<paragraph id="8B833F03CC70FFBC81ABFB75F3BDFAD6" blockId="3.[189,426,1270,1296]" box="[189,426,1270,1296]" pageId="3" pageNumber="4">
<heading id="D0CB886FCC70FFBC81ABFB75F3BDFAD6" bold="true" box="[189,426,1270,1296]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="B948E311CC70FFBC81ABFB75F3BDFAD6" bold="true" box="[189,426,1270,1296]" pageId="3" pageNumber="4">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B833F03CC70FFBC81FCFAA6F34BFAF9" blockId="3.[189,1399,1317,1484]" box="[234,348,1317,1343]" pageId="3" pageNumber="4">
<typeStatus id="548781A1CC70FFBC81FCFAA6F34BFAF9" box="[234,348,1317,1343]" pageId="3" pageNumber="4" type="holotype">
<emphasis id="B948E311CC70FFBC81FCFAA6F34BFAF9" bold="true" box="[234,348,1317,1343]" pageId="3" pageNumber="4">Holotype</emphasis>
</typeStatus>
</paragraph>
<paragraph id="8B833F03CC70FFBC81ABFACAF2E6FA0A" blockId="3.[189,1399,1317,1484]" pageId="3" pageNumber="4">
<materialsCitation id="3B54355ECC70FFBC81ABFACAF2E6FA0A" ID-GBIF-Occurrence="2270457987" collectingDate="2007-06" collectionCode="MNHN" country="Benin" county="Kessounou station" latitude="6.4681945" location="Porto Novo" longLatPrecision="1" longitude="2.615611" municipality="Porto-Novo" pageId="3" pageNumber="4" specimenCode="IU-2017- 209" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectionCode id="ED2DA7C6CC70FFBC81ABFACAF300FAA5" box="[189,279,1353,1379]" country="Benin" httpUri="http://grbio.org/cool/rx6h-yh4d" name="Université National du Bénin" pageId="3" pageNumber="4" type="Herbarium">
<collectingCountry id="F32B7F93CC70FFBC81ABFACAF300FAA5" box="[189,279,1353,1379]" name="Benin" pageId="3" pageNumber="4">BENIN</collectingCountry>
</collectionCode>
<specimenCount id="9D3AF48ACC70FFBC803BFACAF354FAA5" box="[301,323,1353,1379]" pageId="3" pageNumber="4" type="male"></specimenCount>
(
<quantity id="4CC492E6CC70FFBC8044FACBF3A6FAA5" box="[338,433,1352,1379]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="3" pageNumber="4" unit="mm" value="7.5">7.5 mm</quantity>
);
<location id="8EE369D8CC70FFBC80D2FACAF05CFAA5" LSID="urn:lsid:plazi:treatment:03958E15CC70FFB68361FDF6F0B6F868:8EE369D8CC70FFBC80D2FACAF05CFAA5" box="[452,587,1353,1379]" country="Benin" county="Kessounou station" latitude="6.4681945" longLatPrecision="1" longitude="2.615611" municipality="Porto-Novo" name="Porto Novo" pageId="3" pageNumber="4">Porto Novo</location>
city,
<collectingMunicipality id="6BE7A579CC70FFBC839FFACAF103FAA5" box="[649,788,1353,1379]" pageId="3" pageNumber="4">Porto-Novo</collectingMunicipality>
lagoon,
<quantity id="4CC492E6CC70FFBC8261FACBF1FDFAA5" box="[887,1002,1352,1379]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="3" pageNumber="4" unit="cm" value="25.0" valueMax="30.0" valueMin="20.0">2030 cm</quantity>
bottom depth,
<collectingCounty id="62E2478FCC70FFBC858EFACAF765FAA4" box="[1176,1394,1352,1379]" pageId="3" pageNumber="4">Kessounou station</collectingCounty>
, muddy sediment;
<geoCoordinate id="EE0859C4CC70FFBC8084FAE9F028FA40" box="[402,575,1386,1414]" degrees="06" direction="north" minutes="28" orientation="latitude" pageId="3" pageNumber="4" precision="1" seconds="05.5" value="6.4681945">
06°2805.5″
<collectionCode id="ED2DA7C6CC70FFBC8331FAEFF028FA40" box="[551,575,1388,1414]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="3" pageNumber="4" type="Herbarium">N</collectionCode>
</geoCoordinate>
,
<geoCoordinate id="EE0859C4CC70FFBC835AFAE9F0E1FA40" box="[588,758,1386,1414]" degrees="02" direction="east" minutes="36" orientation="longitude" pageId="3" pageNumber="4" precision="1" seconds="56.2" value="2.615611">
02°3656.2″
<collectionCode id="ED2DA7C6CC70FFBC83F7FAEFF0E1FA40" box="[737,758,1388,1414]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="3" pageNumber="4" type="Herbarium">E</collectionCode>
</geoCoordinate>
; water temperature: 28°
<collectionCode id="ED2DA7C6CC70FFBC8502FAEFF63EFA40" box="[1044,1065,1388,1414]" country="Denmark" name="University of Copenhagen" pageId="3" pageNumber="4" type="Herbarium">C</collectionCode>
; water salinity:
<quantity id="4CC492E6CC70FFBC85FEFAEFF718FA40" box="[1256,1295,1388,1414]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="3" pageNumber="4" unit="g" value="4.0">4 g</quantity>
.
<superScript id="7C49924BCC70FFBC8405FAEFF739FABE" attach="left" box="[1299,1326,1386,1414]" fontSize="6" pageId="3" pageNumber="4">L-</superScript>
1;
<date id="FF8219C3CC70FFBC8450FAEFF2ECFA6F" pageId="3" pageNumber="4" value="2007-06">
<collectingDate id="EFC6E02BCC70FFBC8450FAEFF2ECFA6F" pageId="3" pageNumber="4" value="2007-06">
Jun.
<date id="FF8219C3CC70FFBC81ABFA0CF2ECFA6F" box="[189,251,1423,1449]" pageId="3" pageNumber="4" value="2007" year="2007">2007</date>
</collectingDate>
</date>
; collected with live macrophytes (
<taxonomicName id="4C3C4480CC70FFBC839AFA0CF1AEFA6F" authority="Sw." authorityName="Sw." box="[652,953,1423,1449]" class="Liliopsida" family="Poaceae" genus="Paspalum" kingdom="Plantae" order="Poales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="vaginatum">
<emphasis id="B948E311CC70FFBC839AFA0CF193FA6F" box="[652,900,1423,1449]" italics="true" pageId="3" pageNumber="4">Paspalum vaginatum</emphasis>
Sw.
</taxonomicName>
), uprooted by hand;
<collectionCode id="ED2DA7C6CC70FFBC85BFFA0CF714FA6F" box="[1193,1283,1423,1449]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="4">MNHN</collectionCode>
<specimenCode id="DB9A9778CC70FFBC841FFA0CF725FA6F" box="[1289,1330,1423,1449]" collectionCode="IU-" pageId="3" pageNumber="4">IU-</specimenCode>
2017- 209.
</materialsCitation>
</paragraph>
<paragraph id="8B833F03CC70FFBC81FCFA77F370F9C8" blockId="3.[189,1398,1524,1620]" box="[234,359,1524,1550]" pageId="3" pageNumber="4">
<typeStatus id="548781A1CC70FFBC81FCFA77F370F9C8" box="[234,359,1524,1550]" pageId="3" pageNumber="4" type="paratype">
<emphasis id="B948E311CC70FFBC81FCFA77F370F9C8" bold="true" box="[234,359,1524,1550]" pageId="3" pageNumber="4">Paratypes</emphasis>
</typeStatus>
</paragraph>
<paragraph id="8B833F03CC70FFBC81ABF994F3A9F992" blockId="3.[189,1398,1524,1620]" pageId="3" pageNumber="4">
<materialsCitation id="3B54355ECC70FFBC81ABF994F1DEF9F7" ID-GBIF-Occurrence="2270457986" box="[189,969,1558,1585]" collectingDate="2007-06" collectionCode="MNHN" country="Benin" county="Kessounou station" latitude="6.4681945" location="Porto Novo" longLatPrecision="1" longitude="2.615611" municipality="Porto-Novo" pageId="3" pageNumber="4" specimenCode="IU-2017-210" specimenCount="5" specimenCount-male="5" typeStatus="holotype">
<collectionCode id="ED2DA7C6CC70FFBC81ABF994F300F9F7" box="[189,279,1559,1585]" country="Benin" httpUri="http://grbio.org/cool/rx6h-yh4d" name="Université National du Bénin" pageId="3" pageNumber="4" type="Herbarium">
<collectingCountry id="F32B7F93CC70FFBC81ABF994F300F9F7" box="[189,279,1559,1585]" name="Benin" pageId="3" pageNumber="4">BENIN</collectingCountry>
</collectionCode>
<specimenCount id="9D3AF48ACC70FFBC8025F995F36DF9F7" box="[307,378,1558,1585]" pageId="3" pageNumber="4" type="male">5 ♂♂</specimenCount>
; same data as for
<typeStatus id="548781A1CC70FFBC8343F994F0A8F9F7" box="[597,703,1559,1585]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
;
<collectionCode id="ED2DA7C6CC70FFBC83DBF994F130F9F7" box="[717,807,1559,1585]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="4">MNHN</collectionCode>
<specimenCode id="DB9A9778CC70FFBC8226F994F14EF9F7" box="[816,857,1559,1585]" collectionCode="IU-" pageId="3" pageNumber="4">IU-</specimenCode>
2017-210
</materialsCitation>
<materialsCitation id="3B54355ECC70FFBC82F0F995F3ADF992" ID-GBIF-Occurrence="2270457988" collectingDate="2007-06" collectionCode="MNHN" country="Benin" county="Kessounou station" latitude="6.4681945" location="Porto Novo" longLatPrecision="1" longitude="2.615611" municipality="Porto-Novo" pageId="3" pageNumber="4" specimenCode="IU-2017-211" specimenCount="5" specimenCount-female="5" typeStatus="paratype">
<specimenCount id="9D3AF48ACC70FFBC82F0F995F63CF9F7" box="[998,1067,1558,1585]" pageId="3" pageNumber="4" type="female">5 ♀♀</specimenCount>
; same data as for holotype;
<collectionCode id="ED2DA7C6CC70FFBC81ABF9B9F300F992" box="[189,279,1594,1620]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="4">MNHN</collectionCode>
<specimenCode id="DB9A9778CC70FFBC8008F9B9F35FF992" box="[286,328,1594,1620]" collectionCode="IU-" pageId="3" pageNumber="4">
<collectionCode id="ED2DA7C6CC70FFBC8008F9B9F35FF992" box="[286,328,1594,1620]" pageId="3" pageNumber="4">IU-</collectionCode>
</specimenCode>
2017-211
</materialsCitation>
.
</paragraph>
<paragraph id="8B833F03CC70FFBC81ABF9F8F3EEF953" blockId="3.[189,505,1659,1685]" box="[189,505,1659,1685]" pageId="3" pageNumber="4">
<heading id="D0CB886FCC70FFBC81ABF9F8F3EEF953" bold="true" box="[189,505,1659,1685]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="B948E311CC70FFBC81ABF9F8F3EEF953" bold="true" box="[189,505,1659,1685]" pageId="3" pageNumber="4">Other material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B833F03CC70FFBC81ABF929F72EF8CD" blockId="3.[189,1399,1706,1803]" pageId="3" pageNumber="4">
<materialsCitation id="3B54355ECC70FFBC81ABF929F72EF8CD" ID-GBIF-Occurrence="2270457985" collectedFrom="muddy sediment" collectingDate="2007-06" collectingMethod="uprooted by hand" country="Benin" latitude="6.4681945" location="Porto Novo city, Porto-Novo lagoon" longLatPrecision="1" longitude="2.615611" pageId="3" pageNumber="4" specimenCount="50">
<collectingCountry id="F32B7F93CC70FFBC81ABF929F300F902" box="[189,279,1706,1732]" name="Benin" pageId="3" pageNumber="4">BENIN</collectingCountry>
•Around
<specimenCount id="9D3AF48ACC70FFBC8090F929F031F902" box="[390,550,1706,1733]" pageId="3" pageNumber="4" type="generic">50 specimens</specimenCount>
;
<location id="8EE369D8CC70FFBC8339F929F1C6F902" LSID="urn:lsid:plazi:treatment:03958E15CC70FFB68361FDF6F0B6F868:8EE369D8CC70FFBC8339F929F1C6F902" box="[559,977,1706,1732]" country="Benin" latitude="6.4681945" longLatPrecision="1" longitude="2.615611" name="Porto Novo city, Porto-Novo lagoon" pageId="3" pageNumber="4">Porto Novo city, Porto-Novo lagoon</location>
,
<quantity id="4CC492E6CC70FFBC82CFF929F65CF902" box="[985,1099,1706,1732]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="3" pageNumber="4" unit="cm" value="25.0" valueMax="30.0" valueMin="20.0">2030 cm</quantity>
bottom depth,
<location id="8EE369D8CC70FFBC85E2F929F319F921" LSID="urn:lsid:plazi:treatment:03958E15CC70FFB68361FDF6F0B6F868:8EE369D8CC70FFBC85E2F929F319F921" country="Benin" latitude="6.4681945" longLatPrecision="1" longitude="2.615611" name="Kessounou station" pageId="3" pageNumber="4">Kessounou station</location>
,
<collectedFrom id="38207CFECC70FFBC8036F94EF3F9F921" box="[288,494,1741,1767]" pageId="3" pageNumber="4">muddy sediment</collectedFrom>
;
<geoCoordinate id="EE0859C4CC70FFBC80E9F948F0A6F92E" box="[511,689,1739,1768]" degrees="06" direction="north" minutes="28" orientation="latitude" pageId="3" pageNumber="4" precision="1" seconds="05.5" value="6.4681945">06°2805.5″ N</geoCoordinate>
,
<geoCoordinate id="EE0859C4CC70FFBC83D2F948F164F92E" box="[708,883,1739,1768]" degrees="02" direction="east" minutes="36" orientation="longitude" pageId="3" pageNumber="4" precision="1" seconds="56.2" value="2.615611">02°3656.2″ E</geoCoordinate>
; water temperature: 28°C; water salinity:
<quantity id="4CC492E6CC70FFBC81ABF973F2F5F8CD" box="[189,226,1776,1803]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="3" pageNumber="4" unit="g" value="4.0">4 g</quantity>
.
<superScript id="7C49924BCC70FFBC81F0F972F315F93B" attach="left" box="[230,258,1775,1803]" fontSize="6" pageId="3" pageNumber="4">L-</superScript>
1;
<date id="FF8219C3CC70FFBC800FF972F399F8CC" box="[281,398,1776,1803]" pageId="3" pageNumber="4" value="2007-06">
<collectingDate id="EFC6E02BCC70FFBC800FF972F399F8CC" box="[281,398,1776,1803]" pageId="3" pageNumber="4" value="2007-06">Jun. 2007</collectingDate>
</date>
; collected with live macrophytes (
<taxonomicName id="4C3C4480CC70FFBC8237F973F658F8CC" authority="Sw." authorityName="Sw." box="[801,1103,1776,1802]" class="Liliopsida" family="Poaceae" genus="Paspalum" kingdom="Plantae" order="Poales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="vaginatum">
<location id="8EE369D8CC70FFBC8237F973F658F8CC" LSID="urn:lsid:plazi:treatment:03958E15CC70FFB68361FDF6F0B6F868:8EE369D8CC70FFBC8237F973F658F8CC" box="[801,1103,1776,1802]" country="Benin" latitude="6.4681945" longLatPrecision="1" longitude="2.615611" name="Paspalum vaginatum Sw." pageId="3" pageNumber="4">
<emphasis id="B948E311CC70FFBC8237F973F60DF8CC" box="[801,1050,1776,1802]" italics="true" pageId="3" pageNumber="4">Paspalum vaginatum</emphasis>
Sw.
</location>
</taxonomicName>
),
<collectingMethod id="527D4714CC70FFBC8572F973F725F8CC" box="[1124,1330,1776,1802]" pageId="3" pageNumber="4">uprooted by hand</collectingMethod>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3266C88CC70FFB781ABF8B2F10BFA31" lastPageId="8" lastPageNumber="9" pageId="3" pageNumber="4" type="description">
<paragraph id="8B833F03CC70FFBC81ABF8B2F359F88D" blockId="3.[189,334,1841,1867]" box="[189,334,1841,1867]" pageId="3" pageNumber="4">
<heading id="D0CB886FCC70FFBC81ABF8B2F359F88D" bold="true" box="[189,334,1841,1867]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="B948E311CC70FFBC81ABF8B2F359F88D" bold="true" box="[189,334,1841,1867]" pageId="3" pageNumber="4">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B833F03CC70FFBC81FCF8E3F33DF8BC" blockId="3.[189,1399,1888,2020]" box="[234,298,1888,1914]" pageId="3" pageNumber="4">
<emphasis id="B948E311CC70FFBC81FCF8E3F33DF8BC" bold="true" box="[234,298,1888,1914]" pageId="3" pageNumber="4">Male</emphasis>
</paragraph>
<paragraph id="8B833F03CC70FFB881ABF800F3C0FE13" blockId="3.[189,1399,1888,2020]" lastBlockId="7.[189,1399,267,470]" lastPageId="7" lastPageNumber="8" pageId="3" pageNumber="4">
CEPHALON. Lateral cephalic lobes broad, rounded, with anteroventral notch, anteroventral corner rounded. Eyes constricted in center, never divided into two parts (
<figureCitation id="13072386CC70FFBC8248F825F1A9F807" box="[862,958,1958,1985]" captionStart="Fig" captionStartId="4.[189,232,1930,1956]" captionTargetBox="[213,1374,317,1884]" captionTargetId="figure@4.[213,1374,317,1890]" captionTargetPageId="4" captionText="Fig. 2. Quadrivisio laleyei sp. nov., ♂, holotype, 7.5 mm, MNHN IU-2017-209. A. Habitus (scale 1). B. Lower lip (scale 3). C. First antenna (scale 2). D. Second antenna (scale 2). E. Left mandible (scale 4). F. Maxilliped palp (scale 3)." httpUri="https://zenodo.org/record/3258387/files/figure.png" pageId="3" pageNumber="4">Fig. 2A</figureCitation>
). Antenna 1 longer and more slender than antenna 2, sparsely setose (
<figureCitation id="13072386CC70FFBC8325F849F098F822" box="[563,655,1994,2020]" captionStart="Fig" captionStartId="4.[189,232,1930,1956]" captionTargetBox="[213,1374,317,1884]" captionTargetId="figure@4.[213,1374,317,1890]" captionTargetPageId="4" captionText="Fig. 2. Quadrivisio laleyei sp. nov., ♂, holotype, 7.5 mm, MNHN IU-2017-209. A. Habitus (scale 1). B. Lower lip (scale 3). C. First antenna (scale 2). D. Second antenna (scale 2). E. Left mandible (scale 4). F. Maxilliped palp (scale 3)." httpUri="https://zenodo.org/record/3258387/files/figure.png" pageId="3" pageNumber="4">Fig. 2C</figureCitation>
); peduncle much shorter than main flagellum, article 1 a shorter than article 2, article 3 very short (0.2 × length of the article 2); main flagellum with up to 38 articles unequal in length and in width; accessory flagellum well developed composed of six articles. Antenna 2 peduncle longer than the flagellum, article 4 and 5 similar in length (
<figureCitation id="13072386CC76FFBA8517FED1F676FEAA" box="[1025,1121,338,364]" captionStart="Fig" captionStartId="4.[189,232,1930,1956]" captionTargetBox="[213,1374,317,1884]" captionTargetId="figure@4.[213,1374,317,1890]" captionTargetPageId="4" captionText="Fig. 2. Quadrivisio laleyei sp. nov., ♂, holotype, 7.5 mm, MNHN IU-2017-209. A. Habitus (scale 1). B. Lower lip (scale 3). C. First antenna (scale 2). D. Second antenna (scale 2). E. Left mandible (scale 4). F. Maxilliped palp (scale 3)." httpUri="https://zenodo.org/record/3258387/files/figure.png" pageId="5" pageNumber="6">Fig. 2D</figureCitation>
); glandular cone on the peduncular article 2 prominent, reaching to end of peduncular article 3; flagellum with up to 15 articles. Lower lip with weakly-developed inner lobes, more pubescent outer lobes; mandibular lobes rather short and rounding (
<figureCitation id="13072386CC76FFBA80BFFE38F013FE13" box="[425,516,443,469]" captionStart="Fig" captionStartId="4.[189,232,1930,1956]" captionTargetBox="[213,1374,317,1884]" captionTargetId="figure@4.[213,1374,317,1890]" captionTargetPageId="4" captionText="Fig. 2. Quadrivisio laleyei sp. nov., ♂, holotype, 7.5 mm, MNHN IU-2017-209. A. Habitus (scale 1). B. Lower lip (scale 3). C. First antenna (scale 2). D. Second antenna (scale 2). E. Left mandible (scale 4). F. Maxilliped palp (scale 3)." httpUri="https://zenodo.org/record/3258387/files/figure.png" pageId="5" pageNumber="6">Fig. 2B</figureCitation>
). Mandibles asymmetrical. Left mandible incisor process dentate, with four chitinized teeth (
<figureCitation id="13072386CC76FFBA8094FE5DF3C9FE3F" box="[386,478,478,505]" captionStart="Fig" captionStartId="4.[189,232,1930,1956]" captionTargetBox="[213,1374,317,1884]" captionTargetId="figure@4.[213,1374,317,1890]" captionTargetPageId="4" captionText="Fig. 2. Quadrivisio laleyei sp. nov., ♂, holotype, 7.5 mm, MNHN IU-2017-209. A. Habitus (scale 1). B. Lower lip (scale 3). C. First antenna (scale 2). D. Second antenna (scale 2). E. Left mandible (scale 4). F. Maxilliped palp (scale 3)." httpUri="https://zenodo.org/record/3258387/files/figure.png" pageId="5" pageNumber="6">Fig. 2E</figureCitation>
).
<taxonomicName id="4C3C4480CC76FFBA80E5FE5CF0BDFE3F" box="[499,682,479,505]" class="Insecta" family="Drepanidae" genus="Lacinia" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="mobilis">
<emphasis id="B948E311CC76FFBA80E5FE5CF0BDFE3F" box="[499,682,479,505]" italics="true" pageId="5" pageNumber="6">Lacinia mobilis</emphasis>
</taxonomicName>
dentate, with four chitinized teeth, setal row with 6 plumose setae, molar process well developed and columnar, triturative. Palp article 1 shorter than article 2, article 2 slightly longer than article 3 with two setae on outer margin and two distal setae, article 3 rectolinear with two long terminal setae slightly longer than the article. Maxilla 1 and 2 do not present specific characteristics (not drawn). Maxilla 1 inner plate setose along entire inner margin and bearing a submarginal row of setae, outer plate bearing seven stout curved serrate setae, palp with distal article longer than proximal one, distal article with five terminal short stout setae and three subdistal slender setae on distal article. Maxilla 2 inner margin of inner plate of with double row of setae, unequal in length, and third oblique one on posterior face; outer plate with double rows of terminal and subterminal setae; inner and outer plates of maxilliped rather short (
<figureCitation id="13072386CC74FFB88255FEADF18BFE8F" box="[835,924,302,329]" captionStart="Fig" captionStartId="4.[189,232,1930,1956]" captionTargetBox="[213,1374,317,1884]" captionTargetId="figure@4.[213,1374,317,1890]" captionTargetPageId="4" captionText="Fig. 2. Quadrivisio laleyei sp. nov., ♂, holotype, 7.5 mm, MNHN IU-2017-209. A. Habitus (scale 1). B. Lower lip (scale 3). C. First antenna (scale 2). D. Second antenna (scale 2). E. Left mandible (scale 4). F. Maxilliped palp (scale 3)." httpUri="https://zenodo.org/record/3258387/files/figure.png" pageId="7" pageNumber="8">Fig. 2F</figureCitation>
); inner plate reaching to about middle of outer plate and bearing three terminal robust setae; outer plate reaching to about middle length of palp article 2, the ensemble of the distal margin and the distal part of the outer margin bear seven slender setae; palp article 3 slightly shorter than article 2, article 4 shorter than article 3 with a row of fine short setules on inner margin.
</paragraph>
<caption id="DF436F8BCC77FFBB81ABF809F01CF82D" httpUri="https://zenodo.org/record/3258387/files/figure.png" pageId="4" pageNumber="5" startId="4.[189,232,1930,1956]" targetBox="[213,1374,317,1884]" targetPageId="4">
<paragraph id="8B833F03CC77FFBB81ABF809F01CF82D" blockId="4.[189,1398,1930,2027]" pageId="4" pageNumber="5">
<emphasis id="B948E311CC77FFBB81ABF809F31BF862" bold="true" box="[189,268,1930,1956]" pageId="4" pageNumber="5">Fig. 2.</emphasis>
<taxonomicName id="4C3C4480CC77FFBB8003F809F3EFF862" authority="Gnohossou &amp; Piscart &amp; Umr, 2019" authorityName="Gnohossou &amp; Piscart &amp; Umr" authorityYear="2019" box="[277,504,1930,1956]" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="laleyei" status="sp. nov.">
<emphasis id="B948E311CC77FFBB8003F809F3EFF862" box="[277,504,1930,1956]" italics="true" pageId="4" pageNumber="5">Quadrivisio laleyei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B5E6ACC77FFBB8317F808F048F863" box="[513,607,1931,1957]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
, ♂, holotype, 7.5 mm, MNHN IU-2017-209.
<emphasis id="B948E311CC77FFBB8596F809F682F862" bold="true" box="[1152,1173,1930,1956]" pageId="4" pageNumber="5">A</emphasis>
. Habitus (scale 1).
<emphasis id="B948E311CC77FFBB81ABF82DF2C6F80E" bold="true" box="[189,209,1966,1992]" pageId="4" pageNumber="5">B</emphasis>
. Lower lip (scale 3).
<emphasis id="B948E311CC77FFBB80D2F82EF3CEF801" bold="true" box="[452,473,1965,1991]" pageId="4" pageNumber="5">C</emphasis>
. First antenna (scale 2).
<emphasis id="B948E311CC77FFBB83E5F82DF11FF80E" bold="true" box="[755,776,1966,1992]" pageId="4" pageNumber="5">D</emphasis>
. Second antenna (scale 2).
<emphasis id="B948E311CC77FFBB8554F82DF641F80E" bold="true" box="[1090,1110,1966,1992]" pageId="4" pageNumber="5">E</emphasis>
. Left mandible (scale 4).
<emphasis id="B948E311CC77FFBB81ABF852F2D8F82D" bold="true" box="[189,207,2001,2027]" pageId="4" pageNumber="5">F</emphasis>
. Maxilliped palp (scale 3).
</paragraph>
</caption>
<caption id="DF436F8BCC76FFBA81ABF82DF18FF82D" httpUri="https://zenodo.org/record/3258389/files/figure.png" pageId="5" pageNumber="6" startId="5.[189,232,1966,1992]" targetBox="[221,1389,593,1908]" targetPageId="5">
<paragraph id="8B833F03CC76FFBA81ABF82DF18FF82D" blockId="5.[189,1399,1966,2028]" pageId="5" pageNumber="6">
<emphasis id="B948E311CC76FFBA81ABF82DF319F80E" bold="true" box="[189,270,1966,1992]" pageId="5" pageNumber="6">Fig. 3.</emphasis>
<taxonomicName id="4C3C4480CC76FFBA800FF82DF3E8F80E" authority="Gnohossou &amp; Piscart &amp; Umr, 2019" authorityName="Gnohossou &amp; Piscart &amp; Umr" authorityYear="2019" box="[281,511,1966,1992]" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="laleyei" status="sp. nov.">
<emphasis id="B948E311CC76FFBA800FF82DF3E8F80E" box="[281,511,1966,1992]" italics="true" pageId="5" pageNumber="6">Quadrivisio laleyei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B5E6ACC76FFBA831CF82CF07EF80F" box="[522,617,1967,1993]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B948E311CC76FFBA8362F82DF09EF80E" bold="true" box="[628,649,1966,1992]" pageId="5" pageNumber="6">A</emphasis>
.,
<emphasis id="B948E311CC76FFBA83B5F82DF0AFF80E" bold="true" box="[675,696,1966,1992]" pageId="5" pageNumber="6">C</emphasis>
. Holotype, ♂, 7.5 mm, MNHN IU-2017-209, gnathopod.
<emphasis id="B948E311CC76FFBA81ABF851F2C6F82A" bold="true" box="[189,209,2002,2028]" pageId="5" pageNumber="6">B</emphasis>
.,
<emphasis id="B948E311CC76FFBA81F1F851F2EBF82A" bold="true" box="[231,252,2002,2028]" pageId="5" pageNumber="6">D</emphasis>
. Paratype, ♀, 7.0 mm, MNHN IU-2017-211, gnathopod.
</paragraph>
</caption>
<caption id="DF436F8BCC75FFB981ABF809F37BF82D" httpUri="https://zenodo.org/record/3258391/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,232,1930,1956]" targetBox="[323,1276,583,1890]" targetPageId="6">
<paragraph id="8B833F03CC75FFB981ABF809F37BF82D" blockId="6.[189,1398,1930,2027]" pageId="6" pageNumber="7">
<emphasis id="B948E311CC75FFB981ABF809F31EF862" bold="true" box="[189,265,1930,1956]" pageId="6" pageNumber="7">Fig. 4.</emphasis>
<taxonomicName id="4C3C4480CC75FFB98006F809F3F4F862" authority="Gnohossou &amp; Piscart &amp; Umr, 2019" authorityName="Gnohossou &amp; Piscart &amp; Umr" authorityYear="2019" box="[272,483,1930,1956]" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="laleyi" status="sp. nov.">
<emphasis id="B948E311CC75FFB98006F809F3F4F862" box="[272,483,1930,1956]" italics="true" pageId="6" pageNumber="7">Quadrivisio laleyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B5E6ACC75FFB980FFF809F054F862" box="[489,579,1930,1956]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B948E311CC75FFB9835FF809F097F862" bold="true" box="[585,640,1930,1956]" pageId="6" pageNumber="7">AB</emphasis>
.,
<emphasis id="B948E311CC75FFB98383F809F0DBF862" bold="true" box="[661,716,1930,1956]" pageId="6" pageNumber="7">DE</emphasis>
. Holotype, ♂, 7.5 mm, MNHN IU-2017-209.
<emphasis id="B948E311CC75FFB985F0F809F6ECF862" bold="true" box="[1254,1275,1930,1956]" pageId="6" pageNumber="7">C</emphasis>
. Paratype, ♂, 7.6 mm, MNHN IU-2017-210.
<emphasis id="B948E311CC75FFB9837AF82EF096F801" bold="true" box="[620,641,1965,1991]" pageId="6" pageNumber="7">A</emphasis>
. Pereopod 3.
<emphasis id="B948E311CC75FFB98224F82EF151F801" bold="true" box="[818,838,1965,1991]" pageId="6" pageNumber="7">B</emphasis>
. Pereopod 4.
<emphasis id="B948E311CC75FFB982E0F82EF61CF801" bold="true" box="[1014,1035,1965,1991]" pageId="6" pageNumber="7">C</emphasis>
. Pereopod 5.
<emphasis id="B948E311CC75FFB985ABF82EF6C5F801" bold="true" box="[1213,1234,1965,1991]" pageId="6" pageNumber="7">D</emphasis>
. Pereopod 6.
<emphasis id="B948E311CC75FFB981ABF853F2C6F82C" bold="true" box="[189,209,2000,2026]" pageId="6" pageNumber="7">E</emphasis>
. Pereopod 7.
</paragraph>
</caption>
<caption id="DF436F8BCC74FFB881ABF809F6EDF82D" httpUri="https://zenodo.org/record/3258393/files/figure.png" pageId="7" pageNumber="8" startId="7.[189,232,1930,1956]" targetBox="[212,1354,579,1875]" targetPageId="7">
<paragraph id="8B833F03CC74FFB881ABF809F6EDF82D" blockId="7.[189,1398,1930,2027]" pageId="7" pageNumber="8">
<emphasis id="B948E311CC74FFB881ABF809F31EF862" bold="true" box="[189,265,1930,1956]" pageId="7" pageNumber="8">Fig. 5.</emphasis>
<taxonomicName id="4C3C4480CC74FFB88006F809F3F4F862" authority="Gnohossou &amp; Piscart &amp; Umr, 2019" authorityName="Gnohossou &amp; Piscart &amp; Umr" authorityYear="2019" box="[272,483,1930,1956]" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="laleyi" status="sp. nov.">
<emphasis id="B948E311CC74FFB88006F809F3F4F862" box="[272,483,1930,1956]" italics="true" pageId="7" pageNumber="8">Quadrivisio laleyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B5E6ACC74FFB880FCF808F053F863" box="[490,580,1931,1957]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B948E311CC74FFB8835CF809F094F862" bold="true" box="[586,643,1930,1956]" pageId="7" pageNumber="8">AD</emphasis>
.,
<emphasis id="B948E311CC74FFB8838EF809F0BDF862" bold="true" box="[664,682,1930,1956]" pageId="7" pageNumber="8">F</emphasis>
. Holotype, ♂, 7.5 mm, MNHN IU-2017-209.
<emphasis id="B948E311CC74FFB885D5F809F6C0F862" bold="true" box="[1219,1239,1930,1956]" pageId="7" pageNumber="8">E</emphasis>
. Paratype, ♀, 7.0 mm, MNHN IU-2017-211.
<emphasis id="B948E311CC74FFB8833EF82EF02AF801" bold="true" box="[552,573,1965,1991]" pageId="7" pageNumber="8">A</emphasis>
. Male uropod 1 (scale 1).
<emphasis id="B948E311CC74FFB8827DF82DF168F80E" bold="true" box="[875,895,1966,1992]" pageId="7" pageNumber="8">B</emphasis>
. Male uropod 2 (scale 1).
<emphasis id="B948E311CC74FFB885BAF82EF6D6F801" bold="true" box="[1196,1217,1965,1991]" pageId="7" pageNumber="8">C</emphasis>
. Male urosome (scale 1).
<emphasis id="B948E311CC74FFB88026F852F352F82D" bold="true" box="[304,325,2001,2027]" pageId="7" pageNumber="8">D</emphasis>
. Male uropod 3 (scale 1).
<emphasis id="B948E311CC74FFB8836DF852F098F82D" bold="true" box="[635,655,2001,2027]" pageId="7" pageNumber="8">E</emphasis>
. Female uropod 3 (scale 1).
<emphasis id="B948E311CC74FFB882C8F852F1E7F82D" bold="true" box="[990,1008,2001,2027]" pageId="7" pageNumber="8">F</emphasis>
. Male telson (scale 2).
</paragraph>
</caption>
<paragraph id="8B833F03CC7BFFB781ABFE8FF362FCBA" blockId="8.[189,1399,267,892]" pageId="8" pageNumber="9">
THORAX. Coxae 14 equal in depth and deeper than coxa 5. Coxa 1 anteroventral corner not produced forward, lower margin evenly rounded with few small setae and bearing a weak notch horned with one seta at hind corner. Coxae 23 lower margin evenly rounded with few small setae, with notch at hind corner; a seta is inserted in that notch. Coxa 4 posteriorly excavate so as to fit the front margin of coxa 5. Coxae 57 bilobed. Coxa 7 smallest. Coxal gills simples on each pereopod. Gnathopod 1 more slender than gnathopod 2, subchelate (
<figureCitation id="13072386CC7BFFB783F0FE38F151FE13" box="[742,838,443,469]" captionStart="Fig" captionStartId="5.[189,232,1966,1992]" captionTargetBox="[221,1389,593,1908]" captionTargetId="figure@5.[189,1398,583,1927]" captionTargetPageId="5" captionText="Fig. 3. Quadrivisio laleyei sp. nov. A., C. Holotype, ♂, 7.5 mm, MNHN IU-2017-209, gnathopod. B., D. Paratype, ♀, 7.0 mm, MNHN IU-2017-211, gnathopod." httpUri="https://zenodo.org/record/3258389/files/figure.png" pageId="8" pageNumber="9">Fig. 3A</figureCitation>
); basis subequal in length to the depth of coxa 1; carpus subequal to propodus, with hind margin furnished with transverse rows of setae; propodus subrectangular, slightly narrowing distally, palm nearly transverse, very convex and undefined; dactylus curved to fit palm, very stout at base but attenuated toward apex. Gnathopod 2 robust, subchelate; basis shorter than propodus (
<figureCitation id="13072386CC7BFFB7831AFDCBF07FFDA4" box="[524,616,584,610]" captionStart="Fig" captionStartId="5.[189,232,1966,1992]" captionTargetBox="[221,1389,593,1908]" captionTargetId="figure@5.[189,1398,583,1927]" captionTargetPageId="5" captionText="Fig. 3. Quadrivisio laleyei sp. nov. A., C. Holotype, ♂, 7.5 mm, MNHN IU-2017-209, gnathopod. B., D. Paratype, ♀, 7.0 mm, MNHN IU-2017-211, gnathopod." httpUri="https://zenodo.org/record/3258389/files/figure.png" pageId="8" pageNumber="9">Fig. 3C</figureCitation>
); merus with quadrate posterodistal corner bearing two short setae; carpus compressed; propodus strong and stout, widest proximally, proximal hind margin with transverse group of setae, palm very oblique, defined proximally by a tuft of small and long setae and distally by a rounded process adjacent to dactylus hinge, bearing at least four stout short setae; dactylus stout, slightly curved proximally, apically subacute. Pereopods 34 subequal in length and morphologically similar (
<figureCitation id="13072386CC7BFFB78009FD7BF388FCD5" box="[287,415,760,787]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[323,1276,583,1890]" captionTargetId="figure@6.[309,1279,573,1893]" captionTargetPageId="6" captionText="Fig. 4. Quadrivisio laleyi sp. nov. AB., DE. Holotype, ♂, 7.5 mm, MNHN IU-2017-209. C. Paratype, ♂, 7.6 mm, MNHN IU-2017-210. A. Pereopod 3. B. Pereopod 4. C. Pereopod 5. D. Pereopod 6. E. Pereopod 7." httpUri="https://zenodo.org/record/3258391/files/figure.png" pageId="8" pageNumber="9">Fig. 4A, B</figureCitation>
), weakly setose, shorter than subsequent pereopods; basis not expanded. Pereopods 57 increasing length from P5 to P7; basis expanded with posterior margin convex and minutely serrate, with short setae, posterodistal corner subquadrate; merus, carpus and propodus armed with spines only (
<figureCitation id="13072386CC7BFFB781D0FCE1F336FCBA" box="[198,289,866,892]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[323,1276,583,1890]" captionTargetId="figure@6.[309,1279,573,1893]" captionTargetPageId="6" captionText="Fig. 4. Quadrivisio laleyi sp. nov. AB., DE. Holotype, ♂, 7.5 mm, MNHN IU-2017-209. C. Paratype, ♂, 7.6 mm, MNHN IU-2017-210. A. Pereopod 3. B. Pereopod 4. C. Pereopod 5. D. Pereopod 6. E. Pereopod 7." httpUri="https://zenodo.org/record/3258391/files/figure.png" pageId="8" pageNumber="9">Fig. 4C</figureCitation>
, DE).
</paragraph>
<paragraph id="8B833F03CC7BFFB781ABFC20F378FAD8" blockId="8.[189,1399,931,1310]" pageId="8" pageNumber="9">
ABDOMEN. Epimeron 1 posterior and ventral margins smooth. Epimera 23 with a small tooth at posterodistal angle. Urosome posterodorsal margins smooth. Urosomite 1 with 12 short dorsolateral setules (
<figureCitation id="13072386CC7BFFB78008FC6AF36BFBC2" box="[286,380,1001,1028]" captionStart="Fig" captionStartId="7.[189,232,1930,1956]" captionTargetBox="[212,1354,579,1875]" captionTargetId="figure@7.[189,1398,575,1888]" captionTargetPageId="7" captionText="Fig. 5. Quadrivisio laleyi sp. nov. AD., F. Holotype, ♂, 7.5 mm, MNHN IU-2017-209. E. Paratype, ♀, 7.0 mm, MNHN IU-2017-211. A. Male uropod 1 (scale 1). B. Male uropod 2 (scale 1). C. Male urosome (scale 1). D. Male uropod 3 (scale 1). E. Female uropod 3 (scale 1). F. Male telson (scale 2)." httpUri="https://zenodo.org/record/3258393/files/figure.png" pageId="8" pageNumber="9">Fig. 5C</figureCitation>
). Urosomite 2 with a dorsolateral robust setae (one on each side). Uropod 1 peduncle with a basofacial seta and a robust apical seta between rami, longer than rami, of which the inner is slightly the longer (
<figureCitation id="13072386CC7BFFB780B2FBB3F014FB8D" box="[420,515,1072,1099]" captionStart="Fig" captionStartId="7.[189,232,1930,1956]" captionTargetBox="[212,1354,579,1875]" captionTargetId="figure@7.[189,1398,575,1888]" captionTargetPageId="7" captionText="Fig. 5. Quadrivisio laleyi sp. nov. AD., F. Holotype, ♂, 7.5 mm, MNHN IU-2017-209. E. Paratype, ♀, 7.0 mm, MNHN IU-2017-211. A. Male uropod 1 (scale 1). B. Male uropod 2 (scale 1). C. Male urosome (scale 1). D. Male uropod 3 (scale 1). E. Female uropod 3 (scale 1). F. Male telson (scale 2)." httpUri="https://zenodo.org/record/3258393/files/figure.png" pageId="8" pageNumber="9">Fig. 5A</figureCitation>
). Uropod 2 the shortest uropod; peduncle slightly longer than inner ramus; inner ramus slightly longer than outer ramus (
<figureCitation id="13072386CC7BFFB783F9FBD0F158FBA8" box="[751,847,1107,1134]" captionStart="Fig" captionStartId="7.[189,232,1930,1956]" captionTargetBox="[212,1354,579,1875]" captionTargetId="figure@7.[189,1398,575,1888]" captionTargetPageId="7" captionText="Fig. 5. Quadrivisio laleyi sp. nov. AD., F. Holotype, ♂, 7.5 mm, MNHN IU-2017-209. E. Paratype, ♀, 7.0 mm, MNHN IU-2017-211. A. Male uropod 1 (scale 1). B. Male uropod 2 (scale 1). C. Male urosome (scale 1). D. Male uropod 3 (scale 1). E. Female uropod 3 (scale 1). F. Male telson (scale 2)." httpUri="https://zenodo.org/record/3258393/files/figure.png" pageId="8" pageNumber="9">Fig. 5B</figureCitation>
). Uropod 3 peduncle much shorter than rami (around ⅓ and ¼ of rami), reaching to end of inner ramus of uropod 2, rami subequal in length, broad, flat, edged with few short setae, terminally rounding and adorned with longer setae (
<figureCitation id="13072386CC7BFFB785A4FB1AF704FB72" box="[1202,1299,1177,1204]" captionStart="Fig" captionStartId="7.[189,232,1930,1956]" captionTargetBox="[212,1354,579,1875]" captionTargetId="figure@7.[189,1398,575,1888]" captionTargetPageId="7" captionText="Fig. 5. Quadrivisio laleyi sp. nov. AD., F. Holotype, ♂, 7.5 mm, MNHN IU-2017-209. E. Paratype, ♀, 7.0 mm, MNHN IU-2017-211. A. Male uropod 1 (scale 1). B. Male uropod 2 (scale 1). C. Male urosome (scale 1). D. Male uropod 3 (scale 1). E. Female uropod 3 (scale 1). F. Male telson (scale 2)." httpUri="https://zenodo.org/record/3258393/files/figure.png" pageId="8" pageNumber="9">Fig. 5D</figureCitation>
). Telson slightly shorter than uropod 3 peduncle, almost cleft to base, lobes widely divergent, apex truncated, with 23 short stout flagellated spines, outer margins with two sensorial setules, inner margins without setae (
<figureCitation id="13072386CC7BFFB7801EFA80F376FAD8" box="[264,353,1283,1310]" captionStart="Fig" captionStartId="7.[189,232,1930,1956]" captionTargetBox="[212,1354,579,1875]" captionTargetId="figure@7.[189,1398,575,1888]" captionTargetPageId="7" captionText="Fig. 5. Quadrivisio laleyi sp. nov. AD., F. Holotype, ♂, 7.5 mm, MNHN IU-2017-209. E. Paratype, ♀, 7.0 mm, MNHN IU-2017-211. A. Male uropod 1 (scale 1). B. Male uropod 2 (scale 1). C. Male urosome (scale 1). D. Male uropod 3 (scale 1). E. Female uropod 3 (scale 1). F. Male telson (scale 2)." httpUri="https://zenodo.org/record/3258393/files/figure.png" pageId="8" pageNumber="9">Fig. 5F</figureCitation>
).
</paragraph>
<paragraph id="8B833F03CC7BFFB781ABFAC6F30FFA99" blockId="8.[189,280,1349,1375]" box="[189,280,1349,1375]" pageId="8" pageNumber="9">
<heading id="D0CB886FCC7BFFB781ABFAC6F30FFA99" bold="true" box="[189,280,1349,1375]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B948E311CC7BFFB781ABFAC6F30FFA99" bold="true" box="[189,280,1349,1375]" pageId="8" pageNumber="9">Female</emphasis>
</heading>
</paragraph>
<paragraph id="8B833F03CC7BFFB781ABFAF0F10BFA31" blockId="8.[189,1399,1395,1527]" pageId="8" pageNumber="9">
ABDOMEN. Gnathopod 2 sexually dimorphic (
<figureCitation id="13072386CC7BFFB783D9FAF0F146FA48" box="[719,849,1395,1422]" captionStart="Fig" captionStartId="5.[189,232,1966,1992]" captionTargetBox="[221,1389,593,1908]" captionTargetId="figure@5.[189,1398,583,1927]" captionTargetPageId="5" captionText="Fig. 3. Quadrivisio laleyei sp. nov. A., C. Holotype, ♂, 7.5 mm, MNHN IU-2017-209, gnathopod. B., D. Paratype, ♀, 7.0 mm, MNHN IU-2017-211, gnathopod." httpUri="https://zenodo.org/record/3258389/files/figure.png" pageId="8" pageNumber="9">Fig. 3C, D</figureCitation>
); propodus comparatively much smaller; palm evenly and slightly convex, without either the protuberance near the hinge or the membranous cushion. Uropod 3 proportionally much shorter than males but much more spinose (
<figureCitation id="13072386CC7BFFB7854CFA3AF6F5FA12" box="[1114,1250,1465,1492]" captionStart="Fig" captionStartId="7.[189,232,1930,1956]" captionTargetBox="[212,1354,579,1875]" captionTargetId="figure@7.[189,1398,575,1888]" captionTargetPageId="7" captionText="Fig. 5. Quadrivisio laleyi sp. nov. AD., F. Holotype, ♂, 7.5 mm, MNHN IU-2017-209. E. Paratype, ♀, 7.0 mm, MNHN IU-2017-211. A. Male uropod 1 (scale 1). B. Male uropod 2 (scale 1). C. Male urosome (scale 1). D. Male uropod 3 (scale 1). E. Female uropod 3 (scale 1). F. Male telson (scale 2)." httpUri="https://zenodo.org/record/3258393/files/figure.png" pageId="8" pageNumber="9">Fig. 5D, E</figureCitation>
). Oostegites narrow and present from gnathopods to pereopod 6.
</paragraph>
</subSubSection>
<subSubSection id="C3266C88CC7BFFB781ABF99DF760F823" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="8B833F03CC7BFFB781ABF99DF3DEF9FE" blockId="8.[189,457,1566,1592]" box="[189,457,1566,1592]" pageId="8" pageNumber="9">
<heading id="D0CB886FCC7BFFB781ABF99DF3DEF9FE" bold="true" box="[189,457,1566,1592]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B948E311CC7BFFB781ABF99DF3DEF9FE" bold="true" box="[189,457,1566,1592]" pageId="8" pageNumber="9">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B833F03CC7BFFB781ABF9CEF00CF917" blockId="8.[189,1399,1613,1745]" pageId="8" pageNumber="9">
The genus
<taxonomicName id="4C3C4480CC7BFFB7802CF9CEF3D1F9A1" authorityName="Stebbing" authorityYear="1907" box="[314,454,1613,1639]" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B948E311CC7BFFB7802CF9CEF3D1F9A1" box="[314,454,1613,1639]" italics="true" pageId="8" pageNumber="9">Quadrivisio</emphasis>
</taxonomicName>
currently contains eight species:
<taxonomicName id="4C3C4480CC7BFFB78253F9CEF6D7F9A1" authority="(K. H. Barnard, 1940)" authorityName="K. H. Barnard" authorityYear="1940" baseAuthorityName="K. H. Barnard" baseAuthorityYear="1940" box="[837,1216,1613,1639]" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="aviceps">
<emphasis id="B948E311CC7BFFB78253F9CEF1A8F9A1" box="[837,959,1613,1639]" italics="true" pageId="8" pageNumber="9">Q. aviceps</emphasis>
(K.H.
<bibRefCitation id="EFAD42F2CC7BFFB7851BF9CEF6A0F9A1" author="Barnard K. H." box="[1037,1207,1613,1639]" pageId="8" pageNumber="9" pagination="381 - 543" refId="ref5493" refString="Barnard K. H. 1940. Contributions to the crustacean fauna of South Africa, XII. Further additions to the Tanaidacea, Isopoda and Amphipoda, together with keys for identification of the hitherto recorded marine and freshwater species. Annals of the South African Museum 32: 381 - 543." type="journal article" year="1940">Barnard, 1940</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="4C3C4480CC7BFFB785DCF9CEF36FF94C" authority="Stebbing, 1907" authorityName="Stebbing" authorityYear="1907" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bengalensis">
<emphasis id="B948E311CC7BFFB785DCF9CEF760F9A1" box="[1226,1399,1613,1639]" italics="true" pageId="8" pageNumber="9">Q. bengalensis</emphasis>
<bibRefCitation id="EFAD42F2CC7BFFB781ABF9F3F36FF94C" author="Stebbing T. R. R." box="[189,376,1648,1674]" pageId="8" pageNumber="9" pagination="119 - 123" refId="ref5973" refString="Stebbing T. R. R. 1907. The fauna of brackish ponds at Port Canning, Lower Bengal. Part V. Definition of a new genus of Amphipoda, and description of the typical species. Records of the Indian Museum 2: 119 - 123." type="journal article" year="1907">Stebbing, 1907</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C3C4480CC7BFFB7809DF9F3F194F94C" authority="Karaman &amp; Barnard, 1979" authorityName="Karaman &amp; Barnard" authorityYear="1979" box="[395,899,1648,1674]" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bousfieldi">
<emphasis id="B948E311CC7BFFB7809DF9F3F030F94C" box="[395,551,1648,1674]" italics="true" pageId="8" pageNumber="9">Q. bousfieldi</emphasis>
Karaman &amp; Barnard, 1979
</taxonomicName>
;
<taxonomicName id="4C3C4480CC7BFFB78280F9F3F765F94C" authority="Gordon &amp; Monod, 1968" authorityName="Gordon &amp; Monod" authorityYear="1968" box="[918,1394,1648,1674]" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="chevreuxi">
<emphasis id="B948E311CC7BFFB78280F9F3F626F94C" box="[918,1073,1648,1674]" italics="true" pageId="8" pageNumber="9">Q. chevreuxi</emphasis>
Gordon &amp; Monod, 1968
</taxonomicName>
;
<taxonomicName id="4C3C4480CC7BFFB781ABF917F3D1F96B" authority="Asari, 1983" authorityName="Asari" authorityYear="1983" box="[189,454,1683,1710]" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="lobata">
<emphasis id="B948E311CC7BFFB781ABF917F338F96B" box="[189,303,1683,1710]" italics="true" pageId="8" pageNumber="9">Q. lobata</emphasis>
Asari, 1983
</taxonomicName>
;
<taxonomicName id="4C3C4480CC7BFFB780C3F917F109F96B" authority="(Shoemaker, 1933)" authorityName="Shoemaker" authorityYear="1933" baseAuthorityName="Shoemaker" baseAuthorityYear="1933" box="[469,798,1683,1710]" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="lutzi">
<emphasis id="B948E311CC7BFFB780C3F917F038F96B" box="[469,559,1683,1710]" italics="true" pageId="8" pageNumber="9">Q. lutzi</emphasis>
(
<bibRefCitation id="EFAD42F2CC7BFFB78356F910F102F96B" author="Shoemaker C. R." box="[576,789,1683,1709]" pageId="8" pageNumber="9" pagination="1 - 24" refId="ref5949" refString="Shoemaker C. R. 1933. Amphipoda from Florida and the West Indies. American Museum Novitates 598: 1 - 24." type="journal article" year="1933">Shoemaker, 1933</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="4C3C4480CC7BFFB7823BF917F6D0F96B" authority="Hughes &amp; Kaji, 2016" authorityName="Hughes &amp; Kaji" authorityYear="2016" box="[813,1223,1683,1710]" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="meufong">
<emphasis id="B948E311CC7BFFB7823BF917F1AEF96B" box="[813,953,1683,1710]" italics="true" pageId="8" pageNumber="9">Q. meufong</emphasis>
<bibRefCitation id="EFAD42F2CC7BFFB782D5F910F6D0F96B" author="Hughes L. E. &amp; Kaji T." box="[963,1223,1683,1709]" pageId="8" pageNumber="9" pagination="351 - 359" refId="ref5671" refString="Hughes L. E., Kaji T. 2016. Description of a new species of Quadrivisio Stebbing, 1907, from Songkhla Lake, Thailand (Crustacea: Peracarida: Amphipoda: Maeridae). Raffles Bulletin of Zoology 64: 351 - 359." type="journal article" year="2016">Hughes &amp; Kaji, 2016</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C3C4480CC7BFFB78413F917F000F917" authority="Lowry &amp; Springthorpe, 2005" authorityName="Lowry &amp; Springthorpe" authorityYear="2005" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sarina">
<emphasis id="B948E311CC7BFFB78413F917F761F96B" box="[1285,1398,1684,1710]" italics="true" pageId="8" pageNumber="9">Q. sarina</emphasis>
Lowry &amp; Springthorpe, 2005
</taxonomicName>
.
</paragraph>
<paragraph id="8B833F03CC7BFFB781ABF97BF760F823" blockId="8.[189,1399,1784,2021]" pageId="8" pageNumber="9">
<taxonomicName id="4C3C4480CC7BFFB781ABF97BF3A6F8D4" box="[189,433,1784,1810]" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="lalaeyei">
<emphasis id="B948E311CC7BFFB781ABF97BF3A6F8D4" box="[189,433,1784,1810]" italics="true" pageId="8" pageNumber="9">Quadrivisio lalaeyei</emphasis>
</taxonomicName>
differs from most of the known species by several characteristics (see updated key of the genus after
<bibRefCitation id="EFAD42F2CC7BFFB780D2F898F0AAF8F0" author="Hughes L. E. &amp; Kaji T." box="[452,701,1819,1846]" pageId="8" pageNumber="9" pagination="351 - 359" refId="ref5671" refString="Hughes L. E., Kaji T. 2016. Description of a new species of Quadrivisio Stebbing, 1907, from Songkhla Lake, Thailand (Crustacea: Peracarida: Amphipoda: Maeridae). Raffles Bulletin of Zoology 64: 351 - 359." type="journal article" year="2016">Hughes &amp; Kaji 2016</bibRefCitation>
): the lack of carina on pleonites or urosome; the setation of the mandibular palp (
<taxonomicName id="4C3C4480CC7BFFB780AAF8BCF044F89E" authorityName="Karaman &amp; Barnard" authorityYear="1979" box="[444,595,1854,1881]" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bousfieldi">
<emphasis id="B948E311CC7BFFB780AAF8BCF044F89E" box="[444,595,1854,1881]" italics="true" pageId="8" pageNumber="9">Q. bousfieldi</emphasis>
</taxonomicName>
) and telson (
<taxonomicName id="4C3C4480CC7BFFB783E7F8BDF1B7F89E" authorityName="Stebbing" authorityYear="1907" box="[753,928,1854,1880]" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="12" phylum="Arthropoda" rank="species" species="bengalensis">
Q.
<emphasis id="B948E311CC7BFFB78200F8BDF1B7F89E" box="[790,928,1854,1880]" italics="true" pageId="8" pageNumber="9">bengalensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3C4480CC7BFFB782A6F8BCF637F89E" authorityName="Asari" authorityYear="1983" box="[944,1056,1854,1881]" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="lobata">
<emphasis id="B948E311CC7BFFB782A6F8BCF637F89E" box="[944,1056,1854,1881]" italics="true" pageId="8" pageNumber="9">Q. lobata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3C4480CC7BFFB7854AF8BCF6DBF89E" authorityName="Lowry &amp; Springthorpe" authorityYear="2005" box="[1116,1228,1855,1881]" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sarina">
<emphasis id="B948E311CC7BFFB7854AF8BCF6DBF89E" box="[1116,1228,1855,1881]" italics="true" pageId="8" pageNumber="9">Q. sarina</emphasis>
</taxonomicName>
); the shape of pereopod 7 (
<taxonomicName id="4C3C4480CC7BFFB78042F8E1F3D3F8BD" authorityName="Lowry &amp; Springthorpe" authorityYear="2005" box="[340,452,1890,1916]" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sarina">
<emphasis id="B948E311CC7BFFB78042F8E1F3D3F8BD" box="[340,452,1890,1916]" italics="true" pageId="8" pageNumber="9">Q. sarina</emphasis>
</taxonomicName>
), by the number of article on the accessory flagellum of antenna 1 (
<taxonomicName id="4C3C4480CC7BFFB785E4F8E1F778F8BD" baseAuthorityName="K. H. Barnard" baseAuthorityYear="1940" box="[1266,1391,1890,1916]" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="aviceps">
<emphasis id="B948E311CC7BFFB785E4F8E1F778F8BD" box="[1266,1391,1890,1916]" italics="true" pageId="8" pageNumber="9">Q. aviceps</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3C4480CC7BFFB781ABF806F352F859" authorityName="Hughes &amp; Kaji" authorityYear="2016" box="[189,325,1925,1951]" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="meufong">
<emphasis id="B948E311CC7BFFB781ABF806F352F859" box="[189,325,1925,1951]" italics="true" pageId="8" pageNumber="9">Q. meufong</emphasis>
</taxonomicName>
).
<taxonomicName id="4C3C4480CC7BFFB78048F806F059F859" box="[350,590,1925,1951]" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="lalaeyei">
<emphasis id="B948E311CC7BFFB78048F806F059F859" box="[350,590,1925,1951]" italics="true" pageId="8" pageNumber="9">Quadrivisio lalaeyei</emphasis>
</taxonomicName>
closely resembles to
<taxonomicName id="4C3C4480CC7BFFB7825FF806F188F859" baseAuthorityName="Shoemaker" baseAuthorityYear="1933" box="[841,927,1925,1951]" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="lutzi">
<emphasis id="B948E311CC7BFFB7825FF806F188F859" box="[841,927,1925,1951]" italics="true" pageId="8" pageNumber="9">Q. lutzi</emphasis>
</taxonomicName>
but differs by several characters such as the lack of carina on urosomite
<quantity id="4CC492E6CC7BFFB7832EF82BF071F804" box="[568,614,1960,1986]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="8" pageNumber="9" unit="in" value="2.0">2 in</quantity>
females or juveniles, the presence of the dorsolateral robust setae on urosomite 2, the shape of the eyes, the article 2 of the mandibular palp slightly longer than the third.
</paragraph>
</subSubSection>
<subSubSection id="C3266C88CC7AFFB681ABFE88F6E5FEE3" box="[189,1266,266,293]" pageId="9" pageNumber="10" type="key">
<paragraph id="8B833F03CC7AFFB681ABFE88F6E5FEE3" blockId="9.[189,1266,266,293]" box="[189,1266,266,293]" pageId="9" pageNumber="10">
<heading id="D0CB886FCC7AFFB681ABFE88F6E5FEE3" bold="true" box="[189,1266,266,293]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="B948E311CC7AFFB681ABFE88F6E5FEE3" bold="true" box="[189,1266,266,293]" pageId="9" pageNumber="10">
Identification key to known species of
<emphasis id="B948E311CC7AFFB6838FFE89F12FFEE3" bold="true" box="[665,824,266,293]" italics="true" pageId="9" pageNumber="10">
<taxonomicName id="4C3C4480CC7AFFB6838FFE89F130FEE3" authorityName="Stebbing" authorityYear="1907" box="[665,807,266,293]" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">Quadrivisio</taxonomicName>
(
</emphasis>
updated after
<bibRefCitation id="EFAD42F2CC7AFFB682FFFE88F6FEFEE3" author="Hughes L. E. &amp; Kaji T." box="[1001,1257,267,293]" pageId="9" pageNumber="10" pagination="351 - 359" refId="ref5671" refString="Hughes L. E., Kaji T. 2016. Description of a new species of Quadrivisio Stebbing, 1907, from Songkhla Lake, Thailand (Crustacea: Peracarida: Amphipoda: Maeridae). Raffles Bulletin of Zoology 64: 351 - 359." type="journal article" year="2016">Hughes &amp; Kaji 2016</bibRefCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3266C88CC7AFFB681ABFEB9F760FACC" pageId="9" pageNumber="10" type="key">
<key id="FFAD6832CC7AFFB681ABFEB9F760FACC" pageId="9" pageNumber="10">
<paragraph id="8B833F03CC7AFFB681ABFEB9F760FEB0" blockId="9.[189,1399,313,375]" pageId="9" pageNumber="10">
<keyStep id="30C827A6CC7AFFB681ABFEB9F760FEB0" pageId="9" pageNumber="10">
<keyLead id="30CD9C36CC7AFFB681ABFEB9F760FEB0" pageId="9" pageNumber="10">1. Carina absent on pleonites or urosome in males and females .......................................................... 2 Carina present on pleonites or urosome in males and/or females .................................................... 6</keyLead>
</keyStep>
</paragraph>
<keyStep id="30C827A6CC7AFFB681ABFE1DF760FE39" pageId="9" pageNumber="10">
<paragraph id="8B833F03CC7AFFB681ABFE1DF760FE1D" blockId="9.[189,1399,414,511]" pageId="9" pageNumber="10">
<keyLead id="30CD9C36CC7AFFB681ABFE1DF760FE1D" pageId="9" pageNumber="10">
2. Male gnathopod 2 propodus palm without posterodistal shelf ........................................................... ................................................................................................
<taxonomicName id="4C3C4480CC7AFFB682A1FE42F760FE1D" authority="Gordon &amp; Monod, 1968" authorityName="Gordon &amp; Monod" authorityYear="1968" box="[951,1399,449,476]" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="chevreuxi">
<emphasis id="B948E311CC7AFFB682A1FE42F645FE1A" bold="true" box="[951,1106,449,476]" italics="true" pageId="9" pageNumber="10">Q. chevreuxi</emphasis>
Gordon &amp; Monod, 1968
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B833F03CC7AFFB681D7FE66F760FE39" blockId="9.[189,1399,414,511]" box="[193,1399,484,511]" pageId="9" pageNumber="10">
<keyLead id="30CD9C36CC7AFFB681D7FE66F760FE39" box="[193,1399,484,511]" pageId="9" pageNumber="10"> Male gnathopod 2 propodus palm with posterodistal shelf .............................................................. 3</keyLead>
</paragraph>
</keyStep>
<paragraph id="8B833F03CC7AFFB681ABFDA5F760FDA5" blockId="9.[189,1399,550,612]" pageId="9" pageNumber="10">
<keyStep id="30C827A6CC7AFFB681ABFDA5F760FDA5" pageId="9" pageNumber="10">
<keyLead id="30CD9C36CC7AFFB681ABFDA5F760FDA5" pageId="9" pageNumber="10">
3. Male gnathopod 2 dactylus short, not reaching end of palm ..........
<taxonomicName id="4C3C4480CC7AFFB682EEFDA5F761FD86" authority="(K. H. Barnard, 1940)" authorityName="K. H. Barnard" authorityYear="1940" baseAuthorityName="K. H. Barnard" baseAuthorityYear="1940" box="[1016,1398,550,576]" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="aviceps">
<emphasis id="B948E311CC7AFFB682EEFDA5F664FD86" bold="true" box="[1016,1139,550,576]" italics="true" pageId="9" pageNumber="10">Q. aviceps</emphasis>
(K.H.
<bibRefCitation id="EFAD42F2CC7AFFB685D3FDA5F779FD86" author="Barnard K. H." box="[1221,1390,550,576]" pageId="9" pageNumber="10" pagination="381 - 543" refId="ref5493" refString="Barnard K. H. 1940. Contributions to the crustacean fauna of South Africa, XII. Further additions to the Tanaidacea, Isopoda and Amphipoda, together with keys for identification of the hitherto recorded marine and freshwater species. Annals of the South African Museum 32: 381 - 543." type="journal article" year="1940">Barnard, 1940</bibRefCitation>
)
</taxonomicName>
Male gnathopod 2 dactylus subequal to palm length ........................................................................ 4
</keyLead>
</keyStep>
</paragraph>
<keyStep id="30C827A6CC7AFFB681ABFD08F760FD2A" pageId="9" pageNumber="10">
<paragraph id="8B833F03CC7AFFB681ABFD08F760FD0E" blockId="9.[189,1399,650,748]" pageId="9" pageNumber="10">
<keyLead id="30CD9C36CC7AFFB681ABFD08F760FD0E" pageId="9" pageNumber="10">
4. Urosome and telson covered in dense setae; male gnathopod 2 propodus anterior margin produced into lobe .............................................................................................................
<taxonomicName id="4C3C4480CC7AFFB6856FFD2DF760FD0E" authority="Asari, 1983" authorityName="Asari" authorityYear="1983" box="[1145,1399,685,712]" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="lobata">
<emphasis id="B948E311CC7AFFB6856FFD2DF6FFFD0E" bold="true" box="[1145,1256,685,712]" italics="true" pageId="9" pageNumber="10">Q. lobata</emphasis>
Asari, 1983
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B833F03CC7AFFB681D7FD52F760FD2A" blockId="9.[189,1399,650,748]" box="[193,1399,721,748]" pageId="9" pageNumber="10">
<keyLead id="30CD9C36CC7AFFB681D7FD52F760FD2A" box="[193,1399,721,748]" pageId="9" pageNumber="10"> Urosome and telson without dense setae; male gnathopod 2 anterior margin not produced............ 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C827A6CC7AFFB681ABFC91F760FC50" pageId="9" pageNumber="10">
<paragraph id="8B833F03CC7AFFB681ABFC91F760FC96" blockId="9.[189,1399,786,919]" pageId="9" pageNumber="10">
<keyLead id="30CD9C36CC7AFFB681ABFC91F760FC96" pageId="9" pageNumber="10">
5. Posterior margin of pereopod 7 basis regularly convex, telson with robust setae along inner margins of each lobe
<emphasis id="B948E311CC7AFFB68095FCB5F1B5FC89" box="[387,930,822,847]" italics="true" pageId="9" pageNumber="10">..........................................................................</emphasis>
<taxonomicName id="4C3C4480CC7AFFB682BEFCB5F760FC96" authority="Lowry &amp; Springthorpe, 2005" authorityName="Lowry &amp; Springthorpe" authorityYear="2005" box="[936,1399,821,848]" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sarina">
<emphasis id="B948E311CC7AFFB682BEFCB5F60EFC96" bold="true" box="[936,1049,822,848]" italics="true" pageId="9" pageNumber="10">Q. sarina</emphasis>
Lowry &amp; Springthorpe, 2005
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B833F03CC7AFFB681D7FCDAF760FC50" blockId="9.[189,1399,786,919]" pageId="9" pageNumber="10">
<keyLead id="30CD9C36CC7AFFB681D7FCDAF760FC50" pageId="9" pageNumber="10">
Posterior margin of pereopod 7 basis not convex, telson without robust setae along inner margins of each lobe .................................................................................................................
<taxonomicName id="4C3C4480CC7AFFB685B5FCFFF702FC51" authority="Gnohossou &amp; Piscart &amp; Umr, 2019" authorityName="Gnohossou &amp; Piscart &amp; Umr" authorityYear="2019" box="[1187,1301,892,919]" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="laleyei" status="sp. nov.">
<emphasis id="B948E311CC7AFFB685B5FCFFF702FC51" bold="true" box="[1187,1301,892,919]" italics="true" pageId="9" pageNumber="10">Q. laleyei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27B5E6ACC7AFFB6840AFCFFF760FC50" box="[1308,1399,892,918]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<paragraph id="8B833F03CC7AFFB681ABFC3EF760FC3D" blockId="9.[189,1399,957,1019]" pageId="9" pageNumber="10">
<keyStep id="30C827A6CC7AFFB681ABFC3EF760FC3D" pageId="9" pageNumber="10">
<keyLead id="30CD9C36CC7AFFB681ABFC3EF760FC3D" pageId="9" pageNumber="10">
6. Telson with robust setae along inner margins of each lobe ...............
<taxonomicName id="4C3C4480CC7AFFB6851AFC3EF760FC11" authority="Stebbing, 1907" authorityName="Stebbing" authorityYear="1907" box="[1036,1399,957,984]" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="bengalensis">
<emphasis id="B948E311CC7AFFB6851AFC3EF6A9FC1E" bold="true" box="[1036,1214,957,984]" italics="true" pageId="9" pageNumber="10">Q. bengalensis</emphasis>
<bibRefCitation id="EFAD42F2CC7AFFB685D3FC3EF760FC11" author="Stebbing T. R. R." box="[1221,1399,957,983]" pageId="9" pageNumber="10" pagination="119 - 123" refId="ref5973" refString="Stebbing T. R. R. 1907. The fauna of brackish ponds at Port Canning, Lower Bengal. Part V. Definition of a new genus of Amphipoda, and description of the typical species. Records of the Indian Museum 2: 119 - 123." type="journal article" year="1907">Stebbing, 1907</bibRefCitation>
</taxonomicName>
Telson without setae along inner margin of each lobe ...................................................................... 7
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8B833F03CC7AFFB681ABFBA0F760FB99" blockId="9.[189,1399,1058,1120]" pageId="9" pageNumber="10">
<keyStep id="30C827A6CC7AFFB681ABFBA0F760FB99" pageId="9" pageNumber="10">
<keyLead id="30CD9C36CC7AFFB681ABFBA0F760FB99" pageId="9" pageNumber="10">
7. Epimeron 3 margins smooth....................................................................
<taxonomicName id="4C3C4480CC7AFFB68521FBA1F761FBFA" authority="(Shoemaker, 1933)" authorityName="Shoemaker" authorityYear="1933" baseAuthorityName="Shoemaker" baseAuthorityYear="1933" box="[1079,1398,1058,1085]" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="lutzi">
<emphasis id="B948E311CC7AFFB68521FBA1F698FBFB" bold="true" box="[1079,1167,1058,1085]" italics="true" pageId="9" pageNumber="10">Q. lutzi</emphasis>
(
<bibRefCitation id="EFAD42F2CC7AFFB68589FBA1F779FBFA" author="Shoemaker C. R." box="[1183,1390,1058,1084]" pageId="9" pageNumber="10" pagination="1 - 24" refId="ref5949" refString="Shoemaker C. R. 1933. Amphipoda from Florida and the West Indies. American Museum Novitates 598: 1 - 24." type="journal article" year="1933">Shoemaker, 1933</bibRefCitation>
)
</taxonomicName>
Epimeron 3 margins with several teeth ............................................................................................. 8
</keyLead>
</keyStep>
</paragraph>
<keyStep id="30C827A6CC7AFFB681ABFB04F760FACC" pageId="9" pageNumber="10">
<paragraph id="8B833F03CC7AFFB681ABFB04F760FB02" blockId="9.[189,1399,1159,1291]" pageId="9" pageNumber="10">
<keyLead id="30CD9C36CC7AFFB681ABFB04F760FB02" pageId="9" pageNumber="10">
8. Accessory flagellum with 7 articles; mandibular palp articles 23 without setae .............................. .............................................................................................
<taxonomicName id="4C3C4480CC7AFFB6828BFB29F760FB02" authority="Karaman &amp; Barnard, 1979" authorityName="Karaman &amp; Barnard" authorityYear="1979" box="[925,1399,1193,1220]" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="bousfieldi">
<emphasis id="B948E311CC7AFFB6828BFB29F621FB02" bold="true" box="[925,1078,1193,1220]" italics="true" pageId="9" pageNumber="10">Q. bousfieldi</emphasis>
Karaman &amp; Barnard, 1979
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B833F03CC7AFFB681D7FB4DF760FACC" blockId="9.[189,1399,1159,1291]" pageId="9" pageNumber="10">
<keyLead id="30CD9C36CC7AFFB681D7FB4DF760FACC" pageId="9" pageNumber="10">
Accessory flagellum with 910 articles; mandibular palp articles 23 with setae ............................. .......................................................................................................
<taxonomicName id="4C3C4480CC7AFFB682F3FB73F760FACC" authority="Hughes &amp; Haji, 2016" authorityName="Hughes &amp; Haji" authorityYear="2016" box="[997,1399,1264,1290]" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="meufong">
<emphasis id="B948E311CC7AFFB682F3FB73F662FACC" bold="true" box="[997,1141,1264,1290]" italics="true" pageId="9" pageNumber="10">Q. meufong</emphasis>
Hughes &amp; Haji, 2016
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
<subSubSection id="C3266C88CC7AFFB681ABFAB1F0B6F868" pageId="9" pageNumber="10" type="biology_ecology">
<paragraph id="8B833F03CC7AFFB681ABFAB1F3E5FA8D" blockId="9.[189,498,1329,1356]" box="[189,498,1329,1356]" pageId="9" pageNumber="10">
<heading id="D0CB886FCC7AFFB681ABFAB1F3E5FA8D" bold="true" box="[189,498,1329,1356]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="B948E311CC7AFFB681ABFAB1F3E5FA8D" bold="true" box="[189,498,1329,1356]" pageId="9" pageNumber="10">Ecology and distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B833F03CC7AFFB681ABFAE2F0B6F868" blockId="9.[189,1399,1376,1966]" pageId="9" pageNumber="10">
In the Nokoué lagoon,
<taxonomicName id="4C3C4480CC7AFFB680DBFAE2F057FABC" authorityName="Gnohossou &amp; Piscart &amp; Umr" authorityYear="2019" box="[461,576,1376,1403]" class="Malacostraca" family="Melitidae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="laleyei">
<emphasis id="B948E311CC7AFFB680DBFAE2F057FABC" box="[461,576,1376,1403]" italics="true" pageId="9" pageNumber="10">Q. laleyei</emphasis>
</taxonomicName>
was sampled periodically at all five sampling sites visited during this study, down to a water depth of at least
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(
<tableCitation id="C6BE0AB8CC7AFFB683F6FA00F172FA58" box="[736,869,1411,1438]" captionStart-0="Table 1" captionStart-1="Table 2" captionStartId-0="10.[189,258,268,294]" captionStartId-1="11.[189,258,268,294]" captionTargetBox-0="[204,1380,406,1934]" captionTargetBox-1="[313,1251,441,796]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="Table 1. Relationship between abiotic environmental variables and densities of the amphipod Quadrivisio laleyei sp. nov. sampled in 20022003 at five stations of the Nokoué lagoon (basket sampling method; mean of three replicates at each station)." captionText-1="Table 2. Mean abundances of the amphipod Quadrivisio laleyei sp. nov., sampled with four different methods at five stations of the Nokoué lagoon. Bundle (ind.bundle-1) and basket (ind.m-2) methods: 7 monthly samplings in 20022003, n = 3; Macrophytes (ind.macrophyte-1) and Surber (ind.m-2) methods: 2 sampling periods (February and September 2006), (n = 5)." httpUri-0="http://table.plazi.org/id/DF436F8BCC79FFB581ABFE8FF080FEAB" httpUri-1="http://table.plazi.org/id/DF436F8BCC78FFB481ABFE8FF1E4FE56" pageId="9" pageNumber="10" tableUuid-0="DF436F8BCC79FFB581ABFE8FF080FEAB" tableUuid-1="DF436F8BCC78FFB481ABFE8FF1E4FE56">Tables 12</tableCitation>
). Its potential habitat probably extends to the entire lagoon, with a preference for sandy bottoms located near to the entrance of the northern
<collectingRegion id="49F8F1E1CC7AFFB68434FA24F760FA07" box="[1314,1399,1447,1473]" country="Benin" name="Oueme" pageId="9" pageNumber="10">Ouémé</collectingRegion>
and Sô rivers. The levels of abundance observed were highly variable, and associated with the sampling methods (
<tableCitation id="C6BE0AB8CC7AFFB68023FA6EF380F9C1" box="[309,407,1517,1543]" captionStart="Table 2" captionStartId="11.[189,258,268,294]" captionTargetBox="[313,1251,441,796]" captionTargetPageId="11" captionText="Table 2. Mean abundances of the amphipod Quadrivisio laleyei sp. nov., sampled with four different methods at five stations of the Nokoué lagoon. Bundle (ind.bundle-1) and basket (ind.m-2) methods: 7 monthly samplings in 20022003, n = 3; Macrophytes (ind.macrophyte-1) and Surber (ind.m-2) methods: 2 sampling periods (February and September 2006), (n = 5)." httpUri="http://table.plazi.org/id/DF436F8BCC78FFB481ABFE8FF1E4FE56" pageId="9" pageNumber="10" tableUuid="DF436F8BCC78FFB481ABFE8FF1E4FE56">Table 2</tableCitation>
). The basket method showed the greatest sampling efficiency, principally when collectors were filled with local sediment (see
<bibRefCitation id="EFAD42F2CC7AFFB683FEF993F1A3F9ED" author="Gnohossou P." box="[744,948,1552,1579]" pageId="9" pageNumber="10" refId="ref5621" refString="Gnohossou P. 2006. La faune benthique d'une lagune ouest-africaine (le lac Nokoue au Benin), diversite, abondance, variations temporelles et spatiales, place dans la chaine trophique. PhD thesis, Institut National Polytechnique / Ecole Nationale Superieure Agronomique de Toulouse, France." type="book" year="2006">Gnohossou 2006</bibRefCitation>
), demonstrating the close relationship of the species to the substratum (endo-/epibenthic behaviour). Its highest abundance (133 ind.
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) was registered with this method at station Embouchure during the flood event of
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(salinity: 0.0 g.
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; oxygen concentration:
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.
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; depth:
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; sandy bottom). The few specimens collected with bundles (immersed
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below the lagoon surface) suggest this benthic species can also swim in the water column searching for food and new favourable habitats. Within the lagoon, this euryhaline amphipod was found to live in a wide range of salinities (between 0 and
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.
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), disappearing in southern high salinity areas during the low-water season. Furthermore, it shows some tolerance for pollution, as demonstrated by its presence in areas experiencing low oxygen concentration in the water column (down to
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.
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at station Ganvié in
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). It was also observed to have high abundance in the Ahémé and Porto-Novo lagoons during the flooding period (samples collected with Surber net and the macrophytes method).
</paragraph>
</subSubSection>
</treatment>
</document>