274 lines
34 KiB
XML
274 lines
34 KiB
XML
<document ID-DOI="10.5281/zenodo.163451" ID-GBIF-Dataset="f9566778-1df4-44d6-9a4c-849de39a4b6c" ID-GBIF-Taxon="124955159" ID-Zenodo-Dep="163451" checkinTime="1480975261095" checkinUser="plazi" docAuthor="Vincent J. Kalkman" docDate="2016" docId="03938796FFCEFFFB268D8C52FC3DFE4B" docLanguage="en" docName="10.5281zenodo.163451_Kalkman.pdf" docOrigin="Odonatologica 45 (3)" docStyle="DocumentStyle{}" docTitle="Celebargiolestes toli Vincent J. Kalkman 2016, sp. nov." docType="treatment" docVersion="11" lastPageId="21" lastPageNumber="256" masterDocId="FFAAFFEEFFDEFFEE27258934FFCBFF99" masterDocTitle="Revision of the genus Celebargiolestes Kennedy, 1925 (Odonata: Argiolestidae)" masterLastPageNumber="269" masterPageNumber="235" pageId="16" pageNumber="251" updateTime="1643502553511" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the genus Celebargiolestes Kennedy, 1925 (Odonata: Argiolestidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vincent J. Kalkman</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Odonatologica</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2016</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2016-12-01</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>45</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>3</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>235</mods:start>
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<mods:end>269</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.5281/zenodo.163451</mods:identifier>
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<mods:identifier type="GBIF-Dataset">f9566778-1df4-44d6-9a4c-849de39a4b6c</mods:identifier>
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<mods:identifier type="Zenodo-Dep">163451</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="124955159" LSID="urn:lsid:plazi:treatment:03938796FFCEFFFB268D8C52FC3DFE4B" httpUri="http://treatment.plazi.org/id/03938796FFCEFFFB268D8C52FC3DFE4B" lastPageId="21" lastPageNumber="256" pageId="16" pageNumber="251">
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<subSubSection box="[424,770,1382,1410]" pageId="16" pageNumber="251" type="nomenclature">
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||
<paragraph blockId="16.[159,1036,1382,1549]" box="[424,770,1382,1410]" pageId="16" pageNumber="251">
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<heading bold="true" box="[424,770,1382,1410]" centered="true" fontSize="11" level="2" pageId="16" pageNumber="251" reason="0">
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<emphasis bold="true" box="[424,770,1382,1410]" pageId="16" pageNumber="251">
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<taxonomicName authority="Vincent J. Kalkman, 2016" authorityName="Vincent J. Kalkman" authorityYear="2016" box="[424,670,1382,1410]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="251" phylum="Arthropoda" rank="species" species="toli" status="sp. nov.">
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<emphasis bold="true" box="[424,670,1382,1410]" italics="true" pageId="16" pageNumber="251">Celebargiolestes toli</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[677,770,1382,1410]" pageId="16" pageNumber="251" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[393,802,1416,1444]" pageId="16" pageNumber="251" type="description">
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||
<paragraph blockId="16.[159,1036,1382,1549]" box="[393,802,1416,1444]" pageId="16" pageNumber="251">
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<heading box="[393,802,1416,1444]" centered="true" fontSize="11" level="4" pageId="16" pageNumber="251" reason="1">
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(
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<figureCitation box="[399,532,1416,1444]" captionStart="Figures 7 – 14" captionStartId="13.[159,238,1384,1409]" captionTargetBox="[196,998,249,1341]" captionTargetId="figure@13.[196,998,249,1342]" captionTargetPageId="13" captionText="Figures 7 – 14. Head of Celebargiolestes males. 7 – C. cinctus (immature), Ranotongkor, 02 - viii- 1940, JvT 6476; 8 – C. cinctus (old adult), Mapanget, 23 - vi- 1940, JvT 6638; 9 – C. cinctus (‘ red legs’ form), 25 km NE of Luwuk, 30 - i- 1989, JvT 6670; 10 – C. askewi sp. nov., holotype; 11 – C. toli sp. nov., holotype; 12 – C. toli sp. nov., Peleng Island, 05 - viii- 1941, JvT 6453; 13 – C. orri sp. nov., holotype; 14 – C. orri sp. nov., 10 km WNW Palopo, 29 - x- 1993, JvT 11910." httpUri="https://zenodo.org/record/192855/files/figure.png" pageId="16" pageNumber="251">Figs 11–12</figureCitation>
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,
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<figureCitation box="[542,620,1416,1444]" captionStart="Figures 19 – 22" captionStartId="18.[159,238,1470,1495]" captionTargetBox="[212,983,244,1431]" captionTargetId="figure@18.[212,983,244,1431]" captionTargetPageId="18" captionText="Figures 19 – 22. Body of Celebargiolestes males. 19 – C. toli sp. nov., holotype; 20 – C. toli sp. nov., Peleng Island, 05 - viii- 1941, JvT 6453; 21 – C. orri sp. nov., holotype; 22 – C. orri sp. nov., 10 km WNW Palopo, 29 - x- 1993, JvT 11910." httpUri="https://zenodo.org/record/192857/files/figure.png" pageId="16" pageNumber="251">19–20</figureCitation>
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,
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<figureCitation box="[630,708,1416,1444]" captionStart="Figures 28 – 33" captionStartId="26.[159,237,1353,1378]" captionTargetBox="[156,1036,247,1313]" captionTargetId="figure@26.[156,1036,247,1313]" captionTargetPageId="26" captionText="Figures 28 – 33. Genital ligula of Celebargiolestes males. 28 – C. toli sp. nov., Gunung Watuwila, 02 − 04 - xi- 1989, JvT 6636, lateral view, lateral lobe deformed and bent inwards; 29 – C. toli sp. nov., Gunung Watuwila, 02 − 04 - xi- 1989, JvT 6636, ventral view; 30 – C. toli sp. nov., Banggai Archipelago, Peleng Island, 05 - viii- 1941, JvT 6501, lateral view; 31 – C. toli sp. nov., Banggai Archipelago, Peleng Island, 05 - viii- 1941, JvT 6501, ventral view; 32 – C. orri sp. nov., Lore Lindu, 17 - xii- 1985, JvT 6444, lateral view; 33 – C. orri sp. nov., Lore Lindu, 17 - xii- 1985, JvT 6444, ventral view." httpUri="https://zenodo.org/record/192859/files/figure.png" pageId="16" pageNumber="251">28–31</figureCitation>
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,
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<figureCitation box="[718,795,1416,1444]" captionStart="Figures 40 – 45" captionStartId="28.[159,238,1442,1467]" captionTargetBox="[156,1039,522,1404]" captionTargetId="figure@28.[156,1039,522,1404]" captionTargetPageId="28" captionText="Figures 40 – 45. Appendages of Celebargiolestes males. 40 – C. toli sp. nov., holotype, dorsal view; 41 – C. toli sp. nov., holotype, lateral view; 42 – C. toli sp. nov., holotype, ventral view; 43 – C. orri sp. nov., holotype, dorsal view; 44 – C. orri sp. nov., holotype, lateral view; 45 – C. orri sp. nov., JvT 6646, ventral view." httpUri="https://zenodo.org/record/192861/files/figure.png" pageId="16" pageNumber="251">40–42</figureCitation>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="17" lastPageNumber="252" pageId="16" pageNumber="251" type="materials_examined">
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<paragraph blockId="16.[159,1036,1382,1549]" box="[159,360,1462,1488]" pageId="16" pageNumber="251">
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<heading bold="true" box="[159,360,1462,1488]" fontSize="11" level="3" pageId="16" pageNumber="251" reason="0">
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<emphasis bold="true" box="[159,360,1462,1488]" pageId="16" pageNumber="251">Material studied</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="16.[159,1036,1382,1549]" lastBlockId="17.[159,1036,253,1013]" lastPageId="17" lastPageNumber="252" pageId="16" pageNumber="251">
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<materialsCitation ID-GBIF-Occurrence="1413786763" collectedFrom="small and shaded stream in very steep valley, clear running water over boulders, small pools" collectingDate="1989-11-04" collectionCode="RMNH" collectorName="J. van Tol" country="Indonesia" elevation="1050m" lastPageId="17" lastPageNumber="252" latitude="-3.8166666" location="Gunung Watuwila" longLatPrecision="1307" longitude="121.666664" municipality="south of Sanggona" pageId="16" pageNumber="251" specimenCode="89JvT035" specimenCount="1" specimenCount-male="1" stateProvince="Sulawesi Tenggara" typeStatus="holotype">
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<typeStatus box="[159,263,1494,1518]" pageId="16" pageNumber="251">
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<emphasis bold="true" box="[159,263,1494,1518]" pageId="16" pageNumber="251">Holotype</emphasis>
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</typeStatus>
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<specimenCount box="[273,293,1495,1518]" pageId="16" pageNumber="251" type="male">♂</specimenCount>
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.
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<collectingCountry box="[310,417,1494,1518]" name="Indonesia" pageId="16" pageNumber="251">Indonesia</collectingCountry>
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,
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<collectingRegion box="[432,634,1494,1519]" country="Indonesia" name="Sulawesi Tenggara" pageId="16" pageNumber="251">Sulawesi Tenggara</collectingRegion>
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,
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<collectingMunicipality box="[648,856,1494,1518]" pageId="16" pageNumber="251">south of Sanggona</collectingMunicipality>
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,
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<location LSID="urn:lsid:plazi:treatment:03938796FFCEFFFB268D8C52FC3DFE4B:8EE5605BFFCEFFFE24428CE2FF06F994" country="Indonesia" latitude="-3.8166666" longLatPrecision="1307" longitude="121.666664" municipality="south of Sanggona" name="Gunung Watuwila" pageId="16" pageNumber="251" stateProvince="Sulawesi Tenggara">
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Gunung
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<taxonomicName authority=", Sungai Lalonduwasi" authorityName="Sungai Lalonduwasi" class="Insecta" family="Chlorocyphidae" genus="Watuwila" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="251" phylum="Arthropoda" rank="genus">
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Watuwila,
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<location LSID="urn:lsid:plazi:treatment:03938796FFCEFFFB268D8C52FC3DFE4B:8EE5605BFFCEFFFE27FC8CC1FD6AF995" box="[217,673,1525,1549]" country="Indonesia" latitude="-3.8166666" longLatPrecision="1307" longitude="121.666664" municipality="south of Sanggona" name="Sungai Lalonduwasi near Centipede camp" pageId="16" pageNumber="251" stateProvince="Sulawesi Tenggara">Sungai Lalonduwasi near Centipede camp</location>
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</taxonomicName>
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</location>
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,
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<collectedFrom lastPageId="17" lastPageNumber="252" pageId="16" pageNumber="251">small and shaded stream in very steep valley, clear running water over boulders, small pools</collectedFrom>
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,
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<elevation box="[814,903,253,277]" pageId="17" pageNumber="252">1 050 m</elevation>
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(
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<emphasis box="[920,943,254,277]" italics="true" pageId="17" pageNumber="252">ca</emphasis>
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<geoCoordinate box="[951,1031,253,277]" direction="south" orientation="latitude" pageId="17" pageNumber="252" precision="925" value="-3.8166666">03°49’S</geoCoordinate>
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,
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<geoCoordinate box="[159,257,284,308]" direction="east" orientation="longitude" pageId="17" pageNumber="252" precision="925" value="121.666664">121°40’E</geoCoordinate>
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), sample
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<specimenCode box="[355,458,284,308]" pageId="17" pageNumber="252">89JvT035</specimenCode>
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, 02−
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<collectingDate box="[510,627,284,308]" pageId="17" pageNumber="252" value="1989-11-04">
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<date box="[510,627,284,308]" pageId="17" pageNumber="252" value="1989-11-04">04-xi-1989</date>
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</collectingDate>
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,
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<collectorName box="[637,735,284,308]" pageId="17" pageNumber="252">J. van Tol</collectorName>
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leg. (
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<collectionCode box="[792,879,284,308]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum ("Naturalis") [formerly Rijksmuseum van Natuurlijke Historie]" pageId="17" pageNumber="252">RMNH</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph blockId="17.[159,1036,253,1013]" box="[159,344,314,339]" pageId="17" pageNumber="252">
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<emphasis bold="true" box="[159,344,314,339]" pageId="17" pageNumber="252">Further material</emphasis>
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</paragraph>
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<paragraph blockId="17.[159,1036,253,1013]" box="[159,483,345,369]" pageId="17" pageNumber="252">
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<emphasis bold="true" box="[159,483,345,369]" pageId="17" pageNumber="252">Sulawesi Tenggara, mainland</emphasis>
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</paragraph>
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<paragraph blockId="17.[159,1036,253,1013]" pageId="17" pageNumber="252">
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4♂, S of Sanggona, Mokowu river near Mokowu camp, foot of
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<taxonomicName authority="Mts" authorityName="Mts" box="[823,972,376,400]" class="Insecta" family="Chlorocyphidae" genus="Watuwila" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="252" phylum="Arthropoda" rank="genus">Watuwila Mts</taxonomicName>
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, rivulet with ponded areas, clear water, bottom with boulders, some sand, largely shad- ed, 150 m a.s.l., 29−31-x-1989, J. van Tol leg.; 18♂ 3♀, S of Sanggona, Gunung
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<taxonomicName authority=", Sg. Lalonduwasi" authorityName="Sg. Lalonduwasi" box="[159,453,468,492]" class="Insecta" family="Chlorocyphidae" genus="Watuwila" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="252" phylum="Arthropoda" rank="genus">Watuwila, Sg. Lalonduwasi</taxonomicName>
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near Centipede camp, small, shaded stream in very steep valley, clear running water over boulders, small pools, 1 0 50 m a.s.l., 02−04- -xi-1989, J. van Tol leg.; 2♂ 1♀, NE of Kolaka, near Gunung Watowila, Centipede Camp, undisturbed hilly rainforest/narrow stream, 1 100 m a.s.l., 02−05-v-1989, J.P. Duffels leg.
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</paragraph>
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<paragraph blockId="17.[159,1036,253,1013]" box="[159,586,621,645]" pageId="17" pageNumber="252">
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<emphasis bold="true" box="[159,586,621,645]" pageId="17" pageNumber="252">Sulawesi Tengah, Banggai Archipelago</emphasis>
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</paragraph>
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<paragraph blockId="17.[159,1036,253,1013]" box="[159,1036,652,676]" pageId="17" pageNumber="252">4♂, Banggai Archipelago, Peleng [Peleng Island], 05-viii-1941, J.J. van der Starre leg.</paragraph>
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<paragraph blockId="17.[159,1036,253,1013]" box="[159,341,682,706]" pageId="17" pageNumber="252">
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<emphasis bold="true" box="[159,341,682,706]" pageId="17" pageNumber="252">Sulawesi Selatan</emphasis>
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</paragraph>
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<paragraph blockId="17.[159,1036,253,1013]" pageId="17" pageNumber="252">
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1♂, Mountain Tangke Salokko, 1 500 m a.s.l., 15-xi-1931−01-i-1932, G. Heinrich leg.; 1♂,
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<emphasis box="[260,283,745,768]" italics="true" pageId="17" pageNumber="252">ca</emphasis>
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55km over the road N of Wotu, brook with rather quietly flowing water, nearly fully shaded in the forest, but also collected on open spots near the road, ponded sites, bottom with enormous boulders, with pebbles and coarse sand, 920m, 29-iv-1991, J. van Tol leg.; 4♂,
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<emphasis box="[567,590,837,860]" italics="true" pageId="17" pageNumber="252">ca</emphasis>
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40 km over the road N of Wotu, brooklet on steep slope near crossing with road, some ponded sites, full shade by hardly disturbed primary rain forest, width 50−100 cm, clear water, 500 m a.s.l., 29-iv-1991, J. van Tol leg.; 7♂ 2♀, 30 km N of Wotu, Sg Anoa, river through primary forest, waterfall near bridge Wotu-Tentena, upstream ponded sites, small tributaries, boulders, sand, 650 m a.s.l., 23-x-1993, J. van Tol leg.
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</paragraph>
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</subSubSection>
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<subSubSection box="[159,1027,1068,1131]" pageId="17" pageNumber="252" type="etymology">
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<paragraph blockId="17.[159,1027,1068,1131]" box="[159,292,1068,1094]" pageId="17" pageNumber="252">
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<heading bold="true" box="[159,292,1068,1094]" fontSize="11" level="3" pageId="17" pageNumber="252" reason="0">
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<emphasis bold="true" box="[159,292,1068,1094]" pageId="17" pageNumber="252">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="17.[159,1027,1068,1131]" box="[159,1027,1105,1131]" pageId="17" pageNumber="252">The species is dedicated to Jan van Tol (masculine singular genitive noun).</paragraph>
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</subSubSection>
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<subSubSection lastPageId="20" lastPageNumber="255" pageId="17" pageNumber="252" type="description">
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<paragraph blockId="17.[159,1036,1223,1399]" box="[159,349,1223,1250]" pageId="17" pageNumber="252">
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<heading box="[159,349,1223,1250]" fontSize="11" level="5" pageId="17" pageNumber="252" reason="1">
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<emphasis bold="true" box="[159,220,1224,1250]" pageId="17" pageNumber="252">Male</emphasis>
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(holotype)
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</heading>
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</paragraph>
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<paragraph blockId="17.[159,1036,1223,1399]" pageId="17" pageNumber="252">
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Head − Labium completely yellow. Rear of head of normal shape. Front of head, including labrum, mandibles, clypeus and sockets of antennae yelloworange (
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<figureCitation box="[254,337,1335,1362]" captionStart="Figures 7 – 14" captionStartId="13.[159,238,1384,1409]" captionTargetBox="[196,998,249,1341]" captionTargetId="figure@13.[196,998,249,1342]" captionTargetPageId="13" captionText="Figures 7 – 14. Head of Celebargiolestes males. 7 – C. cinctus (immature), Ranotongkor, 02 - viii- 1940, JvT 6476; 8 – C. cinctus (old adult), Mapanget, 23 - vi- 1940, JvT 6638; 9 – C. cinctus (‘ red legs’ form), 25 km NE of Luwuk, 30 - i- 1989, JvT 6670; 10 – C. askewi sp. nov., holotype; 11 – C. toli sp. nov., holotype; 12 – C. toli sp. nov., Peleng Island, 05 - viii- 1941, JvT 6453; 13 – C. orri sp. nov., holotype; 14 – C. orri sp. nov., 10 km WNW Palopo, 29 - x- 1993, JvT 11910." httpUri="https://zenodo.org/record/192855/files/figure.png" pageId="17" pageNumber="252">Fig. 11</figureCitation>
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). Basal two segments of antennae yellow-orange, remainder brown. Top of head black.
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</paragraph>
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<paragraph blockId="17.[159,1036,1453,1554]" lastBlockId="19.[159,1036,253,354]" lastPageId="19" lastPageNumber="254" pageId="17" pageNumber="252">
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Thorax − Sides of prothorax dull orange mottled with black; pronotum dull orange areas with median parts of anterior, middle and posterior lobe blackish. Posterior lobe of pronotum broad, seen from lateral gently sloping and with the outer border steeply raised, lateral corners rounded. Pattern of synthorax as in
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<figureCitation box="[448,566,291,317]" captionStart="Figures 19 – 22" captionStartId="18.[159,238,1470,1495]" captionTargetBox="[212,983,244,1431]" captionTargetId="figure@18.[212,983,244,1431]" captionTargetPageId="18" captionText="Figures 19 – 22. Body of Celebargiolestes males. 19 – C. toli sp. nov., holotype; 20 – C. toli sp. nov., Peleng Island, 05 - viii- 1941, JvT 6453; 21 – C. orri sp. nov., holotype; 22 – C. orri sp. nov., 10 km WNW Palopo, 29 - x- 1993, JvT 11910." httpUri="https://zenodo.org/record/192857/files/figure.png" pageId="19" pageNumber="254">Figure 19</figureCitation>
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; darker areas black, pale areas yellowbrown.
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</paragraph>
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<caption httpUri="https://zenodo.org/record/192857/files/figure.png" pageId="18" pageNumber="253" targetBox="[212,983,244,1431]" targetPageId="18">
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<paragraph blockId="18.[159,1036,1469,1553]" pageId="18" pageNumber="253">
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<emphasis bold="true" box="[159,318,1470,1495]" pageId="18" pageNumber="253">Figures 19–22.</emphasis>
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Body of
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<taxonomicName authorityName="Kennedy" authorityYear="1925" box="[411,587,1469,1493]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="253" phylum="Arthropoda" rank="genus">
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<emphasis box="[411,587,1469,1493]" italics="true" pageId="18" pageNumber="253">Celebargiolestes</emphasis>
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</taxonomicName>
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males. 19 –
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<taxonomicName authority="Vincent J. Kalkman, 2016" authorityName="Vincent J. Kalkman" authorityYear="2016" box="[721,782,1469,1493]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="253" phylum="Arthropoda" rank="species" species="toli" status="sp. nov.">
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<emphasis box="[721,782,1469,1493]" italics="true" pageId="18" pageNumber="253">C. toli</emphasis>
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||
</taxonomicName>
|
||
<taxonomicNameLabel box="[787,873,1469,1494]" pageId="18" pageNumber="253" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype; 20 –
|
||
<taxonomicName authority="Vincent J. Kalkman, 2016" authorityName="Vincent J. Kalkman" authorityYear="2016" box="[159,220,1499,1523]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="253" phylum="Arthropoda" rank="species" species="toli" status="sp. nov.">
|
||
<emphasis box="[159,220,1499,1523]" italics="true" pageId="18" pageNumber="253">C. toli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[226,310,1500,1523]" pageId="18" pageNumber="253" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, Peleng Island, 05-viii-1941, JvT6453; 21 –
|
||
<taxonomicName authority="Vincent J. Kalkman, 2016" authorityName="Vincent J. Kalkman" authorityYear="2016" box="[767,832,1499,1523]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="253" phylum="Arthropoda" rank="species" species="orri" status="sp. nov.">
|
||
<emphasis box="[767,832,1499,1523]" italics="true" pageId="18" pageNumber="253">C. orri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[838,924,1499,1524]" pageId="18" pageNumber="253" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype; 22 –
|
||
<taxonomicName authority="Vincent J. Kalkman, 2016" authorityName="Vincent J. Kalkman" authorityYear="2016" box="[211,276,1528,1552]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="253" phylum="Arthropoda" rank="species" species="orri" status="sp. nov.">
|
||
<emphasis box="[211,276,1528,1552]" italics="true" pageId="18" pageNumber="253">C. orri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[282,367,1530,1553]" pageId="18" pageNumber="253" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, 10 km WNW Palopo, 29-x-1993, JvT11910.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="19.[159,1036,403,615]" pageId="19" pageNumber="254">Legs − Coxae and trochanters dull orange-yellow. Femora yellow with two broad but faint black rings, knees black, tibiae and tarsi yellow-brown. Spines pale orange-brown; femora with respectively 9–10, 9–10, 14–16 spines laterally; tibiae of fore legs with 11–12 smaller and 6 larger spines laterally; tibiae of mid and hind legs with 12–13 and 13 spines respectively laterally.</paragraph>
|
||
<paragraph blockId="19.[159,1036,664,877]" pageId="19" pageNumber="254">Wings − Venation black. Membrane hyaline with a smoky outer border. Fw and Hw with 2 Ax; Fw with 34 Px and Hw with 32 Px. Ac at about ¼ the distance between Ax1 and Ax2. Brace vein absent. Pterostigma dark brown. Up to three rows of cells between anal vein and border of Fw and up to four in Hw. Fields between Ir2 and R3, between R3 and IR3, and between IR3 and R4 at distal end containing 3 or more rows of cells.</paragraph>
|
||
<paragraph blockId="19.[159,1036,925,1362]" pageId="19" pageNumber="254">
|
||
Abdomen − Pattern on abdomen as in
|
||
<figureCitation box="[641,757,925,952]" captionStart="Figures 19 – 22" captionStartId="18.[159,238,1470,1495]" captionTargetBox="[212,983,244,1431]" captionTargetId="figure@18.[212,983,244,1431]" captionTargetPageId="18" captionText="Figures 19 – 22. Body of Celebargiolestes males. 19 – C. toli sp. nov., holotype; 20 – C. toli sp. nov., Peleng Island, 05 - viii- 1941, JvT 6453; 21 – C. orri sp. nov., holotype; 22 – C. orri sp. nov., 10 km WNW Palopo, 29 - x- 1993, JvT 11910." httpUri="https://zenodo.org/record/192857/files/figure.png" pageId="19" pageNumber="254">Figure 19</figureCitation>
|
||
; dark pattern elements dark brown to black, pale pattern elements yellow-brown. Anterior half of S1 yellow, posterior half dark brown, S2 with dorsum black and sides pale brown, S3–7 dorsally largely dark, ventral and the lower lateral areas pale brown, anteriorly on each segment the pale brown reaches to the dorsum forming a ring covering approximately ¹/₅ of the segment. S9–10 black, the latter strongly depressed in the middle, hind border of S10 without modifications. Appendages as shown in
|
||
<figureCitation box="[570,738,1187,1213]" captionStart="Figures 40 – 45" captionStartId="28.[159,238,1442,1467]" captionTargetBox="[156,1039,522,1404]" captionTargetId="figure@28.[156,1039,522,1404]" captionTargetPageId="28" captionText="Figures 40 – 45. Appendages of Celebargiolestes males. 40 – C. toli sp. nov., holotype, dorsal view; 41 – C. toli sp. nov., holotype, lateral view; 42 – C. toli sp. nov., holotype, ventral view; 43 – C. orri sp. nov., holotype, dorsal view; 44 – C. orri sp. nov., holotype, lateral view; 45 – C. orri sp. nov., JvT 6646, ventral view." httpUri="https://zenodo.org/record/192861/files/figure.png" pageId="19" pageNumber="254">Figures 40–42</figureCitation>
|
||
; brown with only base of lower appendages yellow. Lower appendages long, about half the length of upper appendages. Genital ligula as in
|
||
<figureCitation box="[608,775,1261,1288]" captionStart="Figures 28 – 33" captionStartId="26.[159,237,1353,1378]" captionTargetBox="[156,1036,247,1313]" captionTargetId="figure@26.[156,1036,247,1313]" captionTargetPageId="26" captionText="Figures 28 – 33. Genital ligula of Celebargiolestes males. 28 – C. toli sp. nov., Gunung Watuwila, 02 − 04 - xi- 1989, JvT 6636, lateral view, lateral lobe deformed and bent inwards; 29 – C. toli sp. nov., Gunung Watuwila, 02 − 04 - xi- 1989, JvT 6636, ventral view; 30 – C. toli sp. nov., Banggai Archipelago, Peleng Island, 05 - viii- 1941, JvT 6501, lateral view; 31 – C. toli sp. nov., Banggai Archipelago, Peleng Island, 05 - viii- 1941, JvT 6501, ventral view; 32 – C. orri sp. nov., Lore Lindu, 17 - xii- 1985, JvT 6444, lateral view; 33 – C. orri sp. nov., Lore Lindu, 17 - xii- 1985, JvT 6444, ventral view." httpUri="https://zenodo.org/record/192859/files/figure.png" pageId="19" pageNumber="254">Figures 28–31</figureCitation>
|
||
, with shallow and apically broad cleft. Lateral lobes of ligula large with rounded tip but not constricted in the middle.
|
||
</paragraph>
|
||
<paragraph blockId="19.[159,1036,1410,1512]" pageId="19" pageNumber="254">Measurements [mm] − Total length 49, abdomen 39, Fw 36; Fw pterostigma 1.7 (costal length), 2.3 (longest length); Hw pterostigma 1.9 (costal length), 2.5 (longest length).</paragraph>
|
||
<paragraph blockId="20.[159,1036,253,803]" box="[159,289,253,279]" pageId="20" pageNumber="255">
|
||
<heading bold="true" box="[159,289,253,279]" fontSize="11" level="3" pageId="20" pageNumber="255" reason="0">
|
||
<emphasis bold="true" box="[159,289,253,279]" pageId="20" pageNumber="255">Variability</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="20.[159,1036,253,803]" pageId="20" pageNumber="255">
|
||
Head – Specimens from the vicinity of Wotu and from Peleng Island have the anterior third to half of labium black while specimens from the vicinity of Gn
|
||
<taxonomicName box="[232,342,365,391]" class="Insecta" family="Chlorocyphidae" genus="Watuwila" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="236" phylum="Arthropoda" rank="genus">Watuwila</taxonomicName>
|
||
(type area) have a completely yellow labium. Thorax – Fw and Hw with 2 Ax; Fw with 34–40 Px and Hw with 28–36 Px. In specimens from Peleng Island the yellow on the head is more extensive (
|
||
<figureCitation box="[167,253,477,504]" captionStart="Figures 7 – 14" captionStartId="13.[159,238,1384,1409]" captionTargetBox="[196,998,249,1341]" captionTargetId="figure@13.[196,998,249,1342]" captionTargetPageId="13" captionText="Figures 7 – 14. Head of Celebargiolestes males. 7 – C. cinctus (immature), Ranotongkor, 02 - viii- 1940, JvT 6476; 8 – C. cinctus (old adult), Mapanget, 23 - vi- 1940, JvT 6638; 9 – C. cinctus (‘ red legs’ form), 25 km NE of Luwuk, 30 - i- 1989, JvT 6670; 10 – C. askewi sp. nov., holotype; 11 – C. toli sp. nov., holotype; 12 – C. toli sp. nov., Peleng Island, 05 - viii- 1941, JvT 6453; 13 – C. orri sp. nov., holotype; 14 – C. orri sp. nov., 10 km WNW Palopo, 29 - x- 1993, JvT 11910." httpUri="https://zenodo.org/record/192855/files/figure.png" pageId="20" pageNumber="255">Figs 12</figureCitation>
|
||
,
|
||
<figureCitation box="[265,297,478,504]" captionStart="Figures 19 – 22" captionStartId="18.[159,238,1470,1495]" captionTargetBox="[212,983,244,1431]" captionTargetId="figure@18.[212,983,244,1431]" captionTargetPageId="18" captionText="Figures 19 – 22. Body of Celebargiolestes males. 19 – C. toli sp. nov., holotype; 20 – C. toli sp. nov., Peleng Island, 05 - viii- 1941, JvT 6453; 21 – C. orri sp. nov., holotype; 22 – C. orri sp. nov., 10 km WNW Palopo, 29 - x- 1993, JvT 11910." httpUri="https://zenodo.org/record/192857/files/figure.png" pageId="20" pageNumber="255">20</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="20.[159,1036,253,803]" pageId="20" pageNumber="255">
|
||
Abdomen – Seen from ventral the lower appendages show differences in shape between the population from the Gn.
|
||
<taxonomicName box="[690,800,552,578]" class="Insecta" family="Chlorocyphidae" genus="Watuwila" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="236" phylum="Arthropoda" rank="genus">Watuwila</taxonomicName>
|
||
(
|
||
<figureCitation box="[818,902,552,578]" captionStart="Figures 40 – 45" captionStartId="28.[159,238,1442,1467]" captionTargetBox="[156,1039,522,1404]" captionTargetId="figure@28.[156,1039,522,1404]" captionTargetPageId="28" captionText="Figures 40 – 45. Appendages of Celebargiolestes males. 40 – C. toli sp. nov., holotype, dorsal view; 41 – C. toli sp. nov., holotype, lateral view; 42 – C. toli sp. nov., holotype, ventral view; 43 – C. orri sp. nov., holotype, dorsal view; 44 – C. orri sp. nov., holotype, lateral view; 45 – C. orri sp. nov., JvT 6646, ventral view." httpUri="https://zenodo.org/record/192861/files/figure.png" pageId="20" pageNumber="255">Fig. 44</figureCitation>
|
||
) and those from the surroundings of Peleng Island (
|
||
<figureCitation box="[632,716,589,616]" captionStart="Figures 40 – 45" captionStartId="28.[159,238,1442,1467]" captionTargetBox="[156,1039,522,1404]" captionTargetId="figure@28.[156,1039,522,1404]" captionTargetPageId="28" captionText="Figures 40 – 45. Appendages of Celebargiolestes males. 40 – C. toli sp. nov., holotype, dorsal view; 41 – C. toli sp. nov., holotype, lateral view; 42 – C. toli sp. nov., holotype, ventral view; 43 – C. orri sp. nov., holotype, dorsal view; 44 – C. orri sp. nov., holotype, lateral view; 45 – C. orri sp. nov., JvT 6646, ventral view." httpUri="https://zenodo.org/record/192861/files/figure.png" pageId="20" pageNumber="255">Fig. 45</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="20.[159,1036,253,803]" pageId="20" pageNumber="255">
|
||
Measurements [mm] – Total length 44–45, abdomen 35, Fw 32–33, Hw 32– 33 (specimens from Peleng Island). Total length 46–52, abdomen 36–42, Fw 33–36, Hw 33–36 (specimens from the surroundings of Gn.
|
||
<taxonomicName box="[908,1024,701,728]" class="Insecta" family="Chlorocyphidae" genus="Watuwila" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="236" phylum="Arthropoda" rank="genus">Watuwila</taxonomicName>
|
||
). Total length 47–49, abdomen 37–39, Fw 32–34, Hw 32–34 (specimens from the surroundings of Wotu).
|
||
</paragraph>
|
||
<paragraph blockId="20.[159,1036,888,1063]" box="[159,247,888,914]" pageId="20" pageNumber="255">
|
||
<heading bold="true" box="[159,247,888,914]" fontSize="11" level="3" pageId="20" pageNumber="255" reason="0">
|
||
<emphasis bold="true" box="[159,247,888,914]" pageId="20" pageNumber="255">Female</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="20.[159,1036,888,1063]" pageId="20" pageNumber="255">Head − Pattern as in male. Thorax − Pattern on prothorax, synthorax and legs as in male. Fw and Hw with 2 Ax; Fw with 27–32 Px and Hw with 26– 27Px. Pt white with a slight yellow tinge, outer border in older specimens mottled greyish-black.</paragraph>
|
||
<paragraph blockId="20.[159,1036,1112,1363]" pageId="20" pageNumber="255">
|
||
Abdomen − Stockier and shorter than in male. Pattern largely as in male; dark areas dark brown to black, pale parts are yellow-brown. S3–9 dorsally largely dark, ventral and lower lateral areas yellow, the yellow on the anterior of each segment reaches to the dorsum forming a yellow ring covering approximately ¹/₅ to ¹/₁0 of the segment. S10 dorsally black but laterally largely pale brown, cerci brown, base of valvifer pale brown becoming brown to black towards the tip, shaped as in
|
||
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1886" box="[565,677,1337,1363]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="255" phylum="Arthropoda" rank="species" species="cinctus">
|
||
<emphasis box="[565,677,1337,1363]" italics="true" pageId="20" pageNumber="255">C. cinctus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="20.[159,1036,1410,1512]" pageId="20" pageNumber="255">
|
||
Measurements [mm]
|
||
<emphasis bold="true" box="[417,434,1411,1437]" pageId="20" pageNumber="255">−</emphasis>
|
||
Total length 41, abdomen 32, Fw 33; Hw 33 (one female from the surroundings of Wotu). length 40–41, abdomen 31–32, Fw 31–34; Hw 31–34 (surroundings of Gn.
|
||
<taxonomicName box="[625,741,1485,1512]" class="Insecta" family="Chlorocyphidae" genus="Watuwila" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="236" phylum="Arthropoda" rank="genus">Watuwila</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="21" pageNumber="256" type="diagnosis">
|
||
<paragraph blockId="21.[159,1036,253,467]" box="[159,424,253,279]" pageId="21" pageNumber="256">
|
||
<heading bold="true" box="[159,424,253,279]" fontSize="11" level="3" pageId="21" pageNumber="256" reason="0">
|
||
<emphasis bold="true" box="[159,424,253,279]" pageId="21" pageNumber="256">Differential diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="21.[159,1036,253,467]" pageId="21" pageNumber="256">
|
||
Males are distinguished from other species of
|
||
<taxonomicName authorityName="Kennedy" authorityYear="1925" box="[705,884,291,317]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="256" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[705,884,291,317]" italics="true" pageId="21" pageNumber="256">Celebargiolestes</emphasis>
|
||
</taxonomicName>
|
||
by the combination of the following characters: (1) labrum yellow; (2) postclypeus and frons yellow-brown; (3) genital ligula with a shallow apical cleft; (4) lower appendages long, about half the length of the upper appendages and, when seen from lateral, with the tip reaching the level of the upper appendages.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |