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<document id="BDF6C56C9A3FBD897FCF622C9D82FA3F" ID-CLB-Dataset="21939" ID-DOI="10.5852/ejt.2014.97" ID-GBIF-Dataset="37e33fa8-2cb4-4693-86c6-77bb430b5baa" ID-ISSN="2118-9773" ID-Zenodo-Dep="3838926" ID-ZooBank="D58FA727-9A10-4023-B178-9E6CCD1EA457" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590097503938" checkinUser="tatiana" docAuthor="Yoshitomi, Hiroyuki" docDate="2014" docId="03933D4C204AFFA16D4A8778FC0AFA9C" docLanguage="en" docName="ejt-2014-97.pdf.imf" docOrigin="European Journal of Taxonomy 97" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Laius submariniformis Wittmer 1985" docType="treatment" docVersion="9" lastPageNumber="20" masterDocId="FFAA45342058FFB26F7D8673FFDAFF82" masterDocTitle="Comparative morphology of the endophallic structures of the genus Laius (Coleoptera, Melyridae), with the descriptions of three new species" masterLastPageNumber="29" masterPageNumber="1" pageNumber="19" updateTime="1698840803048" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="C7033AB2ABB5E661073768ADBEE93838">Comparative morphology of the endophallic structures of the genus Laius (Coleoptera, Melyridae), with the descriptions of three new species</mods:title>
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<mods:namePart id="D40DEF2A8C37A92BDE80E030B28DD41D">Yoshitomi, Hiroyuki</mods:namePart>
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<mods:title id="2798420AF17BF08A11FF4F01A0FE1FE9">European Journal of Taxonomy</mods:title>
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<mods:date id="3409DA5603976C5B9235A8158CDA55F3">2014</mods:date>
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<mods:number id="3E51C74C6F419D492923EB9F2509F9F8">2014-09-25</mods:number>
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<mods:number id="C241205F0923D1CCDE93854DBAC952CA">97</mods:number>
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<treatment id="03933D4C204AFFA16D4A8778FC0AFA9C" ID-DOI="http://doi.org/10.5281/zenodo.3852586" ID-GBIF-Taxon="164248126" ID-Zenodo-Dep="3852586" LSID="urn:lsid:plazi:treatment:03933D4C204AFFA16D4A8778FC0AFA9C" httpUri="http://treatment.plazi.org/id/03933D4C204AFFA16D4A8778FC0AFA9C" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<subSubSection id="C320DFD1204AFFA06D4A8778FC27FEA4" box="[567,1021,266,294]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="8B858C5A204AFFA06D4A8778FC27FEA4" blockId="18.[567,1021,266,329]" box="[567,1021,266,294]" pageId="18" pageNumber="19">
<heading id="D0CD3B36204AFFA06D4A8778FC27FEA4" box="[567,1021,266,294]" centered="true" fontSize="11" level="2" pageId="18" pageNumber="19" reason="2">
<taxonomicName id="4C3AF7D9204AFFA06D4A8778FC27FEA4" authority="Wittmer, 1985" authorityName="Wittmer" authorityYear="1985" box="[567,1021,266,294]" class="Insecta" family="Melyridae" genus="Laius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="submariniformis">
<emphasis id="B94E5048204AFFA06D4A8778FC97FEA7" bold="true" box="[567,845,266,293]" italics="true" pageId="18" pageNumber="19">Laius submariniformis</emphasis>
<bibRefCitation id="EFABF1AB204AFFA06C298778FC27FEA4" author="Wittmer W." box="[852,1021,267,294]" pageId="18" pageNumber="19" pagination="381 - 390" refId="ref12038" refString="Wittmer W. 1985. 38. Beitrag zur Kenntnis der indo-malaiischen Fauna. Entomologische Arbeiten aus dem Museum G. Frey 33 - 34: 381 - 390." type="journal article" year="1985">Wittmer, 1985</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C320DFD1204AFFA06DDD875DFC49FECA" box="[672,915,302,329]" pageId="18" pageNumber="19" type="description">
<paragraph id="8B858C5A204AFFA06DDD875DFC49FECA" blockId="18.[567,1021,266,329]" box="[672,915,302,329]" pageId="18" pageNumber="19">
<figureCitation id="130190DF204AFFA06DDD875DFD2EFECA" box="[672,756,302,328]" captionStart="Fig" captionStartId="6.[189,231,1895,1921]" captionTargetBox="[198,1346,298,1833]" captionTargetId="figure@6.[189,1398,277,1853]" captionTargetPageId="6" captionText="Fig. 3. Endophallic sclerites of Laius spp. A. Laius baliensis sp. nov. (paratype). B. L. andamanensis sp. nov. (paratype). C. L. rodriguesensis sp. nov. (paratype). D. L. maai Wittmer, 1973 (paratype). E. L. marchei Pic, 1922. F. L. politus Fairmaire, 1880. G. L. sabangensis Wittmer, 1985 (paratype). H. L. sericatus Champion, 1924. I. L. submariniformis Wittmer, 1985 (paratype). J. L. asahinai Nakane, 1955." figureDoi="http://doi.org/10.5281/zenodo.3838932" httpUri="https://zenodo.org/record/3838932/files/figure.png" pageId="18" pageNumber="19">Figs 3I</figureCitation>
,
<figureCitation id="130190DF204AFFA06C7C875DFCFAFECA" box="[769,800,302,328]" captionStart="Fig" captionStartId="11.[189,232,1930,1956]" captionTargetBox="[227,1378,578,1872]" captionTargetId="figure@11.[189,1399,562,1890]" captionTargetPageId="11" captionText="Fig. 6. Dorsal habitus of Laius spp. (left: ♂; right: ♀). A. Laius maai Wittmer, 1973 (paratype). B. L. marchei Pic, 1922. C. L. politus Fairmaire, 1880. D. L. sabangensis Wittmer, 1985 (paratype). E. L. sericatus Champion, 1924. F. L. submariniformis Wittmer, 1985 (paratype). Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3838938" httpUri="https://zenodo.org/record/3838938/files/figure.png" pageId="18" pageNumber="19">6F</figureCitation>
,
<figureCitation id="130190DF204AFFA06C50875CFC9DFECB" box="[813,839,303,329]" captionStart="Fig" captionStartId="13.[189,232,1895,1921]" captionTargetBox="[209,1374,709,1841]" captionTargetId="figure@13.[201,1387,703,1857]" captionTargetPageId="13" captionText="Fig. 7. Left antennae of Laius spp. (above: male; below: female). A. Laius baliensis sp. nov. (paratype). B. L. andamanensis sp. nov. (paratype). C. L. rodriguesensis sp. nov. (paratype). D. L. maai Wittmer, 1973 (paratype). E. L. marchei Pic, 1922. F. L. politus Fairmaire, 1880. G. L. sabangensis Wittmer, 1985 (paratype). H. L. sericatus Champion, 1924. I. L. submariniformis Wittmer, 1985 (paratype)." figureDoi="http://doi.org/10.5281/zenodo.3838940" httpUri="https://zenodo.org/record/3838940/files/figure.png" pageId="18" pageNumber="19">7I</figureCitation>
,
<figureCitation id="130190DF204AFFA06C29875DFCB4FECA" box="[852,878,302,328]" captionStart="Fig" captionStartId="15.[189,232,1895,1921]" captionTargetBox="[213,1376,839,1822]" captionTargetId="figure@15.[206,1381,824,1855]" captionTargetPageId="15" captionText="Fig. 8. Male antennomere III of Laius spp. A. Laius baliensis sp. nov. (paratype). B. L. andamanensis sp. nov. (paratype). C. L. rodriguesensis sp. nov. (paratype). D. L. maai Wittmer, 1973 (paratype). E. L. marchei Pic, 1922. F. L. politus Fairmaire, 1880. G. L. sabangensis Wittmer, 1985 (paratype). H. L. sericatus Champion, 1924. I. L. submariniformis Wittmer, 1985 (paratype)." figureDoi="http://doi.org/10.5281/zenodo.3838942" httpUri="https://zenodo.org/record/3838942/files/figure.png" pageId="18" pageNumber="19">8I</figureCitation>
,
<figureCitation id="130190DF204AFFA06C06875DFC49FECA" box="[891,915,302,328]" captionStart="Fig" captionStartId="16.[189,232,1895,1921]" captionTargetBox="[245,1358,840,1832]" captionTargetId="figure@16.[217,1371,835,1855]" captionTargetPageId="16" captionText="Fig. 9. Aegeagi of Laius spp. A. Laius baliensis sp. nov. (paratype). B. L. andamanensis sp. nov. (paratype). C. L. rodriguesensis sp. nov. (paratype). D. L. maai Wittmer, 1973 (paratype). E. L. marchei Pic, 1922. F. L. politus Fairmaire, 1880. G. L. sabangensis Wittmer, 1985 (paratype). H. L. sericatus Champion, 1924. I. L. submariniformis Wittmer, 1985 (paratype)." figureDoi="http://doi.org/10.5281/zenodo.3838944" httpUri="https://zenodo.org/record/3838944/files/figure.png" pageId="18" pageNumber="19">9I</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C320DFD1204AFFA06FC08703FD60FE08" box="[189,698,367,394]" pageId="18" pageNumber="19" type="reference_group">
<paragraph id="8B858C5A204AFFA06FC08703FD60FE08" blockId="18.[189,698,367,394]" box="[189,698,367,394]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="AB2AAB74204AFFA06FC08703FD60FE08" box="[189,698,367,394]" pageId="18" pageNumber="19">
<treatmentCitation id="0A9BAA4B204AFFA06FC08703FD6CFE08" author="Wittmer W." box="[189,694,367,394]" page="385" pageId="18" pageNumber="19" year="1985">
<taxonomicName id="4C3AF7D9204AFFA06FC08703FD6CFE08" authority="Wittmer, 1985: 385" authorityName="Wittmer" authorityPageNumber="385" authorityYear="1985" box="[189,694,367,394]" class="Insecta" family="Melyridae" genus="Laius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="submariniformis">
<emphasis id="B94E5048204AFFA06FC08703FE12FE0B" box="[189,456,367,393]" italics="true" pageId="18" pageNumber="19">Laius submariniformis</emphasis>
<bibRefCitation id="EFABF1AB204AFFA06EB3871CFD6CFE08" author="Wittmer W." box="[462,694,367,394]" pageId="18" pageNumber="19" pagination="381 - 390" refId="ref12038" refString="Wittmer W. 1985. 38. Beitrag zur Kenntnis der indo-malaiischen Fauna. Entomologische Arbeiten aus dem Museum G. Frey 33 - 34: 381 - 390." type="journal article" year="1985">Wittmer, 1985: 385</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C320DFD1204AFFA06FC087C2FAADFDC2" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="8B858C5A204AFFA06FC087C2FE70FE49" blockId="18.[189,426,433,459]" box="[189,426,433,459]" pageId="18" pageNumber="19">
<heading id="D0CD3B36204AFFA06FC087C2FE70FE49" bold="true" box="[189,426,433,459]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="B94E5048204AFFA06FC087C2FE70FE49" bold="true" box="[189,426,433,459]" pageId="18" pageNumber="19">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B858C5A204AFFA06FC08793FAADFDC2" blockId="18.[189,1399,479,577]" pageId="18" pageNumber="19">
<materialsCitation id="3B528607204AFFA06FC08793FBD4FE7B" ID-GBIF-Occurrence="3761956327" box="[189,1038,479,506]" collectingDate="1990-09-17" collectionCode="EUMJ" country="Malaysia" location="Ken Ito" pageId="18" pageNumber="19" specimenCount="8" specimenCount-female="6" specimenCount-male="2">
<specimenCount id="9D3C47D3204AFFA06FC08793FEDFFE78" box="[189,261,480,506]" pageId="18" pageNumber="19" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D3C47D3204AFFA06E6A87ACFE87FE78" box="[279,349,479,506]" pageId="18" pageNumber="19" type="female">6 ♀♀</specimenCount>
(
<collectionCode id="ED2B149F204AFFA06E0C8793FE1EFE78" box="[369,452,480,506]" collectionName="Japan, Matsuyama, Ehime University" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="18" pageNumber="19">EUMJ</collectionCode>
), “Damai,
<collectingCountry id="F32DCCCA204AFFA06D3687ACFD6AFE7B" box="[587,688,479,505]" name="Malaysia" pageId="18" pageNumber="19">Sarawak</collectingCountry>
<date id="FF84AA9A204AFFA06DC18793FC83FE78" box="[700,857,480,506]" pageId="18" pageNumber="19" value="1990-09-17">
<collectingDate id="EFC05372204AFFA06DC18793FC83FE78" box="[700,857,480,506]" pageId="18" pageNumber="19" value="1990-09-17">17. IX. 1990</collectingDate>
</date>
<location id="8EE5DA81204AFFA06C188793FC18FE78" LSID="urn:lsid:plazi:treatment:03933D4C204AFFA16D4A8778FC0AFA9C:8EE5DA81204AFFA06C188793FC18FE78" box="[869,962,480,506]" country="Malaysia" name="Ken Ito" pageId="18" pageNumber="19">Ken Itô</location>
leg.”;
</materialsCitation>
<materialsCitation id="3B528607204AFFA06B6787ACFE43FD9F" ID-GBIF-Occurrence="3761956325" collectionCode="EUMJ" pageId="18" pageNumber="19" specimenCount="34" specimenCount-female="29" specimenCount-male="5" typeStatus="paratype">
<specimenCount id="9D3C47D3204AFFA06B6787ACFBB8FE78" box="[1050,1122,479,506]" pageId="18" pageNumber="19" type="male">5 ♂♂</specimenCount>
,
<specimenCount id="9D3C47D3204AFFA06B098793FB13FE78" box="[1140,1225,480,506]" pageId="18" pageNumber="19" type="female">29 ♀♀</specimenCount>
(including 10
<typeStatus id="548132F8204AFFA06FC08470FEEBFD9F" box="[189,305,515,541]" pageId="18" pageNumber="19" type="paratype">paratypes</typeStatus>
;
<collectionCode id="ED2B149F204AFFA06E418470FE57FD9F" box="[316,397,515,541]" collectionName="Japan, Matsuyama, Ehime University" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="18" pageNumber="19">EUMJ</collectionCode>
),
</materialsCitation>
<materialsCitation id="3B528607204AFFA06EDD8470FE1EFDC2" ID-GBIF-Occurrence="3761956306" collectionCode="BPBM" country="Philippines" location="Sato" municipality="July" pageId="18" pageNumber="19" specimenCount="1986" specimenCount-female="4" specimenCount-male="1982" stateProvince="Palawan">
“
<collectingCountry id="F32DCCCA204AFFA06ED48470FD83FD9F" box="[425,601,515,541]" name="Philippines" pageId="18" pageNumber="19">PHILIPPINES</collectingCountry>
Sabang, N. of Mt. St. Paul,
<collectingRegion id="49FE42B8204AFFA06CE18470FC25FD9F" box="[924,1023,515,541]" country="Philippines" name="Palawan" pageId="18" pageNumber="19">Palawan</collectingRegion>
<collectingMunicipality id="6BE11620204AFFA06B7B8470FBE2FD9F" box="[1030,1080,515,541]" pageId="18" pageNumber="19">July</collectingMunicipality>
, 11-13,
<specimenCount id="9D3C47D3204AFFA06BEE8470FB2CFD9F" box="[1171,1270,515,541]" pageId="18" pageNumber="19" type="male">1977 M.</specimenCount>
<location id="8EE5DA81204AFFA06B818470FAEAFD9F" LSID="urn:lsid:plazi:treatment:03933D4C204AFFA16D4A8778FC0AFA9C:8EE5DA81204AFFA06B818470FAEAFD9F" box="[1276,1328,515,541]" country="Philippines" municipality="July" name="Sato" pageId="18" pageNumber="19" stateProvince="Palawan">Sato</location>
leg.”;
<specimenCount id="9D3C47D3204AFFA06FC08455FEDBFDC2" box="[189,257,550,577]" pageId="18" pageNumber="19" type="male">5 ♂♂</specimenCount>
,
<specimenCount id="9D3C47D3204AFFA06E738455FE95FDC2" box="[270,335,550,576]" pageId="18" pageNumber="19" type="female">4 ♀♀</specimenCount>
(
<collectionCode id="ED2B149F204AFFA06E228455FE6CFDC2" box="[351,438,550,576]" collectionName="USA, Hawaii, Honolulu, Bernice P. Bishop Museum" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="18" pageNumber="19" type="Museum">BPBM</collectionCode>
),
</materialsCitation>
<materialsCitation id="3B528607204AFFA06EB78455FAADFDC2" ID-GBIF-Occurrence="3761956310" box="[458,1399,550,577]" collectingDate="1957-02-27" country="Philippines" location="Yoshio Kondo" municipality="Sulu I." pageId="18" pageNumber="19" specimenCount="1" stateProvince="Cagayan">
“
<collectingCountry id="F32DCCCA204AFFA06EA98455FD5EFDC2" box="[468,644,550,576]" name="Philippines" pageId="18" pageNumber="19">PHILIPPINES</collectingCountry>
(S.)
<collectingRegion id="49FE42B8204AFFA06DC38455FCFFFDC2" box="[702,805,550,576]" country="Philippines" name="Cagayan" pageId="18" pageNumber="19">Cagayan</collectingRegion>
<collectingMunicipality id="6BE11620204AFFA06C518455FCA0FDC2" box="[812,890,550,576]" pageId="18" pageNumber="19">Sulu I.</collectingMunicipality>
<date id="FF84AA9A204AFFA06CFC8455FBFDFDC3" box="[897,1063,550,577]" pageId="18" pageNumber="19" value="1957-02-27">
<collectingDate id="EFC05372204AFFA06CFC8455FBFDFDC3" box="[897,1063,550,577]" pageId="18" pageNumber="19" value="1957-02-27">Feb. 27. 1957</collectingDate>
</date>
”, “
<location id="8EE5DA81204AFFA06B358455FB2EFDC2" LSID="urn:lsid:plazi:treatment:03933D4C204AFFA16D4A8778FC0AFA9C:8EE5DA81204AFFA06B358455FB2EFDC2" box="[1096,1268,550,576]" country="Philippines" municipality="Sulu I." name="Yoshio Kondo" pageId="18" pageNumber="19" stateProvince="Cagayan">Yoshio Kondo</location>
collector”.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF45DCD2204AFFA06FC081F9FD7FF869" ID-DOI="http://doi.org/10.5281/zenodo.3838946" ID-Zenodo-Dep="3838946" httpUri="https://zenodo.org/record/3838946/files/figure.png" pageId="18" pageNumber="19" startId="18.[189,232,1930,1956]" targetBox="[234,1378,670,1876]" targetPageId="18">
<paragraph id="8B858C5A204AFFA06FC081F9FD7FF869" blockId="18.[189,1399,1930,2027]" pageId="18" pageNumber="19">
<emphasis id="B94E5048204AFFA06FC081F9FEC7F826" bold="true" box="[189,285,1930,1956]" pageId="18" pageNumber="19">Fig. 10.</emphasis>
Larva of
<taxonomicName id="4C3AF7D9204AFFA06EE481F8FD4AF826" authority="Yoshitomi, 2014" authorityName="Yoshitomi" authorityYear="2014" box="[409,656,1930,1956]" class="Insecta" family="Melyridae" genus="Laius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="rodriguesensis" status="sp. nov.">
<emphasis id="B94E5048204AFFA06EE481F8FD4AF826" box="[409,656,1930,1956]" italics="true" pageId="18" pageNumber="19">Laius rodriguesensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27DED33204AFFA06DE781F8FD22F827" box="[666,760,1931,1957]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B94E5048204AFFA06C7F81F9FCCDF826" bold="true" box="[770,791,1930,1956]" pageId="18" pageNumber="19">A</emphasis>
. Dorsal habitus.
<emphasis id="B94E5048204AFFA06C9581F9FC26F826" bold="true" box="[1000,1020,1930,1956]" pageId="18" pageNumber="19">B</emphasis>
. Head in dorsal view.
<emphasis id="B94E5048204AFFA06A6F81F9FAFDF826" bold="true" box="[1298,1319,1930,1956]" pageId="18" pageNumber="19">C</emphasis>
. Head in ventral view.
<emphasis id="B94E5048204AFFA06E0281DDFE4EF84A" bold="true" box="[383,404,1966,1992]" pageId="18" pageNumber="19">D</emphasis>
. Abdominal segment IX in dorsal view.
<emphasis id="B94E5048204AFFA06C0181DDFC4AF84A" bold="true" box="[892,912,1966,1992]" pageId="18" pageNumber="19">E</emphasis>
. Abdominal segment IX in ventral view.
<emphasis id="B94E5048204AFFA06FC081A2FF15F869" bold="true" box="[189,207,2001,2027]" pageId="18" pageNumber="19">F</emphasis>
. Abdominal segment IX in lateral view.
</paragraph>
</caption>
<subSubSection id="C320DFD1204BFFA16FC08778FB3AFBB7" pageId="19" pageNumber="20" type="description">
<paragraph id="8B858C5A204BFFA16FC08778FEB1FEA7" blockId="19.[189,363,267,293]" box="[189,363,267,293]" pageId="19" pageNumber="20">
<heading id="D0CD3B36204BFFA16FC08778FEB1FEA7" bold="true" box="[189,363,267,293]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="B94E5048204BFFA16FC08778FEB1FEA7" bold="true" box="[189,363,267,293]" pageId="19" pageNumber="20">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="8B858C5A204BFFA16F97874FFEF0FED4" blockId="19.[189,1399,316,642]" box="[234,298,316,342]" pageId="19" pageNumber="20">
<emphasis id="B94E5048204BFFA16F97874FFEF0FED4" bold="true" box="[234,298,316,342]" pageId="19" pageNumber="20">Male</emphasis>
</paragraph>
<paragraph id="8B858C5A204BFFA16FC08712FEF8FD03" blockId="19.[189,1399,316,642]" pageId="19" pageNumber="20">
HL/HW 0.75–0.86 (0.80); PW/PL 1.27–1.37 (1.32); EL/EW 1.38–1.50 (1.42); EL/PL 2.64–2.94 (2.79); EW/PW 1.43–1.56 (1.48); TL/EW 2.25–2.49 (2.37). Antennae (
<figureCitation id="130190DF204BFFA16CC987F4FBDFFE23" box="[948,1029,391,417]" captionStart="Fig" captionStartId="13.[189,232,1895,1921]" captionTargetBox="[209,1374,709,1841]" captionTargetId="figure@13.[201,1387,703,1857]" captionTargetPageId="13" captionText="Fig. 7. Left antennae of Laius spp. (above: male; below: female). A. Laius baliensis sp. nov. (paratype). B. L. andamanensis sp. nov. (paratype). C. L. rodriguesensis sp. nov. (paratype). D. L. maai Wittmer, 1973 (paratype). E. L. marchei Pic, 1922. F. L. politus Fairmaire, 1880. G. L. sabangensis Wittmer, 1985 (paratype). H. L. sericatus Champion, 1924. I. L. submariniformis Wittmer, 1985 (paratype)." figureDoi="http://doi.org/10.5281/zenodo.3838940" httpUri="https://zenodo.org/record/3838940/files/figure.png" pageId="19" pageNumber="20">Fig. 7I</figureCitation>
) long; antennomere I long and slightly curved laterally; antennomere III (
<figureCitation id="130190DF204BFFA16DCD87DFFCDBFE44" box="[688,769,428,454]" captionStart="Fig" captionStartId="15.[189,232,1895,1921]" captionTargetBox="[213,1376,839,1822]" captionTargetId="figure@15.[206,1381,824,1855]" captionTargetPageId="15" captionText="Fig. 8. Male antennomere III of Laius spp. A. Laius baliensis sp. nov. (paratype). B. L. andamanensis sp. nov. (paratype). C. L. rodriguesensis sp. nov. (paratype). D. L. maai Wittmer, 1973 (paratype). E. L. marchei Pic, 1922. F. L. politus Fairmaire, 1880. G. L. sabangensis Wittmer, 1985 (paratype). H. L. sericatus Champion, 1924. I. L. submariniformis Wittmer, 1985 (paratype)." figureDoi="http://doi.org/10.5281/zenodo.3838942" httpUri="https://zenodo.org/record/3838942/files/figure.png" pageId="19" pageNumber="20">Fig. 8I</figureCitation>
) oval, deeply concave in dorso-mesal portion, L/W = 0.88; approximate ratio of each antennomeres (n = 1) as 15.0: 1.0: 9.0: 3.3: 3.3: 3.0: 3.3: 3.3: 3.3: 3.3: 5.0. Caudal margin of tergite VII deeply concave. Aedeagus (
<figureCitation id="130190DF204BFFA16CAE8784FBFEFD93" box="[979,1060,503,529]" captionStart="Fig" captionStartId="16.[189,232,1895,1921]" captionTargetBox="[245,1358,840,1832]" captionTargetId="figure@16.[217,1371,835,1855]" captionTargetPageId="16" captionText="Fig. 9. Aegeagi of Laius spp. A. Laius baliensis sp. nov. (paratype). B. L. andamanensis sp. nov. (paratype). C. L. rodriguesensis sp. nov. (paratype). D. L. maai Wittmer, 1973 (paratype). E. L. marchei Pic, 1922. F. L. politus Fairmaire, 1880. G. L. sabangensis Wittmer, 1985 (paratype). H. L. sericatus Champion, 1924. I. L. submariniformis Wittmer, 1985 (paratype)." figureDoi="http://doi.org/10.5281/zenodo.3838944" httpUri="https://zenodo.org/record/3838944/files/figure.png" pageId="19" pageNumber="20">Fig. 9I</figureCitation>
) about
<quantity id="4CC221BF204BFFA16BFD8784FB36FD90" box="[1152,1260,503,530]" metricMagnitude="-3" metricUnit="m" metricValue="1.37" pageId="19" pageNumber="20" unit="mm" value="1.37">1.37 mm</quantity>
, concave at aedeagal apex. Gonoporal piece (
<figureCitation id="130190DF204BFFA16D42846FFD54FDB5" box="[575,654,540,567]" captionStart="Fig" captionStartId="6.[189,231,1895,1921]" captionTargetBox="[198,1346,298,1833]" captionTargetId="figure@6.[189,1398,277,1853]" captionTargetPageId="6" captionText="Fig. 3. Endophallic sclerites of Laius spp. A. Laius baliensis sp. nov. (paratype). B. L. andamanensis sp. nov. (paratype). C. L. rodriguesensis sp. nov. (paratype). D. L. maai Wittmer, 1973 (paratype). E. L. marchei Pic, 1922. F. L. politus Fairmaire, 1880. G. L. sabangensis Wittmer, 1985 (paratype). H. L. sericatus Champion, 1924. I. L. submariniformis Wittmer, 1985 (paratype). J. L. asahinai Nakane, 1955." figureDoi="http://doi.org/10.5281/zenodo.3838932" httpUri="https://zenodo.org/record/3838932/files/figure.png" pageId="19" pageNumber="20">Fig. 3I</figureCitation>
) straight, rather wide, with very short basal projection,
<quantity id="4CC221BF204BFFA16A72846EFAADFDB5" box="[1295,1399,541,567]" metricMagnitude="-4" metricUnit="m" metricValue="7.7" pageId="19" pageNumber="20" unit="mm" value="0.77">0.77 mm</quantity>
in GL; GL/AL 1.51. Ligula (
<figureCitation id="130190DF204BFFA16D778431FD83FDDE" box="[522,601,578,604]" captionStart="Fig" captionStartId="6.[189,231,1895,1921]" captionTargetBox="[198,1346,298,1833]" captionTargetId="figure@6.[189,1398,277,1853]" captionTargetPageId="6" captionText="Fig. 3. Endophallic sclerites of Laius spp. A. Laius baliensis sp. nov. (paratype). B. L. andamanensis sp. nov. (paratype). C. L. rodriguesensis sp. nov. (paratype). D. L. maai Wittmer, 1973 (paratype). E. L. marchei Pic, 1922. F. L. politus Fairmaire, 1880. G. L. sabangensis Wittmer, 1985 (paratype). H. L. sericatus Champion, 1924. I. L. submariniformis Wittmer, 1985 (paratype). J. L. asahinai Nakane, 1955." figureDoi="http://doi.org/10.5281/zenodo.3838932" httpUri="https://zenodo.org/record/3838932/files/figure.png" pageId="19" pageNumber="20">Fig. 3I</figureCitation>
) long, evenly curved; LL
<quantity id="4CC221BF204BFFA16CF48431FC2EFDDE" box="[905,1012,578,604]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="19" pageNumber="20" unit="mm" value="0.37">0.37 mm</quantity>
; LW
<quantity id="4CC221BF204BFFA16B4C8431FB46FDDE" box="[1073,1180,578,604]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="19" pageNumber="20" unit="mm" value="0.14">0.14 mm</quantity>
; LL/LW 2.58; GL/ LL 2.06.
</paragraph>
<paragraph id="8B858C5A204BFFA16F9784D8FE9FFD47" blockId="19.[189,1399,683,860]" box="[234,325,683,709]" pageId="19" pageNumber="20">
<emphasis id="B94E5048204BFFA16F9784D8FE9FFD47" bold="true" box="[234,325,683,709]" pageId="19" pageNumber="20">Female</emphasis>
</paragraph>
<paragraph id="8B858C5A204BFFA16FC084A2FEBAFCD9" blockId="19.[189,1399,683,860]" pageId="19" pageNumber="20">
HL/HW 0.67–0.85 (0.76); PW/PL 1.32–1.45 (1.36); EL/EW 1.33–1.52 (1.39); EL/PL 2.64–2.91 (2.75); EW/PW 1.31–1.59 (1.46); TL/EW 2.15–2.48 (2.31). Antennae (
<figureCitation id="130190DF204BFFA16CC38485FBCBFC93" box="[958,1041,758,785]" captionStart="Fig" captionStartId="13.[189,232,1895,1921]" captionTargetBox="[209,1374,709,1841]" captionTargetId="figure@13.[201,1387,703,1857]" captionTargetPageId="13" captionText="Fig. 7. Left antennae of Laius spp. (above: male; below: female). A. Laius baliensis sp. nov. (paratype). B. L. andamanensis sp. nov. (paratype). C. L. rodriguesensis sp. nov. (paratype). D. L. maai Wittmer, 1973 (paratype). E. L. marchei Pic, 1922. F. L. politus Fairmaire, 1880. G. L. sabangensis Wittmer, 1985 (paratype). H. L. sericatus Champion, 1924. I. L. submariniformis Wittmer, 1985 (paratype)." figureDoi="http://doi.org/10.5281/zenodo.3838940" httpUri="https://zenodo.org/record/3838940/files/figure.png" pageId="19" pageNumber="20">Fig. 7I</figureCitation>
) long; antennomere III short, L/W = 2.14; approximate ratio of each antennomeres (n = 1) as 8.0: 1.0: 4.2: 2.3: 2.0: 2.0: 2.0: 1.8: 1.8: 1.8: 3.3.
</paragraph>
<paragraph id="8B858C5A204BFFA16F9785F6FE78FC1D" blockId="19.[189,1399,901,1077]" box="[234,418,901,927]" pageId="19" pageNumber="20">
<emphasis id="B94E5048204BFFA16F9785F6FE78FC1D" bold="true" box="[234,418,901,927]" pageId="19" pageNumber="20">Measurements</emphasis>
</paragraph>
<paragraph id="8B858C5A204BFFA16FC085D9FB3AFBB7" blockId="19.[189,1399,901,1077]" pageId="19" pageNumber="20">Male (n = 4): TL 4.90–6.33 (5.60) mm; HL 0.88–1.20 (1.02) mm; HW 1.05–1.52 (1.29) mm; PW 1.40– 1.82 (1.60) mm; PL 1.02–1.40 (1.21) mm; EL 3.00–3.78 (3.37) mm; EW 2.00–2.70 (2.37) mm. Female (n = 5): TL 4.95–5.65 (5.30) mm; HL 0.80–1.10 (0.95) mm; HW 1.20–1.32 (1.24) mm; PW 1.45–1.68 (1.58) mm; PL 1.10–1.25 (1.16) mm; EL 3.05–3.30 (3.19) mm; EW 2.10–2.40 (2.30) mm.</paragraph>
</subSubSection>
<subSubSection id="C320DFD1204BFFA16FC0822DFCE7FB28" box="[189,829,1118,1194]" pageId="19" pageNumber="20" type="distribution">
<paragraph id="8B858C5A204BFFA16FC0822DFE8CFBFA" blockId="19.[189,342,1118,1144]" box="[189,342,1118,1144]" pageId="19" pageNumber="20">
<heading id="D0CD3B36204BFFA16FC0822DFE8CFBFA" bold="true" box="[189,342,1118,1144]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="B94E5048204BFFA16FC0822DFE8CFBFA" bold="true" box="[189,342,1118,1144]" pageId="19" pageNumber="20">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B858C5A204BFFA16FC082FCFCE7FB28" blockId="19.[189,829,1167,1194]" box="[189,829,1167,1194]" pageId="19" pageNumber="20">
<collectingCountry id="F32DCCCA204BFFA16FC082FCFE9BFB2B" box="[189,321,1167,1193]" name="Philippines" pageId="19" pageNumber="20">Philippines</collectingCountry>
(
<collectingRegion id="49FE42B8204BFFA16E2D82FCFE62FB2B" box="[336,440,1167,1193]" country="Philippines" name="Palawan" pageId="19" pageNumber="20">Palawan</collectingRegion>
,
<collectingRegion id="49FE42B8204BFFA16EB982FCFE20FB2B" box="[452,506,1167,1193]" country="Philippines" name="Sulu" pageId="19" pageNumber="20">Sulu</collectingRegion>
Isls.),
<collectingCountry id="F32DCCCA204BFFA16D3782FCFD6CFB2B" box="[586,694,1167,1193]" name="Malaysia" pageId="19" pageNumber="20">Malaysia</collectingCountry>
(
<collectingCountry id="F32DCCCA204BFFA16DB882FCFCEBFB28" box="[709,817,1167,1194]" name="Malaysia" pageId="19" pageNumber="20">Sarawak</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C320DFD1204BFFA16FC082A0FC0AFA9C" box="[189,976,1235,1310]" pageId="19" pageNumber="20" type="discussion">
<paragraph id="8B858C5A204BFFA16FC082A0FEF5FB6F" blockId="19.[189,303,1235,1261]" box="[189,303,1235,1261]" pageId="19" pageNumber="20">
<heading id="D0CD3B36204BFFA16FC082A0FEF5FB6F" bold="true" box="[189,303,1235,1261]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="B94E5048204BFFA16FC082A0FEF5FB6F" bold="true" box="[189,303,1235,1261]" pageId="19" pageNumber="20">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B858C5A204BFFA16FC08377FC0AFA9C" blockId="19.[189,976,1284,1310]" box="[189,976,1284,1310]" pageId="19" pageNumber="20">
This is the first record of this species from
<collectingRegion id="49FE42B8204BFFA16DC98377FD30FA9C" box="[692,746,1284,1310]" country="Philippines" name="Sulu" pageId="19" pageNumber="20">Sulu</collectingRegion>
Island and Borneo.
</paragraph>
</subSubSection>
</treatment>
</document>
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