180 lines
18 KiB
XML
180 lines
18 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.4766383" ID-GBIF-Dataset="7dfe533a-40f2-435c-8b08-e9ea9cb1a070" ID-GBIF-Taxon="182165939" ID-ISSN="1854-0392" ID-Zenodo-Dep="4766383" checkinTime="1620958307236" checkinUser="felipe" docAuthor="Stark, Bill P." docDate="2010" docId="039287FB8076DF69FECBF8D32BF8ABB9" docLanguage="en" docName="Illiesia.6.01.1-10.pdf" docOrigin="Illiesia 6 (1)" docStyle="DocumentStyle:3ED4ECBC8873166DC296BE619562C54F.3:Illiesia.2009-.journal_article" docStyleId="3ED4ECBC8873166DC296BE619562C54F" docStyleName="Illiesia.2009-.journal_article" docStyleVersion="3" docTitle="Perlodes kippenhani Stark 2010, sp. n." docType="treatment" docVersion="3" lastPageId="7" lastPageNumber="7" masterDocId="FFABFF838070DF6EFF86FFC32F54AA48" masterDocTitle="Studies On Korean Stoneflies (Insecta: Plecoptera) With Descriptions Of Two New Species" masterLastPageNumber="10" masterPageNumber="1" pageId="6" pageNumber="7" updateTime="1632015763759" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
|
|
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo>
|
|
<mods:title>Studies On Korean Stoneflies (Insecta: Plecoptera) With Descriptions Of Two New Species</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Stark, Bill P.</mods:namePart>
|
|
</mods:name>
|
|
<mods:typeOfResource>text</mods:typeOfResource>
|
|
<mods:relatedItem type="host">
|
|
<mods:titleInfo>
|
|
<mods:title>Illiesia</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part>
|
|
<mods:date>2010</mods:date>
|
|
<mods:detail type="volume">
|
|
<mods:number>6</mods:number>
|
|
</mods:detail>
|
|
<mods:detail type="issue">
|
|
<mods:number>1</mods:number>
|
|
</mods:detail>
|
|
<mods:extent unit="page">
|
|
<mods:start>1</mods:start>
|
|
<mods:end>10</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:classification>journal article</mods:classification>
|
|
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4759533</mods:identifier>
|
|
<mods:identifier type="GBIF-Dataset">7dfe533a-40f2-435c-8b08-e9ea9cb1a070</mods:identifier>
|
|
<mods:identifier type="ISSN">1854-0392</mods:identifier>
|
|
<mods:identifier type="Zenodo-Dep">4759533</mods:identifier>
|
|
</mods:mods>
|
|
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4766383" ID-GBIF-Taxon="182165939" ID-Zenodo-Dep="4766383" LSID="urn:lsid:plazi:treatment:039287FB8076DF69FECBF8D32BF8ABB9" httpUri="http://treatment.plazi.org/id/039287FB8076DF69FECBF8D32BF8ABB9" lastPageId="7" lastPageNumber="7" pageId="6" pageNumber="7">
|
|
<subSubSection box="[333,653,1808,1832]" pageId="6" pageNumber="7" type="nomenclature">
|
|
<paragraph blockId="6.[333,653,1808,1832]" box="[333,653,1808,1832]" pageId="6" pageNumber="7">
|
|
<heading bold="true" box="[333,653,1808,1832]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="7" reason="2">
|
|
<emphasis bold="true" box="[333,653,1808,1832]" pageId="6" pageNumber="7">
|
|
<taxonomicName authority="Stark, 2010" authorityName="Stark" authorityYear="2010" box="[333,575,1808,1832]" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="kippenhani" status="sp. n.">
|
|
<emphasis bold="true" box="[333,575,1808,1832]" italics="true" pageId="6" pageNumber="7">Perlodes kippenhani</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel box="[588,653,1808,1832]" pageId="6" pageNumber="7" rank="species">sp. n.</taxonomicNameLabel>
|
|
</emphasis>
|
|
</heading>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection box="[422,565,1844,1868]" pageId="6" pageNumber="7" type="description">
|
|
<paragraph blockId="6.[422,565,1844,1868]" box="[422,565,1844,1868]" pageId="6" pageNumber="7">
|
|
(
|
|
<figureCitation box="[430,557,1844,1868]" captionStart="Figs" captionStartId="7.[189,241,1752,1776]" captionTargetBox="[299,1387,606,1672]" captionTargetId="figure-198@7.[317,1316,606,1664]" captionTargetPageId="7" captionText="Figs. 19-22. Perlodes kippenhani structures. 19. Head and pronotum. 20. Male terminalia, dorsal. 21. Male terminalia, ventral. 22. Paraproct, lateral." figureDoi="http://doi.org/10.5281/zenodo.4759551" httpUri="https://zenodo.org/record/4759551/files/figure.png" pageId="6" pageNumber="7">Figs. 19-22</figureCitation>
|
|
)
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="6" pageNumber="7" type="materials_examined">
|
|
<paragraph blockId="6.[835,1444,223,1849]" pageId="6" pageNumber="7">
|
|
<emphasis bold="true" box="[835,1070,224,248]" pageId="6" pageNumber="7">Material examined.</emphasis>
|
|
<materialsCitation ID-GBIF-Occurrence="3112257307" collectingDate="2006-05-13" collectionCode="USNM" collectorName="M. Kippenhan" country="South Korea" county="Pyeongchang-gun" location="Odaesan National Park trail to Sangwonsa" municipality="Odaesan National Park" pageId="6" pageNumber="7" specimenCount="1" specimenCount-male="1" stateProvince="Gangwondo" typeStatus="holotype">
|
|
<typeStatus box="[1076,1188,224,248]" pageId="6" pageNumber="7">Holotype</typeStatus>
|
|
<specimenCount box="[1194,1214,223,248]" pageId="6" pageNumber="7" type="male">♂</specimenCount>
|
|
,
|
|
<collectingCountry box="[1226,1439,224,248]" name="South Korea" pageId="6" pageNumber="7">Republic of Korea</collectingCountry>
|
|
,
|
|
<collectingRegion box="[835,984,260,284]" pageId="6" pageNumber="7">Gangwondo</collectingRegion>
|
|
,
|
|
<collectingCounty box="[998,1214,260,284]" pageId="6" pageNumber="7">Pyeongchang-gun</collectingCounty>
|
|
,
|
|
<location LSID="urn:lsid:plazi:treatment:039287FB8076DF69FECBF8D32BF8ABB9:8EE460368076DF68FB4AFEC72B0CAB08" country="South Korea" county="Pyeongchang-gun" municipality="Odaesan National Park" name="Odaesan National Park trail to Sangwonsa" pageId="6" pageNumber="7" stateProvince="Gangwondo">Odaesan National Park trail to Sangwonsa</location>
|
|
,
|
|
<date box="[1122,1268,296,320]" pageId="6" pageNumber="7" value="2006-05-13">
|
|
<collectingDate box="[1122,1268,296,320]" pageId="6" pageNumber="7" value="2006-05-13">13 May 2006</collectingDate>
|
|
</date>
|
|
,
|
|
<collectorName box="[1278,1442,296,320]" pageId="6" pageNumber="7">M. Kippenhan</collectorName>
|
|
(
|
|
<collectionCode box="[843,930,331,355]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="7" type="Museum">USNM</collectionCode>
|
|
).
|
|
</materialsCitation>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="6" pageNumber="7" type="description">
|
|
<paragraph blockId="6.[835,1444,223,1849]" pageId="6" pageNumber="7">
|
|
<emphasis bold="true" box="[835,1019,367,391]" pageId="6" pageNumber="7">Adult habitus.</emphasis>
|
|
General color black with limited areas of yellow pigment. Head dark but with circular interocellar yellow spot and pale M-line forward of median ocellus (
|
|
<figureCitation box="[1037,1122,474,498]" captionStart="Figs" captionStartId="7.[189,241,1752,1776]" captionTargetBox="[299,1387,606,1672]" captionTargetId="figure-198@7.[317,1316,606,1664]" captionTargetPageId="7" captionText="Figs. 19-22. Perlodes kippenhani structures. 19. Head and pronotum. 20. Male terminalia, dorsal. 21. Male terminalia, ventral. 22. Paraproct, lateral." figureDoi="http://doi.org/10.5281/zenodo.4759551" httpUri="https://zenodo.org/record/4759551/files/figure.png" pageId="6" pageNumber="7">Fig. 19</figureCitation>
|
|
); additional pale areas on occiput. Pronotum black with yellow median stripe, thoracic venter mostly black. Femora black, tibiae dark at base but slightly paler for most of length. Wing membrane transparent to pale brown, veins dark brown. Wings with several crossveins between R vein and Costa beyond cord. Gills apparently absent or, if present, collapsed under submentum. Basal two abdominal segments divided by pleural membrane.
|
|
</paragraph>
|
|
<paragraph blockId="6.[835,1444,223,1849]" pageId="6" pageNumber="7">
|
|
<emphasis bold="true" box="[835,904,829,853]" pageId="6" pageNumber="7">Male.</emphasis>
|
|
Forewing length
|
|
<quantity box="[1130,1220,829,853]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="6" pageNumber="7" unit="mm" value="13.0">13 mm</quantity>
|
|
. Abdominal terga without lobes, but terga 9-10 dark with areas of pale pigment (
|
|
<figureCitation box="[975,1083,900,924]" captionStart="Figs" captionStartId="7.[189,241,1752,1776]" captionTargetBox="[299,1387,606,1672]" captionTargetId="figure-198@7.[317,1316,606,1664]" captionTargetPageId="7" captionText="Figs. 19-22. Perlodes kippenhani structures. 19. Head and pronotum. 20. Male terminalia, dorsal. 21. Male terminalia, ventral. 22. Paraproct, lateral." figureDoi="http://doi.org/10.5281/zenodo.4759551" httpUri="https://zenodo.org/record/4759551/files/figure.png" pageId="6" pageNumber="7">Fig. 20</figureCitation>
|
|
); tergum 10 with pale posteromedian area armed with a small patch of short, anteriorly directed, thickened setae; tergum 8 and 9 with wide patches of sensilla basiconica near posterior margins. Paraprocts with slender, dark sclerite along caudolateral margins and much of anterior area occupied by an expansive, eversible membranonous lobe, which extends forward from apex toward posterior margin of tergum 10 (
|
|
<figureCitation captionStart="Figs" captionStartId="7.[189,241,1752,1776]" captionTargetBox="[299,1387,606,1672]" captionTargetId="figure-198@7.[317,1316,606,1664]" captionTargetPageId="7" captionText="Figs. 19-22. Perlodes kippenhani structures. 19. Head and pronotum. 20. Male terminalia, dorsal. 21. Male terminalia, ventral. 22. Paraproct, lateral." figureDoi="http://doi.org/10.5281/zenodo.4759551" httpUri="https://zenodo.org/record/4759551/files/figure.png" pageId="6" pageNumber="7">Figs. 20- 22</figureCitation>
|
|
). Abdominal sternum 7 with a small median lobe projecting dorsad into intersegmental membrane (
|
|
<figureCitation box="[843,926,1291,1315]" captionStart="Figs" captionStartId="7.[189,241,1752,1776]" captionTargetBox="[299,1387,606,1672]" captionTargetId="figure-198@7.[317,1316,606,1664]" captionTargetPageId="7" captionText="Figs. 19-22. Perlodes kippenhani structures. 19. Head and pronotum. 20. Male terminalia, dorsal. 21. Male terminalia, ventral. 22. Paraproct, lateral." figureDoi="http://doi.org/10.5281/zenodo.4759551" httpUri="https://zenodo.org/record/4759551/files/figure.png" pageId="6" pageNumber="7">Fig. 22</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph blockId="6.[835,1444,223,1849]" box="[835,1062,1327,1351]" pageId="6" pageNumber="7">
|
|
<emphasis bold="true" box="[835,929,1327,1351]" pageId="6" pageNumber="7">Female.</emphasis>
|
|
Unknown.
|
|
</paragraph>
|
|
<paragraph blockId="6.[835,1444,223,1849]" box="[835,1042,1362,1387]" pageId="6" pageNumber="7">
|
|
<emphasis bold="true" box="[835,910,1363,1387]" pageId="6" pageNumber="7">Larva.</emphasis>
|
|
Unknown.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="6" pageNumber="7" type="etymology">
|
|
<paragraph blockId="6.[835,1444,223,1849]" pageId="6" pageNumber="7">
|
|
<emphasis bold="true" box="[835,973,1398,1422]" pageId="6" pageNumber="7">Etymology.</emphasis>
|
|
The patronym honors Michael Kippenhan, collector of the
|
|
<typeStatus box="[1161,1266,1434,1458]" pageId="6" pageNumber="7">holotype</typeStatus>
|
|
specimen.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="diagnosis">
|
|
<paragraph blockId="6.[835,1444,223,1849]" lastBlockId="7.[835,1443,224,497]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
|
|
<emphasis bold="true" box="[835,964,1469,1493]" pageId="6" pageNumber="7">Diagnosis.</emphasis>
|
|
<taxonomicName authorityName="Banks" authorityYear="1903" box="[976,1066,1470,1494]" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
|
<emphasis box="[976,1066,1470,1494]" italics="true" pageId="6" pageNumber="7">Perlodes</emphasis>
|
|
</taxonomicName>
|
|
is poorly known in Asia, and only one species,
|
|
<taxonomicName authority="Ra, Kim, Kang & Ham" authorityName="Ra, Kim, Kang & Ham" authorityYear="1994" box="[1042,1439,1505,1530]" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="stigmata">
|
|
<emphasis box="[1042,1165,1506,1529]" italics="true" pageId="6" pageNumber="7">P. stigmata</emphasis>
|
|
Ra, Kim, Kang & Ham
|
|
</taxonomicName>
|
|
, has been reported from
|
|
<collectingCountry box="[1126,1196,1542,1565]" name="South Korea" pageId="6" pageNumber="7">Korea</collectingCountry>
|
|
(Ra et al. 1994). The new species differs from
|
|
<taxonomicName authorityName="Ra, Kim, Kang & Ham" authorityYear="1994" box="[1170,1303,1577,1600]" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="stigmata">
|
|
<emphasis box="[1170,1303,1577,1600]" italics="true" pageId="6" pageNumber="7">P. stigmata</emphasis>
|
|
</taxonomicName>
|
|
in several respects including having the pronotal disks almost entirely black, in lacking submental gills, in having two rather than three basal abdominal segments divided by pleural membrane, and in details of the male paraprocts. Ra et al. (1994) show the paraprocts of
|
|
<taxonomicName authorityName="Ra, Kim, Kang & Ham" authorityYear="1994" box="[874,1005,1790,1814]" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="stigmata">
|
|
<emphasis box="[874,1005,1790,1814]" italics="true" pageId="6" pageNumber="7">P. stigmata</emphasis>
|
|
</taxonomicName>
|
|
as triangular structures projecting posteriorly and crossing near midlength. In
|
|
<taxonomicName authorityName="Stark" authorityYear="2010" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" lastPageId="7" lastPageNumber="8" order="Plecoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="kippenhani">
|
|
<emphasis italics="true" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">P. kippenhani</emphasis>
|
|
</taxonomicName>
|
|
these structures extend upward and terminate on the anteroapical margin with a large membranous bag, or eversible paraproct lobe (EPL of
|
|
<bibRefCitation author="Zwick, P." box="[189,326,331,355]" pageId="7" pageNumber="8" refId="ref4262" refString="Zwick, P. 1997. Rauserella, a new genus of Plecoptera (Perlodidae), with notes on related genera. Pp. 489 - 496 in Landolt, P. & M. Sartori [eds.]. Ephemeroptera & Plecoptera: Biology-Ecology- Systematics. MTL. Fribourg." type="book" year="1997">Zwick 1997</bibRefCitation>
|
|
) similar to those found in at least some other members of
|
|
<taxonomicName authorityName="Banks" authorityYear="1903" box="[428,518,366,390]" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
|
|
<emphasis box="[428,518,366,390]" italics="true" pageId="7" pageNumber="8">Perlodes</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName authority="(Zwick 1997)" baseAuthorityName="Zwick" baseAuthorityYear="1997" class="Insecta" family="Perlodidae" genus="Filchneria" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
|
|
<emphasis box="[591,699,366,390]" italics="true" pageId="7" pageNumber="8">Filchneria</emphasis>
|
|
(
|
|
<bibRefCitation author="Zwick, P." pageId="7" pageNumber="8" refId="ref4262" refString="Zwick, P. 1997. Rauserella, a new genus of Plecoptera (Perlodidae), with notes on related genera. Pp. 489 - 496 in Landolt, P. & M. Sartori [eds.]. Ephemeroptera & Plecoptera: Biology-Ecology- Systematics. MTL. Fribourg." type="book" year="1997">Zwick 1997</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
. No lobe is present on sternum
|
|
<quantity box="[650,696,402,426]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="7" pageNumber="8" unit="in" value="7.0">7 in</quantity>
|
|
<taxonomicName authority="(Pictet)" baseAuthorityName="Pictet" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="microcephalus">
|
|
<emphasis italics="true" pageId="7" pageNumber="8">Perlodes microcephalus</emphasis>
|
|
(Pictet)
|
|
</taxonomicName>
|
|
or
|
|
<taxonomicName authority="(Rambur)" baseAuthorityName="Rambur" box="[482,704,438,462]" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="disbar">
|
|
<emphasis box="[482,580,438,462]" italics="true" pageId="7" pageNumber="8">P. disbar</emphasis>
|
|
(Rambur)
|
|
</taxonomicName>
|
|
(Zwick pers. comm.) and in
|
|
<taxonomicName authorityName="Stark" authorityYear="2010" box="[462,618,473,497]" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kippenhani">
|
|
<emphasis box="[462,618,473,497]" italics="true" pageId="7" pageNumber="8">P. kippenhani</emphasis>
|
|
</taxonomicName>
|
|
this structure projects dorsad and could be easily overlooked. The new species is also similar to one being described by V.A. Teslenko from the Russian Far East (Teslenko pers. comm.) but in her species the posterior margin of tergum 10 extends upward more strongly than in the Korean species and the sclerotized portion of the paraprocts in the Russian species appears to be more extensive than in
|
|
<taxonomicName authorityName="Stark" authorityYear="2010" box="[1041,1188,473,497]" class="Insecta" family="Perlodidae" genus="Perlodes" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kippenhani">
|
|
<emphasis box="[1041,1188,473,497]" italics="true" pageId="7" pageNumber="8">P. kippenhani</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |