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<document id="CBA87E30FAA3B49874698DB84B60F53A" ID="10.11646/zootaxa.4121.3.5" ID-CLB-Dataset="38908" ID-DOI="10.11646/zootaxa.4121.3.5" ID-GBIF-Dataset="a4fdfbad-bf0d-4ec9-862d-78ed02e359bf" ID-ISSN="1175-5326" ID-Zenodo-Dep="264493" ID-ZooBank="D4D78CAD-FBE6-4B7E-950D-DA99EFF9EDFD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1465453745513" checkinUser="plazi" docAuthor="Šťastný, Jaroslav &amp; Hájek, Jiří" docDate="2016" docId="0385BD14FB0D7932F389FB4A62675487" docLanguage="en" docName="zootaxa.4121.3.5.pdf" docOrigin="Zootaxa 4121 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Agabus (Acatodes) abaensis Šťastný &amp; Hájek, 2016, sp. nov." docType="treatment" docVersion="9" lastPageNumber="316" masterDocId="FFBCC56CFB0C7937F31EFFCF617C5707" masterDocTitle="Agabus abaensis sp. nov., a peculiar species of the confinis - group from central China (Coleoptera: Dytiscidae: Agabinae)" masterLastPageNumber="318" masterPageNumber="311" pageNumber="312" updateTime="1698662792877" updateUser="ExternalLinkService">
<mods:mods id="5E52ED225AD0EDAC8C9E2ACD6F4845A3" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4FEB8EA4C385D0E48D5489B922DC57F3">
<mods:title id="BFBFD50B52768155D0A07383F643EBC9">Agabus abaensis sp. nov., a peculiar species of the confinis - group from central China (Coleoptera: Dytiscidae: Agabinae)</mods:title>
</mods:titleInfo>
<mods:name id="CE6384666BF282438E03F2211BB2BD66" type="personal">
<mods:role id="E752889E0B6207E966963B8DB63BFC13">
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</mods:role>
<mods:namePart id="E62E468D0C62CCA2565629F8A77D9226">Šťastný, Jaroslav</mods:namePart>
</mods:name>
<mods:name id="BF0B45D09207CBA641CC51587C7D70E0" type="personal">
<mods:role id="8AE35CDD04DB82552173E867C73F1497">
<mods:roleTerm id="97AB7363A8E849C6C51766B0DCC70381">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CF00EA807F54A5D574B68B309519EC3A">Hájek, Jiří</mods:namePart>
</mods:name>
<mods:typeOfResource id="73C8B58E9A1FB04568B821FAA14736F2">text</mods:typeOfResource>
<mods:relatedItem id="101C794F8E968960684974172050F0BA" type="host">
<mods:titleInfo id="D111AEC6F51E8F65BC2F932E5339733A">
<mods:title id="D95151734260520E6395A0AF41FA636C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="86B232D00B0CB40DDFA2A67AFC16A221">
<mods:date id="422BA722F0AC7046F9DC9564E0F2916F">2016</mods:date>
<mods:detail id="34DCE2190B3C451EA0AC83EF0557A252" type="volume">
<mods:number id="280A03CBA08F5C33547E56225CA0760F">4121</mods:number>
</mods:detail>
<mods:detail id="E94188F45FAD2149A75C8D63C585FF3B" type="issue">
<mods:number id="C6FD8BA423A79F42FFF41DA77B93F663">3</mods:number>
</mods:detail>
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</mods:extent>
</mods:part>
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<mods:classification id="92E6A3A8E54282C3D39C08481BDD8E49">journal article</mods:classification>
<mods:identifier id="DC71792917E6B60AE5B3A98BF0BAB316" type="CLB-Dataset">38908</mods:identifier>
<mods:identifier id="CD40558CF0B28BE302FD259FCB432F96" type="DOI">10.11646/zootaxa.4121.3.5</mods:identifier>
<mods:identifier id="F7749A22E5A533B3BB7D9996AB445755" type="GBIF-Dataset">a4fdfbad-bf0d-4ec9-862d-78ed02e359bf</mods:identifier>
<mods:identifier id="795097E6B21DFB98AE74734642153833" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="F5B1D5FCDBA0C22517ACD232E5F6307F" type="Zenodo-Dep">264493</mods:identifier>
<mods:identifier id="8BF94FBC5997BD5E9165F20339E2E731" type="ZooBank">D4D78CAD-FBE6-4B7E-950D-DA99EFF9EDFD</mods:identifier>
</mods:mods>
<treatment id="0385BD14FB0D7932F389FB4A62675487" ID-DOI="http://doi.org/10.5281/zenodo.6078714" ID-GBIF-Taxon="120826691" ID-Zenodo-Dep="6078714" LSID="urn:lsid:plazi:treatment:0385BD14FB0D7932F389FB4A62675487" httpUri="http://treatment.plazi.org/id/0385BD14FB0D7932F389FB4A62675487" lastPageId="5" lastPageNumber="316" pageId="1" pageNumber="312">
<subSubSection id="C3365F89FB0D7936F389FB4A620D5201" pageId="1" pageNumber="312" type="nomenclature">
<paragraph id="8B930C02FB0D7936F389FB4A63325398" blockId="1.[151,590,1157,1216]" box="[151,590,1157,1183]" pageId="1" pageNumber="312">
<heading id="D0DBBB6EFB0D7936F389FB4A63325398" bold="true" box="[151,590,1157,1183]" fontSize="11" level="1" pageId="1" pageNumber="312" reason="1">
<emphasis id="B958D010FB0D7936F389FB4A63325398" bold="true" box="[151,590,1157,1183]" pageId="1" pageNumber="312">
<taxonomicName id="4C2C7781FB0D7936F389FB4A60955398" ID-CoL="8LZYH" box="[151,489,1157,1183]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="312" phylum="Arthropoda" rank="species" species="abaensis" status="sp. nov." subGenus="Acatodes">
<emphasis id="B958D010FB0D7936F389FB4A618E5398" bold="true" box="[151,242,1157,1183]" italics="true" pageId="1" pageNumber="312">Agabus</emphasis>
(
<emphasis id="B958D010FB0D7936F21DFB4A600C5398" bold="true" box="[259,368,1157,1183]" italics="true" pageId="1" pageNumber="312">Acatodes</emphasis>
)
<emphasis id="B958D010FB0D7936F29FFB4A60955398" bold="true" box="[385,489,1157,1183]" italics="true" pageId="1" pageNumber="312">abaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26B6D6BFB0D7936F2EFFB4A63325398" box="[497,590,1157,1183]" pageId="1" pageNumber="312" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B930C02FB0D7936F389FB68606D53C7" blockId="1.[151,590,1157,1216]" box="[151,273,1191,1216]" pageId="1" pageNumber="312">
(
<figureCitation id="13171087FB0D7936F381FB68607453C7" box="[159,264,1191,1216]" captionStart-0="FIGURES 1 – 2" captionStart-1="FIGURES 3 – 8" captionStartId-0="2.[151,264,1388,1411]" captionStartId-1="4.[151,264,1817,1840]" captionTargetBox-0="[151,1436,303,1361]" captionTargetBox-1="[205,1382,214,1759]" captionTargetId-0="figure@2.[151,1436,303,1363]" captionTargetId-1="figure@4.[151,1436,193,1796]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionText-0="FIGURES 1 – 2. Habitus of Agabus abaensis sp. nov.: 1. male holotype; 2. female paratype." captionText-1="FIGURES 3 – 8. Agabus abaensis sp. nov.: 3. median lobe of aedeagus in lateral view; 4. same in ventral view; 5. paramere; 6. gonocoxa and tergit IX; 7. gonocoxosternum; 8. male protarsus." httpUri-0="https://zenodo.org/record/264494/files/figure.png" httpUri-1="https://zenodo.org/record/264495/files/figure.png" pageId="1" pageNumber="312">Figs. 1–8</figureCitation>
)
</paragraph>
<paragraph id="8B930C02FB0D7936F389FB3F620D5201" blockId="1.[151,881,1264,1286]" box="[151,881,1264,1286]" pageId="1" pageNumber="312">
<treatmentCitationGroup id="AB3C2B2CFB0D7936F389FB3F620D5201" box="[151,881,1264,1286]" pageId="1" pageNumber="312">
<taxonomicName id="4C2C7781FB0D7936F389FB3F619F5201" ID-CoL="QG7" box="[151,227,1264,1286]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="312" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="B958D010FB0D7936F389FB3F619F5201" box="[151,227,1264,1286]" italics="true" pageId="1" pageNumber="312">Agabus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26B6D6BFB0D7936F3F7FB3E605C5201" box="[233,288,1265,1286]" pageId="1" pageNumber="312" rank="species">n. sp.</taxonomicNameLabel>
(
<collectingCountry id="F33B4C92FB0D7936F233FB3F600C5201" box="[301,368,1264,1286]" name="China" pageId="1" pageNumber="312">China</collectingCountry>
):
<treatmentCitation id="0A8D2A13FB0D7936F29EFB3E636C5201" author="Fery" box="[384,528,1264,1286]" page="50" pageId="1" pageNumber="312" year="2011">
<bibRefCitation id="EFBD71F3FB0D7936F29EFB3E60965201" author="Fery" box="[384,490,1264,1286]" pageId="1" pageNumber="312" refString="Fery, H. (2011) Agabus (Acatodes) puetzi sp. n., a new species of the confinis - group from the Qinhai-Tibet Plateau (Coleoptera, Dytiscidae, Agabini). Zootaxa, 3050, 37 - 54." type="journal article" year="2011">Fery 2011</bibRefCitation>
: 50
</treatmentCitation>
, Fig. 39 (drawing of median lobe).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3365F89FB0D7936F389FAFD63E751B4" pageId="1" pageNumber="312" type="materials_examined">
<paragraph id="8B930C02FB0D7936F389FAFD6438524C" blockId="1.[151,1437,1330,2003]" box="[151,1348,1330,1355]" pageId="1" pageNumber="312">
<emphasis id="B958D010FB0D7936F389FAFD6049524C" bold="true" box="[151,309,1330,1355]" pageId="1" pageNumber="312">
<typeStatus id="5497B2A0FB0D7936F389FAFD61AE524C" box="[151,210,1330,1355]" pageId="1" pageNumber="312">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F33B4C92FB0D7936F222FAFD60FE524C" box="[316,386,1330,1355]" name="China" pageId="1" pageNumber="312">China</collectingCountry>
, Gansu province, Dagcanglhamo (= Langmusi), ca.
<geoCoordinate id="EE186AC5FB0D7936F0D4FAFC6550524C" box="[970,1068,1331,1355]" direction="north" orientation="latitude" pageId="1" pageNumber="312" precision="55" value="34.081">34.081N</geoCoordinate>
<geoCoordinate id="EE186AC5FB0D7936F72DFAFD65DE524C" box="[1075,1186,1330,1355]" direction="east" orientation="longitude" pageId="1" pageNumber="312" precision="55" value="102.639">102.639E</geoCoordinate>
,
<quantity id="4CD4A1E7FB0D7936F7B3FAFC6475524D" box="[1197,1289,1331,1355]" metricMagnitude="0" metricUnit="m" metricValue="3.42" pageId="1" pageNumber="312" unit="m" value="3.42">3.420 m</quantity>
a.s.l.
</paragraph>
<paragraph id="8B930C02FB0D7936F3D9FA9963E751B4" blockId="1.[151,1437,1330,2003]" pageId="1" pageNumber="312">
<emphasis id="B958D010FB0D7936F3D9FA9960045268" bold="true" box="[199,376,1366,1391]" pageId="1" pageNumber="312">
<typeStatus id="5497B2A0FB0D7936F3D9FA99607E5268" box="[199,258,1366,1391]" pageId="1" pageNumber="312">Type</typeStatus>
material.
</emphasis>
<typeStatus id="5497B2A0FB0D7936F29DFA9960975268" box="[387,491,1366,1391]" pageId="1" pageNumber="312" type="holotype">Holotype</typeStatus>
Ƌ (
<collectionCode id="ED3D94C7FB0D7936F100FA98630F5268" box="[542,627,1367,1391]" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="312">NMPC</collectionCode>
), labeled: “
<collectingCountry id="F33B4C92FB0D7936F1E6FA98622B5268" box="[760,855,1367,1391]" name="China" pageId="1" pageNumber="312">CHINA</collectingCountry>
: Gansu province / Dagcanglhamo (= Langmusi) /
<quantity id="4CD4A1E7FB0D7936F389FAB560445295" box="[151,312,1402,1427]" metricMagnitude="3" metricUnit="m" metricValue="3.554" metricValueMax="3.6439999999999997" metricValueMin="3.464" pageId="1" pageNumber="312" unit="m" value="3554.0" valueMax="3644.0" valueMin="3464.0">3464–3644 m</quantity>
/
<date id="FF922AC2FB0D7936F244FAB563775294" box="[346,523,1402,1427]" pageId="1" pageNumber="312" value="2005-06-23" valueMax="2005-06-25" valueMin="2005-06-23">23.–25.VI.2005</date>
/ 34°04.6–05.1'N 102°37.7–38.1'E, / J. Hájek, D. Král &amp; J. Růžička leg. // individually collected, grasslands, close valley with loess loam, stream, pools, margin of coniferous forest [printed] //
<typeStatus id="5497B2A0FB0D7936F3AFFA0C603952DC" box="[177,325,1475,1499]" pageId="1" pageNumber="312" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="4C2C7781FB0D7936F27EFA0D634752DC" authority="Acatodes" authorityName="Acatodes" box="[352,571,1474,1499]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="312" phylum="Arthropoda" rank="genus">Agabus (Acatodes)</taxonomicName>
/
<taxonomicName id="4C2C7781FB0D7936F149FA0D63CA52DC" box="[599,694,1474,1499]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="312" phylum="Arthropoda" rank="species" species="abaensis" status="sp. nov.">abaensis</taxonomicName>
<taxonomicNameLabel id="A26B6D6BFB0D7936F1DEFA0C626652DD" box="[704,794,1475,1498]" pageId="1" pageNumber="312" rank="species">sp. nov.</taxonomicNameLabel>
/ J. Šťastný &amp; J. Hájek det. 2013 [red label, printed]”.
<typeStatus id="5497B2A0FB0D7936F389FA28607452F8" box="[151,264,1511,1535]" pageId="1" pageNumber="312" type="paratype">Paratypes</typeStatus>
: 4Ƌ, 14♀, same label data as
<typeStatus id="5497B2A0FB0D7936F14FFA2963CF52F9" box="[593,691,1510,1534]" pageId="1" pageNumber="312" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED3D94C7FB0D7936F1DFFA28626552F8" box="[705,793,1511,1535]" pageId="1" pageNumber="312">BMNH</collectionCode>
,
<collectionCode id="ED3D94C7FB0D7936F03AFA28620452F8" box="[804,888,1511,1535]" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="1" pageNumber="312">IZCAS</collectionCode>
,
<collectionCode id="ED3D94C7FB0D7936F09DFA2862A952F8" box="[899,981,1511,1535]" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="312">NMPC</collectionCode>
,
<collectionCode id="ED3D94C7FB0D7936F0FEFA28655652F8" LSID="urn:lsid:biocol.org:col:13728" box="[992,1066,1511,1535]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13728" name="National Sun Yat-Sen University, Department of Biological Sciences" pageId="1" pageNumber="312">SYSU</collectionCode>
,
<collectionCode id="ED3D94C7FB0D7936F72BFA2865F452F8" box="[1077,1160,1511,1535]" pageId="1" pageNumber="312">ZSMG</collectionCode>
); 1Ƌ, 1♀, “
<collectingCountry id="F33B4C92FB0D7936F61BFA28641F52F8" box="[1285,1379,1511,1535]" name="China" pageId="1" pageNumber="312">CHINA</collectingCountry>
, NW Sichuan,
<quantity id="4CD4A1E7FB0D7936F21AF9C560185125" box="[260,356,1546,1571]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="312" unit="m" value="3500.0">3500 m</quantity>
., /
<quantity id="4CD4A1E7FB0D7936F293F9C560A55124" box="[397,473,1546,1571]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="312" unit="km" value="15.0">15 km</quantity>
NW ABA,
<geoCoordinate id="EE186AC5FB0D7936F171F9C463BD5124" box="[623,705,1547,1571]" direction="north" orientation="latitude" pageId="1" pageNumber="312" precision="555" value="33.0">33.00N</geoCoordinate>
/ /
<geoCoordinate id="EE186AC5FB0D7936F1F4F9C562375124" box="[746,843,1546,1571]" direction="east" orientation="longitude" pageId="1" pageNumber="312" precision="555" value="101.36">101.36E</geoCoordinate>
, border field/road / J.Rejsek leg.,
<date id="FF922AC2FB0D7936F7FBF9C564E45124" box="[1253,1432,1546,1571]" pageId="1" pageNumber="312" value="2001-07-29" valueMax="2001-07-30" valueMin="2001-07-29">29.–30.vii.2001</date>
, [printed]” (
<collectionCode id="ED3D94C7FB0D7936F20AF9E060245140" box="[276,344,1583,1607]" pageId="1" pageNumber="312">JSCL</collectionCode>
); 1Ƌ, “
<collectingCountry id="F33B4C92FB0D7936F2BBF9E0637F5140" box="[421,515,1583,1607]" name="China" pageId="1" pageNumber="312">CHINA</collectingCountry>
, N Sichuan, / pass
<quantity id="4CD4A1E7FB0D7936F1C8F9E062605140" box="[726,796,1582,1607]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="312" unit="km" value="10.0">10 km</quantity>
E SANGARMAI, /
<geoCoordinate id="EE186AC5FB0D7936F71EF9E165295140" box="[1024,1109,1582,1607]" direction="north" orientation="latitude" pageId="1" pageNumber="312" precision="555" value="32.14">32.14N</geoCoordinate>
/
<geoCoordinate id="EE186AC5FB0D7936F740F9E165C05140" box="[1118,1212,1582,1607]" direction="east" orientation="longitude" pageId="1" pageNumber="312" precision="555" value="102.36">102.36E</geoCoordinate>
,
<quantity id="4CD4A1E7FB0D7936F7D8F9E0645C5141" box="[1222,1312,1583,1607]" metricMagnitude="3" metricUnit="m" metricValue="4.3" pageId="1" pageNumber="312" unit="m" value="4300.0">4300 m</quantity>
., / J.Rejsek leg.,
<date id="FF922AC2FB0D7936F3CDF99D600B516C" box="[211,375,1618,1643]" pageId="1" pageNumber="312" value="2001-07-08" valueMax="2001-07-11" valueMin="2001-07-08">8.–11.vii.2001</date>
, / alpine meadows/screes [printed]” (
<collectionCode id="ED3D94C7FB0D7936F033F99C620D516C" box="[813,881,1619,1643]" pageId="1" pageNumber="312">JSCL</collectionCode>
); 1Ƌ, 2♀, “
<collectingCountry id="F33B4C92FB0D7936F0E1F99C6521516C" box="[1023,1117,1619,1643]" name="China" pageId="1" pageNumber="312">CHINA</collectingCountry>
, N Sichuan, /
<quantity id="4CD4A1E7FB0D7936F612F99D6424516C" box="[1292,1368,1618,1643]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="312" unit="km" value="25.0">25 km</quantity>
NNE ZHANGLA,
<geoCoordinate id="EE186AC5FB0D7936F22EF9B960F85188" box="[304,388,1654,1679]" direction="north" orientation="latitude" pageId="1" pageNumber="312" precision="555" value="33.04">33.04N</geoCoordinate>
/
<geoCoordinate id="EE186AC5FB0D7936F293F9B960975188" box="[397,491,1654,1679]" direction="east" orientation="longitude" pageId="1" pageNumber="312" precision="555" value="103.44">103.44E</geoCoordinate>
,
<quantity id="4CD4A1E7FB0D7936F2EEF9B963325189" box="[496,590,1654,1679]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="312" unit="m" value="3500.0">3500 m</quantity>
., / J.Rejsek leg.,
<date id="FF922AC2FB0D7936F013F9B962D75188" box="[781,939,1654,1679]" pageId="1" pageNumber="312" value="2001-08-03" valueMax="2001-08-05" valueMin="2001-08-03">3.–5.viii.2001</date>
[printed]” (
<collectionCode id="ED3D94C7FB0D7936F729F9B865055188" box="[1079,1145,1655,1679]" pageId="1" pageNumber="312">JSCL</collectionCode>
, CHF). Each
<typeStatus id="5497B2A0FB0D7936F600F9B864025188" box="[1310,1406,1655,1679]" pageId="1" pageNumber="312" type="paratype">paratype</typeStatus>
is provided with the respective red printed label.
</paragraph>
</subSubSection>
<subSubSection id="C3365F89FB0D7934F3D9F97160255093" lastPageId="3" lastPageNumber="314" pageId="1" pageNumber="312" type="description">
<paragraph id="8B930C02FB0D7936F3D9F97163285018" blockId="1.[151,1437,1330,2003]" pageId="1" pageNumber="312">
<emphasis id="B958D010FB0D7936F3D9F971635C51D0" bold="true" box="[199,544,1726,1751]" pageId="1" pageNumber="312">
Description of the
<typeStatus id="5497B2A0FB0D7936F2ACF971636751D0" box="[434,539,1726,1751]" pageId="1" pageNumber="312" type="holotype">holotype</typeStatus>
.
</emphasis>
In dorsal view elongate oval, rather narrow (TL/MW: 1.9). Lateral outline weakly curved with a weak discontinuity at level of pronotal base in dorsal aspect (
<figureCitation id="13171087FB0D7936F779F92D65D051FC" box="[1127,1196,1762,1787]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,264,1388,1411]" captionTargetBox="[151,1436,303,1361]" captionTargetId="figure@2.[151,1436,303,1363]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Habitus of Agabus abaensis sp. nov.: 1. male holotype; 2. female paratype." httpUri="https://zenodo.org/record/264494/files/figure.png" pageId="1" pageNumber="312">Fig. 1</figureCitation>
). Broadest in elytral midlength. Entire surface weakly shiny.
</paragraph>
<paragraph id="8B930C02FB0D7935F3D9F8E563DE57FF" blockId="1.[151,1437,1330,2003]" lastBlockId="2.[151,1436,151,248]" lastPageId="2" lastPageNumber="313" pageId="1" pageNumber="312">
<emphasis id="B958D010FB0D7936F3D9F8E560335044" bold="true" box="[199,335,1834,1859]" pageId="1" pageNumber="312">Coloration.</emphasis>
Head black; two posteromedial spots on vertex and narrow anterior margin of clypeus dark brown; labrum brownish; antenna testaceous basally, antennomeres V–XI reddish brown to piceous apically; maxillary and labial palpi testaceous, terminal palpomere of each palpus infuscate apically. Pronotum black with very narrow lateral bead dark rufous to piceous, postero- and anteromedially indistinctly reddish-brown translucent. Elytra dark brown to piceous; base, lateral margin, epipleura and humeral angles diffusely pale rufotestaceous. Ventral surface mainly black; posterior margin of metacoxal process and abdominal ventrites III–VI transparently rufous. Legs rufo-piceous; femora posteromedially piceous to black, dark coloration more extensive on metafemora than on pro- and mesofemora).
</paragraph>
<caption id="DF535C8AFB0E7935F389FAA365465285" box="[151,1082,1388,1411]" httpUri="https://zenodo.org/record/264494/files/figure.png" pageId="2" pageNumber="313" targetBox="[151,1436,303,1361]" targetPageId="2">
<paragraph id="8B930C02FB0E7935F389FAA365465285" blockId="2.[151,1082,1388,1411]" box="[151,1082,1388,1411]" pageId="2" pageNumber="313">
<emphasis id="B958D010FB0E7935F389FAA360465286" bold="true" box="[151,314,1388,1411]" pageId="2" pageNumber="313">FIGURES 1–2.</emphasis>
Habitus of
<taxonomicName id="4C2C7781FB0E7935F2AFFAA363205285" box="[433,604,1388,1410]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="313" phylum="Arthropoda" rank="species" species="abaensis" status="sp. nov.">
<emphasis id="B958D010FB0E7935F2AFFAA363205285" box="[433,604,1388,1410]" italics="true" pageId="2" pageNumber="313">Agabus abaensis</emphasis>
</taxonomicName>
<emphasis id="B958D010FB0E7935F17CFAA263CE5285" bold="true" box="[610,690,1389,1410]" pageId="2" pageNumber="313">
<taxonomicNameLabel id="A26B6D6BFB0E7935F17CFAA263CE5285" box="[610,690,1389,1410]" pageId="2" pageNumber="313" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 1. male holotype; 2. female paratype.
</paragraph>
</caption>
<paragraph id="8B930C02FB0E7935F3D9FA0760BF5176" blockId="2.[151,1437,1480,1901]" pageId="2" pageNumber="313">
<emphasis id="B958D010FB0E7935F3D9FA0763EE52E6" bold="true" box="[199,658,1480,1505]" pageId="2" pageNumber="313">Surface sculpture and setation. Head.</emphasis>
Frons anteriorly between eyes with two shallow oblique transverse grooves containing of irregular row of about seven deeply impressed punctures; along inner side of eyes with row of punctures. Setation absent. Reticulation simple, consisting of weakly impressed, more or less polygonal meshes of irregular size and shape; some meshes opened, occasionally also with central micropuncture; meshes more elongate along eyes.
</paragraph>
<paragraph id="8B930C02FB0E7935F3D9F9B3625E506A" blockId="2.[151,1437,1480,1901]" pageId="2" pageNumber="313">
<emphasis id="B958D010FB0E7935F3D9F9B360355192" bold="true" box="[199,329,1660,1685]" pageId="2" pageNumber="313">Pronotum.</emphasis>
Reticulation consisting of well impressed meshes of varying size and shape; some meshes opened, majority of meshes with 1–2 small distinct punctures inside. Meshes generally larger and shallower medially and smaller and more impressed laterally. Indistinct microreticulation perceptible only on sides of pronotum. Punctation consisting of irregular row of coarse setigerous punctures along margins; punctures coarser along anterior and posterior margin, finer along lateral margin. Punctures along anterior margin slightly sparser medially; posterior row of punctures broadly interrupted medially, but not interrupted sublaterally. Centre of pronotal disc with distinct longitudinal furrow.
</paragraph>
<paragraph id="8B930C02FB0F7934F3D9FF5863205663" blockId="3.[151,1437,151,1940]" pageId="3" pageNumber="314">
<emphasis id="B958D010FB0F7934F3D9FF58606957B7" bold="true" box="[199,277,151,176]" pageId="3" pageNumber="314">Elytra</emphasis>
with distinct double reticulation. Large meshes of varying shape and size well impressed; meshes rather broad (not elongate), on disc polygonal; some meshes with central puncture. Microreticulation presents on entire surface, slightly less distinct anteromesally, consisting of very fine, shallowly impressed polygonal meshes. Punctation consisting of coarse setigerous punctures arranged in sutural, two discal and two lateral longitudinal puncture lines; in second quarter of elytral length with few additional punctures between lines; lines become very irregular in apical quarter.
</paragraph>
<paragraph id="8B930C02FB0F7934F3D9FEA060D955AF" blockId="3.[151,1437,151,1940]" pageId="3" pageNumber="314">
<emphasis id="B958D010FB0F7934F3D9FEA0601D568F" bold="true" box="[199,353,367,392]" pageId="3" pageNumber="314">Ventral side</emphasis>
with almost entire surface reticulated. Genae with moderately large transverse meshes; prosternum with small meshes except for glabrous tectiform medial part; legs, meso- and metaventrite with small transverse meshes; metacoxal plates with double reticulation, large meshes elongate, small meshes almost isodiametric; abdominal ventrites I–V with transverse meshes medially and oblique meshes laterally; abdominal ventrite VI with double reticulation. Punctation consisting of coarse setigerous punctures on gula, prosternum, legs, and abdominal ventrites; on ventrites arranged in transverse lines; fine punctures present on prosternal process and sparsely distributed on metaventrite. Most of ventral surface covered with strioles; strioles short and transverse on metaventrite, longitudinal on metacoxae, and very long, medially transverse and laterally oblique on abdominal ventrites.
</paragraph>
<paragraph id="8B930C02FB0F7934F3D9FD7B64075237" blockId="3.[151,1437,151,1940]" pageId="3" pageNumber="314">
<emphasis id="B958D010FB0F7934F3D9FD7B603A55CA" bold="true" box="[199,326,692,717]" pageId="3" pageNumber="314">Structures</emphasis>
. Head moderately broad, head basal width/maximal pronotal width about 0.63; clypeus with anterior bead narrow and complete. Pronotum broad, maximal pronotal width/pronotal length 2.5, in dorsal view with sides very slightly arcuate, weakly curved before anterior and posterior angles; posterior angles rectangular with obtuse tips, anterior angles acute with tuft of thick golden setae; anterior bead fine, broadly interrupted in middle, lateral bead relatively narrow and well-marked except for anterior angles. Prosternum broadly rounded in cross-section. Prosternum between procoxae tectiform in cross-section, with midline of convexity rounded; blade of prosternal process lanceolate, flat or weakly convex, apex sharply acuminate; lateral beading narrow but distinct, vanishing before apex. Metaventrite with anteromedial process bluntly pointed anteriorly, reaching mesoventral fork beyond midlength of mesocoxae; process with anteromedial impression for reception of prosternal blade almost attaining level of posterior margin of mesocoxae; “metaventral (metasternal) wings” broadly triangular, WC/WS 2.0. Lines of metacoxal processes diverging anteriorly, reaching posterior margin of metaventrite. Abdominal ventrite VI with posterior margin evenly rounded with well-defined beading. Pro- and mesotarsomeres I–III moderately broadly dilated, ventrally with numerous very small, similar sized, slightly ovate suckers on long stalk and some adhesive setae; protarsal claws elongate and relatively straight, strongly curved near apex, anterior claw with ventral tooth (
<figureCitation id="13171087FB0F7934F1C1FB63625F53C3" box="[735,803,1196,1221]" captionStart="FIGURES 3 – 8" captionStartId="4.[151,264,1817,1840]" captionTargetBox="[205,1382,214,1759]" captionTargetId="figure@4.[151,1436,193,1796]" captionTargetPageId="4" captionText="FIGURES 3 – 8. Agabus abaensis sp. nov.: 3. median lobe of aedeagus in lateral view; 4. same in ventral view; 5. paramere; 6. gonocoxa and tergit IX; 7. gonocoxosternum; 8. male protarsus." httpUri="https://zenodo.org/record/264495/files/figure.png" pageId="3" pageNumber="314">Fig. 8</figureCitation>
), somewhat shorter than protarsomere V (protarsomere V is 1.5 times as long as protarsal claws), posterior claw very slightly shorter than anterior one; mesotarsal claws rather short, widened; metatarsomere I being 1.3 times as long as longer (posterior) tibial spur; metatarsomere V long, ca. 2.1 times longer than metatarsomere IV; metatarsal claws equal in length, arcuate.
</paragraph>
<paragraph id="8B930C02FB0F7934F3D9FAF3654E52C7" blockId="3.[151,1437,151,1940]" pageId="3" pageNumber="314">
<emphasis id="B958D010FB0F7934F3D9FAF3600B5252" bold="true" box="[199,375,1340,1365]" pageId="3" pageNumber="314">Male genitalia</emphasis>
. Median lobe robust with large base, in natural position apically turned to right in ventral aspect (
<figureCitation id="13171087FB0F7934F3F0FA906049527F" box="[238,309,1375,1400]" captionStart="FIGURES 3 – 8" captionStartId="4.[151,264,1817,1840]" captionTargetBox="[205,1382,214,1759]" captionTargetId="figure@4.[151,1436,193,1796]" captionTargetPageId="4" captionText="FIGURES 3 – 8. Agabus abaensis sp. nov.: 3. median lobe of aedeagus in lateral view; 4. same in ventral view; 5. paramere; 6. gonocoxa and tergit IX; 7. gonocoxosternum; 8. male protarsus." httpUri="https://zenodo.org/record/264495/files/figure.png" pageId="3" pageNumber="314">Fig. 4</figureCitation>
), apex with subapical dorsal spine in lateral view (
<figureCitation id="13171087FB0F7934F067FA9062C2527F" box="[889,958,1375,1400]" captionStart="FIGURES 3 – 8" captionStartId="4.[151,264,1817,1840]" captionTargetBox="[205,1382,214,1759]" captionTargetId="figure@4.[151,1436,193,1796]" captionTargetPageId="4" captionText="FIGURES 3 – 8. Agabus abaensis sp. nov.: 3. median lobe of aedeagus in lateral view; 4. same in ventral view; 5. paramere; 6. gonocoxa and tergit IX; 7. gonocoxosternum; 8. male protarsus." httpUri="https://zenodo.org/record/264495/files/figure.png" pageId="3" pageNumber="314">Fig. 3</figureCitation>
), ventral groove more or less straight and open over entire length. Parameres (
<figureCitation id="13171087FB0F7934F14DFA4B63E1529A" box="[595,669,1412,1437]" captionStart="FIGURES 3 – 8" captionStartId="4.[151,264,1817,1840]" captionTargetBox="[205,1382,214,1759]" captionTargetId="figure@4.[151,1436,193,1796]" captionTargetPageId="4" captionText="FIGURES 3 – 8. Agabus abaensis sp. nov.: 3. median lobe of aedeagus in lateral view; 4. same in ventral view; 5. paramere; 6. gonocoxa and tergit IX; 7. gonocoxosternum; 8. male protarsus." httpUri="https://zenodo.org/record/264495/files/figure.png" pageId="3" pageNumber="314">Fig. 5</figureCitation>
) stylate, with triangular basal part sclerotised, apex with short appendage; apex and mesal margin densely setose, apically with longer setation.
</paragraph>
<paragraph id="8B930C02FB0F7934F3D9FA036243512B" blockId="3.[151,1437,151,1940]" pageId="3" pageNumber="314">
<emphasis id="B958D010FB0F7934F3D9FA03602C52E2" bold="true" box="[199,336,1484,1509]" pageId="3" pageNumber="314">Variability.</emphasis>
All specimens of the
<typeStatus id="5497B2A0FB0F7934F155FA02630052E2" box="[587,636,1485,1509]" pageId="3" pageNumber="314">type</typeStatus>
series agree well with the
<typeStatus id="5497B2A0FB0F7934F0AEFA03656452E2" box="[944,1048,1484,1509]" pageId="3" pageNumber="314" type="holotype">holotype</typeStatus>
. The specimens show variation in body size, in extent of piceous coloration of lateral beading of the pronotum, in the dark brown coloration on the elytra, and in the sinuation of lateral sides of the pronotum.
</paragraph>
<paragraph id="8B930C02FB0F7934F3D9F9F8646751E7" blockId="3.[151,1437,151,1940]" pageId="3" pageNumber="314">
Females differ from males in the following characters: color of elytra very slightly lighter, meshes of dorsal surface reticulation slightly larger and more deeply impressed than in male, hence elytra appearing rather submatt (
<figureCitation id="13171087FB0F7934F21AF9B06032519F" box="[260,334,1663,1688]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,264,1388,1411]" captionTargetBox="[151,1436,303,1361]" captionTargetId="figure@2.[151,1436,303,1363]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Habitus of Agabus abaensis sp. nov.: 1. male holotype; 2. female paratype." httpUri="https://zenodo.org/record/264494/files/figure.png" pageId="3" pageNumber="314">Fig. 2</figureCitation>
); pro- and mesotarsomeres I–III not dilated and without adhesive setae; protarsal claws thin, without a ventral tooth, posterior protarsal claw slightly shorter then anterior one; mesotarsal claws longer than in male, thin and evenly curved. Gonocoxa and tergite IX as in
<figureCitation id="13171087FB0F7934F071F90862CF51E7" box="[879,947,1735,1760]" captionStart="FIGURES 3 – 8" captionStartId="4.[151,264,1817,1840]" captionTargetBox="[205,1382,214,1759]" captionTargetId="figure@4.[151,1436,193,1796]" captionTargetPageId="4" captionText="FIGURES 3 – 8. Agabus abaensis sp. nov.: 3. median lobe of aedeagus in lateral view; 4. same in ventral view; 5. paramere; 6. gonocoxa and tergit IX; 7. gonocoxosternum; 8. male protarsus." httpUri="https://zenodo.org/record/264495/files/figure.png" pageId="3" pageNumber="314">Fig. 6</figureCitation>
, gonocoxosternum as in
<figureCitation id="13171087FB0F7934F7CFF908646B51E7" box="[1233,1303,1735,1760]" captionStart="FIGURES 3 – 8" captionStartId="4.[151,264,1817,1840]" captionTargetBox="[205,1382,214,1759]" captionTargetId="figure@4.[151,1436,193,1796]" captionTargetPageId="4" captionText="FIGURES 3 – 8. Agabus abaensis sp. nov.: 3. median lobe of aedeagus in lateral view; 4. same in ventral view; 5. paramere; 6. gonocoxa and tergit IX; 7. gonocoxosternum; 8. male protarsus." httpUri="https://zenodo.org/record/264495/files/figure.png" pageId="3" pageNumber="314">Fig. 7</figureCitation>
.
</paragraph>
<paragraph id="8B930C02FB0F7934F3D9F92360255093" blockId="3.[151,1437,151,1940]" pageId="3" pageNumber="314">
<emphasis id="B958D010FB0F7934F3D9F92360065002" bold="true" box="[199,378,1772,1797]" pageId="3" pageNumber="314">Measurements</emphasis>
.
<typeStatus id="5497B2A0FB0F7934F290F92360815002" box="[398,509,1772,1797]" pageId="3" pageNumber="314" type="holotype">Holotype</typeStatus>
: TL =
<quantity id="4CD4A1E7FB0F7934F143F92363C15003" box="[605,701,1772,1796]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="3" pageNumber="314" unit="mm" value="7.9">7.9 mm</quantity>
, TL-h =
<quantity id="4CD4A1E7FB0F7934F028F92362EA5003" box="[822,918,1772,1796]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="3" pageNumber="314" unit="mm" value="7.4">7.4 mm</quantity>
, MW =
<quantity id="4CD4A1E7FB0F7934F716F92365145003" box="[1032,1128,1772,1796]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="3" pageNumber="314" unit="mm" value="4.2">4.2 mm</quantity>
, TL-h/MW = 1.76; male
<typeStatus id="5497B2A0FB0F7934F389F8DE6076502E" box="[151,266,1809,1833]" pageId="3" pageNumber="314" type="paratype">paratypes</typeStatus>
: TL =
<quantity id="4CD4A1E7FB0F7934F274F8C06086502F" box="[362,506,1807,1832]" metricMagnitude="-3" metricUnit="m" metricValue="8.05" metricValueMax="8.5" metricValueMin="7.6" pageId="3" pageNumber="314" unit="mm" value="8.05" valueMax="8.5" valueMin="7.6">7.6–8.5 mm</quantity>
(8.08±
<quantity id="4CD4A1E7FB0F7934F14DF8DF63C9502F" box="[595,693,1808,1832]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="314" unit="mm" value="0.3">0.3 mm</quantity>
), TL-h = 7.0–
<quantity id="4CD4A1E7FB0F7934F074F8DF62BB502F" box="[874,967,1808,1832]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="3" pageNumber="314" unit="mm" value="7.7">7.7 mm</quantity>
(7.33±
<quantity id="4CD4A1E7FB0F7934F701F8DF65EE502F" box="[1055,1170,1808,1832]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="314" unit="mm" value="0.22">0.22 mm</quantity>
), MW =
<quantity id="4CD4A1E7FB0F7934F612F8DF64E0502F" box="[1292,1436,1808,1832]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="4.4" metricValueMin="4.1" pageId="3" pageNumber="314" unit="mm" value="4.25" valueMax="4.4" valueMin="4.1">4.1–4.4 mm</quantity>
(4.19±
<quantity id="4CD4A1E7FB0F7934F3FFF8FB602D504B" box="[225,337,1844,1868]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="314" unit="mm" value="0.09">0.09 mm</quantity>
), TL-h/MW = 1.70–1.83; (1.53±
<quantity id="4CD4A1E7FB0F7934F1C8F8FB623B504B" box="[726,839,1844,1869]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="3" pageNumber="314" unit="mm" value="0.38">0.38 mm</quantity>
); female
<typeStatus id="5497B2A0FB0F7934F0A7F8FA6557504A" box="[953,1067,1845,1869]" pageId="3" pageNumber="314" type="paratype">paratypes</typeStatus>
: TL =
<quantity id="4CD4A1E7FB0F7934F79BF8FB646F504B" box="[1157,1299,1844,1868]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="8.2" metricValueMin="7.8" pageId="3" pageNumber="314" unit="mm" value="8.0" valueMax="8.2" valueMin="7.8">7.8–8.2 mm</quantity>
(8.01±
<quantity id="4CD4A1E7FB0F7934F679F8FB61B45077" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="314" unit="mm" value="0.16">0.16 mm</quantity>
), TL-h =
<quantity id="4CD4A1E7FB0F7934F25BF89760A85077" box="[325,468,1880,1904]" metricMagnitude="-3" metricUnit="m" metricValue="7.25" metricValueMax="7.4" metricValueMin="7.1" pageId="3" pageNumber="314" unit="mm" value="7.25" valueMax="7.4" valueMin="7.1">7.1–7.4 mm</quantity>
(7.26±
<quantity id="4CD4A1E7FB0F7934F132F89863E65077" box="[556,666,1879,1904]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="314" unit="mm" value="0.11">0.11 mm</quantity>
), MW =
<quantity id="4CD4A1E7FB0F7934F00FF89762DC5077" box="[785,928,1880,1904]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.1" metricValueMin="3.9" pageId="3" pageNumber="314" unit="mm" value="4.0" valueMax="4.1" valueMin="3.9">3.9–4.1 mm</quantity>
(4.04±
<quantity id="4CD4A1E7FB0F7934F0E9F898651B5077" box="[1015,1127,1879,1904]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="314" unit="mm" value="0.06">0.06 mm</quantity>
), TL-h/MW = 1.76–1.85; (1.80±
<quantity id="4CD4A1E7FB0F7934F3FEF8B360305093" box="[224,332,1916,1940]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="314" unit="mm" value="0.02">0.02 mm</quantity>
).
</paragraph>
</subSubSection>
<caption id="DF535C8AFB087933F389F8D662635049" httpUri="https://zenodo.org/record/264495/files/figure.png" pageId="4" pageNumber="315" targetBox="[205,1382,214,1759]" targetPageId="4">
<paragraph id="8B930C02FB087933F389F8D662635049" blockId="4.[151,1436,1817,1870]" pageId="4" pageNumber="315">
<emphasis id="B958D010FB087933F389F8D660465028" bold="true" box="[151,314,1817,1840]" pageId="4" pageNumber="315">FIGURES 3–8.</emphasis>
<taxonomicName id="4C2C7781FB087933F25EF8D560905037" box="[320,492,1818,1840]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="315" phylum="Arthropoda" rank="species" species="abaensis" status="sp. nov.">
<emphasis id="B958D010FB087933F25EF8D560905037" box="[320,492,1818,1840]" italics="true" pageId="4" pageNumber="315">Agabus abaensis</emphasis>
</taxonomicName>
<emphasis id="B958D010FB087933F2ECF8D5633E5028" bold="true" box="[498,578,1818,1839]" pageId="4" pageNumber="315">
<taxonomicNameLabel id="A26B6D6BFB087933F2ECF8D5633E5028" box="[498,578,1818,1839]" pageId="4" pageNumber="315" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 3. median lobe of aedeagus in lateral view; 4. same in ventral view; 5. paramere; 6. gonocoxa and tergit IX; 7. gonocoxosternum; 8. male protarsus.
</paragraph>
</caption>
<subSubSection id="C3365F89FB097932F3D9FF586545551F" pageId="5" pageNumber="316" type="diagnosis">
<paragraph id="8B930C02FB097932F3D9FF586545551F" blockId="5.[151,1436,151,896]" pageId="5" pageNumber="316">
<emphasis id="B958D010FB097932F3D9FF5860AD57B7" bold="true" box="[199,465,151,176]" pageId="5" pageNumber="316">Differential diagnosis.</emphasis>
With the presence of subapical dorsal spine on the median lobe of male, the new species is without any doubt classified in the subgenus
<taxonomicName id="4C2C7781FB097932F004FF7365FC57D3" authority="Nilsson (2000)" authorityName="Nilsson" authorityYear="2000" box="[794,1152,188,213]" class="Insecta" family="Dytiscidae" genus="Acatodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="316" phylum="Arthropoda" rank="genus">
<emphasis id="B958D010FB097932F004FF7362FD57D3" box="[794,897,188,212]" italics="true" pageId="5" pageNumber="316">Acatodes</emphasis>
sensu
<bibRefCitation id="EFBD71F3FB097932F0CCFF7365FC57D3" author="Nilsson" box="[978,1152,188,213]" pageId="5" pageNumber="316" refString="Nilsson, A. N. (2000) A new view of the generic classification of the Agabus - group of genera of the Agabini, aimed at solving the problem with a paraphyletic Agabus (Coleoptera: Dytiscidae). Koleopterologische Rundschau, 70, 17 - 36." type="journal article" year="2000">Nilsson (2000)</bibRefCitation>
</taxonomicName>
. More precisely,
<taxonomicName id="4C2C7781FB097932F656FF73618557FF" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="316" phylum="Arthropoda" rank="species" species="abaensis" status="sp. nov.">
<emphasis id="B958D010FB097932F656FF73618557FF" italics="true" pageId="5" pageNumber="316">Agabus abaensis</emphasis>
</taxonomicName>
<emphasis id="B958D010FB097932F3E1FF10602457FF" bold="true" box="[255,344,223,248]" pageId="5" pageNumber="316">
<taxonomicNameLabel id="A26B6D6BFB097932F3E1FF10602457FF" box="[255,344,223,248]" pageId="5" pageNumber="316" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is assigned to the
<taxonomicName id="4C2C7781FB097932F13AFF2F63E757FF" box="[548,667,224,248]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="316" phylum="Arthropoda" rank="species" species="confinis">
<emphasis id="B958D010FB097932F13AFF2F63E757FF" box="[548,667,224,248]" italics="true" pageId="5" pageNumber="316">A. confinis</emphasis>
</taxonomicName>
species group as defined by
<bibRefCitation id="EFBD71F3FB097932F0C2FF2F650857FF" author="Larson" box="[988,1140,224,248]" pageId="5" pageNumber="316" refString="Larson, D. J. (1989) Revision of North American Agabus Leach (Coleoptera: Dytiscidae): introduction, key to species groups, and classification of the ambiguus -, tristis -, and arcticus - groups. The Canadian Entomologist, 121, 861 - 919. http: // dx. doi. org / 10.4039 / Ent 121861 - 10" type="journal article" year="1989">Larson (1989</bibRefCitation>
,
<bibRefCitation id="EFBD71F3FB097932F761FF2F65C457FF" author="Larson" box="[1151,1208,224,248]" pageId="5" pageNumber="316" refString="Larson, D. J. (1991) Revision of North American Agabus Leach (Coleoptera: Dytiscidae): elongatus -, zetterstedti -, and confinis - groups. The Canadian Entomologist, 123, 1239 - 1317. http: // dx. doi. org / 10.4039 / Ent 1231239 - 6" type="journal article" year="1991">1991</bibRefCitation>
) and
<bibRefCitation id="EFBD71F3FB097932F7EBFF1064E057FF" author="Nilsson" box="[1269,1436,223,248]" pageId="5" pageNumber="316" refString="Nilsson, A. N. (1990) Revisional notes on selected East Palaearctic species of Agabus Leach (Coleoptera, Dytiscidae). Entomologisk Tidskrift, 111, 149 - 161." type="journal article" year="1990">Nilsson (1990)</bibRefCitation>
based on: 1) narrow anterior bead of clypeus; 2) pronotum without broad yellow lateral stripes; 3) posterior angles of pronotum rectangular; 4) apex of prosternal process elongate and pointed; 5) metaventrite with anteromedial emargination long, reaching level of posterior margin of mesocoxae; 6) male pro- and mesotarsomeres I–III moderately dilated, ventrally with numerous small elongate adhesive setae; 7) metatibia impunctate on posteroventral margin; 8) metatarsomere V more than 1.5 times longer than metatarsomere IV; 9) shape of base of male median lobe; and 10) paramere stylate.
<taxonomicName id="4C2C7781FB097932F195FE77623B56D7" box="[651,839,440,464]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="316" phylum="Arthropoda" rank="species" species="abaensis" status="sp. nov.">
<emphasis id="B958D010FB097932F195FE77623B56D7" box="[651,839,440,464]" italics="true" pageId="5" pageNumber="316">Agabus abaensis</emphasis>
</taxonomicName>
<emphasis id="B958D010FB097932F050FE7862DB56D7" bold="true" box="[846,935,439,464]" pageId="5" pageNumber="316">
<taxonomicNameLabel id="A26B6D6BFB097932F050FE7862DB56D7" box="[846,935,439,464]" pageId="5" pageNumber="316" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from all other species in the group by the combination of the following characters: elytral sculpture with considerably large, distinctly double reticulation in both sexes, male anterior protarsal claw dentate, and shape of male median lobe.
</paragraph>
</subSubSection>
<subSubSection id="C3365F89FB097932F3D9FDEB62B5543F" pageId="5" pageNumber="316" type="etymology">
<paragraph id="8B930C02FB097932F3D9FDEB637A5583" blockId="5.[151,1436,151,896]" pageId="5" pageNumber="316">
<emphasis id="B958D010FB097932F3D9FDEB6032553A" bold="true" box="[199,334,548,573]" pageId="5" pageNumber="316">Etymology.</emphasis>
The new species is named after its area of occurrence—the Aba region (formally Ngawa Tibetan and Qiang Autonomous Prefecture) in Sichuan, bordering on Gansu province in the north; the specific epithet is an adjective in the nominative case.
</paragraph>
<paragraph id="8B930C02FB097932F3D9FD4062B5543F" blockId="5.[151,1436,151,896]" pageId="5" pageNumber="316">
<emphasis id="B958D010FB097932F3D9FD40608555AF" bold="true" box="[199,505,655,680]" pageId="5" pageNumber="316">Collecting circumstances.</emphasis>
At the
<typeStatus id="5497B2A0FB097932F151FD5F630355AF" box="[591,639,656,680]" pageId="5" pageNumber="316">type</typeStatus>
locality,
<taxonomicName id="4C2C7781FB097932F1F6FD5F62DA55AF" box="[744,934,656,680]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="316" phylum="Arthropoda" rank="species" species="abaensis" status="sp. nov.">
<emphasis id="B958D010FB097932F1F6FD5F62DA55AF" box="[744,934,656,680]" italics="true" pageId="5" pageNumber="316">Agabus abaensis</emphasis>
</taxonomicName>
<emphasis id="B958D010FB097932F0B0FD40657555AF" bold="true" box="[942,1033,655,680]" pageId="5" pageNumber="316">
<taxonomicNameLabel id="A26B6D6BFB097932F0B0FD40657555AF" box="[942,1033,655,680]" pageId="5" pageNumber="316" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was collected in pools in grassland (
<figureCitation id="13171087FB097932F381FD7B619855CB" box="[159,228,692,717]" captionStart="FIGURE 9" captionStartId="5.[151,250,1827,1849]" captionTargetBox="[151,1436,964,1805]" captionTargetId="figure@5.[151,1436,949,1806]" captionTargetPageId="5" captionText="FIGURE 9. Type locality of Agabus abaensis sp. nov.: pools near Langmusi." httpUri="https://zenodo.org/record/264496/files/figure.png" pageId="5" pageNumber="316">Fig. 9</figureCitation>
). The new species was found in a species rich
<taxonomicName id="4C2C7781FB097932F1F0FD7B621A55CA" box="[750,870,692,717]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="316" phylum="Arthropoda" rank="family">Dytiscidae</taxonomicName>
association, together with
<taxonomicName id="4C2C7781FB097932F78DFD7B61AE55F7" authority="Harold, 1880" authorityName="Harold" authorityYear="1880" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="316" phylum="Arthropoda" rank="species" species="brandti">
<emphasis id="B958D010FB097932F78DFD7B644355CB" box="[1171,1343,692,716]" italics="true" pageId="5" pageNumber="316">Agabus brandti</emphasis>
Harold, 1880
</taxonomicName>
,
<taxonomicName id="4C2C7781FB097932F3C2FD17637C55F7" authority="Guignot, 1952" authorityName="Guignot" authorityYear="1952" box="[220,512,727,752]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="316" phylum="Arthropoda" rank="species" species="lobonyx">
<emphasis id="B958D010FB097932F3C2FD17602955F7" box="[220,341,728,752]" italics="true" pageId="5" pageNumber="316">A. lobonyx</emphasis>
Guignot, 1952
</taxonomicName>
,
<taxonomicName id="4C2C7781FB097932F115FD16628355F7" authority="Falkenstrom, 1936" authorityName="Falkenstrom" authorityYear="1936" box="[523,1023,727,752]" class="Insecta" family="Dytiscidae" genus="Boreonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="316" phylum="Arthropoda" rank="species" species="emmerichi">
<emphasis id="B958D010FB097932F115FD16626D55F7" box="[523,785,728,752]" italics="true" pageId="5" pageNumber="316">Boreonectes emmerichi</emphasis>
(Falkenström, 1936)
</taxonomicName>
,
<taxonomicName id="4C2C7781FB097932F714FD1760315412" authority="Toledo &amp; Mazzoldi, 1996" authorityName="Toledo &amp; Mazzoldi" authorityYear="1996" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="316" phylum="Arthropoda" rank="species" species="nanpingensis">
<emphasis id="B958D010FB097932F714FD17645755E8" box="[1034,1323,728,752]" italics="true" pageId="5" pageNumber="316">Hydroporus nanpingensis</emphasis>
Toledo &amp; Mazzoldi, 1996
</taxonomicName>
,
<taxonomicName id="4C2C7781FB097932F245FD32621C5413" authority="Schaller, 1783" authorityName="Schaller" authorityYear="1783" box="[347,864,764,789]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="316" phylum="Arthropoda" rank="species" species="impressopunctatus">
<emphasis id="B958D010FB097932F245FD3263E15413" box="[347,669,765,788]" italics="true" pageId="5" pageNumber="316">Hygrotus impressopunctatus</emphasis>
(Schaller, 1783)
</taxonomicName>
,
<taxonomicName id="4C2C7781FB097932F071FD3265EA5413" authority="Fery, 2003" authorityName="Fery" authorityYear="2003" box="[879,1174,764,789]" class="Insecta" family="Dytiscidae" genus="Hygrotus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="316" phylum="Arthropoda" rank="species" species="zigetangco">
<emphasis id="B958D010FB097932F071FD3265715413" box="[879,1037,765,788]" italics="true" pageId="5" pageNumber="316">H. zigetangco</emphasis>
Fery, 2003
</taxonomicName>
,
<taxonomicName id="4C2C7781FB097932F7BAFD3361AE543F" authority="Sharp, 1878" authorityName="Sharp" authorityYear="1878" class="Insecta" family="Dytiscidae" genus="Ilybius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="316" phylum="Arthropoda" rank="species" species="cinctus">
<emphasis id="B958D010FB097932F7BAFD3364355413" box="[1188,1353,764,788]" italics="true" pageId="5" pageNumber="316">Ilybius cinctus</emphasis>
Sharp, 1878
</taxonomicName>
,
<taxonomicName id="4C2C7781FB097932F3C3FCEE60A4543F" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" box="[221,472,799,824]" class="Insecta" family="Dytiscidae" genus="Ilybius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="316" phylum="Arthropoda" rank="species" species="obtusus">
<emphasis id="B958D010FB097932F3C3FCEE6035543F" box="[221,329,800,824]" italics="true" pageId="5" pageNumber="316">I. obtusus</emphasis>
Sharp, 1882
</taxonomicName>
and
<taxonomicName id="4C2C7781FB097932F110FCEF62B9543F" authority="Zaitzev, 1908" authorityName="Zaitzev" authorityYear="1908" box="[526,965,799,824]" class="Insecta" family="Dytiscidae" genus="Nebrioporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="316" phylum="Arthropoda" rank="species" species="formaster">
<emphasis id="B958D010FB097932F110FCEF626C543F" box="[526,784,800,824]" italics="true" pageId="5" pageNumber="316">Nebrioporus formaster</emphasis>
(Zaitzev, 1908)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3365F89FB097932F3D9FC8B62675487" pageId="5" pageNumber="316" type="distribution">
<paragraph id="8B930C02FB097932F3D9FC8B62675487" blockId="5.[151,1436,151,896]" pageId="5" pageNumber="316">
<emphasis id="B958D010FB097932F3D9FC8B601C545A" bold="true" box="[199,352,836,861]" pageId="5" pageNumber="316">Distribution.</emphasis>
So far known from several localities in south-western Gansu and northern Sichuan provinces, at the eastern border of the Tibetan plateau,
<collectingCountry id="F33B4C92FB097932F176FCA863D75487" box="[616,683,871,896]" name="China" pageId="5" pageNumber="316">China</collectingCountry>
(
<figureCitation id="13171087FB097932F1A5FCA862725487" box="[699,782,871,896]" captionStart="FIGURE 10" captionStartId="6.[151,250,1207,1229]" captionTargetBox="[184,1384,195,1178]" captionTargetId="figure@6.[181,1386,193,1184]" captionTargetPageId="6" captionText="FIGURE 10. Distributional map of Agabus abaensis sp. nov." httpUri="https://zenodo.org/record/264497/files/figure.png" pageId="5" pageNumber="316">Fig. 10</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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