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<document ID-DOI="10.5252/geodiversitas2021v43a2" ID-GBIF-Dataset="a2545208-1888-44e9-82d3-b429219cd070" ID-ISSN="1638-9395" ID-Zenodo-Dep="4486479" ID-ZooBank="urn:lsid:zoobank.org:pub:6EF0DFAC-609D-407D-B4CC-CB985C3295FC" checkinTime="1612186952565" checkinUser="felipe" docAuthor="Devillez, Julien &amp; Charbonnier, Sylvain" docDate="2021" docId="297587890769FFF0106301BFFD0DAA51" docLanguage="en" docName="Geodiversitas.43.2.25-73.pdf" docOrigin="Geodiversitas 43 (2)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.3:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="3" docTitle="Eryma westphali Schweigert, Dietl &amp; Roper 2000" docType="treatment" docVersion="11" lastPageNumber="42" masterDocId="D54CFFF10778FFE3133E020AFF98AF25" masterDocTitle="Review of the Late Jurassic erymoid lobsters (Crustacea: Decapoda)" masterLastPageNumber="73" masterPageNumber="25" pageNumber="40" updateTime="1675447118536" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title>Review of the Late Jurassic erymoid lobsters (Crustacea: Decapoda)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Devillez, Julien</mods:namePart>
<mods:affiliation>Muséum national dHistoire naturelle, Paris Centre de Recherche en Paléontologie Paris (CR 2 P, UMR 7207), Sorbonne Université, MNHN, UPMC, CNRS, 57 rue Cuvier, F- 75231 Parix cedex 05 Paris (France)</mods:affiliation>
<mods:nameIdentifier type="email">julien.devillez@edu.mnhn.fr</mods:nameIdentifier>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Charbonnier, Sylvain</mods:namePart>
<mods:affiliation>Muséum national dHistoire naturelle, Paris Centre de Recherche en Paléontologie Paris (CR 2 P, UMR 7207), Sorbonne Université, MNHN, UPMC, CNRS, 57 rue Cuvier, F- 75231 Parix cedex 05 Paris (France)</mods:affiliation>
<mods:nameIdentifier type="email">sylvain.charbonnier@mnhn.fr</mods:nameIdentifier>
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<mods:title>Geodiversitas</mods:title>
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<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-01-28</mods:number>
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<mods:detail type="volume">
<mods:number>43</mods:number>
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<mods:number>2</mods:number>
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<mods:start>25</mods:start>
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<mods:identifier type="DOI">10.5252/geodiversitas2021v43a2</mods:identifier>
<mods:identifier type="GBIF-Dataset">a2545208-1888-44e9-82d3-b429219cd070</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4498476" ID-GBIF-Taxon="177207990" ID-Zenodo-Dep="4498476" LSID="urn:lsid:plazi:treatment:297587890769FFF0106301BFFD0DAA51" httpUri="http://treatment.plazi.org/id/297587890769FFF0106301BFFD0DAA51" lastPageId="19" lastPageNumber="42" pageId="17" pageNumber="40">
<subSubSection box="[861,1407,948,975]" pageId="17" pageNumber="40" type="nomenclature">
<paragraph blockId="17.[861,1407,948,1007]" box="[861,1407,948,975]" pageId="17" pageNumber="40">
<taxonomicName authority="Schweigert, Dietl &amp; Roper, 2000" authorityName="Schweigert, Dietl &amp; Roper" authorityYear="2000" box="[861,1407,948,975]" class="Malacostraca" family="Erymidae" genus="Eryma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="40" phylum="Arthropoda" rank="species" species="westphali">
<emphasis bold="true" box="[861,1042,948,975]" italics="true" pageId="17" pageNumber="40">Eryma westphali</emphasis>
<bibRefCitation author="SCHWEIGERT G. &amp; DIETL G. &amp; ROPER M." box="[1049,1407,949,975]" pageId="17" pageNumber="40" pagination="1 - 25" refId="ref35446" refString="SCHWEIGERT G., DIETL G. &amp; ROPER M. 2000. - Die Panzerkrebse des Familie Erymidae van Straelen (Crustacea, Decapoda) aus dem Nusplinger Plattenkalk (Ober-Kimmeridgium, Schwabische Alb) im Vergleich mit frankischen Vorkommen. Stuttgarter Beitrage zur Naturkunde B (285): 1 - 25. https: // www. biodiversitylibrary. org / page / 59557644" type="journal article" year="2000">Schweigert, Dietl &amp; Röper, 2000</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection box="[1074,1194,980,1007]" pageId="17" pageNumber="40" type="description">
<paragraph blockId="17.[861,1407,948,1007]" box="[1074,1194,980,1007]" pageId="17" pageNumber="40">
(
<figureCitation box="[1083,1145,980,1007]" captionStart="FIG" captionStartId="18.[132,143,1894,1911]" captionTargetBox="[132,1455,215,1880]" captionTargetId="figure-28@18.[698,1267,769,1326]" captionTargetPageId="18" captionText="FIG. 7. — Eryma major Oppel, 1861, Eryma veltheimii (Münster, 1839) and Eryma westphali Schweigert, Dietl &amp; Röper, 2000 from the Kimmeridgian of Germany: A, holotype SMNS 3682 of E. major from Egesheimer;B, specimen SMNS 67655 of E. major from Gerhausen; C, specimen SMNS 64371 of E. major from Nusplingen; D-F, holotype BSPG AS VII 186 of E. veltheimii from Kehlheim: part (D), counter-part (E), details of telson and uropods (F); G-J, holotype SMNS 24227 of E. westphali from Nusplingen: general view (G), schema of the carapace (H), schema of left chela (I), schema of right chela (J); C, specimen SMNS 63733 from Nusplingen. Abbreviations: a, branchiocardiac; as, antennal spine; b, antennal groove; b1, hepatic groove; c, postcervical; d, gastro-orbital groove; e1e, cervical groove; ip, intercalated plate. Black arrows show the spines. Photographs and line drawings: J. Devillez. Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4486491" httpUri="https://zenodo.org/record/4486491/files/figure.png" pageId="17" pageNumber="40">Fig. 7</figureCitation>
G-J)
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="40" type="reference_group">
<paragraph blockId="17.[813,1457,1037,1115]" pageId="17" pageNumber="40">
<taxonomicName authority="Schweigert, Dietl &amp; Roper, 2000: 6" authorityName="Schweigert, Dietl &amp; Roper" authorityPageNumber="6" authorityYear="2000" box="[813,1292,1037,1062]" class="Malacostraca" family="Erymidae" genus="Eryma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="40" phylum="Arthropoda" rank="species" species="westphali">
<emphasis box="[813,960,1037,1061]" italics="true" pageId="17" pageNumber="40">Eryma westphali</emphasis>
<bibRefCitation author="SCHWEIGERT G. &amp; DIETL G. &amp; ROPER M." box="[964,1292,1037,1062]" pageId="17" pageNumber="40" pagination="1 - 25" refId="ref35446" refString="SCHWEIGERT G., DIETL G. &amp; ROPER M. 2000. - Die Panzerkrebse des Familie Erymidae van Straelen (Crustacea, Decapoda) aus dem Nusplinger Plattenkalk (Ober-Kimmeridgium, Schwabische Alb) im Vergleich mit frankischen Vorkommen. Stuttgarter Beitrage zur Naturkunde B (285): 1 - 25. https: // www. biodiversitylibrary. org / page / 59557644" type="journal article" year="2000">Schweigert, Dietl &amp; Röper, 2000: 6</bibRefCitation>
</taxonomicName>
, pl. 3, figs 1-3.
<emphasis box="[1430,1457,1038,1061]" italics="true" pageId="17" pageNumber="40"></emphasis>
<bibRefCitation author="DIETL G. &amp; SCHWEIGERT G." box="[813,1104,1064,1088]" pageId="17" pageNumber="40" refId="ref31494" refString="DIETL G. &amp; SCHWEIGERT G. 2001. - Im Reich der Meerengel: der Nusplinger Plattenkalk und seine Fossilien. F. Pfeil, Munchen, 144 p." type="book" year="2001">Dietl &amp; Schweigert 2001: 53</bibRefCitation>
, 110, fig. 85.
<emphasis box="[1246,1271,1065,1088]" italics="true" pageId="17" pageNumber="40"></emphasis>
<bibRefCitation author="GARASSINO A. &amp; SCHWEIGERT G." pageId="17" pageNumber="40" pagination="1 - 64" refId="ref32867" refString="GARASSINO A. &amp; SCHWEIGERT G. 2006. - The Upper Jurassic Solnhofen decapod crustacean fauna: review of the types from old descriptions. Part I. Infraorders Astacidea, Thalassinidea and Palinura. Memorie della Societa italiana di Scienze naturali e del Museo civico di Storia naturale di Milano 34 (1): 1 - 64. https: // www. biodiversitylibrary. org / page / 58176047" type="journal article" year="2006">Garassino &amp; Schweigert 2006: 8</bibRefCitation>
.
<emphasis box="[978,1003,1091,1114]" italics="true" pageId="17" pageNumber="40"></emphasis>
<bibRefCitation author="FELDMANN R. M. &amp; TITUS A. L." box="[1010,1290,1090,1115]" pageId="17" pageNumber="40" pagination="63 - 68" refId="ref32116" refString="FELDMANN R. M. &amp; TITUS A. L. 2006. - Eryma jungostrix n. sp. (Decapoda; Erymidae) from the Redwater Shale of the Stump Formation (Jurassic; Oxfordian) of Utah. Journal of Crustacean Biology 26 (1): 63 - 68. https: // doi. org / 10.1651 / S- 2644.1" type="journal article" year="2006">Feldmann &amp; Titus 2006: 64</bibRefCitation>
.
</paragraph>
<paragraph blockId="17.[813,1456,1138,1188]" pageId="17" pageNumber="40">
<taxonomicName authority="Schweitzer et al. 2010" box="[813,982,1138,1162]" class="Malacostraca" family="Erymidae" genus="Galicia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="40" phylum="Arthropoda" rank="species" species="westphali">
<emphasis box="[813,982,1138,1162]" italics="true" pageId="17" pageNumber="40">Galicia westphali</emphasis>
</taxonomicName>
<bibRefCitation author="SCHWEITZER C. E. &amp; FELDMANN R. M. &amp; GARASSINO A. &amp; KARASAWA H. &amp; SCHWEIGERT G." box="[1014,1281,1138,1162]" pageId="17" pageNumber="40" pagination="1 - 222" refId="ref35549" refString="SCHWEITZER C. E., FELDMANN R. M., GARASSINO A., KARASAWA H. &amp; SCHWEIGERT G. 2010. - Systematic list of fossil decapod crustacean species. Crustaceana Monographs 10: 1 - 222. https: // doi. org / 10.1163 / ej. 9789004178915. i- 222" type="journal article" year="2010">
Schweitzer
<emphasis box="[1129,1180,1138,1162]" italics="true" pageId="17" pageNumber="40">et al.</emphasis>
2010: 25
</bibRefCitation>
.
<emphasis box="[1296,1322,1139,1162]" italics="true" pageId="17" pageNumber="40"></emphasis>
<bibRefCitation author="HYZNY M. &amp; SCHLOGL J. &amp; CHARBONNIER S. &amp; SCHWEIGERT G. &amp; LEAU L. &amp; GOUTTENOIRE M." pageId="17" pageNumber="40" pagination="371 - 384" refId="ref33235" refString="HYZNY M., SCHLOGL J., CHARBONNIER S., SCHWEIGERT G., RUL- LEAU L. &amp; GOUTTENOIRE M. 2015. - Intraspecific variation and taphonomy of a new erymid lobster (Crustacea: Decapoda) from the Middle Jurassic of Belmont (Beaujolais, France). Geobios 48: 371 - 384. https: // doi. org / 10.1016 / j. geobios. 2015.07.006" type="journal article" year="2015">
Hyžný
<emphasis box="[1405,1456,1138,1162]" italics="true" pageId="17" pageNumber="40">et al.</emphasis>
2015: 375
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="40" type="materials_examined">
<paragraph blockId="17.[813,1439,1212,1331]" pageId="17" pageNumber="40">
<materialsCitation ID-GBIF-Occurrence="3754821308" collectionCode="SMNS" country="Germany" location="Nusplingen" pageId="17" pageNumber="40" specimenCode="SMNS 24227" specimenCount="1" stateProvince="Baden-Wurttemberg" typeStatus="holotype">
TYPE MATERIAL. —
<typeStatus box="[1006,1099,1212,1236]" pageId="17" pageNumber="40">Holotype</typeStatus>
<specimenCode box="[1105,1246,1212,1236]" pageId="17" pageNumber="40">SMNS 24227</specimenCode>
. TYPE LOCALITY. —
<location LSID="urn:lsid:plazi:treatment:297587890769FFF0106301BFFD0DAA51:A40360440769FFF210D506E6FBC7AA21" box="[1003,1119,1260,1284]" country="Germany" name="Nusplingen" pageId="17" pageNumber="40" stateProvince="Baden-Wurttemberg">Nusplingen</location>
,
<collectingRegion box="[1130,1332,1260,1284]" country="Germany" name="Baden-Wuerttemberg" pageId="17" pageNumber="40">Baden-Württemberg</collectingRegion>
,
<collectingCountry box="[1342,1435,1260,1284]" name="Germany" pageId="17" pageNumber="40">Germany</collectingCountry>
. TYPE AGE. —
<geologicalTimeScale box="[950,1096,1307,1331]" pageId="17" pageNumber="40">Kimmeridgian</geologicalTimeScale>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="19" lastPageNumber="42" pageId="17" pageNumber="40" type="description">
<paragraph blockId="17.[811,1457,1364,1709]" box="[813,957,1364,1390]" pageId="17" pageNumber="40">DESCRIPTION</paragraph>
<paragraph blockId="17.[811,1457,1364,1709]" box="[813,909,1396,1422]" pageId="17" pageNumber="40">
<emphasis box="[813,909,1396,1422]" italics="true" pageId="17" pageNumber="40">Carapace</emphasis>
</paragraph>
<paragraph blockId="17.[811,1457,1364,1709]" pageId="17" pageNumber="40">Short, spiny rostrum; fusiform intercalated plate; deep, wide cervical groove, joined to dorsal margin and to antennal groove; deep antennal groove; short, sub-horizontal gastroorbital groove; postcervical groove slightly curved forward, not joined to dorsal margin, joined to branchiocardiac groove at carapace mid-height, with a short ventral extension; branchiocardiac groove dorsally deep, narrowing in its ventral part, not joined to dorsal margin, joined to hepatic groove; hepatic groove concavo-convex.</paragraph>
<paragraph blockId="17.[813,1003,1746,1772]" box="[813,1003,1746,1772]" pageId="17" pageNumber="40">
<heading box="[813,1003,1746,1772]" fontSize="11" level="2" pageId="17" pageNumber="40" reason="8">
<emphasis box="[813,1003,1746,1772]" italics="true" pageId="17" pageNumber="40">Pleon and uropods</emphasis>
</heading>
</paragraph>
<paragraph blockId="17.[812,1457,1778,1899]" pageId="17" pageNumber="40">Somites with subtriangular pleurites, with a bulge on their basis; wide rounded uropods; uropodal endopods with a longitudinal carina; uropodal exopods with a small spine on external margin.</paragraph>
<paragraph blockId="17.[813,1022,1937,1963]" box="[813,1022,1937,1963]" pageId="17" pageNumber="40">
<heading box="[813,1022,1937,1963]" fontSize="11" level="2" pageId="17" pageNumber="40" reason="8">
<emphasis box="[813,1022,1937,1963]" italics="true" pageId="17" pageNumber="40">Thoracic appendages</emphasis>
</heading>
</paragraph>
<paragraph blockId="17.[811,1455,1969,2027]" pageId="17" pageNumber="40">Chelate P1; P1 propodus subrectangular, narrow and strongly elongated; P1 fingers shorter than propodus, slender, almost</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4486491" ID-Zenodo-Dep="4486491" httpUri="https://zenodo.org/record/4486491/files/figure.png" pageId="18" pageNumber="41" startId="18.[132,143,1894,1911]" targetBox="[132,1455,215,1880]" targetPageId="18">
<paragraph blockId="18.[132,1457,1894,2025]" pageId="18" pageNumber="41">
FIG. 7. —
<taxonomicName authority="Oppel, 1861" authorityName="Oppel" authorityYear="1861" box="[212,426,1894,1911]" class="Malacostraca" family="Erymidae" genus="Eryma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="41" phylum="Arthropoda" rank="species" species="major">
<emphasis box="[212,316,1894,1911]" italics="true" pageId="18" pageNumber="41">Eryma major</emphasis>
<bibRefCitation author="OPPEL A." box="[321,426,1894,1911]" pageId="18" pageNumber="41" pagination="355 - 361" refId="ref34602" refString="OPPEL A. 1861. - Die Arten der Gattungen Eryma, Pseudastacus, Magila und Etallonia. Jahreshefte des Vereins fur vaterlandische Naturkunde in Wurttemberg 17: 355 - 361. https: // www. biodiversitylibrary. org / page / 8899023" type="journal article" year="1861">Oppel, 1861</bibRefCitation>
</taxonomicName>
,
<taxonomicName authority="(Munster, 1839)" baseAuthorityName="Munster" baseAuthorityYear="1839" box="[434,703,1894,1911]" class="Malacostraca" family="Erymidae" genus="Eryma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="41" phylum="Arthropoda" rank="species" species="veltheimii">
<emphasis box="[434,569,1894,1911]" italics="true" pageId="18" pageNumber="41">Eryma veltheimii</emphasis>
(
<bibRefCitation author="MUNSTER G. &amp; GRAF ZU" box="[578,700,1894,1911]" pageId="18" pageNumber="41" pagination="1 - 88" refId="ref34482" refString="MUNSTER G. &amp; GRAF ZU 1839. - Decapoda Macrura. Abbildung und Beschreibung der Fossilen langschwanzigen Krebse in den Kalkschiefern von Bayern. Beitrage zur Petrefaktenkunde 2: 1 - 88." type="journal article" year="1839">Münster, 1839</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName authority="Schweigert, Dietl &amp; Roper, 2000" authorityName="Schweigert, Dietl &amp; Roper" authorityYear="2000" box="[746,1153,1894,1911]" class="Malacostraca" family="Erymidae" genus="Eryma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="41" phylum="Arthropoda" rank="species" species="westphali">
<emphasis box="[746,881,1894,1911]" italics="true" pageId="18" pageNumber="41">Eryma westphali</emphasis>
<bibRefCitation author="SCHWEIGERT G. &amp; DIETL G. &amp; ROPER M." box="[886,1153,1894,1911]" pageId="18" pageNumber="41" pagination="1 - 25" refId="ref35446" refString="SCHWEIGERT G., DIETL G. &amp; ROPER M. 2000. - Die Panzerkrebse des Familie Erymidae van Straelen (Crustacea, Decapoda) aus dem Nusplinger Plattenkalk (Ober-Kimmeridgium, Schwabische Alb) im Vergleich mit frankischen Vorkommen. Stuttgarter Beitrage zur Naturkunde B (285): 1 - 25. https: // www. biodiversitylibrary. org / page / 59557644" type="journal article" year="2000">Schweigert, Dietl &amp; Röper, 2000</bibRefCitation>
</taxonomicName>
from the Kimmeridgian of Germany:
<emphasis bold="true" box="[132,145,1917,1934]" pageId="18" pageNumber="41">A</emphasis>
, holotype SMNS 3682 of
<taxonomicName authorityName="Oppel" authorityYear="1861" box="[354,422,1917,1934]" class="Malacostraca" family="Erymidae" genus="Eryma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="41" phylum="Arthropoda" rank="species" species="major">
<emphasis box="[354,422,1917,1934]" italics="true" pageId="18" pageNumber="41">E. major</emphasis>
</taxonomicName>
from Egesheimer;
<emphasis bold="true" box="[575,588,1917,1934]" pageId="18" pageNumber="41">B</emphasis>
, specimen SMNS 67655 of
<taxonomicName authorityName="Oppel" authorityYear="1861" box="[816,884,1917,1934]" class="Malacostraca" family="Erymidae" genus="Eryma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="41" phylum="Arthropoda" rank="species" species="major">
<emphasis box="[816,884,1917,1934]" italics="true" pageId="18" pageNumber="41">E. major</emphasis>
</taxonomicName>
from Gerhausen;
<emphasis bold="true" box="[1030,1044,1917,1934]" pageId="18" pageNumber="41">C</emphasis>
, specimen SMNS 64371 of
<taxonomicName authorityName="Oppel" authorityYear="1861" box="[1271,1339,1917,1934]" class="Malacostraca" family="Erymidae" genus="Eryma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="41" phylum="Arthropoda" rank="species" species="major">
<emphasis box="[1271,1339,1917,1934]" italics="true" pageId="18" pageNumber="41">E. major</emphasis>
</taxonomicName>
from Nusplingen;
<emphasis bold="true" box="[174,206,1940,1957]" pageId="18" pageNumber="41">D -F</emphasis>
, holotype BSPG AS VII 186 of
<taxonomicName baseAuthorityName="Munster" baseAuthorityYear="1839" box="[467,567,1940,1957]" class="Malacostraca" family="Erymidae" genus="Eryma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="41" phylum="Arthropoda" rank="species" species="veltheimii">
<emphasis box="[467,567,1940,1957]" italics="true" pageId="18" pageNumber="41">E. veltheimii</emphasis>
</taxonomicName>
from Kehlheim: part (
<emphasis bold="true" box="[748,762,1940,1957]" pageId="18" pageNumber="41">D</emphasis>
), counter-part (
<emphasis bold="true" box="[892,904,1940,1957]" pageId="18" pageNumber="41">E</emphasis>
), details of telson and uropods (
<emphasis bold="true" box="[1175,1186,1940,1957]" pageId="18" pageNumber="41">F</emphasis>
);
<emphasis bold="true" box="[1201,1233,1940,1957]" pageId="18" pageNumber="41">G -J</emphasis>
, holotype SMNS 24227 of
<taxonomicName authorityName="Schweigert, Dietl &amp; Roper" authorityYear="2000" box="[132,232,1962,1979]" class="Malacostraca" family="Erymidae" genus="Eryma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="41" phylum="Arthropoda" rank="species" species="westphali">
<emphasis box="[132,232,1962,1979]" italics="true" pageId="18" pageNumber="41">E. westphali</emphasis>
</taxonomicName>
from Nusplingen: general view (
<emphasis bold="true" box="[502,516,1962,1979]" pageId="18" pageNumber="41">G</emphasis>
), schema of the carapace (
<emphasis bold="true" box="[744,758,1962,1979]" pageId="18" pageNumber="41">H</emphasis>
), schema of left chela (
<emphasis bold="true" box="[952,958,1962,1979]" pageId="18" pageNumber="41">I</emphasis>
), schema of right chela (
<emphasis bold="true" box="[1163,1173,1962,1979]" pageId="18" pageNumber="41">J</emphasis>
);
<emphasis bold="true" box="[1189,1203,1962,1979]" pageId="18" pageNumber="41">C</emphasis>
, specimen SMNS 63733 from Nusplingen. Abbreviations:
<emphasis bold="true" box="[361,372,1985,2002]" pageId="18" pageNumber="41">a</emphasis>
, branchiocardiac;
<emphasis bold="true" box="[526,547,1985,2002]" pageId="18" pageNumber="41">as</emphasis>
, antennal spine;
<emphasis bold="true" box="[687,698,1985,2002]" pageId="18" pageNumber="41">b</emphasis>
, antennal groove;
<emphasis bold="true" box="[852,871,1985,2005]" pageId="18" pageNumber="41">
b
<subScript attach="right" box="[864,871,1993,2005]" fontSize="5" pageId="18" pageNumber="41">1</subScript>
</emphasis>
, hepatic groove;
<emphasis bold="true" box="[1015,1026,1985,2002]" pageId="18" pageNumber="41">c</emphasis>
, postcervical;
<emphasis bold="true" box="[1146,1157,1985,2002]" pageId="18" pageNumber="41">d</emphasis>
, gastro-orbital groove;
<emphasis bold="true" box="[1353,1382,1985,2005]" pageId="18" pageNumber="41">
e
<subScript attach="both" box="[1363,1370,1993,2005]" fontSize="5" pageId="18" pageNumber="41">1</subScript>
e
</emphasis>
, cervical groove;
<emphasis bold="true" box="[200,216,2008,2025]" pageId="18" pageNumber="41">ip</emphasis>
, intercalated plate. Black arrows show the spines. Photographs and line drawings: J. Devillez. Scale bars: 1 cm.
</paragraph>
</caption>
<paragraph blockId="19.[130,778,215,306]" pageId="19" pageNumber="42">straight or slightly curved inward, with a terminal hook; occlusal margin without teeth; P1 carpus short, subtriangular; thin P2-P5; P2-P3 chelate.</paragraph>
<paragraph blockId="19.[132,294,344,370]" box="[132,294,344,370]" pageId="19" pageNumber="42">
<heading box="[132,294,344,370]" fontSize="11" level="2" pageId="19" pageNumber="42" reason="8">
<emphasis box="[132,294,344,370]" italics="true" pageId="19" pageNumber="42">Ornamentation</emphasis>
</heading>
</paragraph>
<paragraph blockId="19.[130,777,377,563]" pageId="19" pageNumber="42">Carapace with a dense, heterogeneous ornamentation; gastric and cardiac regions covered by tubercles; antennal, hepatic, and branchial regions covered by small rounded depressions; presence of an orbital spine; tergites and pleurites densely covered by small rounded depressions; presence of a strong antennal spine; smooth P1-P5.</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="42" type="discussion">
<paragraph blockId="19.[130,777,602,1396]" box="[132,262,602,628]" pageId="19" pageNumber="42">DISCUSSION</paragraph>
<paragraph blockId="19.[130,777,602,1396]" pageId="19" pageNumber="42">
This species is only known by three specimens. It is assigned to
<taxonomicName authorityName="Meyer" authorityYear="1840" box="[160,228,667,692]" class="Malacostraca" family="Erymidae" genus="Eryma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis box="[160,228,667,692]" italics="true" pageId="19" pageNumber="42">Eryma</emphasis>
</taxonomicName>
because of its typical carapace groove pattern (short gastro-orbital groove, postcervical and branchiocardiac grooves joined at carapace mid-height, hepatic groove concavo-convex) and the shape of the P1 chelae (elongated propodus, subrectangular, with slender fingers progressively narrowing to their distal extremity).
</paragraph>
<paragraph blockId="19.[130,777,602,1396]" pageId="19" pageNumber="42">
Among the erymid lobsters,
<taxonomicName authorityName="Schweigert, Dietl &amp; Roper" authorityYear="2000" box="[475,651,858,884]" class="Malacostraca" family="Erymidae" genus="Eryma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="42" phylum="Arthropoda" rank="species" species="westphali">
<emphasis box="[475,651,858,884]" italics="true" pageId="19" pageNumber="42">Eryma westphali</emphasis>
</taxonomicName>
is the only species with heterochelous P1 chelae on a same specimen. Left propodus is more elongated and narrower than the right one, and the left fingers are straight and short while the right ones are curved and slightly longer (
<figureCitation box="[522,620,986,1013]" captionStart="FIG" captionStartId="18.[132,143,1894,1911]" captionTargetBox="[132,1455,215,1880]" captionTargetId="figure-28@18.[698,1267,769,1326]" captionTargetPageId="18" captionText="FIG. 7. — Eryma major Oppel, 1861, Eryma veltheimii (Münster, 1839) and Eryma westphali Schweigert, Dietl &amp; Röper, 2000 from the Kimmeridgian of Germany: A, holotype SMNS 3682 of E. major from Egesheimer;B, specimen SMNS 67655 of E. major from Gerhausen; C, specimen SMNS 64371 of E. major from Nusplingen; D-F, holotype BSPG AS VII 186 of E. veltheimii from Kehlheim: part (D), counter-part (E), details of telson and uropods (F); G-J, holotype SMNS 24227 of E. westphali from Nusplingen: general view (G), schema of the carapace (H), schema of left chela (I), schema of right chela (J); C, specimen SMNS 63733 from Nusplingen. Abbreviations: a, branchiocardiac; as, antennal spine; b, antennal groove; b1, hepatic groove; c, postcervical; d, gastro-orbital groove; e1e, cervical groove; ip, intercalated plate. Black arrows show the spines. Photographs and line drawings: J. Devillez. Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4486491" httpUri="https://zenodo.org/record/4486491/files/figure.png" pageId="19" pageNumber="42">Fig. 7I, J</figureCitation>
). The strongly elongated and narrow P1 propodus with relatively short fingers of
<taxonomicName authorityName="Schweigert, Dietl &amp; Roper" authorityYear="2000" box="[245,373,1050,1076]" class="Malacostraca" family="Erymidae" genus="Eryma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="42" phylum="Arthropoda" rank="species" species="westphali">
<emphasis box="[245,373,1050,1076]" italics="true" pageId="19" pageNumber="42">E. westphali</emphasis>
</taxonomicName>
is characteristic of the species. Only
<taxonomicName authority="(Munster, 1839)" baseAuthorityName="Munster" baseAuthorityYear="1839" box="[134,498,1082,1108]" class="Malacostraca" family="Erymidae" genus="Eryma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="42" phylum="Arthropoda" rank="species" species="veltheimii">
<emphasis box="[134,311,1082,1108]" italics="true" pageId="19" pageNumber="42">Eryma veltheimii</emphasis>
(
<bibRefCitation author="MUNSTER G. &amp; GRAF ZU" box="[327,490,1082,1108]" pageId="19" pageNumber="42" pagination="1 - 88" refId="ref34482" refString="MUNSTER G. &amp; GRAF ZU 1839. - Decapoda Macrura. Abbildung und Beschreibung der Fossilen langschwanzigen Krebse in den Kalkschiefern von Bayern. Beitrage zur Petrefaktenkunde 2: 1 - 88." type="journal article" year="1839">Münster, 1839</bibRefCitation>
)
</taxonomicName>
have P1 with a very long propodus, but it is clearly wider.
</paragraph>
<paragraph blockId="19.[130,777,602,1396]" pageId="19" pageNumber="42">
In
<taxonomicName authorityName="Schweigert, Dietl &amp; Roper" authorityYear="2000" box="[184,309,1146,1172]" class="Malacostraca" family="Erymidae" genus="Eryma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="42" phylum="Arthropoda" rank="species" species="westphali">
<emphasis box="[184,309,1146,1172]" italics="true" pageId="19" pageNumber="42">E. westphali</emphasis>
</taxonomicName>
the branchial region is dorsally shorter than in
<taxonomicName box="[164,275,1179,1204]" class="Malacostraca" family="Erymidae" genus="Eryma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="42" phylum="Arthropoda" rank="species" species="georgeii">
<emphasis box="[164,275,1179,1204]" italics="true" pageId="19" pageNumber="42">E. georgeii</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Feldmann &amp; Titus" authorityYear="2006" box="[291,427,1179,1204]" class="Malacostraca" family="Erymidae" genus="Eryma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="42" phylum="Arthropoda" rank="species" species="jungostrix">
<emphasis box="[291,427,1179,1204]" italics="true" pageId="19" pageNumber="42">E. jungostrix</emphasis>
</taxonomicName>
,
<taxonomicName baseAuthorityName="Etallon" baseAuthorityYear="1861" box="[444,530,1178,1204]" class="Malacostraca" family="Erymidae" genus="Eryma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="42" phylum="Arthropoda" rank="species" species="lerasi">
<emphasis box="[444,530,1178,1204]" italics="true" pageId="19" pageNumber="42">E. lerasi</emphasis>
</taxonomicName>
,
<taxonomicName baseAuthorityName="Schlotheim" baseAuthorityYear="1822" box="[546,718,1178,1204]" class="Malacostraca" family="Erymidae" genus="Eryma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="42" phylum="Arthropoda" rank="species" species="modestiforme">
<emphasis box="[546,718,1178,1204]" italics="true" pageId="19" pageNumber="42">E. modestiforme</emphasis>
</taxonomicName>
, and
<taxonomicName authorityName="Erymoidea" baseAuthorityName="Meyer" baseAuthorityYear="1835" box="[134,267,1211,1236]" class="Malacostraca" family="Erymidae" genus="Eryma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="42" phylum="Arthropoda" rank="species" species="ventrosum">
<emphasis box="[134,267,1211,1236]" italics="true" pageId="19" pageNumber="42">E. ventrosum</emphasis>
</taxonomicName>
. Contrary to
<taxonomicName baseAuthorityName="Etallon" baseAuthorityYear="1861" box="[414,496,1210,1236]" class="Malacostraca" family="Erymidae" genus="Eryma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="42" phylum="Arthropoda" rank="species" species="lerasi">
<emphasis box="[414,496,1210,1236]" italics="true" pageId="19" pageNumber="42">E. lerasi</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Trautschold" authorityYear="1866" box="[549,767,1210,1236]" class="Malacostraca" family="Erymidae" genus="Eryma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="42" phylum="Arthropoda" rank="species" species="quadriverrucatum">
<emphasis box="[549,767,1210,1236]" italics="true" pageId="19" pageNumber="42">E. quadriverrucatum</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Schweigert, Dietl &amp; Roper" authorityYear="2000" box="[132,260,1242,1268]" class="Malacostraca" family="Erymidae" genus="Eryma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="42" phylum="Arthropoda" rank="species" species="westphali">
<emphasis box="[132,260,1242,1268]" italics="true" pageId="19" pageNumber="42">E. westphali</emphasis>
</taxonomicName>
exhibits a ventral extension of the postcervical groove. Moreover, the dichotomy of the ornamentation of the carapace between the gastric and cardiac regions, covered by tubercles, and the remaining regions of
<taxonomicName authorityName="Schweigert, Dietl &amp; Roper" authorityYear="2000" box="[554,679,1338,1364]" class="Malacostraca" family="Erymidae" genus="Eryma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="42" phylum="Arthropoda" rank="species" species="westphali">
<emphasis box="[554,679,1338,1364]" italics="true" pageId="19" pageNumber="42">E. westphali</emphasis>
</taxonomicName>
, covered by small depressions, is unique among the genus.
</paragraph>
</subSubSection>
</treatment>
</document>