294 lines
34 KiB
XML
294 lines
34 KiB
XML
<document id="333FC6AD454DB0148033B8783155572E" ID-DOI="10.5281/zenodo.4509152" ID-ISSN="2345-7600" ID-Zenodo-Dep="4509152" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612548437016" checkinUser="carolina" docAuthor="ski, P. Jałoszy" docDate="2009" docId="03FEC075FF85C163FC79FC69FEBAFE11" docLanguage="en" docName="RafflesBZool.57.2.297-303.pdf" docOrigin="Raffles Bulletin of Zoology 57 (2)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Horaeomorphus minor ski 2009, new species" docType="treatment" docVersion="4" lastPageNumber="300" masterDocId="FFC7B80DFF84C167FFE6FFB7FFCAFFA4" masterDocTitle="Four New Species Of Horaeomorphus Schaufuss From The Oriental Region (Coleoptera: Staphylinidae, Scydmaeninae)" masterLastPageNumber="303" masterPageNumber="297" pageNumber="298" updateTime="1698867913614" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
|
||
<mods:mods id="89BCE9D683D6731847AA23E5F01FC64D" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="82FBFE4C547F2C8C4C432EE4D9692C05">
|
||
<mods:title id="53EA23E1CBE1FDBB05C24ACE4CC8984D">Four New Species Of Horaeomorphus Schaufuss From The Oriental Region (Coleoptera: Staphylinidae, Scydmaeninae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="D273DB7AC49AEC26EA5F50F8CE65A8C0" type="personal">
|
||
<mods:role id="263F29FBFE9D9823F51DF7EE5FFF2CBE">
|
||
<mods:roleTerm id="DC78F6DFA38B160391FA4A720D952663">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="9509DB3CF2982166C5179F26BFE8070B">ski, P. Jałoszy</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="0739AC2ECFEC828A1D59C9F4B45E6A4B">text</mods:typeOfResource>
|
||
<mods:relatedItem id="4A8DAA01D491E59BE61382A9AEEE3E98" type="host">
|
||
<mods:titleInfo id="285F63804543946D16BC465418203F0F">
|
||
<mods:title id="B9960B3F07C97B1EB2C24B5BBE51DE05">Raffles Bulletin of Zoology</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="BB5632DC47A2A077A6FE7A53DEB24464">
|
||
<mods:date id="D4B89E77FC2BC326B559C01A6FCAD5F2">2009</mods:date>
|
||
<mods:detail id="7EC728E1EB941D16AAA777B46467BF05" type="pubDate">
|
||
<mods:number id="5F4E7F51173E668AEDC21CE9C33E6C4A">2009-08-31</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="AE51D18DB75E12B17275F2B9FCB4719C" type="volume">
|
||
<mods:number id="AD1F71DA0CF967E03DBACB01DD895110">57</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="4D69C62E5A9A85BAF457A707DA61C7B6" type="issue">
|
||
<mods:number id="E5C3DA65455DA3CE5FE97ABE355DC1B2">2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="387A597BB91ED39071DCF6E8AC50DBB2" unit="page">
|
||
<mods:start id="20B4B2132BF258AB0EE9E07DF33263DC">297</mods:start>
|
||
<mods:end id="F12EB0C5C63D3B16D91425F53BDD20BC">303</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="EF1BE8FB86060E5E41459554714CC0D2">journal article</mods:classification>
|
||
<mods:identifier id="5AF5DB59ED580A1B64EED02499C8B9C5" type="DOI">10.5281/zenodo.4509152</mods:identifier>
|
||
<mods:identifier id="2847E6FB2CD377829F6ADCB108D68A51" type="ISSN">2345-7600</mods:identifier>
|
||
<mods:identifier id="DCB8B8FD04C7287D41FA7E38A6BEC686" type="Zenodo-Dep">4509152</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03FEC075FF85C163FC79FC69FEBAFE11" LSID="urn:lsid:plazi:treatment:03FEC075FF85C163FC79FC69FEBAFE11" httpUri="http://treatment.plazi.org/id/03FEC075FF85C163FC79FC69FEBAFE11" lastPageId="4" lastPageNumber="300" pageId="1" pageNumber="298">
|
||
<subSubSection id="C34D22E8FF85C166FC79FC69FAFEFC52" box="[927,1332,990,1014]" pageId="1" pageNumber="298" type="nomenclature">
|
||
<paragraph id="8BE87163FF85C166FC79FC69FAFEFC52" blockId="1.[927,1332,990,1045]" box="[927,1332,990,1014]" pageId="1" pageNumber="298">
|
||
<heading id="D0A0C60FFF85C166FC79FC69FAFEFC52" bold="true" box="[927,1332,990,1014]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="298" reason="2">
|
||
<emphasis id="B923AD71FF85C166FC79FC69FAFEFC52" bold="true" box="[927,1332,990,1014]" pageId="1" pageNumber="298">
|
||
<taxonomicName id="4C570AE0FF85C166FC79FC69FB6AFC52" ID-CoL="3MNKS" authority="ski, 2009" authorityName="ski" authorityYear="2009" box="[927,1184,990,1014]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="298" phylum="Arthropoda" rank="species" species="minor" status="sp. nov.">
|
||
<emphasis id="B923AD71FF85C166FC79FC69FB6AFC52" bold="true" box="[927,1184,990,1014]" italics="true" pageId="1" pageNumber="298">Horaeomorphus minor</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A210100AFF85C166FB49FC69FAFEFC52" box="[1199,1332,990,1014]" pageId="1" pageNumber="298" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34D22E8FF85C166FBF0FC49FB76FBB1" box="[1046,1212,1022,1045]" pageId="1" pageNumber="298" type="description">
|
||
<paragraph id="8BE87163FF85C166FBF0FC49FB76FBB1" blockId="1.[927,1332,990,1045]" box="[1046,1212,1022,1045]" pageId="1" pageNumber="298">
|
||
(
|
||
<figureCitation id="136C6DE6FF85C166FBF8FC49FBB2FBB1" box="[1054,1144,1022,1045]" captionStart="Fig" captionStartId="2.[134,169,1165,1186]" captionTargetBox="[134,1466,358,1140]" captionTargetId="figure-89@2.[130,1472,338,1154]" captionTargetPageId="2" captionText="Fig. 1. Dorsal habitus of holotype males: a, Horaeomorphus imitator, new species; b, Horaeomorphus minor, new species; c, Horaeomorphus fakfakensis, new species; d, Horaeomorphus samosirensis, new species (d). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4509154" httpUri="https://zenodo.org/record/4509154/files/figure.png" pageId="1" pageNumber="298">Figs. 1b</figureCitation>
|
||
,
|
||
<figureCitation id="136C6DE6FF85C166FB60FC49FB7EFBB1" box="[1158,1204,1022,1045]" captionStart="Fig" captionStartId="2.[134,169,2060,2081]" captionTargetBox="[140,1473,1299,2029]" captionTargetId="figure-84@2.[122,1500,1284,2048]" captionTargetPageId="2" captionText="Fig. 2. Aedeagus in ventral (a, c) and lateral (b, d) views: a, b, Horaeomorphus imitator, new species; c, d, Horaeomorphus minor, new species. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4509156" httpUri="https://zenodo.org/record/4509156/files/figure.png" pageId="1" pageNumber="298">2c,d</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34D22E8FF85C166FCCCFB8AFC19FAF1" pageId="1" pageNumber="298" type="materials_examined">
|
||
<paragraph id="8BE87163FF85C166FCCCFB8AFC19FAF1" blockId="1.[810,1449,1085,1365]" pageId="1" pageNumber="298">
|
||
Material examined. –
|
||
<materialsCitation id="3B3F7B3EFF85C166FBE7FB8AFC19FB61" collectionCode="SMNS" elevation="1700" pageId="1" pageNumber="298" specimenCount="2" typeStatus="holotype">
|
||
<emphasis id="B923AD71FF85C166FBE7FB8AFBAEFBF6" bold="true" box="[1025,1124,1085,1106]" pageId="1" pageNumber="298">
|
||
<typeStatus id="54ECCFC1FF85C166FBE7FB8AFBAAFBF6" box="[1025,1120,1085,1106]" pageId="1" pageNumber="298" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Male, two labels: “MINDANAO,
|
||
<quantity id="4CAFDC86FF85C166FCCCFBEDFCABFBCB" box="[810,865,1114,1135]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="298" unit="km" value="30.0">30km</quantity>
|
||
NW of \ MARAMAG, 13.-17. MAY \ BAGONGSILANG,
|
||
<quantity id="4CAFDC86FF85C166FCCCFBC1FCBBFB2F" box="[810,881,1142,1163]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="298" unit="m" value="1700.0">
|
||
<elevation id="007A9650FF85C166FCCCFBC1FCBBFB2F" box="[810,881,1142,1163]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="298" unit="m" value="1700.0">1700 m</elevation>
|
||
</quantity>
|
||
\ Bolm lgt., 1996” [yellow, printed], “
|
||
<taxonomicName id="4C570AE0FF85C166FB31FBC0FA62FB28" box="[1239,1448,1143,1164]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="298" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B923AD71FF85C166FB31FBC0FA62FB28" box="[1239,1448,1143,1164]" italics="true" pageId="1" pageNumber="298">HORAEOMORPHUS</emphasis>
|
||
</taxonomicName>
|
||
\
|
||
<taxonomicName id="4C570AE0FF85C166FCDFFB24FCB8FB0C" authorityName="ski" authorityYear="2009" box="[825,882,1171,1192]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="298" phylum="Arthropoda" rank="species" species="minor">
|
||
<emphasis id="B923AD71FF85C166FCDFFB24FCB8FB0C" box="[825,882,1171,1192]" italics="true" pageId="1" pageNumber="298">minor</emphasis>
|
||
</taxonomicName>
|
||
m. \
|
||
<typeStatus id="54ECCFC1FF85C166FC4AFB24FBF1FB0C" box="[940,1083,1171,1192]" pageId="1" pageNumber="298" type="holotype">HOLOTYPUS</typeStatus>
|
||
\ det. P. JAŁOSZY SKI, ‘08” [red, printed] (
|
||
<collectionCode id="ED46E9A6FF85C166FC60FB07FC06FB61" box="[902,972,1200,1221]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="1" pageNumber="298">SMNS</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B3F7B3EFF85C166FC39FB07FCB5FB46" collectionCode="SMNS" elevation="1700" pageId="1" pageNumber="298" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
|
||
<emphasis id="B923AD71FF85C166FC39FB07FB86FB61" bold="true" box="[991,1100,1200,1221]" pageId="1" pageNumber="298">
|
||
<typeStatus id="54ECCFC1FF85C166FC39FB07FB8DFB61" box="[991,1095,1200,1221]" pageId="1" pageNumber="298" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<specimenCount id="9D51BAEAFF85C166FBB3FB07FB51FB61" box="[1109,1179,1200,1221]" pageId="1" pageNumber="298" type="male">1 male</specimenCount>
|
||
, same data as for holotype (
|
||
<collectionCode id="ED46E9A6FF85C166FCD7FB7AFCB2FB46" box="[817,888,1229,1250]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="1" pageNumber="298">SMNS</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B3F7B3EFF85C166FC6BFB7AFC04FAF1" collectionCode="SMNS" elevation="1700" pageId="1" pageNumber="298" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
|
||
<specimenCount id="9D51BAEAFF85C166FC6BFB7AFC1EFB46" box="[909,980,1229,1250]" pageId="1" pageNumber="298" type="male">1 male</specimenCount>
|
||
, “MINDANAO, 1-3. May \ MISAMIS OCC.,
|
||
<quantity id="4CAFDC86FF85C166FCCCFB5DFCB8FB5B" box="[810,882,1258,1279]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="298" unit="m" value="1700.0">
|
||
<elevation id="007A9650FF85C166FCCCFB5DFCB8FB5B" box="[810,882,1258,1279]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="298" unit="m" value="1700.0">1700 m</elevation>
|
||
</quantity>
|
||
\ DON VICTORIANO \ Bolm lgt., 1996” [yellow, printed] (
|
||
<collectionCode id="ED46E9A6FF85C166FCD7FAB1FCBDFABF" box="[817,887,1286,1307]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="1" pageNumber="298">SMNS</collectionCode>
|
||
);
|
||
<typeStatus id="54ECCFC1FF85C166FC6EFAB1FC2EFABF" box="[904,996,1286,1307]" pageId="1" pageNumber="298" type="paratype">paratypes</typeStatus>
|
||
additionally provided with identification labels analogous to that on the
|
||
<typeStatus id="54ECCFC1FF85C166FBD3FA94FB5BFA9C" box="[1077,1169,1315,1336]" pageId="1" pageNumber="298" type="holotype">holotype</typeStatus>
|
||
, but yellow and annotated “
|
||
<typeStatus id="54ECCFC1FF85C166FCD5FAF7FC0FFAF1" box="[819,965,1344,1365]" pageId="1" pageNumber="298" type="paratype">PARATYPUS</typeStatus>
|
||
”
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34D22E8FF85C164FCCCFAC9FBD9FC31" lastPageId="3" lastPageNumber="300" pageId="1" pageNumber="298" type="diagnosis">
|
||
<paragraph id="8BE87163FF85C166FCCCFAC9FB61F971" blockId="1.[810,1449,1406,1749]" pageId="1" pageNumber="298">
|
||
<emphasis id="B923AD71FF85C166FCCCFAC9FC0CFA32" bold="true" box="[810,966,1406,1430]" italics="true" pageId="1" pageNumber="298">Diagnosis. –</emphasis>
|
||
This is one of the smallest species of
|
||
<taxonomicName id="4C570AE0FF85C166FCCCFA28FC29FA12" box="[810,995,1439,1462]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="298" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B923AD71FF85C166FCCCFA28FC29FA12" box="[810,995,1439,1462]" italics="true" pageId="1" pageNumber="298">Horaeomorphus</emphasis>
|
||
</taxonomicName>
|
||
, remarkable by not reaching
|
||
<quantity id="4CAFDC86FF85C166FAA2FA29FA4DFA11" box="[1348,1415,1438,1461]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="298" unit="mm" value="2.0">2 mm</quantity>
|
||
in length; it differs from all congeners also in the following combination of characters: lateral margins of pronotum slightly, but distinctly raised in posterior part to form rounded lateral longitudinal carinae; base of pronotum without groove, with two large lateral pits and very small median pit; elytra broadest near anterior third and very strongly narrowing from broadest place toward apices; legs in males non-modified; aedeagus with strongly asymmetrical, massive internal sac.
|
||
</paragraph>
|
||
<paragraph id="8BE87163FF85C166FCCCF949FB65F8D1" blockId="1.[810,1449,1790,1909]" pageId="1" pageNumber="298">
|
||
<emphasis id="B923AD71FF85C166FCCCF949FC4AF8B2" bold="true" box="[810,896,1790,1814]" italics="true" pageId="1" pageNumber="298">Male. –</emphasis>
|
||
Body (
|
||
<figureCitation id="136C6DE6FF85C166FC33F949FBEEF8B1" box="[981,1060,1790,1813]" captionStart="Fig" captionStartId="2.[134,169,1165,1186]" captionTargetBox="[134,1466,358,1140]" captionTargetId="figure-89@2.[130,1472,338,1154]" captionTargetPageId="2" captionText="Fig. 1. Dorsal habitus of holotype males: a, Horaeomorphus imitator, new species; b, Horaeomorphus minor, new species; c, Horaeomorphus fakfakensis, new species; d, Horaeomorphus samosirensis, new species (d). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4509154" httpUri="https://zenodo.org/record/4509154/files/figure.png" pageId="1" pageNumber="298">Fig. 1b</figureCitation>
|
||
) very small (length
|
||
<quantity id="4CAFDC86FF85C166FAEFF949FA6FF8B1" box="[1289,1445,1790,1813]" metricMagnitude="-3" metricUnit="m" metricValue="1.855" metricValueMax="1.93" metricValueMin="1.78" pageId="1" pageNumber="298" unit="mm" value="1.855" valueMax="1.93" valueMin="1.78">1.78-1.93 mm</quantity>
|
||
, mean
|
||
<quantity id="4CAFDC86FF85C166FC8DF8A9FC05F891" box="[875,975,1822,1845]" metricMagnitude="-3" metricUnit="m" metricValue="1.87" pageId="1" pageNumber="298" unit="mm" value="1.87">1.87 mm</quantity>
|
||
), slender, flattened, moderately dark brown, vestiture slightly lighter than cuticle; setae on pronotum slightly darker than those on elytra.
|
||
</paragraph>
|
||
<paragraph id="8BE87163FF85C164FCCCF829FD8BFDD1" blockId="1.[810,1449,1950,2069]" lastBlockId="3.[111,750,190,629]" lastPageId="3" lastPageNumber="300" pageId="1" pageNumber="298">
|
||
Head broadest at moderately large, moderately convex eyes, length
|
||
<quantity id="4CAFDC86FF85C166FC9CF809FBD3F871" box="[890,1049,1982,2005]" metricMagnitude="-4" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="1" pageNumber="298" unit="mm" value="0.265" valueMax="0.28" valueMin="0.25">0.25-0.28 mm</quantity>
|
||
(mean
|
||
<quantity id="4CAFDC86FF85C166FB92F809FB14F871" box="[1140,1246,1982,2005]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="1" pageNumber="298" unit="mm" value="0.27">0.27 mm</quantity>
|
||
), width
|
||
<quantity id="4CAFDC86FF85C166FAD9F809FC9EF851" metricMagnitude="-4" metricUnit="m" metricValue="3.65" metricValueMax="3.8" metricValueMin="3.5" pageId="1" pageNumber="298" unit="mm" value="0.365" valueMax="0.38" valueMin="0.35">0.35-0.38 mm</quantity>
|
||
(mean
|
||
<quantity id="4CAFDC86FF85C166FC48F869FBD3F851" box="[942,1049,2014,2037]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="1" pageNumber="298" unit="mm" value="0.37">0.37 mm</quantity>
|
||
); tempora as long as eye in dorsal view, regularly, weakly rounded up to posterior 3/4, where they are strongly bent toward occipital constriction; vertex weakly convex, with pair of small but deep pits located near posterior margins of supraantennal tubercles; frons weakly convex; supraantennal tubercles very distinct, strongly raised, well delimited from frons but indistinctly delimited from vertex. Punctures on vertex very fine, barely noticeable under magnification 80×; median part of frons bears dense, moderately large, distinct punctures unevenly distributed and separated by spaces 1–3× puncture diameters; setae moderately long, sparse, suberect to erect. Antennae moderately slender, shorter than half length of body, length 1.75–
|
||
<quantity id="4CAFDC86FF87C164FF13FDA9FE9DFD91" box="[245,343,542,565]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="300" unit="mm" value="0.8">0.80 mm</quantity>
|
||
(mean
|
||
<quantity id="4CAFDC86FF87C164FE4BFDA9FDD9FD91" box="[429,531,542,565]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" pageId="3" pageNumber="300" unit="mm" value="0.78">0.78 mm</quantity>
|
||
); relative lengths of antennomeres (shortest antennomere II as 1): 0.83; 1; 1.07; 0.93; 0.8; 0.77; 0.73; 0.70; 0.97; 0.83; 1.5.
|
||
</paragraph>
|
||
<caption id="DF2821EBFF86C165FF60FB3AFBF1FB1B" ID-DOI="http://doi.org/10.5281/zenodo.4509154" ID-Zenodo-Dep="4509154" httpUri="https://zenodo.org/record/4509154/files/figure.png" pageId="2" pageNumber="299" startId="2.[134,169,1165,1186]" targetBox="[134,1466,358,1140]" targetPageId="2">
|
||
<paragraph id="8BE87163FF86C165FF60FB3AFBF1FB1B" blockId="2.[134,1471,1165,1215]" pageId="2" pageNumber="299">
|
||
Fig. 1. Dorsal habitus of holotype males: a,
|
||
<taxonomicName id="4C570AE0FF86C165FDF9FB3AFCC7FB06" authority="ski, 2009" authorityName="ski" authorityYear="2009" box="[543,781,1165,1186]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="imitator" status="sp. nov.">
|
||
<emphasis id="B923AD71FF86C165FDF9FB3AFCC7FB06" box="[543,781,1165,1186]" italics="true" pageId="2" pageNumber="299">Horaeomorphus imitator</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A210100AFF86C165FCFEFB3AFC46FB06" box="[792,908,1165,1186]" pageId="2" pageNumber="299" rank="species">new species</taxonomicNameLabel>
|
||
; b,
|
||
<taxonomicName id="4C570AE0FF86C165FC4DFB3AFB4CFB06" authority="ski, 2009" authorityName="ski" authorityYear="2009" box="[939,1158,1165,1186]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="minor" status="sp. nov.">
|
||
<emphasis id="B923AD71FF86C165FC4DFB3AFB4CFB06" box="[939,1158,1165,1186]" italics="true" pageId="2" pageNumber="299">Horaeomorphus minor</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A210100AFF86C165FB76FB3AFACFFB06" box="[1168,1285,1165,1186]" pageId="2" pageNumber="299" rank="species">new species</taxonomicNameLabel>
|
||
; c,
|
||
<taxonomicName id="4C570AE0FF86C165FAC4FB3AFF3BFB1B" authority="ski, 2009" authorityName="ski" authorityYear="2009" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="fakfakensis" status="sp. nov.">
|
||
<emphasis id="B923AD71FF86C165FAC4FB3AFF3BFB1B" italics="true" pageId="2" pageNumber="299">Horaeomorphus fakfakensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A210100AFF86C165FF19FB1DFEBDFB1B" box="[255,375,1194,1215]" pageId="2" pageNumber="299" rank="species">new species</taxonomicNameLabel>
|
||
; d,
|
||
<taxonomicName id="4C570AE0FF86C165FE7AFB1DFD76FB1B" authority="ski, 2009" authorityName="ski" authorityYear="2009" box="[412,700,1194,1215]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="samosirensis" status="sp. nov.">
|
||
<emphasis id="B923AD71FF86C165FE7AFB1DFD76FB1B" box="[412,700,1194,1215]" italics="true" pageId="2" pageNumber="299">Horaeomorphus samosirensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A210100AFF86C165FD2CFB1DFCF5FB1B" box="[714,831,1194,1215]" pageId="2" pageNumber="299" rank="species">new species</taxonomicNameLabel>
|
||
(d). Scale bar = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF2821EBFF86C165FF60F7BBFE68F79A" ID-DOI="http://doi.org/10.5281/zenodo.4509156" ID-Zenodo-Dep="4509156" httpUri="https://zenodo.org/record/4509156/files/figure.png" pageId="2" pageNumber="299" startId="2.[134,169,2060,2081]" targetBox="[140,1473,1299,2029]" targetPageId="2">
|
||
<paragraph id="8BE87163FF86C165FF60F7BBFE68F79A" blockId="2.[134,1472,2060,2110]" pageId="2" pageNumber="299">
|
||
Fig. 2. Aedeagus in ventral (a, c) and lateral (b, d) views: a, b,
|
||
<taxonomicName id="4C570AE0FF86C165FD10F7BBFC22F785" authority="ski, 2009" authorityName="ski" authorityYear="2009" box="[758,1000,2060,2081]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="imitator" status="sp. nov.">
|
||
<emphasis id="B923AD71FF86C165FD10F7BBFC22F785" box="[758,1000,2060,2081]" italics="true" pageId="2" pageNumber="299">Horaeomorphus imitator</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A210100AFF86C165FC10F7BBFBA4F785" box="[1014,1134,2060,2081]" pageId="2" pageNumber="299" rank="species">new species</taxonomicNameLabel>
|
||
; c, d,
|
||
<taxonomicName id="4C570AE0FF86C165FB4AF7BBFA40F785" authority="ski, 2009" authorityName="ski" authorityYear="2009" box="[1196,1418,2060,2081]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="299" phylum="Arthropoda" rank="species" species="minor" status="sp. nov.">
|
||
<emphasis id="B923AD71FF86C165FB4AF7BBFA40F785" box="[1196,1418,2060,2081]" italics="true" pageId="2" pageNumber="299">Horaeomorphus minor</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A210100AFF86C165FA7EF7BBFF04F79A" pageId="2" pageNumber="299" rank="species">new species</taxonomicNameLabel>
|
||
. Scale bar = 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE87163FF87C164FF89FD29FEF7FBD1" blockId="3.[111,750,670,1141]" pageId="3" pageNumber="300">
|
||
Pronotum inversely subtrapezoidal, elongate, broadest at anterior third, length
|
||
<quantity id="4CAFDC86FF87C164FEB7FD09FE21FD71" box="[337,491,702,725]" metricMagnitude="-4" metricUnit="m" metricValue="5.15" metricValueMax="5.3" metricValueMin="5.0" pageId="3" pageNumber="300" unit="mm" value="0.515" valueMax="0.53" valueMin="0.5">0.50–0.53 mm</quantity>
|
||
(mean
|
||
<quantity id="4CAFDC86FF87C164FDDAFD09FD54FD71" box="[572,670,702,725]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="3" pageNumber="300" unit="mm" value="0.52">0.52 mm</quantity>
|
||
), width at base
|
||
<quantity id="4CAFDC86FF87C164FF2EFD69FEE5FD51" box="[200,303,734,757]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="3" pageNumber="300" unit="mm" value="0.33">0.33 mm</quantity>
|
||
, maximum width
|
||
<quantity id="4CAFDC86FF87C164FE18FD69FD55FD51" box="[510,671,734,757]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" metricValueMax="4.5" metricValueMin="4.3" pageId="3" pageNumber="300" unit="mm" value="0.44" valueMax="0.45" valueMin="0.43">0.43–0.45 mm</quantity>
|
||
(mean
|
||
<quantity id="4CAFDC86FF87C164FF89FD49FF1BFCB1" box="[111,209,766,789]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="3" pageNumber="300" unit="mm" value="0.43">0.43 mm</quantity>
|
||
); anterior margin rounded; lateral margins rounded near broadest place, then nearly straight up to indistinct subbasal constriction separating short posterior collar; lateral margins in posterior third to half raised, forming low and rounded lateral carinae; hind angles blunt, obtuse; posterior margin weakly arcuate; base of pronotum with pair of large lateral pits and very small, indistinct median pit. Punctures very small, moderately sharply marked, but well visible under magnification 40×, unevenly distributed, those on median part of disc are separated by spaces 1–3× as long as puncture diameters; setation moderately dense and long, suberect.
|
||
</paragraph>
|
||
<paragraph id="8BE87163FF87C164FF89FB29FAFBFDF1" blockId="3.[111,749,1182,1269]" lastBlockId="3.[810,1449,190,597]" pageId="3" pageNumber="300">
|
||
Elytra strongly elongate, drop-shaped, about as convex as pronotum, broadest near anterior third and strongly narrowing toward apices, length
|
||
<quantity id="4CAFDC86FF87C164FE13FB69FD56FB51" box="[501,668,1246,1269]" metricMagnitude="-3" metricUnit="m" metricValue="1.08" metricValueMax="1.13" metricValueMin="1.03" pageId="3" pageNumber="300" unit="mm" value="1.08" valueMax="1.13" valueMin="1.03">1.03–1.13 mm</quantity>
|
||
(mean
|
||
<quantity id="4CAFDC86FF87C164FCCCFF09FC44FF71" box="[810,910,190,213]" metricMagnitude="-3" metricUnit="m" metricValue="1.08" pageId="3" pageNumber="300" unit="mm" value="1.08">1.08 mm</quantity>
|
||
), width
|
||
<quantity id="4CAFDC86FF87C164FC00FF09FB48FF71" box="[998,1154,190,213]" metricMagnitude="-4" metricUnit="m" metricValue="6.25" metricValueMax="6.5" metricValueMin="6.0" pageId="3" pageNumber="300" unit="mm" value="0.625" valueMax="0.65" valueMin="0.6">0.60–0.65 mm</quantity>
|
||
(mean
|
||
<quantity id="4CAFDC86FF87C164FB32FF09FAF2FF71" box="[1236,1336,190,213]" metricMagnitude="-4" metricUnit="m" metricValue="6.3" pageId="3" pageNumber="300" unit="mm" value="0.63">0.63 mm</quantity>
|
||
), EI 1.69– 1.73. Base of elytra arcuate so that humeri are projected anteriorly, humeral calli weakly marked, delimited from adsutural region by short and shallow, very broad basal impressions; basal foveae indiscernible; apices of elytra separately rounded. Punctures in anterior 2/3 of elytra much more distinct than those on pronotum, large, deep and relatively sharply marked; in median part of each elytron separated by spaces distinctly shorter than puncture diameters, punctures are gradually smaller and shallower toward apices and lateral margins of elytra. Vestiture slightly longer and distinctly more suberect than that on pronotum. Metathoracic wings long, functional.
|
||
</paragraph>
|
||
<paragraph id="8BE87163FF87C164FCCCFDC9FC34FD31" blockId="3.[810,1022,638,661]" box="[810,1022,638,661]" pageId="3" pageNumber="300">Legs non-modified.</paragraph>
|
||
<paragraph id="8BE87163FF87C164FCCCFD09FA48FCF1" blockId="3.[810,1449,702,853]" pageId="3" pageNumber="300">
|
||
Aedeagus (
|
||
<figureCitation id="136C6DE6FF87C164FC4FFD09FBEFFD71" box="[937,1061,702,725]" captionStart="Fig" captionStartId="2.[134,169,2060,2081]" captionTargetBox="[140,1473,1299,2029]" captionTargetId="figure-84@2.[122,1500,1284,2048]" captionTargetPageId="2" captionText="Fig. 2. Aedeagus in ventral (a, c) and lateral (b, d) views: a, b, Horaeomorphus imitator, new species; c, d, Horaeomorphus minor, new species. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4509156" httpUri="https://zenodo.org/record/4509156/files/figure.png" pageId="3" pageNumber="300">Figs. 2c, d</figureCitation>
|
||
)
|
||
<quantity id="4CAFDC86FF87C164FBDEFD09FB56FD71" box="[1080,1180,702,725]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="3" pageNumber="300" unit="mm" value="0.45">0.45 mm</quantity>
|
||
in length; median lobe moderately slender, with subtrapezoidal apical part; internal sac strongly asymmetrical, darkly sclerotized and complicated; parameres slender, not exceeding apex of median lobe, each with 4–5 apical and subapical setae.
|
||
</paragraph>
|
||
<paragraph id="8BE87163FF87C164FCCCFCC9FBD9FC31" blockId="3.[810,1043,894,918]" box="[810,1043,894,918]" pageId="3" pageNumber="300">
|
||
<emphasis id="B923AD71FF87C164FCCCFCC9FC53FC32" bold="true" box="[810,921,894,918]" italics="true" pageId="3" pageNumber="300">Female. –</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34D22E8FF87C164FCCCFC09FC64FC51" pageId="3" pageNumber="300" type="etymology">
|
||
<paragraph id="8BE87163FF87C164FCCCFC09FC64FC51" blockId="3.[810,1449,958,1013]" pageId="3" pageNumber="300">
|
||
<emphasis id="B923AD71FF87C164FCCCFC09FC71FC72" bold="true" box="[810,955,958,982]" italics="true" pageId="3" pageNumber="300">Etymology. –</emphasis>
|
||
The name
|
||
<taxonomicName id="4C570AE0FF87C164FBDCFC09FBB0FC71" authorityName="ski" authorityYear="2009" box="[1082,1146,958,981]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="300" phylum="Arthropoda" rank="species" species="minor">
|
||
<emphasis id="B923AD71FF87C164FBDCFC09FBB0FC71" box="[1082,1146,958,981]" italics="true" pageId="3" pageNumber="300">minor</emphasis>
|
||
</taxonomicName>
|
||
refers to the small body of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34D22E8FF87C164FCCCFBA9FB1AFBF1" pageId="3" pageNumber="300" type="distribution">
|
||
<paragraph id="8BE87163FF87C164FCCCFBA9FB1AFBF1" blockId="3.[810,1448,1054,1109]" pageId="3" pageNumber="300">
|
||
<emphasis id="B923AD71FF87C164FCCCFBA9FC14FB92" bold="true" box="[810,990,1054,1078]" italics="true" pageId="3" pageNumber="300">Distribution. –</emphasis>
|
||
<collectingCountry id="F34031F3FF87C164FC0CFBA9FBB9FB91" box="[1002,1139,1054,1077]" name="Philippines" pageId="3" pageNumber="300">Philippines</collectingCountry>
|
||
: Provinces
|
||
<collectingRegion id="4993BF81FF87C164FAE4FBA9FAB9FB91" box="[1282,1395,1054,1077]" country="Philippines" name="Bukidnon" pageId="3" pageNumber="300">Bukidnon</collectingRegion>
|
||
and
|
||
<collectingRegion id="4993BF81FF87C164FCCCFB89FBCDFBF1" box="[810,1031,1086,1109]" country="Philippines" name="Misamis Occidental" pageId="3" pageNumber="300">Misamis Occidental</collectingRegion>
|
||
: Mindanao Island.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34D22E8FF87C163FCCCFBC9FEBAFE11" lastPageId="4" lastPageNumber="301" pageId="3" pageNumber="300" type="discussion">
|
||
<paragraph id="8BE87163FF87C163FCCCFBC9FEBAFE11" blockId="3.[810,1450,1150,1269]" lastBlockId="4.[133,772,190,437]" lastPageId="4" lastPageNumber="301" pageId="3" pageNumber="300">
|
||
<emphasis id="B923AD71FF87C164FCCCFBC9FC61FB32" bold="true" box="[810,939,1150,1174]" italics="true" pageId="3" pageNumber="300">Remarks. –</emphasis>
|
||
This small, slender and flattened species can be easily distinguished from any other congeners by the asymmetrical and very large internal sac of the aedeagus. The only other species with asymmetrical internal sacs are
|
||
<taxonomicName id="4C570AE0FF80C163FF63FF09FEB2FF71" authority="Jaloszy" authorityName="Jaloszy" box="[133,376,190,213]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="301" phylum="Arthropoda" rank="species" species="deformatus">
|
||
<emphasis id="B923AD71FF80C163FF63FF09FEEAFF71" box="[133,288,190,213]" italics="true" pageId="4" pageNumber="301">H. deformatus</emphasis>
|
||
Jałoszy
|
||
</taxonomicName>
|
||
ski, 2006 from W
|
||
<collectingCountry id="F34031F3FF80C163FDB7FF09FD79FF71" box="[593,691,190,213]" name="Malaysia" pageId="4" pageNumber="301">Malaysia</collectingCountry>
|
||
(Kuala
|
||
<collectingRegion id="4993BF81FF80C163FF63FF69FEC0FF51" box="[133,266,222,245]" country="Malaysia" name="Terengganu" pageId="4" pageNumber="301">Terengganu</collectingRegion>
|
||
), and
|
||
<taxonomicName id="4C570AE0FF80C163FEB7FF69FD5BFF51" authority="Jaloszy" authorityName="Jaloszy" box="[337,657,222,245]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="301" phylum="Arthropoda" rank="species" species="pseudosabahensis">
|
||
<emphasis id="B923AD71FF80C163FEB7FF69FDF2FF51" box="[337,568,222,245]" italics="true" pageId="4" pageNumber="301">H. pseudosabahensis</emphasis>
|
||
Jałoszy
|
||
</taxonomicName>
|
||
ski, 2006 from
|
||
<collectingRegion id="4993BF81FF80C163FF25FF49FECEFEB1" box="[195,260,254,277]" country="Malaysia" name="Sabah" pageId="4" pageNumber="301">Sabah</collectingRegion>
|
||
and
|
||
<collectingCountry id="F34031F3FF80C163FED8FF49FE57FEB1" box="[318,413,254,277]" name="Malaysia" pageId="4" pageNumber="301">Sarawak</collectingCountry>
|
||
. The latter species has distinctly different body form (much larger and strongly convex).
|
||
<taxonomicName id="4C570AE0FF80C163FF63FE89FE78FEF1" box="[133,434,318,341]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="301" phylum="Arthropoda" rank="species" species="deformatus">
|
||
<emphasis id="B923AD71FF80C163FF63FE89FE78FEF1" box="[133,434,318,341]" italics="true" pageId="4" pageNumber="301">Horaeomorphus deformatus</emphasis>
|
||
</taxonomicName>
|
||
is relatively similar to
|
||
<taxonomicName id="4C570AE0FF80C163FD43FE89FCCEFEF1" authorityName="ski" authorityYear="2009" box="[677,772,318,341]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="301" phylum="Arthropoda" rank="species" species="minor">
|
||
<emphasis id="B923AD71FF80C163FD43FE89FCCEFEF1" box="[677,772,318,341]" italics="true" pageId="4" pageNumber="301">H. minor</emphasis>
|
||
</taxonomicName>
|
||
in general body shape, but it can be easily distinguished on the basis of strongly modified metatrochanters and metafemora in males.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |