351 lines
38 KiB
XML
351 lines
38 KiB
XML
<document id="2F7F51588F24D061ED3549985BD0868F" ID-DOI="10.5281/zenodo.189961" ID-GBIF-Dataset="f4918554-8847-4b2e-89af-ea92badaad40" ID-ISSN="1175-5326" ID-Zenodo-Dep="189961" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461106560087" checkinUser="plazi" docAuthor="Colloff, Matthew J. & Perdomo, Giselle" docDate="2009" docId="03FEC052FFEDDC62569F5B85FEBD0ACA" docLanguage="en" docName="zt02217p036.pdf" docOrigin="Zootaxa 2217" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Crotonia ramus Womersley 1957" docType="treatment" docVersion="9" lastPageNumber="30" masterDocId="FFC7B82AFFF7DC7C56085F5DFFFD0B69" masterDocTitle="New species of Crotonia (Acari: Oribatida: Camisiidae) from Nothofagus and Eucalyptus forests in Victoria, Australia, with a redescription of the fossil species Crotonia ramus (Womersley, 1957)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="27" updateTime="1698593351658" updateUser="plazi">
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<mods:mods id="EB691AA33C9F2D6846904FFE310EB1E3" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="899574D55B8291969D518E155B589872">
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<mods:title id="16C0BBBD126E428B46D666A5304208A4">New species of Crotonia (Acari: Oribatida: Camisiidae) from Nothofagus and Eucalyptus forests in Victoria, Australia, with a redescription of the fossil species Crotonia ramus (Womersley, 1957)</mods:title>
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<mods:name id="ABD87860B61A67E0C3D34CE75FCA8524" type="personal">
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<mods:roleTerm id="4FB0D8224926A0B24BB6A16DCEC8CFAC">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="85D5995AD78E2F16D8F169A5CF4762F3">Colloff, Matthew J.</mods:namePart>
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</mods:name>
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<mods:name id="91BC01E3F5F0E632374C15BBA1F7B292" type="personal">
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<mods:role id="029304F35DCA2CA67C37CD7D3F519FA3">
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<mods:roleTerm id="4F22AB2AF5007546C6C8A90DC68377B9">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="565C0FFCEB73E343084A1A8472C1F30C">Perdomo, Giselle</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="A7C71B32A66444349F03A46799EFC501">text</mods:typeOfResource>
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<mods:titleInfo id="CA8429CD3F0A42BD65710E55813E96B4">
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<mods:title id="7F08864FFB0B7548A1791FD047B205DE">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="7049F425C02D1CBE864ACEC9D76DE0A6">
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<mods:date id="2DFF30D1B35C24518F8AEEFCC0EB374F">2009</mods:date>
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<mods:detail id="AB3B8861E5D7285919ECD42DB0400F30" type="volume">
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<mods:number id="FF9DF0C278204A344051837961D903A1">2217</mods:number>
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</mods:detail>
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<mods:extent id="EEB80935F97468991E69CA872C67E0D6" unit="page">
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<mods:start id="FC10B4EB89462D54FBE003DC430686AB">1</mods:start>
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<mods:end id="75620A4DB583CDA57A5235CC5BF551C8">36</mods:end>
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<mods:classification id="A8155A6DED940EA41FAEA7D3D98B019E">journal article</mods:classification>
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<mods:identifier id="6A0F26A3A74C1B6D7DAC0055EC299217" type="DOI">10.5281/zenodo.189961</mods:identifier>
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<mods:identifier id="78B411B3BBB81FE5D624CE741D8D3FB3" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="BCCF86ECF894F072963DE59ADE8DA507" type="Zenodo-Dep">189961</mods:identifier>
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<treatment id="03FEC052FFEDDC62569F5B85FEBD0ACA" ID-DOI="http://doi.org/10.5281/zenodo.6225406" ID-GBIF-Taxon="119623402" ID-Zenodo-Dep="6225406" LSID="urn:lsid:plazi:treatment:03FEC052FFEDDC62569F5B85FEBD0ACA" httpUri="http://treatment.plazi.org/id/03FEC052FFEDDC62569F5B85FEBD0ACA" lastPageId="30" lastPageNumber="30" pageId="26" pageNumber="27">
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<subSubSection id="C34D22CFFFEDDC66569F5B85FEC80E7F" pageId="26" pageNumber="27" type="nomenclature">
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<paragraph id="8BE87144FFEDDC66569F5B85FDAE0F9B" blockId="26.[151,595,1240,1302]" box="[151,595,1240,1267]" pageId="26" pageNumber="27">
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<heading id="D0A0C628FFEDDC66569F5B85FDAE0F9B" bold="true" box="[151,595,1240,1267]" fontSize="11" level="1" pageId="26" pageNumber="27" reason="1">
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<emphasis id="B923AD56FFEDDC66569F5B85FDAE0F9B" bold="true" box="[151,595,1240,1267]" pageId="26" pageNumber="27">
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†
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<taxonomicName id="4C570AC7FFEDDC6656AD5B84FDAE0F9B" authority="Womersley, 1957" authorityName="Womersley" authorityYear="1957" box="[165,595,1240,1267]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="ramus">
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<emphasis id="B923AD56FFEDDC6656AD5B84FE980F9A" bold="true" box="[165,357,1241,1267]" italics="true" pageId="26" pageNumber="27">Crotonia ramus</emphasis>
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(
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<bibRefCitation id="EFC60CB5FFEDDC66577C5B85FDB60F9B" author="Womersley" box="[372,587,1240,1266]" pageId="26" pageNumber="36" refString="Womersley, H. (1957) A fossil mite (Acronothrus ramus n. sp.) from Cainozoic resin at Allendale, Victoria. Proceedings of the Royal Society of Victoria, 69, 21 - 23." type="journal article" year="1957">Womersley, 1957</bibRefCitation>
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)
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BE87144FFEDDC66569F5BA1FEC80E7F" blockId="26.[151,595,1240,1302]" box="[151,309,1276,1302]" pageId="26" pageNumber="27">
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(
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<figureCitation id="136C6DC1FFEDDC6656975BA1FEFF0E7F" box="[159,258,1276,1302]" captionStart="FIGURE 16" captionStartId="27.[151,255,1856,1880]" captionTargetBox="[207,1379,491,1815]" captionTargetId="figure@27.[195,1392,482,1831]" captionTargetPageId="27" captionText="FIGURE 16. Crotonia ramus (Womersley, 1957). Holotype female a) dorsal; b) ventral." httpUri="https://zenodo.org/record/189973/files/figure.png" pageId="26" pageNumber="27">Figs. 16</figureCitation>
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,
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<figureCitation id="136C6DC1FFEDDC6657065BA1FED10E7F" box="[270,300,1276,1302]" captionStart="FIGURE 19" captionStartId="30.[151,257,1379,1403]" captionTargetBox="[169,1413,494,1336]" captionTargetId="figure@30.[151,1436,476,1352]" captionTargetPageId="30" captionText="FIGURE 19. Distribution map of Crotonia spp. in Victoria. Stippled area is the distribution of the broad vegetation group categorised as ‘ wet or damp forests’ (Department of Sustainability and Environment, 2005). NP = National Park; SF = State Forest. Black circles are towns and cities, provided for spatial reference. Inset: 30 - year means of annual rainfall (source: Australian Bureau of Meteorology, www. bom. gov. au)." httpUri="https://zenodo.org/record/189976/files/figure.png" pageId="26" pageNumber="27">19</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C34D22CFFFEDDC66569F5A14FD760E99" pageId="26" pageNumber="27" type="reference_group">
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<paragraph id="8BE87144FFEDDC66569F5A14FD3B0E09" blockId="26.[151,802,1353,1408]" box="[151,710,1353,1376]" pageId="26" pageNumber="27">
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<taxonomicName id="4C570AC7FFEDDC66569F5A14FDDA0E09" authority="Womersley, 1957" authorityName="Womersley" authorityYear="1957" box="[151,551,1353,1376]" class="Arachnida" family="Crotoniidae" genus="Acronothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="ramus">
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<emphasis id="B923AD56FFEDDC66569F5A14FE980E09" box="[151,357,1353,1376]" italics="true" pageId="26" pageNumber="27">Acronothrus ramus</emphasis>
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<bibRefCitation id="EFC60CB5FFEDDC6657635A14FDDA0E09" author="Womersley" box="[363,551,1353,1376]" pageId="26" pageNumber="36" refString="Womersley, H. (1957) A fossil mite (Acronothrus ramus n. sp.) from Cainozoic resin at Allendale, Victoria. Proceedings of the Royal Society of Victoria, 69, 21 - 23." type="journal article" year="1957">Womersley, 1957</bibRefCitation>
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</taxonomicName>
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, p. 22,
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<figureCitation id="136C6DC1FFEDDC66547D5A14FD3C0E09" box="[629,705,1353,1376]" captionStart="FIGURE 11" captionStartId="20.[151,255,1578,1602]" captionTargetBox="[278,1294,414,1534]" captionTargetId="figure@20.[229,1357,390,1553]" captionTargetPageId="20" captionText="FIGURE 11. Crotonia blacki sp. nov. protonymph a) dorsal; b) ventral" httpUri="https://zenodo.org/record/189969/files/figure.png" pageId="26" pageNumber="27">Fig. 11</figureCitation>
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.
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</paragraph>
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<paragraph id="8BE87144FFEDDC66569F5A34FCDF0EE9" blockId="26.[151,802,1353,1408]" box="[151,802,1385,1408]" pageId="26" pageNumber="27">
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<taxonomicName id="4C570AC7FFEDDC66569F5A34FDE80EE9" authority="Womersley, 1957" authorityName="Womersley" authorityYear="1957" box="[151,533,1385,1408]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="ramus">
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<emphasis id="B923AD56FFEDDC66569F5A34FEBD0EE9" box="[151,320,1385,1408]" italics="true" pageId="26" pageNumber="27">Crotonia ramus</emphasis>
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(
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<bibRefCitation id="EFC60CB5FFEDDC6657455A34FDF00EE9" author="Womersley" box="[333,525,1385,1408]" pageId="26" pageNumber="36" refString="Womersley, H. (1957) A fossil mite (Acronothrus ramus n. sp.) from Cainozoic resin at Allendale, Victoria. Proceedings of the Royal Society of Victoria, 69, 21 - 23." type="journal article" year="1957">Womersley, 1957</bibRefCitation>
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)
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</taxonomicName>
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:
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<bibRefCitation id="EFC60CB5FFEDDC6654175A34FD300EE9" author="Wallwork" box="[543,717,1385,1408]" pageId="26" pageNumber="36" refString="Wallwork, J. A. (1977) On the genus Crotonia Thorell 1876 (Acari: Cryptostigmata) Acarologia, 19, 513 - 539." type="journal article" year="1977">Wallwork, 1977</bibRefCitation>
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, p. 525.
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</paragraph>
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<paragraph id="8BE87144FFEDDC66569F5AF2FD760E99" blockId="26.[151,1437,1455,2003]" pageId="26" pageNumber="27">
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<emphasis id="B923AD56FFEDDC66569F5AF2FED50EA0" bold="true" box="[151,296,1455,1481]" pageId="26" pageNumber="27">Dimensions</emphasis>
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.
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<typeStatus id="54ECCFE6FFEDDC66573F5AEDFE540EA3" box="[311,425,1456,1482]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
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: a damaged female with several dorsal setae broken off; length 705 breadth 362; ratio of length of prodorsum to total length: 0.4.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34D22CFFFEDDC6056CD5AA0FDE40BB2" lastPageId="28" lastPageNumber="29" pageId="26" pageNumber="27" type="description">
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<paragraph id="8BE87144FFEDDC6656CD5AA0FC590DB1" blockId="26.[151,1437,1455,2003]" pageId="26" pageNumber="27">
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<emphasis id="B923AD56FFEDDC6656CD5AA0FEDC0D7E" bold="true" box="[197,289,1533,1559]" pageId="26" pageNumber="27">Female</emphasis>
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.
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<emphasis id="B923AD56FFEDDC6657395AA0FE4B0D7F" box="[305,438,1533,1558]" italics="true" pageId="26" pageNumber="27">Prodorsum</emphasis>
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: rostrum rounded, not prominent; rostral setae (
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<emphasis id="B923AD56FFEDDC6655F45AA0FBE90D7F" box="[1020,1044,1533,1558]" italics="true" pageId="26" pageNumber="27">ro</emphasis>
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) short, spiniform, 15 (
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<figureCitation id="136C6DC1FFEDDC66532F5AA0FA820D7E" box="[1319,1407,1533,1559]" captionStart="FIGURE 16" captionStartId="27.[151,255,1856,1880]" captionTargetBox="[207,1379,491,1815]" captionTargetId="figure@27.[195,1392,482,1831]" captionTargetPageId="27" captionText="FIGURE 16. Crotonia ramus (Womersley, 1957). Holotype female a) dorsal; b) ventral." httpUri="https://zenodo.org/record/189973/files/figure.png" pageId="26" pageNumber="27">Fig. 16</figureCitation>
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a). Lamellar setae (
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<emphasis id="B923AD56FFEDDC6657565979FE890D54" box="[350,372,1572,1597]" italics="true" pageId="26" pageNumber="27">le</emphasis>
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) 130, recurved, barbed. Lamellar apophyses 58, half as long as their mutual distance; extending anteriorly as far as bases of rostral setae. Interlamellar apophyses twice as long as broad; interlamellar setae (
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<emphasis id="B923AD56FFEDDC665777592CFE680DE3" box="[383,405,1649,1674]" italics="true" pageId="26" pageNumber="27">in</emphasis>
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) broken off. Prodorsal ridges extending two thirds of the distance between interlamellar and lamellar apophyses. With inter-bothridial ridge in the form of a shallow curve; cuticle posterior of ridge smooth. Median field of muscle sigilla present. Prodorsum porose.
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</paragraph>
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<paragraph id="8BE87144FFEDDC6756CD59B8FE760AC1" blockId="26.[151,1437,1455,2003]" lastBlockId="27.[151,1437,152,424]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
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<emphasis id="B923AD56FFEDDC6656CD59B8FEB70D97" box="[197,330,1765,1790]" italics="true" pageId="26" pageNumber="27">Notogaster</emphasis>
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: ratio of length to breadth 1.54; broadest between
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<emphasis id="B923AD56FFEDDC6655A759B8FC310D97" box="[943,972,1765,1790]" italics="true" pageId="26" pageNumber="27">cp</emphasis>
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and
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<emphasis id="B923AD56FFEDDC66520F59B8FBE90D97" box="[1031,1044,1765,1790]" italics="true" pageId="26" pageNumber="27">e</emphasis>
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2 (
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<figureCitation id="136C6DC1FFEDDC66522659B8FB7A0D96" box="[1070,1159,1765,1791]" captionStart="FIGURE 16" captionStartId="27.[151,255,1856,1880]" captionTargetBox="[207,1379,491,1815]" captionTargetId="figure@27.[195,1392,482,1831]" captionTargetPageId="27" captionText="FIGURE 16. Crotonia ramus (Womersley, 1957). Holotype female a) dorsal; b) ventral." httpUri="https://zenodo.org/record/189973/files/figure.png" pageId="26" pageNumber="27">Fig. 16</figureCitation>
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a). Dorsosejugal suture discrete, simple. With 13 pairs of notogastral setae;
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<emphasis id="B923AD56FFEDDC6655075852FCE10C41" box="[783,796,1807,1832]" italics="true" pageId="26" pageNumber="27">c</emphasis>
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3,
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<emphasis id="B923AD56FFEDDC66553D5852FCC00C41" box="[821,829,1807,1832]" italics="true" pageId="26" pageNumber="27">f</emphasis>
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1 and
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<emphasis id="B923AD56FFEDDC66558A5852FC6C0C41" box="[898,913,1807,1832]" italics="true" pageId="26" pageNumber="27">h</emphasis>
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1 broken off. Pre-notogastral shield bearing relatively long, 72, setiform setae
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<emphasis id="B923AD56FFEDDC66544F5864FDA90C3B" box="[583,596,1849,1874]" italics="true" pageId="26" pageNumber="27">c</emphasis>
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1 and separated from notogastral shield by transverse hyaline strip. Apophyses of setae
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<emphasis id="B923AD56FFEDDC665780583FFE680C12" box="[392,405,1890,1915]" italics="true" pageId="26" pageNumber="27">c</emphasis>
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3 very well-developed, 32, twice as long as broad. Notogastral shield discrete, porose; bordered laterally by two narrow strips lacking pori, extending posteriorly almost as far as setae
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<emphasis id="B923AD56FFEDDC66532D58D1FAD00CCC" box="[1317,1325,1932,1957]" italics="true" pageId="26" pageNumber="27">f</emphasis>
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2. Region posterior of setae
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<emphasis id="B923AD56FFEDDC66576158E8FE8C0CA7" box="[361,369,1973,1998]" italics="true" pageId="26" pageNumber="27">f</emphasis>
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2 with sparse tubercles. Lateral hyaline strip (suprapleural scissure) bearing alveoli of setae
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<emphasis id="B923AD56FFECDC67569F5FC5FF4E0BD8" box="[151,179,152,177]" italics="true" pageId="27" pageNumber="28">cp</emphasis>
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, and
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<emphasis id="B923AD56FFECDC6756FC5FC5FEFC0BD8" box="[244,257,152,177]" italics="true" pageId="27" pageNumber="28">e</emphasis>
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2. Tubercles of setae
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<emphasis id="B923AD56FFECDC6757FE5FC5FE030BD8" box="[502,510,152,177]" italics="true" pageId="27" pageNumber="28">f</emphasis>
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2 squat, projecting beyond lateral margin. Setae
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<emphasis id="B923AD56FFECDC67523C5FC5FBC10BD8" box="[1076,1084,152,177]" italics="true" pageId="27" pageNumber="28">f</emphasis>
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2 spiniform, barbed, short, 23. Setae
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<emphasis id="B923AD56FFECDC6756E85F9CFF000BB3" box="[224,253,193,218]" italics="true" pageId="27" pageNumber="28">cp</emphasis>
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and
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<emphasis id="B923AD56FFECDC6757315F9CFEBB0BB3" box="[313,326,193,218]" italics="true" pageId="27" pageNumber="28">e</emphasis>
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2 and subequal, 58; setae
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<emphasis id="B923AD56FFECDC6754715F9CFD750BB3" box="[633,648,193,218]" italics="true" pageId="27" pageNumber="28">d</emphasis>
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2 slightly shorter, 47, but well-developed, setiform. Opisthosomal gland opening
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<emphasis id="B923AD56FFECDC6757455FB6FE890A6D" box="[333,372,235,260]" italics="true" pageId="27" pageNumber="28">gla</emphasis>
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positioned ca. 35 anterior of
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<emphasis id="B923AD56FFECDC6754E85FB6FD150A6D" box="[736,744,235,260]" italics="true" pageId="27" pageNumber="28">f</emphasis>
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2. Apophyses of setae
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<emphasis id="B923AD56FFECDC6755F05FB6FBFD0A6D" box="[1016,1024,235,260]" italics="true" pageId="27" pageNumber="28">f</emphasis>
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1 emerging from dorsal surface of apophyses of setae
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<emphasis id="B923AD56FFECDC6757745E48FE770A47" box="[380,394,277,302]" italics="true" pageId="27" pageNumber="28">h</emphasis>
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2, projecting posteriolaterally, adjacent and parallel with those of
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<emphasis id="B923AD56FFECDC6752815E48FB650A47" box="[1161,1176,277,302]" italics="true" pageId="27" pageNumber="28">h</emphasis>
|
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1, and subequal, 20, to them. Setae
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<emphasis id="B923AD56FFECDC6757235E63FECE0A3E" box="[299,307,318,343]" italics="true" pageId="27" pageNumber="28">f</emphasis>
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1,
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<emphasis id="B923AD56FFECDC6757445E63FEA60A3E" box="[332,347,318,343]" italics="true" pageId="27" pageNumber="28">h</emphasis>
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1 and
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<emphasis id="B923AD56FFECDC6757965E63FE500A3E" box="[414,429,318,343]" italics="true" pageId="27" pageNumber="28">h</emphasis>
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3 broken off
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<typeStatus id="54ECCFE6FFECDC67544B5E63FD560A31" box="[579,683,318,344]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
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specimen. Setae
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<emphasis id="B923AD56FFECDC6755755E63FC710A3E" box="[893,908,318,343]" italics="true" pageId="27" pageNumber="28">h</emphasis>
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2 short, 30, spinose, barbed; their apophyses relatively long, 65, parallel for most of their length, slightly diverging apically; caudal margin between them an inverted U-shape.
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</paragraph>
|
||
<caption id="DF2821CCFFECDC67569F581DFBBA0C31" box="[151,1095,1856,1880]" httpUri="https://zenodo.org/record/189973/files/figure.png" pageId="27" pageNumber="28" targetBox="[207,1379,491,1815]" targetPageId="27">
|
||
<paragraph id="8BE87144FFECDC67569F581DFBBA0C31" blockId="27.[151,1095,1856,1880]" box="[151,1095,1856,1880]" pageId="27" pageNumber="28">
|
||
<emphasis id="B923AD56FFECDC67569F581DFEDB0C31" bold="true" box="[151,294,1856,1880]" pageId="27" pageNumber="28">FIGURE 16.</emphasis>
|
||
<taxonomicName id="4C570AC7FFECDC675725581CFD560C31" authority="Womersley, 1957" authorityName="Womersley" authorityYear="1957" box="[301,683,1857,1880]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="ramus">
|
||
<emphasis id="B923AD56FFECDC675725581CFE2B0C31" box="[301,470,1857,1880]" italics="true" pageId="27" pageNumber="28">Crotonia ramus</emphasis>
|
||
(Womersley, 1957)
|
||
</taxonomicName>
|
||
. Holotype female a) dorsal; b) ventral.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE87144FFECDC6056CD58DCFDE40BB2" blockId="27.[151,1436,1921,2024]" lastBlockId="28.[151,1436,152,219]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
|
||
<emphasis id="B923AD56FFECDC6756CD58DCFEF10CF3" box="[197,268,1921,1946]" italics="true" pageId="27" pageNumber="28">Ven t er</emphasis>
|
||
: epimeres porose (
|
||
<figureCitation id="136C6DC1FFECDC6757F958DCFDBA0CF2" box="[497,583,1921,1947]" captionStart="FIGURE 16" captionStartId="27.[151,255,1856,1880]" captionTargetBox="[207,1379,491,1815]" captionTargetId="figure@27.[195,1392,482,1831]" captionTargetPageId="27" captionText="FIGURE 16. Crotonia ramus (Womersley, 1957). Holotype female a) dorsal; b) ventral." httpUri="https://zenodo.org/record/189973/files/figure.png" pageId="27" pageNumber="28">Fig. 16</figureCitation>
|
||
b); epimeral setae smooth, spiniform, formula 3-2-3-3; setae
|
||
<emphasis id="B923AD56FFECDC67531E58DCFACF0CF3" box="[1302,1330,1921,1946]" italics="true" pageId="27" pageNumber="28">3c</emphasis>
|
||
on welldeveloped tubercles. Genital plates sub-circular. Each plate 95 long, 56 broad with eight spiniform setae. Anal plate 30 broad, 165 long, with three pairs of stout setiform setae, 22; Ventral margin of notogaster surrounding anal plates U-shaped. Setae
|
||
<emphasis id="B923AD56FFEBDC6057EC5FC5FE0E0BD8" box="[484,499,152,177]" italics="true" pageId="28" pageNumber="29">p</emphasis>
|
||
2 and
|
||
<emphasis id="B923AD56FFEBDC60543D5FC5FDB90BD8" box="[565,580,152,177]" italics="true" pageId="28" pageNumber="29">p</emphasis>
|
||
3 curved, subequal, 24,
|
||
<emphasis id="B923AD56FFEBDC6055505FC5FC9B0BD8" box="[856,870,152,177]" italics="true" pageId="28" pageNumber="29">p</emphasis>
|
||
1 60, recurved, stout, barbed, on well-developed tubercles, 15 very close together.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF2821CCFFEBDC60569F5AA8FEAF0D3A" httpUri="https://zenodo.org/record/189974/files/figure.png" pageId="28" pageNumber="29" targetBox="[230,1360,297,1483]" targetPageId="28">
|
||
<paragraph id="8BE87144FFEBDC60569F5AA8FEAF0D3A" blockId="28.[151,1437,1525,1619]" pageId="28" pageNumber="29">
|
||
<emphasis id="B923AD56FFEBDC60569F5AA8FED50D64" bold="true" box="[151,296,1525,1549]" pageId="28" pageNumber="29">FIGURE 17.</emphasis>
|
||
Posteriolateral views of a)
|
||
<taxonomicName id="4C570AC7FFEBDC60545B5AABFCEF0D64" box="[595,786,1526,1549]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="momitoi" status="sp. nov.">
|
||
<emphasis id="B923AD56FFEBDC60545B5AABFCEF0D64" box="[595,786,1526,1549]" italics="true" pageId="28" pageNumber="29">Crotonia momitoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A210102DFFEBDC6055125AA8FC8D0D64" box="[794,880,1525,1549]" pageId="28" pageNumber="29" rank="species">
|
||
<emphasis id="B923AD56FFEBDC6055125AA8FC970D64" bold="true" box="[794,874,1525,1549]" pageId="28" pageNumber="29">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, male paratype; b)
|
||
<taxonomicName id="4C570AC7FFEBDC60524C5AABFB100D64" box="[1092,1261,1526,1549]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="blacki" status="sp. nov.">
|
||
<emphasis id="B923AD56FFEBDC60524C5AABFB100D64" box="[1092,1261,1526,1549]" italics="true" pageId="28" pageNumber="29">Crotonia blacki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A210102DFFEBDC6052FD5AA8FAB60D64" box="[1269,1355,1525,1549]" pageId="28" pageNumber="29" rank="species">
|
||
<emphasis id="B923AD56FFEBDC6052FD5AA8FAB90D64" bold="true" box="[1269,1348,1525,1549]" pageId="28" pageNumber="29">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
female paratype; c)
|
||
<taxonomicName id="4C570AC7FFEBDC60572C5944FDE80D59" box="[292,533,1561,1584]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="gadubanudi" status="sp. nov.">
|
||
<emphasis id="B923AD56FFEBDC60572C5944FDE80D59" box="[292,533,1561,1584]" italics="true" pageId="28" pageNumber="29">Crotonia gadubanudi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B923AD56FFEBDC6054155945FD8A0D59" bold="true" box="[541,631,1560,1584]" pageId="28" pageNumber="29">
|
||
<taxonomicNameLabel id="A210102DFFEBDC6054155945FD8A0D59" box="[541,631,1560,1584]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; d)
|
||
<taxonomicName id="4C570AC7FFEBDC6054AE5944FCA40D59" box="[678,857,1561,1584]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="alpina" status="sp. nov.">
|
||
<emphasis id="B923AD56FFEBDC6054AE5944FCA40D59" box="[678,857,1561,1584]" italics="true" pageId="28" pageNumber="29">Crotonia alpina</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B923AD56FFEBDC6055695945FC460D59" bold="true" box="[865,955,1560,1584]" pageId="28" pageNumber="29">
|
||
<taxonomicNameLabel id="A210102DFFEBDC6055695945FC460D59" box="[865,955,1560,1584]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; e)
|
||
<taxonomicName id="4C570AC7FFEBDC6055E15944FB510D59" box="[1001,1196,1561,1584]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="cornuta" status="sp. nov.">
|
||
<emphasis id="B923AD56FFEBDC6055E15944FB510D59" box="[1001,1196,1561,1584]" italics="true" pageId="28" pageNumber="29">Crotonia cornuta</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B923AD56FFEBDC6052BC5945FAF30D59" bold="true" box="[1204,1294,1560,1584]" pageId="28" pageNumber="29">
|
||
<taxonomicNameLabel id="A210102DFFEBDC6052BC5945FAF30D59" box="[1204,1294,1560,1584]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; f)
|
||
<taxonomicName id="4C570AC7FFEBDC6053315944FF0B0D3B" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="victoriae" status="sp. nov.">
|
||
<emphasis id="B923AD56FFEBDC6053315944FF0B0D3B" italics="true" pageId="28" pageNumber="29">Crotonia victoriae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A210102DFFEBDC6056F45966FEAF0D3A" box="[252,338,1595,1619]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C34D22CFFFEBDC6056CD59DDFB010C5C" pageId="28" pageNumber="29" type="materials_examined">
|
||
<paragraph id="8BE87144FFEBDC6056CD59DDFB010C5C" blockId="28.[151,1437,1664,2007]" pageId="28" pageNumber="29">
|
||
<emphasis id="B923AD56FFEBDC6056CD59DDFD5E0DF3" bold="true" box="[197,675,1664,1690]" pageId="28" pageNumber="29">Material Examined and Locality Data</emphasis>
|
||
.
|
||
<typeStatus id="54ECCFE6FFEBDC6054BA59DCFCE20DF2" box="[690,799,1665,1691]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
|
||
female from block of fossilized resin (retinite) of late Tertiary age, probably Pliocene, near Ballarat in the Victorian goldfields. This species was in the Department of Geology, University of Melbourne, transferred to Department of Invertebrate Palaeontology, Museum Victoria from whence it was loaned. Permission was given to the first author to remount the
|
||
<typeStatus id="54ECCFE6FFEBDC6052D959A9FAC50C67" box="[1233,1336,1780,1806]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
|
||
because the notogaster was obscured by debris. Unfortunately, the specimen was lost during the process.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34D22CFFFEBDC6156CD581CFC380BB1" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29" type="discussion">
|
||
<paragraph id="8BE87144FFEBDC6156CD581CFC380BB1" blockId="28.[151,1437,1664,2007]" lastBlockId="29.[151,1436,152,220]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
|
||
<emphasis id="B923AD56FFEBDC6056CD581CFEBD0C32" bold="true" box="[197,320,1857,1883]" pageId="28" pageNumber="29">Remarks.</emphasis>
|
||
<bibRefCitation id="EFC60CB5FFEBDC605740581FFDDA0C35" author="Womersley" box="[328,551,1858,1884]" pageId="28" pageNumber="36" refString="Womersley, H. (1957) A fossil mite (Acronothrus ramus n. sp.) from Cainozoic resin at Allendale, Victoria. Proceedings of the Royal Society of Victoria, 69, 21 - 23." type="journal article" year="1957">Womersley (1957)</bibRefCitation>
|
||
illustrated this species with the caudal tritonymphal exuvium intact. This was removed during remounting and cleaning by the first author to reveal short spiniform adult setae
|
||
<emphasis id="B923AD56FFEBDC6053345834FAB90CEB" box="[1340,1348,1897,1922]" italics="true" pageId="28" pageNumber="29">f</emphasis>
|
||
2 and
|
||
<emphasis id="B923AD56FFEBDC60538D5834FA690CEB" box="[1413,1428,1897,1922]" italics="true" pageId="28" pageNumber="29">h</emphasis>
|
||
2 rather than the long, curved tritonymphal setae illustrated in Womersley’s
|
||
<figureCitation id="136C6DC1FFEBDC60523C58CFFB600CC5" box="[1076,1181,1938,1964]" captionStart="FIGURE 1" captionStartId="3.[151,255,1725,1749]" captionTargetBox="[212,1366,476,1675]" captionTargetId="figure@3.[199,1387,461,1700]" captionTargetPageId="3" captionText="FIGURE 1. Crotonia alpina sp. nov. holotype female a) dorsal; b) ventral." httpUri="https://zenodo.org/record/189962/files/figure.png" pageId="28" pageNumber="29">Figure 1</figureCitation>
|
||
, as well as the more elongated nature of the caudal region and apophyses. Setae
|
||
<emphasis id="B923AD56FFEBDC60556158E4FC8B0CBB" box="[873,886,1977,2002]" italics="true" pageId="28" pageNumber="29">c</emphasis>
|
||
3 appear to have been broken off the
|
||
<typeStatus id="54ECCFE6FFEBDC60533C58E4FA600CBA" box="[1332,1437,1977,2003]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
|
||
specimen prior to the original description, but the presence of massive protuberant apophyses strongly suggest that setae
|
||
<emphasis id="B923AD56FFEADC6157045FE2FEE40BB1" box="[268,281,191,216]" italics="true" pageId="29" pageNumber="30">c</emphasis>
|
||
3 were elongated, flagelliform and probably longer than
|
||
<emphasis id="B923AD56FFEADC6155A25FE2FC4A0BB1" box="[938,951,191,216]" italics="true" pageId="29" pageNumber="30">c</emphasis>
|
||
1.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF2821CCFFEADC61569F586FFB5F0C05" httpUri="https://zenodo.org/record/189975/files/figure.png" pageId="29" pageNumber="30" targetBox="[304,1277,294,1804]" targetPageId="29">
|
||
<paragraph id="8BE87144FFEADC61569F586FFB5F0C05" blockId="29.[151,1436,1842,1900]" pageId="29" pageNumber="30">
|
||
<emphasis id="B923AD56FFEADC61569F586FFEDA0C23" bold="true" box="[151,295,1842,1866]" pageId="29" pageNumber="30">FIGURE 18.</emphasis>
|
||
Details of bothridia of a)
|
||
<taxonomicName id="4C570AC7FFEADC615448586FFD000C20" box="[576,765,1842,1865]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="momitoi" status="sp. nov.">
|
||
<emphasis id="B923AD56FFEADC615448586FFD000C20" box="[576,765,1842,1865]" italics="true" pageId="29" pageNumber="30">Crotonia momitoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A210102DFFEADC61550D586FFCA70C20" box="[773,858,1842,1866]" pageId="29" pageNumber="30" rank="species">
|
||
<emphasis id="B923AD56FFEADC61550D586FFCA90C23" bold="true" box="[773,852,1842,1866]" pageId="29" pageNumber="30">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
; b)
|
||
<taxonomicName id="4C570AC7FFEADC61558F586FFBD30C20" box="[903,1070,1842,1865]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="blacki" status="sp. nov.">
|
||
<emphasis id="B923AD56FFEADC61558F586FFBD30C20" box="[903,1070,1842,1865]" italics="true" pageId="29" pageNumber="30">Crotonia blacki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A210102DFFEADC61523E586FFB770C20" box="[1078,1162,1842,1866]" pageId="29" pageNumber="30" rank="species">
|
||
<emphasis id="B923AD56FFEADC61523E586FFB780C23" bold="true" box="[1078,1157,1842,1866]" pageId="29" pageNumber="30">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
; c)
|
||
<taxonomicName id="4C570AC7FFEADC6152BE586FFA610C20" box="[1206,1436,1842,1865]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="gadubanudi" status="sp. nov.">
|
||
<emphasis id="B923AD56FFEADC6152BE586FFA610C20" box="[1206,1436,1842,1865]" italics="true" pageId="29" pageNumber="30">Crotonia gadubanudi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B923AD56FFEADC61569F5809FF110C05" bold="true" box="[151,236,1876,1900]" pageId="29" pageNumber="30">
|
||
<taxonomicNameLabel id="A210102DFFEADC61569F5809FF110C05" box="[151,236,1876,1900]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; d)
|
||
<taxonomicName id="4C570AC7FFEADC61571F5808FE3D0C05" box="[279,448,1877,1900]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="alpina" status="sp. nov.">
|
||
<emphasis id="B923AD56FFEADC61571F5808FE3D0C05" box="[279,448,1877,1900]" italics="true" pageId="29" pageNumber="30">Crotonia alpina</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B923AD56FFEADC6157C05809FDE00C05" bold="true" box="[456,541,1876,1900]" pageId="29" pageNumber="30">
|
||
<taxonomicNameLabel id="A210102DFFEADC6157C05809FDE00C05" box="[456,541,1876,1900]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; e)
|
||
<taxonomicName id="4C570AC7FFEADC61544E5808FD020C05" box="[582,767,1877,1900]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="cornuta" status="sp. nov.">
|
||
<emphasis id="B923AD56FFEADC61544E5808FD020C05" box="[582,767,1877,1900]" italics="true" pageId="29" pageNumber="30">Crotonia cornuta</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A210102DFFEADC61550E5809FCA60C05" box="[774,859,1876,1900]" pageId="29" pageNumber="30" rank="species">
|
||
<emphasis id="B923AD56FFEADC61550E5809FCA60C05" bold="true" box="[774,859,1876,1900]" pageId="29" pageNumber="30">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
; f)
|
||
<taxonomicName id="4C570AC7FFEADC6155895808FBBB0C05" box="[897,1094,1877,1900]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="victoriae" status="sp. nov.">
|
||
<emphasis id="B923AD56FFEADC6155895808FBBB0C05" box="[897,1094,1877,1900]" italics="true" pageId="29" pageNumber="30">Crotonia victoriae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A210102DFFEADC6152455809FB5F0C05" box="[1101,1186,1876,1900]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C34D22CFFFEADC6256CD58C7FEBD0ACA" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="reference_group">
|
||
<paragraph id="8BE87144FFEADC6256CD58C7FEBD0ACA" blockId="29.[151,1436,1946,2011]" lastBlockId="30.[151,1436,152,419]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
|
||
<taxonomicName id="4C570AC7FFEADC6156CD58C7FE830CDA" box="[197,382,1946,1971]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="ramus">
|
||
<emphasis id="B923AD56FFEADC6156CD58C7FE830CDA" box="[197,382,1946,1971]" italics="true" pageId="29" pageNumber="30">Crotonia ramus</emphasis>
|
||
</taxonomicName>
|
||
differs from other members of the genus by the following combination of characters: 1) it is by far the smallest member of the genus - the next longest (812) being
|
||
<taxonomicName id="4C570AC7FFEADC615206589CFA8B0CB2" ID-CoL="ZRGQ" authority="Wallwork, 1977" authorityName="Wallwork" authorityYear="1977" box="[1038,1398,1985,2011]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="chiloensis">
|
||
<emphasis id="B923AD56FFEADC615206589CFB570CB3" box="[1038,1194,1985,2010]" italics="true" pageId="29" pageNumber="30">C. chiloensis</emphasis>
|
||
<bibRefCitation id="EFC60CB5FFEADC6152BA589CFA8B0CB2" author="Wallwork" box="[1202,1398,1985,2011]" pageId="29" pageNumber="36" refString="Wallwork, J. A. (1977) On the genus Crotonia Thorell 1876 (Acari: Cryptostigmata) Acarologia, 19, 513 - 539." type="journal article" year="1977">Wallwork, 1977</bibRefCitation>
|
||
</taxonomicName>
|
||
; 2) setae
|
||
<emphasis id="B923AD56FFE9DC6256D65FC5FF100BD8" box="[222,237,152,177]" italics="true" pageId="30" pageNumber="31">d</emphasis>
|
||
2 are long, well-developed, almost as long as
|
||
<emphasis id="B923AD56FFE9DC6255225FC5FCBA0BD8" box="[810,839,152,177]" italics="true" pageId="30" pageNumber="31">cp</emphasis>
|
||
and
|
||
<emphasis id="B923AD56FFE9DC62558E5FC5FC6E0BD8" box="[902,915,152,177]" italics="true" pageId="30" pageNumber="31">e</emphasis>
|
||
2; 3) setae
|
||
<emphasis id="B923AD56FFE9DC6252115FC5FBDC0BD8" box="[1049,1057,152,177]" italics="true" pageId="30" pageNumber="31">f</emphasis>
|
||
2 are spiniform, barbed; 4) the notogastral shield is porose, with two narrow lateral strips lacking tubercles; 5) setae
|
||
<emphasis id="B923AD56FFE9DC6252945F9CFB560BB3" box="[1180,1195,193,218]" italics="true" pageId="30" pageNumber="31">p</emphasis>
|
||
1 are stout, reflexed, barbed and positioned adjacent to each other; 6) with the epimeral setal formula 3-2-3-3. It is morphologically most similar to
|
||
<taxonomicName id="4C570AC7FFE9DC6257585E4FFE2E0A42" ID-CoL="ZRHB" box="[336,467,274,299]" class="Arachnida" family="Crotoniidae" genus="Crotonia" kingdom="Animalia" order="Sarcoptiformes" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="momitoi" status="sp. nov.">
|
||
<emphasis id="B923AD56FFE9DC6257585E4FFE2E0A42" box="[336,467,274,299]" italics="true" pageId="30" pageNumber="31">C. momitoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B923AD56FFE9DC6257D25E4CFDC50A42" bold="true" box="[474,568,273,299]" pageId="30" pageNumber="31">
|
||
<taxonomicNameLabel id="A210102DFFE9DC6257D25E4CFDC50A42" box="[474,568,273,299]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from which it differs by its smaller size, in the shape of the apophyses of setae
|
||
<emphasis id="B923AD56FFE9DC6256D55E64FF110A3B" box="[221,236,313,338]" italics="true" pageId="30" pageNumber="31">h</emphasis>
|
||
2, the morphology of setae
|
||
<emphasis id="B923AD56FFE9DC6254315E64FDBC0A3B" box="[569,577,313,338]" italics="true" pageId="30" pageNumber="31">f</emphasis>
|
||
2 and
|
||
<emphasis id="B923AD56FFE9DC6254805E64FD6A0A3B" box="[648,663,313,338]" italics="true" pageId="30" pageNumber="31">p</emphasis>
|
||
1, the absence of narrow lateral strips of small tubercles on the notogastral shield, a less prominent rostrum and rostral setae and the presence of barbed ornamentation on the lamellar setae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |