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<document id="A9A74006D4DD04A54BDAE125E95BD624" ID-DOI="10.5281/zenodo.5352700" ID-ISSN="2345-7600" ID-Zenodo-Dep="5352700" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630208580763" checkinUser="tatiana" docAuthor="Polhemus, Dan A. &amp; Polhemus, John T." docDate="2013" docId="03FE7253F512FFE8FF12FB78FDC3FAB1" docLanguage="en" docName="61rbz665-686.pdf" docOrigin="Raffles Bulletin of Zoology 61 (2)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Heleocoris malayensis Polhemus &amp; Polhemus 2013, new species" docType="treatment" docVersion="3" lastPageNumber="678" masterDocId="FFC70A2BF519FFE5FFFEFFA4FFFDFFE7" masterDocTitle="Guide To The Aquatic Heteroptera Of Singapore And Peninsular Malaysia. Xi. Infraorder Nepomorpha- Families Naucoridae And Aphelocheiridae" masterLastPageNumber="686" masterPageNumber="665" pageNumber="676" updateTime="1699186152900" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="7D84D3FD9DC8B93F44E4359AAD00EE7A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="24257E9DA0FDF8E05AF0FC955D415823">
<mods:title id="43FB4BEA2A25A6FD4B6EFF4303914D38">Guide To The Aquatic Heteroptera Of Singapore And Peninsular Malaysia. Xi. Infraorder Nepomorpha- Families Naucoridae And Aphelocheiridae</mods:title>
</mods:titleInfo>
<mods:name id="E0B814A08698397B2C68CC31C29A39F7" type="personal">
<mods:role id="A5CCED90BB3BCA3EE5FB8D8393B26AA7">
<mods:roleTerm id="1B8545682167985B34C555BC2E282F2F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="73DAF346A149ADC077F59AB66D957363">Polhemus, Dan A.</mods:namePart>
</mods:name>
<mods:name id="811094DCC797743F86C4368B478F050D" type="personal">
<mods:role id="B5B4FF0A2398A356AA88AE8A44830327">
<mods:roleTerm id="C2B3302931D9161FA3A0B0907A7C3BDA">Author</mods:roleTerm>
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<mods:namePart id="857227699704C73B50F1AF453C239131">Polhemus, John T.</mods:namePart>
</mods:name>
<mods:typeOfResource id="7D31410C95B366B02AB046250949D697">text</mods:typeOfResource>
<mods:relatedItem id="EC313B5A7945BC0342A79652D1B2FB88" type="host">
<mods:titleInfo id="1E51D49026F94D0BE5511D96460CC218">
<mods:title id="0C1F062B25E6903D484EA112C62EB229">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="B6993E5A3584477DEC2EA57A77217482">
<mods:date id="25189C6BF9F17EAA788896F02E7C4314">2013</mods:date>
<mods:detail id="E416217B5C140F72E2D532C42C25A629" type="pubDate">
<mods:number id="40DCFAD8FB507A24A194FDBB0E7A4841">2013-08-30</mods:number>
</mods:detail>
<mods:detail id="433987BA756E598D869F91A1086644DA" type="volume">
<mods:number id="93D6805558E1B61F9CC355402407F875">61</mods:number>
</mods:detail>
<mods:detail id="12CBB06E4CA502DD0DF5C21247C3027F" type="issue">
<mods:number id="DF7A0BE1583D9CD9A8C4B32EC705DBFF">2</mods:number>
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<mods:extent id="F7EF7FD73DA43C3D14DC59578306202E" unit="page">
<mods:start id="6330C7918637E42427B9583603486656">665</mods:start>
<mods:end id="0F42B7FE3ADF84CE13DF410C78B1569C">686</mods:end>
</mods:extent>
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<mods:classification id="28DCE9923F0BB1D95E3217795178759C">journal article</mods:classification>
<mods:identifier id="B802640FED3D84A19E4CCD1BDF39FCEF" type="DOI">10.5281/zenodo.5352700</mods:identifier>
<mods:identifier id="B9F6F42AE33F00EE0FE8A92F93F1F906" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="C6541F47E33A187EE4BE616685BDCEF9" type="Zenodo-Dep">5352700</mods:identifier>
</mods:mods>
<treatment id="03FE7253F512FFE8FF12FB78FDC3FAB1" LSID="urn:lsid:plazi:treatment:03FE7253F512FFE8FF12FB78FDC3FAB1" httpUri="http://treatment.plazi.org/id/03FE7253F512FFE8FF12FB78FDC3FAB1" lastPageId="13" lastPageNumber="678" pageId="11" pageNumber="676">
<subSubSection id="C34D90CEF512FFEEFF12FB78FD8AFB13" box="[236,631,1244,1269]" pageId="11" pageNumber="676" type="nomenclature">
<paragraph id="8BE8C345F512FFEEFF12FB78FD8AFB13" blockId="11.[236,631,1244,1301]" box="[236,631,1244,1269]" pageId="11" pageNumber="676">
<heading id="D0A07429F512FFEEFF12FB78FD8AFB13" bold="true" box="[236,631,1244,1269]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="676" reason="2">
<emphasis id="B9231F57F512FFEEFF12FB78FD8AFB13" bold="true" box="[236,631,1244,1269]" pageId="11" pageNumber="676">
<taxonomicName id="4C57B8C6F512FFEEFF12FB78FE1EFB12" authority="Polhemus &amp; Polhemus, 2013" authorityName="Polhemus &amp; Polhemus" authorityYear="2013" box="[236,483,1244,1269]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="676" phylum="Arthropoda" rank="species" species="malayensis" status="sp. nov.">
<emphasis id="B9231F57F512FFEEFF12FB78FE1EFB12" bold="true" box="[236,483,1244,1269]" italics="true" pageId="11" pageNumber="676">Heleocoris malayensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A210A22CF512FFEEFE0CFB79FD8AFB13" box="[498,631,1244,1269]" pageId="11" pageNumber="676" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34D90CEF512FFEEFED4FB59FA8EFC52" pageId="11" pageNumber="676" type="description">
<paragraph id="8BE8C345F512FFEEFED4FB59FDC4FAF2" blockId="11.[236,631,1244,1301]" box="[298,569,1277,1301]" pageId="11" pageNumber="676">
(
<figureCitation id="136CDFC0F512FFEEFECCFB59FE71FAF2" box="[306,396,1277,1301]" captionStart="Figs" captionStartId="10.[136,180,1861,1883]" captionTargetBox="[139,775,714,1834]" captionTargetId="figure-8@10.[117,790,706,1848]" captionTargetPageId="10" captionText="Figs. 24–28. Heleocoris species, male phallotheca. 24. Heleocoris malayensis, new species, specimen from Malaysia, Terengganu, Sungai Brang. 25. Heleocoris montandoni Lundblad, specimen from Singapore, Nee Soon swamp forest. 26. Heleocoris nebulosus Montandon, specimen from Indonesia, Bali, Melangit River. 27. Heleocoris ovatus Montandon, specimen from Vietnam, Lam Dong Prov., Pongour Falls, CL 3091. 28. Heleocoris strabus Montandon, specimen from Thailand, Chiang Mai Prov., Huay Hia Creek, CL 2198." figureDoi="http://doi.org/10.5281/zenodo.5352718" httpUri="https://zenodo.org/record/5352718/files/figure.png" pageId="11" pageNumber="676">Figs. 24</figureCitation>
,
<figureCitation id="136CDFC0F512FFEEFE67FB59FE4BFAF2" box="[409,438,1277,1301]" captionStart="Figs" captionStartId="11.[812,856,2004,2026]" captionTargetBox="[893,1369,1020,1970]" captionTargetId="figure-13@11.[880,1386,1000,1976]" captionTargetPageId="11" captionText="Figs. 29, 30. Heleocoris species, male proctigers. 29. Heleocoris montandoni Lundblad, specimen from Singapore, Nee Soon swamp forest. 30. Heleocoris malayensis new species, specimen from Malaysia, Terengganu, Sungai Brang." figureDoi="http://doi.org/10.5281/zenodo.5352720" httpUri="https://zenodo.org/record/5352720/files/figure.png" pageId="11" pageNumber="676">30</figureCitation>
,
<figureCitation id="136CDFC0F512FFEEFE3CFB59FE22FAF2" box="[450,479,1277,1301]" captionStart="Figs" captionStartId="12.[835,879,1091,1113]" captionTargetBox="[834,1469,233,1050]" captionTargetId="figure-8@12.[819,1486,216,1072]" captionTargetPageId="12" captionText="Figs. 31, 32. Heleocoris species, female terminal abdomen, ventral view, showing shape of subgenital plate. 31. Heleocoris montandoni Lundblad, specimen from Vietnam, Lam Dong Prov., CL 3094. 32. Heleocoris malayensis, new species, specimen from Malaysia, Perak, Kerunai River, CL 2078." figureDoi="http://doi.org/10.5281/zenodo.5352722" httpUri="https://zenodo.org/record/5352722/files/figure.png" pageId="11" pageNumber="676">32</figureCitation>
,
<figureCitation id="136CDFC0F512FFEEFE15FB59FDF5FAF2" box="[491,520,1277,1301]" captionStart="Fig" captionStartId="12.[836,871,2061,2083]" captionTargetBox="[836,1474,1300,2038]" captionTargetId="figure-494@12.[836,1476,1299,2038]" captionTargetPageId="12" captionText="Fig. 33. Heleocoris malayensis, new species, male, colour photo of dorsal habitus.Specimen from Malaysia, Terengganu, SungaiBrang." figureDoi="http://doi.org/10.5281/zenodo.5352724" httpUri="https://zenodo.org/record/5352724/files/figure.png" pageId="11" pageNumber="676">33</figureCitation>
,
<figureCitation id="136CDFC0F512FFEEFDEBFB59FDCCFAF2" box="[533,561,1277,1301]" captionStart="Figs" captionStartId="13.[812,856,2002,2024]" captionTargetBox="[892,1329,785,1964]" captionTargetId="figure-226@13.[872,1393,776,1974]" captionTargetPageId="13" captionText="Figs. 34, 35. Heleocoris species, male left abdominal paratergites. 34. Heleocoris malayensis, new species, specimen from Malaysia, Terengganu, Sungai Brang. 35. Heleocoris ovatus Montandon, specimen from Vietnam, Lam Dong Prov., Pongour Falls." figureDoi="http://doi.org/10.5281/zenodo.5352726" httpUri="https://zenodo.org/record/5352726/files/figure.png" pageId="11" pageNumber="676">34</figureCitation>
)
</paragraph>
<paragraph id="8BE8C345F512FFEEFCD2FF19FAE0FD32" blockId="11.[812,1451,189,725]" pageId="11" pageNumber="676">posterior margin broadly dark, width across eyes/length = 4.20/1.40; eyes dark red, shining, roughly teardrop-shaped when viewed from above with well developed lateral flange, tapering anteriorly, width/length = 0.70/1.20, inner margins straight and slightly convergent anteriorly, lateral margins broadly curving, separated from vertex by shallow furrows, anterior/posterior interocular width = 1.70/2.20; posterior margin of vertex nearly straight, not produced behind eyes; anteclypeus broadly rounded, not projecting ahead of eyes, anterior margin projecting beyond rostrum for 0.42 length of rostrum, bearing a pair (1+1) of transversely ovate dark brown pits widely spaced to either side of longitudinal midline, these pits bearing short, pale setae; labrum golden, roughly triangular, broadly rounded, apex blunt; rostrum evenly tapering to apex, dark yellow, terminal segment dark brown; antennae with all segments thickened, not extending to lateral margin of head, segment III longest.</paragraph>
<paragraph id="8BE8C345F512FFEEFCD2FD59FA8EFC52" blockId="11.[812,1451,765,949]" pageId="11" pageNumber="676">
<emphasis id="B9231F57F512FFEEFCD2FD59FC60FCF2" bold="true" box="[812,925,765,789]" pageId="11" pageNumber="676">Pronotum</emphasis>
yellowish brown, irregularly dotted with numerous small, dark brown spots, lateral margins translucent golden, posterior margin broadly grayish brown; dorsal surface broadly and evenly domed, not depressed medially behind vertex, width/length (midline) = 6.15/3.60, lateral margins not explanate, posterolateral angles truncate, rounded.
</paragraph>
</subSubSection>
<subSubSection id="C34D90CEF512FFEEFF8FFA9EFD70F902" pageId="11" pageNumber="676" type="materials_examined">
<paragraph id="8BE8C345F512FFEEFF8FFA9EFD70F902" blockId="11.[113,754,1338,1765]" pageId="11" pageNumber="676">
<emphasis id="B9231F57F512FFEEFF8FFA9EFEC7FAB7" bold="true" box="[113,314,1338,1360]" pageId="11" pageNumber="676">
<emphasis id="B9231F57F512FFEEFF8FFA9EFEC9FAB7" bold="true" box="[113,308,1338,1360]" italics="true" pageId="11" pageNumber="676">Material examined</emphasis>
.
</emphasis>
—
<materialsCitation id="3B3FC918F512FFEEFE9BFA9FFE7EFA6C" collectingDate="1985-08-18" collectionCode="T, JTPC" country="Malaysia" location="Tapah" municipality="Sungai Cheroh" pageId="11" pageNumber="676" specimenCount="1" stateProvince="Pahang" typeStatus="holotype">
<typeStatus id="54EC7DE7F512FFEEFE9BFA9FFE3EFAB6" box="[357,451,1339,1361]" pageId="11" pageNumber="676" type="holotype">Holotype</typeStatus>
, male,
<collectingCountry id="F34083D5F512FFEEFDF0FA9FFD72FAB7" box="[526,655,1339,1360]" name="Malaysia" pageId="11" pageNumber="676">MALAYSIA</collectingCountry>
,
<emphasis id="B9231F57F512FFEEFD62FA9FFD16FAB7" bold="true" box="[668,747,1339,1360]" pageId="11" pageNumber="676">
<collectingRegion id="49930DA7F512FFEEFD62FA9FFD16FAB7" box="[668,747,1339,1360]" country="Malaysia" name="Pahang" pageId="11" pageNumber="676">Pahang</collectingRegion>
</emphasis>
,
<collectingMunicipality id="6B8C593FF512FFEEFF8FFAFCFEFEFA89" box="[113,259,1368,1390]" pageId="11" pageNumber="676">Sungai Cheroh</collectingMunicipality>
,
<quantity id="4CAF6EA0F512FFEEFEF3FAFCFEC2FA89" box="[269,319,1368,1390]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="676" unit="km" value="6.0">6 km</quantity>
E. of
<location id="8E88959EF512FFEEFE87FAFCFE4BFA89" LSID="urn:lsid:plazi:treatment:03FE7253F512FFE8FF12FB78FDC3FAB1:8E88959EF512FFEEFE87FAFCFE4BFA89" box="[377,438,1368,1390]" country="Malaysia" municipality="Sungai Cheroh" name="Tapah" pageId="11" pageNumber="676" stateProvince="Pahang">Tapah</location>
,
<date id="FFE9E585F512FFEEFE3EFAFCFDC2FA89" box="[448,575,1368,1390]" pageId="11" pageNumber="676" value="1985-08-18">
<collectingDate id="EFAD1C6DF512FFEEFE3EFAFCFDC2FA89" box="[448,575,1368,1390]" pageId="11" pageNumber="676" value="1985-08-18">18 Aug.1985</collectingDate>
</date>
, CL 2072, coll. D. A. &amp; J.
<collectionCode id="ED465B80F512FFEEFF43FAD1FF30FA6D" box="[189,205,1397,1418]" name="Tavera, Department of Geology and Geophysics" pageId="11" pageNumber="676">T</collectionCode>
. Polhemus (
<collectionCode id="ED465B80F512FFEEFEBFFAD1FE81FA6C" box="[321,380,1397,1419]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33646" lsid="urn:lsid:biocol.org:col:33646" name="Colorado Entomological Museum (formerly John T. Polhemus collection)" pageId="11" pageNumber="676">JTPC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B3FC918F512FFEEFE72FAD1FD70FA4F" collectingDate="1985-08-18" collectionCode="JTPC, ZRC" country="Malaysia" location="Tapah" municipality="Sungai Cheroh" pageId="11" pageNumber="676" specimenCount="35" specimenCount-female="18" specimenCount-male="17" stateProvince="Pahang" typeStatus="paratype">
<typeStatus id="54EC7DE7F512FFEEFE72FAD1FE10FA6D" box="[396,493,1397,1418]" pageId="11" pageNumber="676" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="F34083D5F512FFEEFE08FAD1FD88FA6D" box="[502,629,1397,1418]" name="Malaysia" pageId="11" pageNumber="676">MALAYSIA</collectingCountry>
,
<emphasis id="B9231F57F512FFEEFD81FAD1FD31FA6D" bold="true" box="[639,716,1397,1418]" pageId="11" pageNumber="676">
<collectingRegion id="49930DA7F512FFEEFD81FAD1FD31FA6D" box="[639,716,1397,1418]" country="Malaysia" name="Pahang" pageId="11" pageNumber="676">Pahang</collectingRegion>
</emphasis>
:
<specimenCount id="9D5108CCF512FFEEFD27FAD1FF56FA4F" pageId="11" pageNumber="676" type="male">17 males</specimenCount>
,
<specimenCount id="9D5108CCF512FFEEFF48FA36FEDFFA4F" box="[182,290,1426,1448]" pageId="11" pageNumber="676" type="female">18 females</specimenCount>
, same data as holotype (
<collectionCode id="ED465B80F512FFEEFDF1FA36FDB5FA40" box="[527,584,1426,1447]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33646" lsid="urn:lsid:biocol.org:col:33646" name="Colorado Entomological Museum (formerly John T. Polhemus collection)" pageId="11" pageNumber="676">JTPC</collectionCode>
,
<collectionCode id="ED465B80F512FFEEFDADFA36FD7BFA4F" box="[595,646,1426,1448]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="676">ZRC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3FC918F512FFEEFD69FA36FE06FA22" collectionCode="ZRC" collectorName="L. C. Fong" country="Malaysia" location="Taman Negara" pageId="11" pageNumber="676" specimenCount="1" specimenCount-female="1" stateProvince="Pahang" typeStatus="paratype">
<specimenCount id="9D5108CCF512FFEEFD69FA36FD13FA4F" box="[663,750,1426,1448]" pageId="11" pageNumber="676" type="female">1 female</specimenCount>
,
<location id="8E88959EF512FFEEFF8FFA0BFEFEFA23" LSID="urn:lsid:plazi:treatment:03FE7253F512FFE8FF12FB78FDC3FAB1:8E88959EF512FFEEFF8FFA0BFEFEFA23" box="[113,259,1455,1476]" country="Malaysia" name="Taman Negara" pageId="11" pageNumber="676" stateProvince="Pahang">Taman Negara</location>
, coll.
<collectorName id="26A2A693F512FFEEFEBCFA0BFE4DFA23" box="[322,432,1455,1476]" pageId="11" pageNumber="676">L. C. Fong</collectorName>
(
<collectionCode id="ED465B80F512FFEEFE3EFA0BFE09FA22" box="[448,500,1455,1477]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="676">ZRC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3FC918F512FFEEFDF9FA0BFDDEFA06" collectingDate="1992-11-11" collectionCode="ZRC" collectorName="J. Cramphorn" country="Malaysia" location="Sungai Kla" municipality="Raub" pageId="11" pageNumber="676" specimenCount="1" specimenCount-female="1" stateProvince="Pahang" typeStatus="paratype">
<specimenCount id="9D5108CCF512FFEEFDF9FA0BFD9DFA23" box="[519,608,1454,1476]" pageId="11" pageNumber="676" type="female">1 female</specimenCount>
,
<collectingMunicipality id="6B8C593FF512FFEEFD93FA0AFD5FFA23" box="[621,674,1454,1476]" pageId="11" pageNumber="676">Raub</collectingMunicipality>
,
<location id="8E88959EF512FFEEFD50FA0AFF6BFA06" LSID="urn:lsid:plazi:treatment:03FE7253F512FFE8FF12FB78FDC3FAB1:8E88959EF512FFEEFD50FA0AFF6BFA06" country="Malaysia" municipality="Raub" name="Sungai Kla" pageId="11" pageNumber="676" stateProvince="Pahang">Sungai Kla</location>
,
<date id="FFE9E585F512FFEEFF5EFA6FFEE0FA07" box="[160,285,1483,1504]" pageId="11" pageNumber="676" value="1992-11-11">
<collectingDate id="EFAD1C6DF512FFEEFF5EFA6FFEE0FA07" box="[160,285,1483,1504]" pageId="11" pageNumber="676" value="1992-11-11">11 Nov.1992</collectingDate>
</date>
, coll.
<collectorName id="26A2A693F512FFEEFEA6FA68FE26FA06" box="[344,475,1483,1505]" pageId="11" pageNumber="676">J. Cramphorn</collectorName>
(
<collectionCode id="ED465B80F512FFEEFE16FA6FFDE1FA06" box="[488,540,1483,1505]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="676">ZRC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B3FC918F512FFEEFDD2FA6FFE2FF9FC" collectingDate="1985-08-19" collectionCode="T, JTPC" collectorName="Polhemus" country="Malaysia" location="Grik" municipality="Sungai Kerunai" pageId="11" pageNumber="676" specimenCount="35" specimenCount-female="16" specimenCount-male="19" stateProvince="Perak" typeStatus="paratype">
<emphasis id="B9231F57F512FFEEFDD2FA6FFD94FA07" bold="true" box="[556,617,1483,1504]" pageId="11" pageNumber="676">
<collectingRegion id="49930DA7F512FFEEFDD2FA6FFD94FA07" box="[556,617,1483,1504]" country="Malaysia" name="Perak" pageId="11" pageNumber="676">Perak</collectingRegion>
</emphasis>
:
<specimenCount id="9D5108CCF512FFEEFD88FA6FFD32FA06" box="[630,719,1483,1505]" pageId="11" pageNumber="676" type="male">19 males</specimenCount>
,
<specimenCount id="9D5108CCF512FFEEFD27FA6FFF43FA19" pageId="11" pageNumber="676" type="female">16 females</specimenCount>
,
<collectingMunicipality id="6B8C593FF512FFEEFF36FA4CFE9DFA19" box="[200,352,1512,1534]" pageId="11" pageNumber="676">Sungai Kerunai</collectingMunicipality>
,
<quantity id="4CAF6EA0F512FFEEFE94FA4CFE66FA19" box="[362,411,1512,1534]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="11" pageNumber="676" unit="km" value="9.0">9 km</quantity>
N. of
<location id="8E88959EF512FFEEFE29FA4CFDF8FA19" LSID="urn:lsid:plazi:treatment:03FE7253F512FFE8FF12FB78FDC3FAB1:8E88959EF512FFEEFE29FA4CFDF8FA19" box="[471,517,1512,1534]" country="Malaysia" municipality="Sungai Kerunai" name="Grik" pageId="11" pageNumber="676" stateProvince="Perak">Grik</location>
,
<date id="FFE9E585F512FFEEFDF1FA4CFD73FA19" box="[527,654,1512,1534]" pageId="11" pageNumber="676" value="1985-08-19">
<collectingDate id="EFAD1C6DF512FFEEFDF1FA4CFD73FA19" box="[527,654,1512,1534]" pageId="11" pageNumber="676" value="1985-08-19">19 Aug.1985</collectingDate>
</date>
, CL 2078, coll. D. A. &amp; J.
<collectionCode id="ED465B80F512FFEEFEF5F9A1FEE6F9FD" box="[267,283,1541,1562]" name="Tavera, Department of Geology and Geophysics" pageId="11" pageNumber="676">T</collectionCode>
.
<collectorName id="26A2A693F512FFEEFEDBF9A1FE79F9FC" box="[293,388,1541,1563]" pageId="11" pageNumber="676">Polhemus</collectorName>
(
<collectionCode id="ED465B80F512FFEEFE6FF9A1FE31F9FC" box="[401,460,1541,1563]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33646" lsid="urn:lsid:biocol.org:col:33646" name="Colorado Entomological Museum (formerly John T. Polhemus collection)" pageId="11" pageNumber="676">JTPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3FC918F512FFEEFE22F9A1FD63F9DF" collectingDate="1997-02-17" collectionCode="ZRC" collectorName="H. K. Lua" country="Malaysia" location="Gerik" municipality="Sungai Kenderong" pageId="11" pageNumber="676" specimenCount="1" specimenCount-female="1" stateProvince="Perak" typeStatus="paratype">
<specimenCount id="9D5108CCF512FFEEFE22F9A1FDCEF9FC" box="[476,563,1541,1563]" pageId="11" pageNumber="676" type="female">1 female</specimenCount>
,
<collectingMunicipality id="6B8C593FF512FFEEFDC3F9A1FD0CF9FC" box="[573,753,1541,1563]" pageId="11" pageNumber="676">Sungai Kenderong</collectingMunicipality>
at
<location id="8E88959EF512FFEEFF74F986FF3EF9DF" LSID="urn:lsid:plazi:treatment:03FE7253F512FFE8FF12FB78FDC3FAB1:8E88959EF512FFEEFF74F986FF3EF9DF" box="[138,195,1570,1592]" country="Malaysia" municipality="Sungai Kenderong" name="Gerik" pageId="11" pageNumber="676" stateProvince="Perak">Gerik</location>
,
<date id="FFE9E585F512FFEEFF30F986FEB4F9DF" box="[206,329,1570,1592]" pageId="11" pageNumber="676" value="1997-02-17">
<collectingDate id="EFAD1C6DF512FFEEFF30F986FEB4F9DF" box="[206,329,1570,1592]" pageId="11" pageNumber="676" value="1997-02-17">17 Feb.1997</collectingDate>
</date>
, coll.
<collectorName id="26A2A693F512FFEEFE7BF986FE17F9D0" box="[389,490,1570,1591]" pageId="11" pageNumber="676">H. K. Lua</collectorName>
, LHK0324 (
<collectionCode id="ED465B80F512FFEEFD9AF986FD65F9DF" box="[612,664,1570,1592]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="676">ZRC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3FC918F512FFEEFD57F986FF4EF996" collectingDate="1997-02-16" collectionCode="ZRC" collectorName="H. K. Lua" country="Malaysia" location="Jalong" municipality="Sungai Korbu" pageId="11" pageNumber="676" specimenCount="1" specimenCount-male="1" stateProvince="Perak" typeStatus="paratype">
<specimenCount id="9D5108CCF512FFEEFD57F986FD10F9DF" box="[681,749,1570,1592]" pageId="11" pageNumber="676" type="male">1 male</specimenCount>
,
<collectingMunicipality id="6B8C593FF512FFEEFF8FF99AFF04F9B3" box="[113,249,1598,1620]" pageId="11" pageNumber="676">Sungai Korbu</collectingMunicipality>
at
<location id="8E88959EF512FFEEFEE4F99AFEA6F9B3" LSID="urn:lsid:plazi:treatment:03FE7253F512FFE8FF12FB78FDC3FAB1:8E88959EF512FFEEFEE4F99AFEA6F9B3" box="[282,347,1598,1620]" country="Malaysia" municipality="Sungai Korbu" name="Jalong" pageId="11" pageNumber="676" stateProvince="Perak">Jalong</location>
,
<date id="FFE9E585F512FFEEFE98F99AFE1FF9B3" box="[358,482,1598,1620]" pageId="11" pageNumber="676" value="1997-02-16">
<collectingDate id="EFAD1C6DF512FFEEFE98F99AFE1FF9B3" box="[358,482,1598,1620]" pageId="11" pageNumber="676" value="1997-02-16">16 Feb.1997</collectingDate>
</date>
, coll.
<collectorName id="26A2A693F512FFEEFDE1F99BFD78F9B3" box="[543,645,1599,1620]" pageId="11" pageNumber="676">H. K. Lua</collectorName>
, LHK0321 (
<collectionCode id="ED465B80F512FFEEFF86F9FFFF51F996" box="[120,172,1627,1649]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="676">ZRC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3FC918F512FFEEFF40F9FFFED0F997" box="[190,301,1627,1648]" country="Malaysia" location="Trengganu" pageId="11" pageNumber="676" specimenCount="1" stateProvince="Perak" typeStatus="paratype">
<emphasis id="B9231F57F512FFEEFF40F9FFFED0F997" bold="true" box="[190,301,1627,1648]" pageId="11" pageNumber="676">
<location id="8E88959EF512FFEEFF40F9FFFED0F997" LSID="urn:lsid:plazi:treatment:03FE7253F512FFE8FF12FB78FDC3FAB1:8E88959EF512FFEEFF40F9FFFED0F997" box="[190,301,1627,1648]" country="Malaysia" name="Trengganu" pageId="11" pageNumber="676" stateProvince="Perak">Trengganu</location>
</emphasis>
</materialsCitation>
:
<materialsCitation id="3B3FC918F512FFEEFEC2F9FFFD67F969" collectingDate="1995-05-16" collectionCode="ZRC" collectorName="B. Tan &amp; G. Sumita" country="Malaysia" location="Sekayu" municipality="Sungai Trengganu" pageId="11" pageNumber="676" specimenCount="1" specimenCount-male="1" stateProvince="Perak" typeStatus="paratype">
<specimenCount id="9D5108CCF512FFEEFEC2F9FFFE7CF996" box="[316,385,1627,1649]" pageId="11" pageNumber="676" type="male">1 male</specimenCount>
,
<collectingMunicipality id="6B8C593FF512FFEEFE72F9FFFDC0F996" box="[396,573,1627,1649]" pageId="11" pageNumber="676">Sungai Trengganu</collectingMunicipality>
tributary,
<location id="8E88959EF512FFEEFD5AF9FFFD10F996" LSID="urn:lsid:plazi:treatment:03FE7253F512FFE8FF12FB78FDC3FAB1:8E88959EF512FFEEFD5AF9FFFD10F996" box="[676,749,1627,1649]" country="Malaysia" municipality="Sungai Trengganu" name="Sekayu" pageId="11" pageNumber="676" stateProvince="Perak">Sekayu</location>
,
<date id="FFE9E585F512FFEEFF8FF9DCFF07F969" box="[113,250,1656,1678]" pageId="11" pageNumber="676" value="1995-05-16">
<collectingDate id="EFAD1C6DF512FFEEFF8FF9DCFF07F969" box="[113,250,1656,1678]" pageId="11" pageNumber="676" value="1995-05-16">16 May 1995</collectingDate>
</date>
, coll.
<collectorName id="26A2A693F512FFEEFEC5F9DDFE83F969" box="[315,382,1657,1678]" pageId="11" pageNumber="676">B. Tan</collectorName>
&amp;
<collectorName id="26A2A693F512FFEEFE5DF9DCFDF7F969" box="[419,522,1656,1678]" pageId="11" pageNumber="676">G. Sumita</collectorName>
, TG06 (
<collectionCode id="ED465B80F512FFEEFDA1F9DCFD6EF969" box="[607,659,1656,1678]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="676">ZRC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3FC918F512FFEEFD59F9DCFE66F92F" collectingDate="1992-03-18" collectionCode="ZRC" country="Malaysia" location="Sekayu Waterfall Park" municipality="Sungai Trengganu" pageId="11" pageNumber="676" specimenCount="1" specimenCount-male="1" stateProvince="Perak" typeStatus="paratype">
<specimenCount id="9D5108CCF512FFEEFD59F9DCFD10F969" box="[679,749,1656,1678]" pageId="11" pageNumber="676" type="male">1 male</specimenCount>
, tributary of
<collectingMunicipality id="6B8C593FF512FFEEFF1CF931FE6DF94D" box="[226,400,1685,1707]" pageId="11" pageNumber="676">Sungai Trengganu</collectingMunicipality>
downriver of
<location id="8E88959EF512FFEEFDE8F931FD10F94C" LSID="urn:lsid:plazi:treatment:03FE7253F512FFE8FF12FB78FDC3FAB1:8E88959EF512FFEEFDE8F931FD10F94C" box="[534,749,1685,1707]" country="Malaysia" municipality="Sungai Trengganu" name="Sekayu Waterfall Park" pageId="11" pageNumber="676" stateProvince="Perak">Sekayu Waterfall Park</location>
,
<date id="FFE9E585F512FFEEFF8FF916FF10F920" box="[113,237,1714,1735]" pageId="11" pageNumber="676" value="1992-03-18">
<collectingDate id="EFAD1C6DF512FFEEFF8FF916FF10F920" box="[113,237,1714,1735]" pageId="11" pageNumber="676" value="1992-03-18">18 Mar.1992</collectingDate>
</date>
, LHK179c (
<collectionCode id="ED465B80F512FFEEFE9FF916FE68F92F" box="[353,405,1714,1736]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="676">ZRC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3FC918F512FFEEFE5AF916FD74F902" collectingDate="1998-10-21" collectionCode="ZRC" collectorName="H. K. Lua" country="Malaysia" location="Sungai Brang" municipality="Sekayu" pageId="11" pageNumber="676" specimenCount="13" specimenCount-female="5" specimenCount-male="8" stateProvince="Perak" typeStatus="paratype">
<specimenCount id="9D5108CCF512FFEEFE5AF916FE13F92F" box="[420,494,1714,1736]" pageId="11" pageNumber="676" type="male">8 males</specimenCount>
,
<specimenCount id="9D5108CCF512FFEEFE09F916FDA8F92F" box="[503,597,1714,1736]" pageId="11" pageNumber="676" type="female">5 females</specimenCount>
,
<collectingMunicipality id="6B8C593FF512FFEEFDA3F916FD5BF92F" box="[605,678,1714,1736]" pageId="11" pageNumber="676">Sekayu</collectingMunicipality>
,
<location id="8E88959EF512FFEEFD50F916FF53F903" LSID="urn:lsid:plazi:treatment:03FE7253F512FFE8FF12FB78FDC3FAB1:8E88959EF512FFEEFD50F916FF53F903" country="Malaysia" municipality="Sekayu" name="Sungai Brang" pageId="11" pageNumber="676" stateProvince="Perak">Sungai Brang</location>
,
<date id="FFE9E585F512FFEEFF47F96BFEC9F903" box="[185,308,1743,1764]" pageId="11" pageNumber="676" value="1998-10-21">
<collectingDate id="EFAD1C6DF512FFEEFF47F96BFEC9F903" box="[185,308,1743,1764]" pageId="11" pageNumber="676" value="1998-10-21">21 Oct.1998</collectingDate>
</date>
, coll.
<collectorName id="26A2A693F512FFEEFE91F96BFE29F903" box="[367,468,1743,1764]" pageId="11" pageNumber="676">H. K. Lua</collectorName>
, LHK0401 (
<collectionCode id="ED465B80F512FFEEFDB0F96BFD7FF902" box="[590,642,1743,1765]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="676">ZRC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34D90CEF512FFE8FF8FF8A9FE27FD31" lastPageId="13" lastPageNumber="678" pageId="11" pageNumber="676" type="description">
<paragraph id="8BE8C345F512FFEEFF8FF8A9FD1FF823" blockId="11.[113,754,1804,1989]" pageId="11" pageNumber="676">
<emphasis id="B9231F57F512FFEEFF8FF8A9FDFFF8C3" bold="true" box="[113,514,1804,1829]" pageId="11" pageNumber="676">
<emphasis id="B9231F57F512FFEEFF8FF8A9FF0DF8C2" bold="true" box="[113,240,1805,1829]" italics="true" pageId="11" pageNumber="676">Description</emphasis>
. — Macropterous form
</emphasis>
: Of moderate size for genus, general body form ovate, widest across embolium of hemelytra (
<figureCitation id="136CDFC0F512FFEEFF14F8E9FEC5F882" box="[234,312,1869,1893]" captionStart="Fig" captionStartId="12.[836,871,2061,2083]" captionTargetBox="[836,1474,1300,2038]" captionTargetId="figure-494@12.[836,1476,1299,2038]" captionTargetPageId="12" captionText="Fig. 33. Heleocoris malayensis, new species, male, colour photo of dorsal habitus.Specimen from Malaysia, Terengganu, SungaiBrang." figureDoi="http://doi.org/10.5281/zenodo.5352724" httpUri="https://zenodo.org/record/5352724/files/figure.png" pageId="11" pageNumber="676">Fig. 33</figureCitation>
), basic colouration pale yellowish brown, with scutellum, hemelytra and wing membrane dark brown. Male length 11.00 mm; maximum width (across embolia)
<quantity id="4CAF6EA0F512FFEEFD3DF829FF63F823" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="11" pageNumber="676" unit="mm" value="7.4">7.40 mm</quantity>
; female length
<quantity id="4CAF6EA0F512FFEEFEBBF809FE48F823" box="[325,437,1965,1989]" metricMagnitude="-2" metricUnit="m" metricValue="1.09" pageId="11" pageNumber="676" unit="mm" value="10.9">10.90 mm</quantity>
, maximum width
<quantity id="4CAF6EA0F512FFEEFD85F809FD23F823" box="[635,734,1965,1989]" metricMagnitude="-3" metricUnit="m" metricValue="7.35" pageId="11" pageNumber="676" unit="mm" value="7.35">7.35 mm</quantity>
.
</paragraph>
<caption id="DF2893CDF512FFEEFCD2F870FB67F7A7" ID-DOI="http://doi.org/10.5281/zenodo.5352720" ID-Zenodo-Dep="5352720" httpUri="https://zenodo.org/record/5352720/files/figure.png" pageId="11" pageNumber="676" startId="11.[812,856,2004,2026]" targetBox="[893,1369,1020,1970]" targetPageId="11">
<paragraph id="8BE8C345F512FFEEFCD2F870FB67F7A7" blockId="11.[812,1453,2004,2113]" pageId="11" pageNumber="676">
Figs. 29, 30.
<taxonomicName id="4C57B8C6F512FFEEFC4FF872FB96F80D" box="[945,1131,2004,2026]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="676" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9231F57F512FFEEFC4FF872FBE7F80D" box="[945,1050,2006,2026]" italics="true" pageId="11" pageNumber="676">Heleocoris</emphasis>
species
</taxonomicName>
, male proctigers. 29.
<taxonomicName id="4C57B8C6F512FFEEFABDF872FBFCF7E0" authority="Lundblad" authorityName="Lundblad" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="676" phylum="Arthropoda" rank="species" species="montandoni">
<emphasis id="B9231F57F512FFEEFABDF872FC63F7E1" italics="true" pageId="11" pageNumber="676">Heleocoris montandoni</emphasis>
Lundblad
</taxonomicName>
, specimen from Singapore, Nee Soon swamp forest. 30.
<taxonomicName id="4C57B8C6F512FFEEFC63F7ABFB7FF7C4" authority="Polhemus &amp; Polhemus, 2013" authorityName="Polhemus &amp; Polhemus" authorityYear="2013" box="[925,1154,2063,2083]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="676" phylum="Arthropoda" rank="species" species="malayensis" status="sp. nov.">
<emphasis id="B9231F57F512FFEEFC63F7ABFB7FF7C4" box="[925,1154,2063,2083]" italics="true" pageId="11" pageNumber="676">Heleocoris malayensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A210A22CF512FFEEFB72F7ABFAF5F7C3" box="[1164,1288,2062,2084]" pageId="11" pageNumber="676" rank="species">new species</taxonomicNameLabel>
, specimen from Malaysia, Terengganu, Sungai Brang.
</paragraph>
</caption>
<paragraph id="8BE8C345F512FFE9FF8FF849FDBEFED2" blockId="11.[113,753,2029,2117]" lastBlockId="12.[137,776,188,309]" lastPageId="12" lastPageNumber="677" pageId="11" pageNumber="676">
<emphasis id="B9231F57F512FFEEFF8FF849FF53F7E2" bold="true" box="[113,174,2029,2053]" pageId="11" pageNumber="676">Head</emphasis>
pale yellowish brown, with scattered dark brown dots medially and 3 ovate, evenly spaced dark patches along lateral margins of vertex adjacent to each inner eye margin;
<emphasis id="B9231F57F515FFE9FF77FF18FF04FF33" bold="true" box="[137,249,188,212]" pageId="12" pageNumber="677">Scutellum</emphasis>
mottled medium to dark brown, set with numerous tiny pale asperities, extreme apex pale yellow, width/length (midline) = 4.00/1.92, basal margin broadly posteriorly concave, lateral margins broadly sinuate.
</paragraph>
<paragraph id="8BE8C345F515FFE9FF77FEF9FF15FD52" blockId="12.[137,776,349,693]" pageId="12" pageNumber="677">
<emphasis id="B9231F57F515FFE9FF77FEF9FEFFFE92" bold="true" box="[137,258,349,373]" pageId="12" pageNumber="677">Hemelytra</emphasis>
dark blackish-brown set with numerous tiny pale aperities, outer half of embolium translucent golden yellow, inner margin of this pale area forming a straight line angling from inner anterior base of embolium backward and outward to outer posterior terminus; clavus and corium well defined, membrane highly reduced, obscure; tips of hemelytra broadly rounded, extending to apex of genital segment; embolium demarcated by broadly arcuate furrow along inner margin, furrow marking posterior margin obscure, basal half of lateral margin bearing very short, posteriorly angling reddish-brown spinules.
</paragraph>
<paragraph id="8BE8C345F515FFE9FF77FD78FDF4FBD2" blockId="12.[137,776,732,1077]" pageId="12" pageNumber="677">
<emphasis id="B9231F57F515FFE9FF77FD78FF05FD13" bold="true" box="[137,248,732,756]" pageId="12" pageNumber="677">Abdomen</emphasis>
with lateral portions of segments II–IV exposed when viewed dorsally, all visible paratergites pale golden yellow, dark brown at extreme posterolateral angles, these posterolateral angles symmetrical, produced into short, curving spinose projections on paratergites II–V, lateral margins immediately ahead of each projection inwardly notched, these notches each bearing 3 short, stout spines plus acuminate tufts of very long golden setae, remaining lateral margins of paratergites set with very short reddish posteriorly angling brown spinules; lateral margin of left paratergite VI bearing a hooked process (
<figureCitation id="136CDFC0F515FFE9FE55FBB9FE01FBD2" box="[427,508,1053,1077]" captionStart="Figs" captionStartId="13.[812,856,2002,2024]" captionTargetBox="[892,1329,785,1964]" captionTargetId="figure-226@13.[872,1393,776,1974]" captionTargetPageId="13" captionText="Figs. 34, 35. Heleocoris species, male left abdominal paratergites. 34. Heleocoris malayensis, new species, specimen from Malaysia, Terengganu, Sungai Brang. 35. Heleocoris ovatus Montandon, specimen from Vietnam, Lam Dong Prov., Pongour Falls." figureDoi="http://doi.org/10.5281/zenodo.5352726" httpUri="https://zenodo.org/record/5352726/files/figure.png" pageId="12" pageNumber="677">Fig. 34</figureCitation>
).
</paragraph>
<caption id="DF2893CDF515FFE9FCBDFBE7FB84FB2B" ID-DOI="http://doi.org/10.5281/zenodo.5352722" ID-Zenodo-Dep="5352722" httpUri="https://zenodo.org/record/5352722/files/figure.png" pageId="12" pageNumber="677" startId="12.[835,879,1091,1113]" targetBox="[834,1469,233,1050]" targetPageId="12">
<paragraph id="8BE8C345F515FFE9FCBDFBE7FB84FB2B" blockId="12.[835,1474,1091,1228]" pageId="12" pageNumber="677">
Figs. 31, 32.
<taxonomicName id="4C57B8C6F515FFE9FC41FBE0FB8BFBBE" box="[959,1142,1091,1113]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="677" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9231F57F515FFE9FC41FBE0FBD5FBBF" box="[959,1064,1092,1112]" italics="true" pageId="12" pageNumber="677">Heleocoris</emphasis>
species
</taxonomicName>
, female terminal abdomen, ventral view, showing shape of subgenital plate. 31.
<taxonomicName id="4C57B8C6F515FFE9FB1DFBC5FC5FFB74" authority="Lundblad" authorityName="Lundblad" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="677" phylum="Arthropoda" rank="species" species="montandoni">
<emphasis id="B9231F57F515FFE9FB1DFBC5FA3CFB92" box="[1251,1473,1121,1141]" italics="true" pageId="12" pageNumber="677">Heleocoris montandoni</emphasis>
Lundblad
</taxonomicName>
, specimen from Vietnam, Lam Dong Prov., CL 3094. 32.
<taxonomicName id="4C57B8C6F515FFE9FC99FB3FFBBFFB48" authority="Polhemus &amp; Polhemus, 2013" authorityName="Polhemus &amp; Polhemus" authorityYear="2013" box="[871,1090,1179,1199]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="677" phylum="Arthropoda" rank="species" species="malayensis" status="sp. nov.">
<emphasis id="B9231F57F515FFE9FC99FB3FFBBFFB48" box="[871,1090,1179,1199]" italics="true" pageId="12" pageNumber="677">Heleocoris malayensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A210A22CF515FFE9FBB0FB3EFB39FB57" box="[1102,1220,1178,1200]" pageId="12" pageNumber="677" rank="species">new species</taxonomicNameLabel>
, specimen from Malaysia, Perak, Kerunai River, CL 2078.
</paragraph>
</caption>
<paragraph id="8BE8C345F515FFE9FF77FBF9FF1DF992" blockId="12.[137,776,1116,1653]" pageId="12" pageNumber="677">
<emphasis id="B9231F57F515FFE9FF77FBF9FEC7FB93" bold="true" box="[137,314,1116,1141]" pageId="12" pageNumber="677">Ventral surface</emphasis>
pale brown, generally lacking setae, with posterolateral sections of propleuron and central sections of abdominal ventrites I–VII covered with thick recumbent gold hydrofuge pile; propleura not reflexed posteriorly, barely covering basolateral portions of mesosternal plate, anterolateral angles bearing depressed, brown, ovate hydrostatic sense organs present below lateral eye margins; mesosternal plate broadly tumescent centrally, anterior margin of this tumescence transversely folded and raised, entire medial tumescence bearing long, scattered, fine golden setae; metasternal plate elongate, posterior section elongate and triangular with acute posterior apex, longitudinal midline raised into a sharp carina on basal half, this carina becoming widened and rounded on posterior half. Abdominal sternites and parasternites delineated by distinct sutures; all parasternites dull brown, each with 1–3 elongate glabrous patches.
</paragraph>
<paragraph id="8BE8C345F515FFE8FF77F939FD16FE31" blockId="12.[137,776,1693,2101]" lastBlockId="13.[113,753,190,470]" lastPageId="13" lastPageNumber="678" pageId="12" pageNumber="677">
<emphasis id="B9231F57F515FFE9FF77F939FF42F952" bold="true" box="[137,191,1693,1717]" pageId="12" pageNumber="677">Legs</emphasis>
pale yellow; fore coxa unmodified; fore trochanter bearing roughly circular patch of very short, densely packed reddish brown setae centrally; anterior femur uniformly pale yellow, bearing a thick fringe of short, dense gold setae along anterior margin, posterior margin lacking setae; anterior tibia slender, straight, slightly expanded distally, broadly grooved along inner face, ventral margin bearing a distally expanding fringe of short, dense, reddish-brown setae along entire length; anterior tarsi two segmented, bearing two moderately large golden-brown claws, ventral tarsal surfaces bearing thick hair pads similar to that on anterior tibia; middle and hind coxae each bearing a single glabrous tubercle distally; middle and hind femora ventrally with a longitudinal row of tightly packed, elongate, reddish spinules, additional spinules of this same
<typeStatus id="54EC7DE7F514FFE8FF50FF7BFF21FF10" box="[174,220,223,247]" pageId="13" pageNumber="678">type</typeStatus>
present ventrally on basal part of anterior margin, posterior margin of middle femur also bearing thick fringe of short, dense, reddish-golden setae; middle tibiae bearing numerous short, stout brown spines, ventral surface with thick pad of dark golden setae on distal four-fifths, tarsal segments of middle leg bearing similar hair pads ventrally; hind tibiae with scattered moderately long, stout dark brown spines, posterior margin with fringe of long gold swimming hairs.
</paragraph>
<caption id="DF2893CDF515FFE9FCBAF7A9FA3EF7A7" ID-DOI="http://doi.org/10.5281/zenodo.5352724" ID-Zenodo-Dep="5352724" httpUri="https://zenodo.org/record/5352724/files/figure.png" pageId="12" pageNumber="677" startId="12.[836,871,2061,2083]" targetBox="[836,1474,1300,2038]" targetPageId="12">
<paragraph id="8BE8C345F515FFE9FCBAF7A9FA3EF7A7" blockId="12.[836,1475,2061,2112]" pageId="12" pageNumber="677">
Fig. 33.
<taxonomicName id="4C57B8C6F515FFE9FC6DF7AAFB91F7C5" authority="Polhemus &amp; Polhemus, 2013" authorityName="Polhemus &amp; Polhemus" authorityYear="2013" box="[915,1132,2062,2082]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="677" phylum="Arthropoda" rank="species" species="malayensis" status="sp. nov.">
<emphasis id="B9231F57F515FFE9FC6DF7AAFB91F7C5" box="[915,1132,2062,2082]" italics="true" pageId="12" pageNumber="677">Heleocoris malayensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A210A22CF515FFE9FB89F7AAFB10F7C4" box="[1143,1261,2061,2083]" pageId="12" pageNumber="677" rank="species">new species</taxonomicNameLabel>
, male, colour photo of dorsal habitus. Specimen from Malaysia, Terengganu, Sungai Brang.
</paragraph>
</caption>
<paragraph id="8BE8C345F514FFE8FF8FFE5AFD81FD71" blockId="13.[113,753,510,662]" pageId="13" pageNumber="678">
<emphasis id="B9231F57F514FFE8FF8FFE5AFEE7FDF1" bold="true" box="[113,282,510,534]" pageId="13" pageNumber="678">Male genitalia</emphasis>
with proctiger elongate, apex acute (
<figureCitation id="136CDFC0F514FFE8FD3BFE5AFF6DFDD1" captionStart="Figs" captionStartId="11.[812,856,2004,2026]" captionTargetBox="[893,1369,1020,1970]" captionTargetId="figure-13@11.[880,1386,1000,1976]" captionTargetPageId="11" captionText="Figs. 29, 30. Heleocoris species, male proctigers. 29. Heleocoris montandoni Lundblad, specimen from Singapore, Nee Soon swamp forest. 30. Heleocoris malayensis new species, specimen from Malaysia, Terengganu, Sungai Brang." figureDoi="http://doi.org/10.5281/zenodo.5352720" httpUri="https://zenodo.org/record/5352720/files/figure.png" pageId="13" pageNumber="678">Fig. 30</figureCitation>
); parameres highly reduced, elongate ovate (
<figureCitation id="136CDFC0F514FFE8FD70FDBAFD19FDD1" box="[654,740,542,566]" captionStart="Figs" captionStartId="10.[136,180,1861,1883]" captionTargetBox="[139,775,714,1834]" captionTargetId="figure-8@10.[117,790,706,1848]" captionTargetPageId="10" captionText="Figs. 24–28. Heleocoris species, male phallotheca. 24. Heleocoris malayensis, new species, specimen from Malaysia, Terengganu, Sungai Brang. 25. Heleocoris montandoni Lundblad, specimen from Singapore, Nee Soon swamp forest. 26. Heleocoris nebulosus Montandon, specimen from Indonesia, Bali, Melangit River. 27. Heleocoris ovatus Montandon, specimen from Vietnam, Lam Dong Prov., Pongour Falls, CL 3091. 28. Heleocoris strabus Montandon, specimen from Thailand, Chiang Mai Prov., Huay Hia Creek, CL 2198." figureDoi="http://doi.org/10.5281/zenodo.5352718" httpUri="https://zenodo.org/record/5352718/files/figure.png" pageId="13" pageNumber="678">Fig. 24</figureCitation>
); male phallotheca asymmetrical, with apex expanded into a rounded lobe (
<figureCitation id="136CDFC0F514FFE8FEEDFDFAFE95FD91" box="[275,360,606,630]" captionStart="Figs" captionStartId="10.[136,180,1861,1883]" captionTargetBox="[139,775,714,1834]" captionTargetId="figure-8@10.[117,790,706,1848]" captionTargetPageId="10" captionText="Figs. 24–28. Heleocoris species, male phallotheca. 24. Heleocoris malayensis, new species, specimen from Malaysia, Terengganu, Sungai Brang. 25. Heleocoris montandoni Lundblad, specimen from Singapore, Nee Soon swamp forest. 26. Heleocoris nebulosus Montandon, specimen from Indonesia, Bali, Melangit River. 27. Heleocoris ovatus Montandon, specimen from Vietnam, Lam Dong Prov., Pongour Falls, CL 3091. 28. Heleocoris strabus Montandon, specimen from Thailand, Chiang Mai Prov., Huay Hia Creek, CL 2198." figureDoi="http://doi.org/10.5281/zenodo.5352718" httpUri="https://zenodo.org/record/5352718/files/figure.png" pageId="13" pageNumber="678">Fig. 24</figureCitation>
).
<emphasis id="B9231F57F514FFE8FE80FDFAFD69FD91" bold="true" box="[382,660,606,630]" pageId="13" pageNumber="678">Female subgenital plate</emphasis>
roughly trapezoidal, posterior broadly concave (
<figureCitation id="136CDFC0F514FFE8FDE0FDDAFD92FD71" box="[542,623,638,662]" captionStart="Figs" captionStartId="12.[835,879,1091,1113]" captionTargetBox="[834,1469,233,1050]" captionTargetId="figure-8@12.[819,1486,216,1072]" captionTargetPageId="12" captionText="Figs. 31, 32. Heleocoris species, female terminal abdomen, ventral view, showing shape of subgenital plate. 31. Heleocoris montandoni Lundblad, specimen from Vietnam, Lam Dong Prov., CL 3094. 32. Heleocoris malayensis, new species, specimen from Malaysia, Perak, Kerunai River, CL 2078." figureDoi="http://doi.org/10.5281/zenodo.5352722" httpUri="https://zenodo.org/record/5352722/files/figure.png" pageId="13" pageNumber="678">Fig. 32</figureCitation>
).
</paragraph>
<paragraph id="8BE8C345F514FFE8FF8FFD1AFE27FD31" blockId="13.[113,474,702,726]" box="[113,474,702,726]" pageId="13" pageNumber="678">
<emphasis id="B9231F57F514FFE8FF8FFD1AFEA7FD31" bold="true" box="[113,346,702,726]" pageId="13" pageNumber="678">Brachypterous form</emphasis>
: Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C34D90CEF514FFE8FF8FFD59FB18FDC9" pageId="13" pageNumber="678" type="distribution">
<paragraph id="8BE8C345F514FFE8FF8FFD59FF35FCD1" blockId="13.[113,753,765,822]" pageId="13" pageNumber="678">
<emphasis id="B9231F57F514FFE8FF8FFD59FEFDFCF1" bold="true" box="[113,256,765,790]" pageId="13" pageNumber="678">
<emphasis id="B9231F57F514FFE8FF8FFD59FF04FCF1" bold="true" box="[113,249,765,790]" italics="true" pageId="13" pageNumber="678">Distribution</emphasis>
.
</emphasis>
— Peninsular
<collectingCountry id="F34083D5F514FFE8FE52FD5AFDF2FCF1" box="[428,527,766,790]" name="Malaysia" pageId="13" pageNumber="678">Malaysia</collectingCountry>
(below the Isthmus of Kra).
</paragraph>
<paragraph id="8BE8C345F514FFE8FCD2FF1BFB18FDC9" blockId="13.[812,1452,190,558]" pageId="13" pageNumber="678">
<materialsCitation id="3B3FC918F514FFE8FCD2FF1BFBA8FF17" collectingDate="2001-03-14" collectionCode="T" collectorName="Hwy." latitude="13.964722" longLatPrecision="21" longitude="108.763336" pageId="13" pageNumber="678" specimenCount="365">
<collectorName id="26A2A693F514FFE8FCD2FF1BFC9DFF33" box="[812,864,191,212]" pageId="13" pageNumber="678">Hwy.</collectorName>
<quantity id="4CAF6EA0F514FFE8FC9BFF1AFC36FF33" box="[869,971,190,212]" metricMagnitude="4" metricUnit="m" metricValue="1.9365" pageId="13" pageNumber="678" unit="m" value="19365.0">
19,
<specimenCount id="9D5108CCF514FFE8FC74FF1AFC36FF33" box="[906,971,190,212]" pageId="13" pageNumber="678" type="generic">365 m</specimenCount>
</quantity>
.,
<geoCoordinate id="EE63A582F514FFE8FC27FF1AFBB1FF33" box="[985,1100,190,212]" degrees="13" direction="north" minutes="57" orientation="latitude" pageId="13" pageNumber="678" precision="15" seconds="53" value="13.964722">13°57'53&quot;N</geoCoordinate>
,
<geoCoordinate id="EE63A582F514FFE8FBA8FF1AFB2EFF33" box="[1110,1235,190,212]" degrees="108" direction="east" minutes="45" orientation="longitude" pageId="13" pageNumber="678" precision="15" seconds="48" value="108.763336">108°45'48&quot;E</geoCoordinate>
, water temp. 22°C,
<date id="FFE9E585F514FFE8FA6DFF1AFC76FF17" pageId="13" pageNumber="678" value="2001-03-14">
<collectingDate id="EFAD1C6DF514FFE8FA6DFF1AFC76FF17" pageId="13" pageNumber="678" value="2001-03-14">14 Mar.2001</collectingDate>
</date>
, CL 4293, coll. J.
<collectionCode id="ED465B80F514FFE8FBBBFF7FFBA8FF17" box="[1093,1109,219,240]" name="Tavera, Department of Geology and Geophysics" pageId="13" pageNumber="678">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B3FC918F514FFE8FB9EFF7FFB5FFEA0" collectingDate="1985-11-16" collectionCode="USNM, T" collectorName="Polhemus &amp; P. Nguyen &amp; Mae Mao River, S." country="Thailand" elevation="500" location="Fang Horticultural Station" municipality="Fang Dist." pageId="13" pageNumber="678" specimenCount="21" specimenCount-female="10" specimenCount-male="11" stateProvince="Chiang Mai">
<collectorName id="26A2A693F514FFE8FB9EFF7FFB42FF16" box="[1120,1215,219,241]" pageId="13" pageNumber="678">Polhemus</collectorName>
&amp;
<collectorName id="26A2A693F514FFE8FB1CFF7FFAB5FF17" box="[1250,1352,219,240]" pageId="13" pageNumber="678">P. Nguyen</collectorName>
(
<collectionCode id="ED465B80F514FFE8FAA9FF7FFA5FFF16" box="[1367,1442,219,241]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="678" type="Museum">USNM</collectionCode>
).
<collectingCountry id="F34083D5F514FFE8FCD2FF5CFC53FEEA" box="[812,942,248,269]" name="Thailand" pageId="13" pageNumber="678">THAILAND</collectingCountry>
,
<emphasis id="B9231F57F514FFE8FC45FF5CFB80FEEA" bold="true" box="[955,1149,248,270]" pageId="13" pageNumber="678">
<collectingRegion id="49930DA7F514FFE8FC45FF5CFB80FEEA" box="[955,1149,248,270]" country="Thailand" name="Chiang Mai" pageId="13" pageNumber="678">Chiang Mai Prov.</collectingRegion>
</emphasis>
:
<specimenCount id="9D5108CCF514FFE8FB70FF5CFB11FEE9" box="[1166,1260,248,270]" pageId="13" pageNumber="678" type="male">11 males</specimenCount>
,
<specimenCount id="9D5108CCF514FFE8FB04FF5CFA91FEE9" box="[1274,1388,248,270]" pageId="13" pageNumber="678" type="female">10 females</specimenCount>
,
<collectingMunicipality id="6B8C593FF514FFE8FA84FF5CFCA0FECC" pageId="13" pageNumber="678">Fang Dist.</collectingMunicipality>
,
<collectorName id="26A2A693F514FFE8FC98FEB1FBE2FECD" box="[870,1055,277,299]" pageId="13" pageNumber="678">Mae Mao River, S.</collectorName>
of
<location id="8E88959EF514FFE8FBBEFEB1FAC2FECC" LSID="urn:lsid:plazi:treatment:03FE7253F512FFE8FF12FB78FDC3FAB1:8E88959EF514FFE8FBBEFEB1FAC2FECC" box="[1088,1343,277,299]" country="Thailand" municipality="Fang Dist." name="Fang Horticultural Station" pageId="13" pageNumber="678" stateProvince="Chiang Mai">Fang Horticultural Station</location>
,
<quantity id="4CAF6EA0F514FFE8FAB7FEB0FA74FECD" box="[1353,1417,276,298]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="678" unit="m" value="500.0">
<elevation id="007A2476F514FFE8FAB7FEB0FA74FECD" box="[1353,1417,276,298]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="678" unit="m" value="500.0">500 m</elevation>
</quantity>
,
<date id="FFE9E585F514FFE8FA6DFEB1FC71FEA0" pageId="13" pageNumber="678" value="1985-11-16">
<collectingDate id="EFAD1C6DF514FFE8FA6DFEB1FC71FEA0" pageId="13" pageNumber="678" value="1985-11-16">16 Nov.1985</collectingDate>
</date>
, CL 2200, coll. D. A. &amp; J.
<collectionCode id="ED465B80F514FFE8FB6CFE96FB5FFEA0" box="[1170,1186,306,327]" name="Tavera, Department of Geology and Geophysics" pageId="13" pageNumber="678">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B3FC918F514FFE8FB52FE95FC30FE67" collectingDate="1985-11-19" collectionCode="JTPC, T" collectorName="Polhemus" country="Thailand" location="10 km NW of Mae Rim" pageId="13" pageNumber="678" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Chiang Mai">
<collectorName id="26A2A693F514FFE8FB52FE95FAF6FEA0" box="[1196,1291,305,327]" pageId="13" pageNumber="678">Polhemus</collectorName>
(
<collectionCode id="ED465B80F514FFE8FAE6FE96FAAEFEAF" box="[1304,1363,306,328]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33646" lsid="urn:lsid:biocol.org:col:33646" name="Colorado Entomological Museum (formerly John T. Polhemus collection)" pageId="13" pageNumber="678">JTPC</collectionCode>
);
<specimenCount id="9D5108CCF514FFE8FA9AFE96FA55FEA0" box="[1380,1448,305,327]" pageId="13" pageNumber="678" type="male">1 male</specimenCount>
,
<specimenCount id="9D5108CCF514FFE8FCD2FEEAFC7FFE83" box="[812,898,334,356]" pageId="13" pageNumber="678" type="female">1 female</specimenCount>
, stream
<location id="8E88959EF514FFE8FC2DFEEAFB43FE83" LSID="urn:lsid:plazi:treatment:03FE7253F512FFE8FF12FB78FDC3FAB1:8E88959EF514FFE8FC2DFEEAFB43FE83" box="[979,1214,334,356]" country="Thailand" name="10 km NW of Mae Rim" pageId="13" pageNumber="678" stateProvince="Chiang Mai">
<quantity id="4CAF6EA0F514FFE8FC2DFEEAFBECFE83" box="[979,1041,334,356]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="678" unit="km" value="10.0">10 km</quantity>
NW of Mae Rim
</location>
,
<date id="FFE9E585F514FFE8FB36FEEAFABAFE83" box="[1224,1351,334,356]" pageId="13" pageNumber="678" value="1985-11-19">
<collectingDate id="EFAD1C6DF514FFE8FB36FEEAFABAFE83" box="[1224,1351,334,356]" pageId="13" pageNumber="678" value="1985-11-19">19 Nov.1985</collectingDate>
</date>
, CL 2204, coll. D.A. &amp; J.
<collectionCode id="ED465B80F514FFE8FC43FECFFC30FE67" box="[957,973,363,384]" name="Tavera, Department of Geology and Geophysics" pageId="13" pageNumber="678">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B3FC918F514FFE8FC2AFECFFB6AFE13" collectingDate="1998-10-24" collectionCode="USNM, T" collectorName="Polhemus &amp; Division" country="Myanmar" county="River" elevation="520" latitude="20.729933" location="Thazi" longLatPrecision="1" longitude="96.49128" municipality="Kalaw" pageId="13" pageNumber="678" specimenCount="21" specimenCount-female="11" specimenCount-male="10" stateProvince="Shan">
<collectorName id="26A2A693F514FFE8FC2AFECFFBCEFE66" box="[980,1075,363,385]" pageId="13" pageNumber="678">Polhemus</collectorName>
(
<collectionCode id="ED465B80F514FFE8FBC1FECFFB77FE66" box="[1087,1162,363,385]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="678" type="Museum">USNM</collectionCode>
).
<collectingCountry id="F34083D5F514FFE8FB66FECFFB0EFE67" box="[1176,1267,363,384]" name="Myanmar" pageId="13" pageNumber="678">BURMA</collectingCountry>
,
<emphasis id="B9231F57F514FFE8FB05FECEFA75FE66" bold="true" box="[1275,1416,362,385]" pageId="13" pageNumber="678">
<collectingRegion id="49930DA7F514FFE8FB05FECEFAD2FE67" box="[1275,1327,362,384]" country="Myanmar" name="Shan" pageId="13" pageNumber="678">Shan</collectingRegion>
<collectorName id="26A2A693F514FFE8FACAFECFFA75FE66" box="[1332,1416,363,385]" pageId="13" pageNumber="678">Division</collectorName>
</emphasis>
:
<specimenCount id="9D5108CCF514FFE8FA6DFECFFC9BFE79" pageId="13" pageNumber="678" type="male">10 males</specimenCount>
,
<specimenCount id="9D5108CCF514FFE8FC8EFE2CFC27FE79" box="[880,986,392,414]" pageId="13" pageNumber="678" type="female">11 females</specimenCount>
,
<collectingRegion id="49930DA7F514FFE8FC1AFE2CFBD1FE7A" box="[996,1068,392,413]" country="Myanmar" name="Magway" pageId="13" pageNumber="678">Magwe</collectingRegion>
<collectingCounty id="6289BBC9F514FFE8FBCCFE2CFB9AFE79" box="[1074,1127,392,414]" pageId="13" pageNumber="678">River</collectingCounty>
at confluence with clear tributary in sandstone bed,
<locationDeviation id="9A5ADA67F514FFE8FC26FE01FB5CFE5C" box="[984,1185,421,443]" location="Kalaw" pageId="13" pageNumber="678">
<quantity id="4CAF6EA0F514FFE8FC26FE01FBE9FE5C" box="[984,1044,421,443]" metricMagnitude="4" metricUnit="m" metricValue="2.6" pageId="13" pageNumber="678" unit="km" value="26.0">26 km</quantity>
NW of Kalaw
</locationDeviation>
along
<collectingMunicipality id="6B8C593FF514FFE8FB1FFE01FADDFE5C" box="[1249,1312,421,443]" pageId="13" pageNumber="678">Kalaw</collectingMunicipality>
to
<location id="8E88959EF514FFE8FAC0FE01FA88FE5C" LSID="urn:lsid:plazi:treatment:03FE7253F512FFE8FF12FB78FDC3FAB1:8E88959EF514FFE8FAC0FE01FA88FE5C" box="[1342,1397,421,443]" country="Myanmar" county="River" latitude="20.729933" longLatPrecision="1" longitude="96.49128" municipality="Kalaw" name="Thazi" pageId="13" pageNumber="678" stateProvince="Shan">Thazi</location>
road,
<quantity id="4CAF6EA0F514FFE8FCD2FE65FC94FE30" box="[812,873,449,471]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="13" pageNumber="678" unit="m" value="520.0">
<elevation id="007A2476F514FFE8FCD2FE65FC94FE30" box="[812,873,449,471]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="13" pageNumber="678" unit="m" value="520.0">520 m</elevation>
</quantity>
,
<geoCoordinate id="EE63A582F514FFE8FC8FFE65FC17FE30" box="[881,1002,449,471]" degrees="20" direction="north" minutes="43.796" orientation="latitude" pageId="13" pageNumber="678" precision="1" value="20.729933">20°43.796'N</geoCoordinate>
,
<geoCoordinate id="EE63A582F514FFE8FC0CFE65FB94FE30" box="[1010,1129,449,471]" degrees="96" direction="east" minutes="29.477" orientation="longitude" pageId="13" pageNumber="678" precision="1" value="96.49128">96°29.477'E</geoCoordinate>
, water temp. 29°C (main river),
<date id="FFE9E585F514FFE8FA6AFE66FC7AFE14" pageId="13" pageNumber="678" value="1998-10-24">
<collectingDate id="EFAD1C6DF514FFE8FA6AFE66FC7AFE14" pageId="13" pageNumber="678" value="1998-10-24">24 Oct.1998</collectingDate>
</date>
, CL 4023, coll. D. A. &amp; J.
<collectionCode id="ED465B80F514FFE8FB79FE7BFB6AFE13" box="[1159,1175,479,500]" name="Tavera, Department of Geology and Geophysics" pageId="13" pageNumber="678">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B3FC918F514FFE8FB5EFE7AFB1CFDC9" collectionCode="USNM, JTPC" collectorName="Polhemus &amp; D. Dudgeon" country="Hong Kong" location="Lam Tsuen River" pageId="13" pageNumber="678" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
<collectorName id="26A2A693F514FFE8FB5EFE7AFB02FE13" box="[1184,1279,478,500]" pageId="13" pageNumber="678">Polhemus</collectorName>
(
<collectionCode id="ED465B80F514FFE8FAF5FE7AFAABFE13" box="[1291,1366,478,500]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="678" type="Museum">USNM</collectionCode>
).
<collectingCountry id="F34083D5F514FFE8FA98FE7AFC88FDF7" name="Hong Kong" pageId="13" pageNumber="678">HONG KONG</collectingCountry>
:
<specimenCount id="9D5108CCF514FFE8FC81FE5FFC39FDF6" box="[895,964,507,529]" pageId="13" pageNumber="678" type="male">1 male</specimenCount>
,
<specimenCount id="9D5108CCF514FFE8FC2EFE5FFBCDFDF6" box="[976,1072,507,529]" pageId="13" pageNumber="678" type="female">2 females</specimenCount>
,
<location id="8E88959EF514FFE8FBC2FE5FFB1AFDF6" LSID="urn:lsid:plazi:treatment:03FE7253F512FFE8FF12FB78FDC3FAB1:8E88959EF514FFE8FBC2FE5FFB1AFDF6" box="[1084,1255,507,529]" country="Hong Kong" name="Lam Tsuen River" pageId="13" pageNumber="678">Lam Tsuen River</location>
, lower course, from trailing vegetation, coll.
<collectorName id="26A2A693F514FFE8FBE5FDBCFB6CFDC9" box="[1051,1169,536,558]" pageId="13" pageNumber="678">D. Dudgeon</collectorName>
(
<collectionCode id="ED465B80F514FFE8FB61FDBCFB27FDC9" box="[1183,1242,536,558]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33646" lsid="urn:lsid:biocol.org:col:33646" name="Colorado Entomological Museum (formerly John T. Polhemus collection)" pageId="13" pageNumber="678">JTPC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34D90CEF514FFE8FCD2FDF2FDC3FAB1" pageId="13" pageNumber="678" type="diagnosis">
<paragraph id="8BE8C345F514FFE8FCD2FDF2FD41FB91" blockId="13.[812,1452,598,718]" lastBlockId="13.[113,753,862,1142]" pageId="13" pageNumber="678">
<emphasis id="B9231F57F514FFE8FCD2FDF2FC55FD89" bold="true" box="[812,936,598,622]" pageId="13" pageNumber="678">
<emphasis id="B9231F57F514FFE8FCD2FDF2FC5DFD89" bold="true" box="[812,928,598,622]" italics="true" pageId="13" pageNumber="678">Diagnosis</emphasis>
.
</emphasis>
— Length 10.2–11.0 mm, maximum width (across abdomen) 7.0–
<quantity id="4CAF6EA0F514FFE8FBD7FDD2FB7FFD6A" box="[1065,1154,630,654]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="13" pageNumber="678" unit="mm" value="7.4">7.4 mm</quantity>
, general body form ovate. Head and pronotum dull medium brown finely maculated with dark brown or black; scutellum blackish brown, extreme
<emphasis id="B9231F57F514FFE8FF8FFCFAFF0EFC91" bold="true" box="[113,243,862,886]" pageId="13" pageNumber="678">
<emphasis id="B9231F57F514FFE8FF8FFCFAFF11FC91" bold="true" box="[113,236,862,886]" italics="true" pageId="13" pageNumber="678">Discussion</emphasis>
.
</emphasis>
— The larger of the two
<taxonomicName id="4C57B8C6F514FFE8FDDEFCFBFD0DFC91" box="[544,752,862,886]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="678" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9231F57F514FFE8FDDEFCFBFD65FC91" box="[544,664,863,886]" italics="true" pageId="13" pageNumber="678">Heleocoris</emphasis>
species
</taxonomicName>
currently known to occur in Peninsular
<collectingCountry id="F34083D5F514FFE8FDA6FCDAFD38FC71" box="[600,709,894,918]" name="Malaysia" pageId="13" pageNumber="678">Malaysia</collectingCountry>
,
<taxonomicName id="4C57B8C6F514FFE8FD28FCDBFF15FC51" authorityName="Polhemus &amp; Polhemus" authorityYear="2013" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="678" phylum="Arthropoda" rank="species" species="malayensis">
<emphasis id="B9231F57F514FFE8FD28FCDBFD14FC71" box="[726,745,895,918]" italics="true" pageId="13" pageNumber="678">H</emphasis>
.
<emphasis id="B9231F57F514FFE8FF8FFC3BFF15FC51" box="[113,232,927,950]" italics="true" pageId="13" pageNumber="678">malayensis</emphasis>
</taxonomicName>
may be recognised by its body size in excess of
<quantity id="4CAF6EA0F514FFE8FF8FFC1AFF3DFC31" box="[113,192,958,982]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="13" pageNumber="678" unit="mm" value="11.0">11 mm</quantity>
; the generally dark hemelytra with the anterolateral half of the embolium broadly pale yellow (
<figureCitation id="136CDFC0F514FFE8FD9CFC7AFD4AFC11" box="[610,695,990,1014]" captionStart="Fig" captionStartId="12.[836,871,2061,2083]" captionTargetBox="[836,1474,1300,2038]" captionTargetId="figure-494@12.[836,1476,1299,2038]" captionTargetPageId="12" captionText="Fig. 33. Heleocoris malayensis, new species, male, colour photo of dorsal habitus.Specimen from Malaysia, Terengganu, SungaiBrang." figureDoi="http://doi.org/10.5281/zenodo.5352724" httpUri="https://zenodo.org/record/5352724/files/figure.png" pageId="13" pageNumber="678">Fig. 33</figureCitation>
); the hooked process on the lateral margin of male abdominal left paratergite VI (
<figureCitation id="136CDFC0F514FFE8FEE7FBBAFE96FBD1" box="[281,363,1054,1078]" captionStart="Figs" captionStartId="13.[812,856,2002,2024]" captionTargetBox="[892,1329,785,1964]" captionTargetId="figure-226@13.[872,1393,776,1974]" captionTargetPageId="13" captionText="Figs. 34, 35. Heleocoris species, male left abdominal paratergites. 34. Heleocoris malayensis, new species, specimen from Malaysia, Terengganu, Sungai Brang. 35. Heleocoris ovatus Montandon, specimen from Vietnam, Lam Dong Prov., Pongour Falls." figureDoi="http://doi.org/10.5281/zenodo.5352726" httpUri="https://zenodo.org/record/5352726/files/figure.png" pageId="13" pageNumber="678">Fig. 34</figureCitation>
), the shape of the male phallotheca (
<figureCitation id="136CDFC0F514FFE8FF87FB9AFF34FBB1" box="[121,201,1086,1110]" captionStart="Figs" captionStartId="10.[136,180,1861,1883]" captionTargetBox="[139,775,714,1834]" captionTargetId="figure-8@10.[117,790,706,1848]" captionTargetPageId="10" captionText="Figs. 24–28. Heleocoris species, male phallotheca. 24. Heleocoris malayensis, new species, specimen from Malaysia, Terengganu, Sungai Brang. 25. Heleocoris montandoni Lundblad, specimen from Singapore, Nee Soon swamp forest. 26. Heleocoris nebulosus Montandon, specimen from Indonesia, Bali, Melangit River. 27. Heleocoris ovatus Montandon, specimen from Vietnam, Lam Dong Prov., Pongour Falls, CL 3091. 28. Heleocoris strabus Montandon, specimen from Thailand, Chiang Mai Prov., Huay Hia Creek, CL 2198." figureDoi="http://doi.org/10.5281/zenodo.5352718" httpUri="https://zenodo.org/record/5352718/files/figure.png" pageId="13" pageNumber="678">Fig. 24</figureCitation>
); the elongate shape of the male proctiger (
<figureCitation id="136CDFC0F514FFE8FD6BFB9AFD18FBB1" box="[661,741,1086,1110]" captionStart="Figs" captionStartId="11.[812,856,2004,2026]" captionTargetBox="[893,1369,1020,1970]" captionTargetId="figure-13@11.[880,1386,1000,1976]" captionTargetPageId="11" captionText="Figs. 29, 30. Heleocoris species, male proctigers. 29. Heleocoris montandoni Lundblad, specimen from Singapore, Nee Soon swamp forest. 30. Heleocoris malayensis new species, specimen from Malaysia, Terengganu, Sungai Brang." figureDoi="http://doi.org/10.5281/zenodo.5352720" httpUri="https://zenodo.org/record/5352720/files/figure.png" pageId="13" pageNumber="678">Fig. 30</figureCitation>
), and the shape of the female subgenital plate (
<figureCitation id="136CDFC0F514FFE8FDA0FBFAFD52FB91" box="[606,687,1118,1142]" captionStart="Figs" captionStartId="12.[835,879,1091,1113]" captionTargetBox="[834,1469,233,1050]" captionTargetId="figure-8@12.[819,1486,216,1072]" captionTargetPageId="12" captionText="Figs. 31, 32. Heleocoris species, female terminal abdomen, ventral view, showing shape of subgenital plate. 31. Heleocoris montandoni Lundblad, specimen from Vietnam, Lam Dong Prov., CL 3094. 32. Heleocoris malayensis, new species, specimen from Malaysia, Perak, Kerunai River, CL 2078." figureDoi="http://doi.org/10.5281/zenodo.5352722" httpUri="https://zenodo.org/record/5352722/files/figure.png" pageId="13" pageNumber="678">Fig. 32</figureCitation>
).
</paragraph>
<paragraph id="8BE8C345F514FFE8FF8FFB3AFDC3FAB1" blockId="13.[113,753,1182,1366]" pageId="13" pageNumber="678">
Previous records of
<taxonomicName id="4C57B8C6F514FFE8FEBFFB3BFD6EFB51" authority="Montandon" authorityName="Montandon" authorityYear="1897" box="[321,659,1182,1206]" class="Insecta" family="Naucoridae" genus="Laccocoris" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="678" phylum="Arthropoda" rank="species" species="nervicus">
<emphasis id="B9231F57F514FFE8FEBFFB3BFDE9FB51" box="[321,532,1183,1206]" italics="true" pageId="13" pageNumber="678">Laccocoris nervicus</emphasis>
Montandon
</taxonomicName>
from the Peninsular Malaysian states of
<collectingRegion id="49930DA7F514FFE8FE4AFB1AFE0DFB31" box="[436,496,1214,1238]" country="Malaysia" name="Perak" pageId="13" pageNumber="678">Perak</collectingRegion>
and
<collectingRegion id="49930DA7F514FFE8FDDFFB1AFD82FB31" box="[545,639,1214,1238]" country="Malaysia" name="Selangor" pageId="13" pageNumber="678">Selangor</collectingRegion>
(Fernando &amp; Cheng, 1974) are referable to this species, whereas
<taxonomicName id="4C57B8C6F514FFE8FD25FB7BFF36FAF1" authorityName="Montandon" authorityYear="1897" class="Insecta" family="Naucoridae" genus="Laccocoris" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="678" phylum="Arthropoda" rank="species" species="nervicus">
<emphasis id="B9231F57F514FFE8FD25FB7BFF36FAF1" italics="true" pageId="13" pageNumber="678">L. nervicus</emphasis>
</taxonomicName>
is endemic to the mountains of Sumatra. Previous records of
<taxonomicName id="4C57B8C6F514FFE8FF19FABBFE2FFAD1" authority="Montandon" authorityName="Montandon" authorityYear="1897" box="[231,466,1310,1334]" class="Insecta" family="Naucoridae" genus="Heleocoris" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="678" phylum="Arthropoda" rank="species" species="ovatus">
<emphasis id="B9231F57F514FFE8FF19FABBFEB2FAD1" box="[231,335,1311,1334]" italics="true" pageId="13" pageNumber="678">H. ovatus</emphasis>
Montandon
</taxonomicName>
from Peninsular
<collectingCountry id="F34083D5F514FFE8FD71FABAFD0CFAD1" box="[655,753,1310,1334]" name="Malaysia" pageId="13" pageNumber="678">Malaysia</collectingCountry>
are also represent this species (see below).
</paragraph>
</subSubSection>
</treatment>
</document>
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