585 lines
70 KiB
XML
585 lines
70 KiB
XML
<document id="48DAEF0484986FF11FC41276E77090A2" ID="10.11646/zootaxa.4173.3.2" ID-CLB-Dataset="38076" ID-DOI="10.11646/zootaxa.4173.3.2" ID-GBIF-Dataset="b5d9a5ac-47cd-4159-aede-94c05cfcfc50" ID-ISSN="1175-5326" ID-Zenodo-Dep="159361" ID-ZooBank="0E56B22B-B86E-4750-8904-00FD1B2F49B3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1475647930173" checkinUser="plazi" docAuthor="Freyhof, Jörg, Abdullah, Younis Sabir, Ararat, Korsh, Ibrahim, Hamad & Geiger, Matthias F." docDate="2016" docId="03E587B34F37721B21BFFE5A6CABF978" docLanguage="en" docName="zootaxa.4173.3.2.pdf" docOrigin="Zootaxa 4173 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eidinemacheilus proudlovei Freyhof, Abdullah, Ararat, Ibrahim & Geiger, 2016, new species" docType="treatment" docVersion="9" lastPageNumber="234" masterDocId="FFDCFFCB4F3472122128FFED6E2DFFDE" masterDocTitle="Eidinemacheilus proudlovei, a new subterranean loach from Iraqi Kurdistan (Teleostei; Nemacheilidae)" masterLastPageNumber="236" masterPageNumber="225" pageNumber="228" updateTime="1698429163261" updateUser="ExternalLinkService">
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<mods:titleInfo id="D03D80975298B5766DB0134DF7E408D1">
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<mods:title id="F257003CC4B10C6EAC8CC59E748B29D0">Eidinemacheilus proudlovei, a new subterranean loach from Iraqi Kurdistan (Teleostei; Nemacheilidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="182F4CE3CB9C138E1B965F37971A1F68" type="personal">
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<mods:namePart id="4188EA9D3E8F742C02E072B2BC05BD7F">Freyhof, Jörg</mods:namePart>
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<mods:roleTerm id="2714A81070A34357C511E69C1DB33B39">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C39DAA08E38ED08E31308F2D9EE5158A">Abdullah, Younis Sabir</mods:namePart>
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</mods:name>
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<mods:roleTerm id="BFEEC90E095C39BA84D6DAC2CDE1FD42">Author</mods:roleTerm>
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<mods:namePart id="97E4B0EC5B3F41FF41674D0EA4980809">Ararat, Korsh</mods:namePart>
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<mods:roleTerm id="D093DB47A19232153E7861B69E8673CF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7C2C76B3E7F10B60914616617A1A74FC">Ibrahim, Hamad</mods:namePart>
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</mods:name>
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<mods:roleTerm id="8A90F9EEFE85ACFA03232BFEA9434E4A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="008299ECE1F98C8947BA609197DE26ED">Geiger, Matthias F.</mods:namePart>
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<mods:title id="1FE903809FFBA5CC7F3A90BD24BFAC93">Zootaxa</mods:title>
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<mods:date id="916140BD7F28D0EF78CBA88B7B17A791">2016</mods:date>
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<mods:detail id="32D7BD8DBCC034A7193B57CCF8F3CA2B" type="volume">
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<mods:number id="BE148AE44C360477A51C710595E3AB71">4173</mods:number>
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<treatment id="03E587B34F37721B21BFFE5A6CABF978" ID-DOI="http://doi.org/10.5281/zenodo.5693941" ID-GBIF-Taxon="124557000" ID-Zenodo-Dep="5693941" LSID="urn:lsid:plazi:treatment:03E587B34F37721B21BFFE5A6CABF978" httpUri="http://treatment.plazi.org/id/03E587B34F37721B21BFFE5A6CABF978" lastPageId="9" lastPageNumber="234" pageId="3" pageNumber="228">
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<subSubSection id="C356652E4F37721121BFFE5A6F3FFE2D" pageId="3" pageNumber="228" type="nomenclature">
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<paragraph id="8BF336A54F37721121BFFE5A6CA4FE0C" blockId="3.[151,649,439,499]" box="[151,649,439,466]" pageId="3" pageNumber="228">
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<heading id="D0BB81C94F37721121BFFE5A6CA4FE0C" bold="true" box="[151,649,439,466]" fontSize="11" level="1" pageId="3" pageNumber="228" reason="1">
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<emphasis id="B938EAB74F37721121BFFE5A6CA4FE0C" bold="true" box="[151,649,439,466]" pageId="3" pageNumber="228">
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<taxonomicName id="4C4C4D264F37721121BFFE5A6FC7FE0F" ID-CoL="6DYJL" box="[151,490,439,465]" class="Actinopterygii" family="Nemacheilidae" genus="Eidinemacheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="228" phylum="Chordata" rank="species" species="proudlovei" status="sp. nov.">
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<emphasis id="B938EAB74F37721121BFFE5A6FC7FE0F" bold="true" box="[151,490,439,465]" italics="true" pageId="3" pageNumber="228">Eidinemacheilus proudlovei</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A20B57CC4F37721120D1FE556CA4FE0C" box="[505,649,440,466]" pageId="3" pageNumber="228" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF336A54F37721121BFFE376F3FFE2D" blockId="3.[151,649,439,499]" box="[151,274,474,499]" pageId="3" pageNumber="228">
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(
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<figureCitation id="13772A204F37721121B7FE376F24FE2D" box="[159,265,474,499]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStartId-0="3.[151,249,1562,1584]" captionStartId-1="5.[151,249,1483,1505]" captionStartId-2="5.[151,249,2008,2030]" captionStartId-3="6.[151,249,1254,1276]" captionStartId-4="7.[151,249,1966,1988]" captionStartId-5="8.[151,249,1952,1974]" captionTargetBox-0="[151,1435,695,1540]" captionTargetBox-1="[151,1434,193,1461]" captionTargetBox-2="[151,1436,1616,1986]" captionTargetBox-3="[418,1176,176,1230]" captionTargetBox-4="[222,1360,193,1930]" captionTargetBox-5="[176,1406,193,1918]" captionTargetId-0="figure@3.[151,1435,695,1540]" captionTargetId-1="figure@5.[151,1435,193,1462]" captionTargetId-2="figure@5.[151,1436,1616,1986]" captionTargetId-3="figure@6.[418,1176,175,1231]" captionTargetId-4="figure@7.[222,1360,193,1932]" captionTargetId-5="figure@8.[176,1408,193,1920]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionText-0="FIGURE 2. Eidinemacheilus proudlovei, ZFMK-ICH 102318, holotype, 53.6 mm SL; Iraqi Kurdistan: Ephemeral spring in Tabeen Gorge." captionText-1="FIGURE 3. Eidinemacheilus proudlovei, paratypes, FSJF 3542, 48.5, 45.5 mm SL; Iraqi Kurdistan: Ephemeral spring in Tabeen Gorge." captionText-2="FIGURE 4. Eidinemacheilus proudlovei, alive, not preserved, 45 mm SL; Iraqi Kurdistan: Ephemeral spring in Tabeen Gorge." captionText-3="FIGURE 5. Eidinemacheilus proudlovei, FSJF 3542, paratype, 48.5 SL; right pectoral fin, dorsal view showing many small unculi; Iraqi Kurdistan: Ephemeral spring in Tabeen Gorge." captionText-4="FIGURE 6. Eidinemacheilus proudlovei, ZFMK-ICH 102318, holotype, 53.6 mm SL; lateral, and ventral view of head showing large mouth; Iraqi Kurdistan: Ephemeral spring in Tabeen Gorge." captionText-5="FIGURE 7. Eidinemacheilus proudlovei, FSJF 3542, paratype, 48.5 SL; Iraqi Kurdistan: Ephemeral spring in Tabeen Gorge; E. smithi, FSJF 3541, 34 mm SL; Iran: Natural well-like opening being the origin of the stream Kay-e-Ru; bony swim bladder capsule, dorsal, ventral, posterior and anterior views." httpUri-0="https://zenodo.org/record/159362/files/figure.png" httpUri-1="https://zenodo.org/record/159363/files/figure.png" httpUri-2="https://zenodo.org/record/159364/files/figure.png" httpUri-3="https://zenodo.org/record/159365/files/figure.png" httpUri-4="https://zenodo.org/record/159366/files/figure.png" httpUri-5="https://zenodo.org/record/159367/files/figure.png" pageId="3" pageNumber="228">Figs. 2–7</figureCitation>
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)
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||
</paragraph>
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</subSubSection>
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<subSubSection id="C356652E4F37721121BFFDCF6F36F96D" pageId="3" pageNumber="228" type="materials_examined">
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<paragraph id="8BF336A54F37721121BFFDCF6D5CFD5D" blockId="3.[151,1436,546,643]" pageId="3" pageNumber="228">
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<emphasis id="B938EAB74F37721121BFFDCF6F26FDE5" bold="true" box="[151,267,546,571]" pageId="3" pageNumber="228">Holotype.</emphasis>
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ZFMK-ICH
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<date id="FFF210654F377211208AFDCE6FD4FDE5" box="[418,505,547,571]" pageId="3" pageNumber="228" value="18-10-23">102318</date>
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,
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<quantity id="4CB49B404F377211232DFDCF6C47FDE4" box="[517,618,546,571]" metricMagnitude="-2" metricUnit="m" metricValue="5.36" pageId="3" pageNumber="228" unit="mm" value="53.6">53.6 mm</quantity>
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SL;
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||
<materialsCitation id="3B243CF84F377211238AFDCE6D5CFD5D" ID-GBIF-Occurrence="1316260495" collectingDate="2016-03-31" country="Iraq" county="Kanishok" elevation="800" latitude="35.83428" location="Sulaimani prov." longLatPrecision="1" longitude="45.106148" municipality="Piramagrun mountain" pageId="3" pageNumber="228" specimenCount="1">
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||
<collectingCountry id="F35B76354F377211238AFDCE6CFEFDE4" box="[674,723,547,570]" name="Iraq" pageId="3" pageNumber="228">Iraq</collectingCountry>
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||
:
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||
<location id="8E93607E4F37721123F7FDCF6DB8FDE4" LSID="urn:lsid:plazi:treatment:03E587B34F37721B21BFFE5A6CABF978:8E93607E4F37721123F7FDCF6DB8FDE4" box="[735,917,546,571]" country="Iraq" county="Kanishok" latitude="35.83428" longLatPrecision="1" longitude="45.106148" municipality="Piramagrun mountain" name="Sulaimani prov." pageId="3" pageNumber="228">Sulaimani prov.</location>
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||
:
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||
<location id="8E93607E4F3772112288FDCF6A34FDE5" LSID="urn:lsid:plazi:treatment:03E587B34F37721B21BFFE5A6CABF978:8E93607E4F3772112288FDCF6A34FDE5" box="[928,1049,546,571]" country="Iraq" county="Kanishok" latitude="35.83428" longLatPrecision="1" longitude="45.106148" municipality="Piramagrun mountain" name="Ephemeral" pageId="3" pageNumber="228">Ephemeral</location>
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spring in the
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<location id="8E93607E4F3772112590FDCF6B78FDE5" LSID="urn:lsid:plazi:treatment:03E587B34F37721B21BFFE5A6CABF978:8E93607E4F3772112590FDCF6B78FDE5" box="[1208,1365,546,571]" country="Iraq" county="Kanishok" latitude="35.83428" longLatPrecision="1" longitude="45.106148" municipality="Piramagrun mountain" name="Tabeen Gorge" pageId="3" pageNumber="228">Tabeen Gorge</location>
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at the slope the northernmost
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<collectingMunicipality id="6B97ACDF4F3772112099FDAB6C80FD81" box="[433,685,582,607]" pageId="3" pageNumber="228">Piramagrun mountain</collectingMunicipality>
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||
, about
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||
<elevation id="0061D1964F3772112221FDAA6D7BFD80" box="[777,854,583,607]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="228" unit="m" value="800.0">
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<quantity id="4CB49B404F3772112221FDAA6D7BFD80" box="[777,854,583,607]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="228" unit="m" value="800.0">800 m</quantity>
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</elevation>
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east of the village
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<collectingCounty id="62924E294F3772112561FDAB6A9AFD81" box="[1097,1207,582,607]" pageId="3" pageNumber="228">Kanishok</collectingCounty>
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,
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<geoCoordinate id="EE7850624F37721125E0FDAB6B4EFD81" box="[1224,1379,582,607]" direction="north" orientation="latitude" pageId="3" pageNumber="228" precision="1" value="35.83428">35°50'3.40"N</geoCoordinate>
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,
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||
<geoCoordinate id="EE7850624F377211245DFDAB6F24FD5D" direction="east" orientation="longitude" pageId="3" pageNumber="228" precision="1" value="45.106148">45° 6'22.14"E</geoCoordinate>
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||
; Younis Sabir Abdullah & Korsh Ararat,
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||
<collectingDate id="EFB6E98D4F37721123F0FD866D40FD5D" box="[728,877,618,643]" pageId="3" pageNumber="228" value="2016-03-31">
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||
<date id="FFF210654F37721123F0FD866D40FD5D" box="[728,877,618,643]" pageId="3" pageNumber="228" value="2016-03-31">31 Mar 2016</date>
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||
</collectingDate>
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||
.
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||
</materialsCitation>
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||
</paragraph>
|
||
<caption id="DF33662D4F37721121BFF9F76F01F991" httpUri="https://zenodo.org/record/159362/files/figure.png" pageId="3" pageNumber="228" targetBox="[151,1435,695,1540]" targetPageId="3">
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<paragraph id="8BF336A54F37721121BFF9F76F01F991" blockId="3.[151,1436,1562,1615]" pageId="3" pageNumber="228">
|
||
<emphasis id="B938EAB74F37721121BFF9F76F39F9F1" bold="true" box="[151,276,1562,1584]" pageId="3" pageNumber="228">FIGURE 2.</emphasis>
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<taxonomicName id="4C4C4D264F3772112034F9F76C17F9EE" box="[284,570,1562,1584]" class="Actinopterygii" family="Nemacheilidae" genus="Eidinemacheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="228" phylum="Chordata" rank="species" species="proudlovei">
|
||
<emphasis id="B938EAB74F3772112034F9F76C17F9EE" box="[284,570,1562,1584]" italics="true" pageId="3" pageNumber="228">Eidinemacheilus proudlovei</emphasis>
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||
</taxonomicName>
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, ZFMK-ICH 102318, holotype, 53.6 mm SL; Iraqi Kurdistan: Ephemeral spring in Tabeen Gorge.
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||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF336A54F37721121EFF99B6F36F96D" blockId="3.[151,1436,1654,2003]" pageId="3" pageNumber="228">
|
||
<materialsCitation id="3B243CF84F37721121EFF99B6BB8F951" ID-GBIF-Occurrence="1316260494" box="[199,1429,1654,1679]" collectionCode="FSJF" pageId="3" pageNumber="228" specimenCode="FSJF 3542, 2" specimenCount="1" typeStatus="paratype">
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||
<emphasis id="B938EAB74F37721121EFF99B6F6BF951" bold="true" box="[199,326,1654,1679]" pageId="3" pageNumber="228">
|
||
<typeStatus id="54F788074F37721121EFF99B6F6CF951" box="[199,321,1654,1679]" pageId="3" pageNumber="228" type="paratype">Paratypes</typeStatus>
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||
.
|
||
</emphasis>
|
||
<specimenCode id="DBEA9EDE4F3772112065F99A6FC8F951" box="[333,485,1654,1679]" pageId="3" pageNumber="228">
|
||
<collectionCode id="ED5DAE604F3772112065F99A6FAAF951" box="[333,391,1655,1679]" pageId="3" pageNumber="228">FSJF</collectionCode>
|
||
3542, 2
|
||
</specimenCode>
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||
, 45.5,
|
||
<quantity id="4CB49B404F3772112318F99B6CB8F950" box="[560,661,1654,1679]" metricMagnitude="-2" metricUnit="m" metricValue="4.85" pageId="3" pageNumber="228" unit="mm" value="48.5">48.5 mm</quantity>
|
||
SL; same data as holotype; Genbank accession number:
|
||
<accessionNumber id="941FAB464F377211243EF99A6BB8F951" box="[1302,1429,1655,1679]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KX774390" pageId="3" pageNumber="228">KX774390</accessionNumber>
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</materialsCitation>
|
||
-
|
||
<accessionNumber id="941FAB464F37721121BFF9766F3AF96D" box="[151,279,1691,1715]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KX774392" pageId="3" pageNumber="228">KX774392</accessionNumber>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C356652E4F37721121EFF9536A36F89D" pageId="3" pageNumber="228" type="diagnosis">
|
||
<paragraph id="8BF336A54F37721121EFF9536A36F89D" blockId="3.[151,1436,1654,2003]" pageId="3" pageNumber="228">
|
||
<emphasis id="B938EAB74F37721121EFF9536F6CF909" bold="true" box="[199,321,1726,1751]" pageId="3" pageNumber="228">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C4C4D264F377211207AF9536CB8F909" box="[338,661,1726,1751]" class="Actinopterygii" family="Nemacheilidae" genus="Eidinemacheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="228" phylum="Chordata" rank="species" species="proudlovei">
|
||
<emphasis id="B938EAB74F377211207AF9536CB8F909" box="[338,661,1726,1751]" italics="true" pageId="3" pageNumber="228">Eidinemacheilus proudlovei</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="4C4C4D264F377211229FF9526A09F909" box="[951,1060,1726,1751]" class="Actinopterygii" family="Nemacheilidae" genus="Eidinemacheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="228" phylum="Chordata" rank="species" species="smithi">
|
||
<emphasis id="B938EAB74F377211229FF9526A09F909" box="[951,1060,1726,1751]" italics="true" pageId="3" pageNumber="228">E. smithi</emphasis>
|
||
</taxonomicName>
|
||
, the only other troglomorphic nemacheilid loach species in the Middle East, and the only other species in
|
||
<taxonomicName id="4C4C4D264F3772112539F90F6AE1F925" box="[1041,1228,1762,1787]" class="Actinopterygii" family="Nemacheilidae" genus="Eidinemacheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="228" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAB74F3772112539F90F6AE1F925" box="[1041,1228,1762,1787]" italics="true" pageId="3" pageNumber="228">Eidinemacheilus</emphasis>
|
||
</taxonomicName>
|
||
known so far, by having 8+8 or 8+7 branched caudal-fin rays (vs. 7+7), no adipose keel on the caudal peduncle (vs. present), enlarged jaws (not enlarged) and a fully developed head canal system (reduced).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C356652E4F37721421EFF8A36DD9FABF" lastPageId="6" lastPageNumber="231" pageId="3" pageNumber="228" type="description">
|
||
<paragraph id="8BF336A54F37721621EFF8A36C65FEBA" blockId="3.[151,1436,1654,2003]" lastBlockId="4.[151,1437,151,357]" lastPageId="4" lastPageNumber="229" pageId="3" pageNumber="228">
|
||
<emphasis id="B938EAB74F37721121EFF8A36F7CF8B9" bold="true" box="[199,337,1870,1895]" pageId="3" pageNumber="228">Description</emphasis>
|
||
. See
|
||
<figureCitation id="13772A204F37721120B8F8A36C3AF8B9" box="[400,535,1870,1895]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="3.[151,249,1562,1584]" captionStartId-1="5.[151,249,1483,1505]" captionStartId-2="5.[151,249,2008,2030]" captionTargetBox-0="[151,1435,695,1540]" captionTargetBox-1="[151,1434,193,1461]" captionTargetBox-2="[151,1436,1616,1986]" captionTargetId-0="figure@3.[151,1435,695,1540]" captionTargetId-1="figure@5.[151,1435,193,1462]" captionTargetId-2="figure@5.[151,1436,1616,1986]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="5" captionText-0="FIGURE 2. Eidinemacheilus proudlovei, ZFMK-ICH 102318, holotype, 53.6 mm SL; Iraqi Kurdistan: Ephemeral spring in Tabeen Gorge." captionText-1="FIGURE 3. Eidinemacheilus proudlovei, paratypes, FSJF 3542, 48.5, 45.5 mm SL; Iraqi Kurdistan: Ephemeral spring in Tabeen Gorge." captionText-2="FIGURE 4. Eidinemacheilus proudlovei, alive, not preserved, 45 mm SL; Iraqi Kurdistan: Ephemeral spring in Tabeen Gorge." httpUri-0="https://zenodo.org/record/159362/files/figure.png" httpUri-1="https://zenodo.org/record/159363/files/figure.png" httpUri-2="https://zenodo.org/record/159364/files/figure.png" pageId="3" pageNumber="228">Figures 2–4</figureCitation>
|
||
for general appearance and
|
||
<tableCitation id="C6CE031E4F377211227FF8A36D86F8B9" box="[855,939,1870,1895]" captionStart="TABLE 1" captionStartId="4.[151,236,404,427]" captionTargetBox="[159,1324,486,1474]" captionTargetPageId="4" captionText="TABLE 1. Morphometric data of Eidinemacheilus proudlovei (ZFMK-ICH 102318, holotype; FSJF 3542; paratypes; n = 3)." httpUri="http://table.plazi.org/id/DF33662D4F30721621BFFE796EFCFE15" pageId="3" pageNumber="228" tableUuid="DF33662D4F30721621BFFE796EFCFE15">Table 1</tableCitation>
|
||
for morphometric data of
|
||
<typeStatus id="54F788074F37721125F0F8A36B17F8B9" box="[1240,1338,1870,1895]" pageId="3" pageNumber="228" type="holotype">holotype</typeStatus>
|
||
and two
|
||
<typeStatus id="54F788074F37721121BFF89E6F2BF855" box="[151,262,1907,1931]" pageId="3" pageNumber="228" type="paratype">paratypes</typeStatus>
|
||
. Medium sized, moderately stout species with large head. Body deepest at, or slightly behind nape, profile of back straight, or body depth slightly decreasing towards dorsal-fin origin, and slowly decreasing below dorsal-fin base and towards caudal-fin base. No hump at nape. Greatest body width at pectoral-fin base. Head conical, flattened on ventral surface. Caudal peduncle compressed laterally, 1.8–1.9 times longer than deep. Pectoral fin reaching approximately 60–70% of distance from pectoral-fin origin to pelvic-fin origin. Pelvic axillary lobe absent. Pelvic-fin origin below unbranched or first branched dorsal-fin ray. Pelvic fin not reaching to anus. Anal-fin origin shortly behind anus. Anal-fin origin at vertical of middle between dorsal- and caudal-fin origins. No adipose keel on caudal peduncle. Margin of dorsal fin straight. Caudal fin moderately emarginated. Largest known specimen
|
||
<quantity id="4CB49B404F307216209DFEA16C34FEBA" box="[437,537,332,357]" metricMagnitude="-2" metricUnit="m" metricValue="5.36" pageId="4" pageNumber="229" unit="mm" value="53.6">53.6 mm</quantity>
|
||
SL.
|
||
</paragraph>
|
||
<caption id="DF33662D4F30721621BFFE796EFCFE15" ID-Table-UUID="DF33662D4F30721621BFFE796EFCFE15" httpUri="http://table.plazi.org/id/DF33662D4F30721621BFFE796EFCFE15" pageId="4" pageNumber="229" targetBox="[159,1324,486,1474]" targetIsTable="true" targetPageId="4">
|
||
<paragraph id="8BF336A54F30721621BFFE796EFCFE15" blockId="4.[151,1436,403,459]" pageId="4" pageNumber="229">
|
||
<emphasis id="B938EAB74F30721621BFFE796F24FE75" bold="true" box="[151,265,404,427]" pageId="4" pageNumber="229">TABLE 1.</emphasis>
|
||
Morphometric data of
|
||
<taxonomicName id="4C4C4D264F3072162327FE7E6D11FE75" box="[527,828,403,427]" class="Actinopterygii" family="Nemacheilidae" genus="Eidinemacheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="229" phylum="Chordata" rank="species" species="proudlovei">
|
||
<emphasis id="B938EAB74F3072162327FE7E6D11FE75" box="[527,828,403,427]" italics="true" pageId="4" pageNumber="229">Eidinemacheilus proudlovei</emphasis>
|
||
</taxonomicName>
|
||
(ZFMK-ICH 102318, holotype; FSJF 3542; paratypes; n=3).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF336A54F307216221DFE0B6ACFFA1C" pageId="4" pageNumber="229">
|
||
<table id="F94CC4054F308DED21B7FE0B6B01FA1C" box="[159,1324,486,1474]" gridcols="4" gridrows="25" pageId="4" pageNumber="229">
|
||
<tr id="357C34E74F308DED21B7FE0B6B01FE22" box="[159,1324,486,508]" gridrow="0" pageId="4" pageNumber="229" rowspan-0="1">
|
||
<th id="76AD5D9B4F308DED221DFE0B6DA3FE22" box="[821,910,486,508]" gridcol="1" gridrow="0" pageId="4" pageNumber="229">holotype</th>
|
||
<th id="76AD5D9B4F308DED252BFE0B6A77FE22" box="[1027,1114,486,508]" gridcol="2" gridrow="0" pageId="4" pageNumber="229">paratype</th>
|
||
<th id="76AD5D9B4F308DED25FDFE0B6B01FE22" box="[1237,1324,486,508]" gridcol="3" gridrow="0" pageId="4" pageNumber="229">paratype</th>
|
||
</tr>
|
||
<tr id="357C34E74F308DED21B7FDE36B01FDFA" box="[159,1324,526,548]" gridrow="1" pageId="4" pageNumber="229">
|
||
<th id="76AD5D9B4F308DED21B7FDE36CA7FDFA" box="[159,650,526,548]" gridcol="0" gridrow="1" pageId="4" pageNumber="229">SL (mm)</th>
|
||
<td id="76AD5D9B4F308DED221DFDE36DA3FDFA" box="[821,910,526,548]" gridcol="1" gridrow="1" pageId="4" pageNumber="229">53.6</td>
|
||
<td id="76AD5D9B4F308DED252BFDE36A77FDFA" box="[1027,1114,526,548]" gridcol="2" gridrow="1" pageId="4" pageNumber="229">45.5</td>
|
||
<td id="76AD5D9B4F308DED25FDFDE36B01FDFA" box="[1237,1324,526,548]" gridcol="3" gridrow="1" pageId="4" pageNumber="229">48.5</td>
|
||
</tr>
|
||
<tr id="357C34E74F308DED21B7FDDB6B01FD92" box="[159,1324,566,588]" gridrow="2" pageId="4" pageNumber="229" rowspan-1="1" rowspan-2="1" rowspan-3="1">
|
||
<th id="76AD5D9B4F308DED21B7FDDB6CA7FD92" box="[159,650,566,588]" gridcol="0" gridrow="2" pageId="4" pageNumber="229">In percent of standard length</th>
|
||
</tr>
|
||
<tr id="357C34E74F308DED21B7FDB36B01FDAA" box="[159,1324,606,628]" gridrow="3" pageId="4" pageNumber="229">
|
||
<th id="76AD5D9B4F308DED21B7FDB36CA7FDAA" box="[159,650,606,628]" gridcol="0" gridrow="3" pageId="4" pageNumber="229">Head length</th>
|
||
<td id="76AD5D9B4F308DED221DFDB36DA3FDAA" box="[821,910,606,628]" gridcol="1" gridrow="3" pageId="4" pageNumber="229">28.9</td>
|
||
<td id="76AD5D9B4F308DED252BFDB36A77FDAA" box="[1027,1114,606,628]" gridcol="2" gridrow="3" pageId="4" pageNumber="229">26.4</td>
|
||
<td id="76AD5D9B4F308DED25FDFDB36B01FDAA" box="[1237,1324,606,628]" gridcol="3" gridrow="3" pageId="4" pageNumber="229">27.0</td>
|
||
</tr>
|
||
<tr id="357C34E74F308DED21B7FD6A6B01FD43" box="[159,1324,647,669]" gridrow="4" pageId="4" pageNumber="229">
|
||
<th id="76AD5D9B4F308DED21B7FD6A6CA7FD43" box="[159,650,647,669]" gridcol="0" gridrow="4" pageId="4" pageNumber="229">Body depth at dorsal-fin origin</th>
|
||
<td id="76AD5D9B4F308DED221DFD6A6DA3FD43" box="[821,910,647,669]" gridcol="1" gridrow="4" pageId="4" pageNumber="229">14.4</td>
|
||
<td id="76AD5D9B4F308DED252BFD6A6A77FD43" box="[1027,1114,647,669]" gridcol="2" gridrow="4" pageId="4" pageNumber="229">15.6</td>
|
||
<td id="76AD5D9B4F308DED25FDFD6A6B01FD43" box="[1237,1324,647,669]" gridcol="3" gridrow="4" pageId="4" pageNumber="229">17.1</td>
|
||
</tr>
|
||
<tr id="357C34E74F308DED21B7FD426B01FD1B" box="[159,1324,687,709]" gridrow="5" pageId="4" pageNumber="229">
|
||
<th id="76AD5D9B4F308DED21B7FD426CA7FD1B" box="[159,650,687,709]" gridcol="0" gridrow="5" pageId="4" pageNumber="229">Predorsal length</th>
|
||
<td id="76AD5D9B4F308DED221DFD426DA3FD1B" box="[821,910,687,709]" gridcol="1" gridrow="5" pageId="4" pageNumber="229">57.0</td>
|
||
<td id="76AD5D9B4F308DED252BFD426A77FD1B" box="[1027,1114,687,709]" gridcol="2" gridrow="5" pageId="4" pageNumber="229">35.5</td>
|
||
<td id="76AD5D9B4F308DED25FDFD426B01FD1B" box="[1237,1324,687,709]" gridcol="3" gridrow="5" pageId="4" pageNumber="229">57.0</td>
|
||
</tr>
|
||
<tr id="357C34E74F308DED21B7FD3A6B01FD33" box="[159,1324,727,749]" gridrow="6" pageId="4" pageNumber="229">
|
||
<th id="76AD5D9B4F308DED21B7FD3A6CA7FD33" box="[159,650,727,749]" gridcol="0" gridrow="6" pageId="4" pageNumber="229">Postdorsal length</th>
|
||
<td id="76AD5D9B4F308DED221DFD3A6DA3FD33" box="[821,910,727,749]" gridcol="1" gridrow="6" pageId="4" pageNumber="229">34.0</td>
|
||
<td id="76AD5D9B4F308DED252BFD3A6A77FD33" box="[1027,1114,727,749]" gridcol="2" gridrow="6" pageId="4" pageNumber="229">34.7</td>
|
||
<td id="76AD5D9B4F308DED25FDFD3A6B01FD33" box="[1237,1324,727,749]" gridcol="3" gridrow="6" pageId="4" pageNumber="229">36.6</td>
|
||
</tr>
|
||
<tr id="357C34E74F308DED21B7FCED6B01FCC8" box="[159,1324,768,790]" gridrow="7" pageId="4" pageNumber="229">
|
||
<th id="76AD5D9B4F308DED21B7FCED6CA7FCC8" box="[159,650,768,790]" gridcol="0" gridrow="7" pageId="4" pageNumber="229">Prepelvic length</th>
|
||
<td id="76AD5D9B4F308DED221DFCED6DA3FCC8" box="[821,910,768,790]" gridcol="1" gridrow="7" pageId="4" pageNumber="229">55.8</td>
|
||
<td id="76AD5D9B4F308DED252BFCED6A77FCC8" box="[1027,1114,768,790]" gridcol="2" gridrow="7" pageId="4" pageNumber="229">57.2</td>
|
||
<td id="76AD5D9B4F308DED25FDFCED6B01FCC8" box="[1237,1324,768,790]" gridcol="3" gridrow="7" pageId="4" pageNumber="229">56.8</td>
|
||
</tr>
|
||
<tr id="357C34E74F308DED21B7FCC56B01FCE0" box="[159,1324,808,830]" gridrow="8" pageId="4" pageNumber="229">
|
||
<th id="76AD5D9B4F308DED21B7FCC56CA7FCE0" box="[159,650,808,830]" gridcol="0" gridrow="8" pageId="4" pageNumber="229">Preanal length</th>
|
||
<td id="76AD5D9B4F308DED221DFCC56DA3FCE0" box="[821,910,808,830]" gridcol="1" gridrow="8" pageId="4" pageNumber="229">75.7</td>
|
||
<td id="76AD5D9B4F308DED252BFCC56A77FCE0" box="[1027,1114,808,830]" gridcol="2" gridrow="8" pageId="4" pageNumber="229">77.8</td>
|
||
<td id="76AD5D9B4F308DED25FDFCC56B01FCE0" box="[1237,1324,808,830]" gridcol="3" gridrow="8" pageId="4" pageNumber="229">78.0</td>
|
||
</tr>
|
||
<tr id="357C34E74F308DED21B7FCBD6B01FCB8" box="[159,1324,848,870]" gridrow="9" pageId="4" pageNumber="229">
|
||
<th id="76AD5D9B4F308DED21B7FCBD6CA7FCB8" box="[159,650,848,870]" gridcol="0" gridrow="9" pageId="4" pageNumber="229">Distance between pectoral and pelvic-fin origins</th>
|
||
<td id="76AD5D9B4F308DED221DFCBD6DA3FCB8" box="[821,910,848,870]" gridcol="1" gridrow="9" pageId="4" pageNumber="229">30.2</td>
|
||
<td id="76AD5D9B4F308DED252BFCBD6A77FCB8" box="[1027,1114,848,870]" gridcol="2" gridrow="9" pageId="4" pageNumber="229">31.7</td>
|
||
<td id="76AD5D9B4F308DED25FDFCBD6B01FCB8" box="[1237,1324,848,870]" gridcol="3" gridrow="9" pageId="4" pageNumber="229">32.7</td>
|
||
</tr>
|
||
<tr id="357C34E74F308DED21B7FC956B01FC50" box="[159,1324,888,910]" gridrow="10" pageId="4" pageNumber="229">
|
||
<th id="76AD5D9B4F308DED21B7FC956CA7FC50" box="[159,650,888,910]" gridcol="0" gridrow="10" pageId="4" pageNumber="229">Distance between pelvic and anal-fin origins</th>
|
||
<td id="76AD5D9B4F308DED221DFC956DA3FC50" box="[821,910,888,910]" gridcol="1" gridrow="10" pageId="4" pageNumber="229">20.3</td>
|
||
<td id="76AD5D9B4F308DED252BFC956A77FC50" box="[1027,1114,888,910]" gridcol="2" gridrow="10" pageId="4" pageNumber="229">20.9</td>
|
||
<td id="76AD5D9B4F308DED25FDFC956B01FC50" box="[1237,1324,888,910]" gridcol="3" gridrow="10" pageId="4" pageNumber="229">19.9</td>
|
||
</tr>
|
||
<tr id="357C34E74F308DED21B7FC4C6B01FC69" box="[159,1324,929,951]" gridrow="11" pageId="4" pageNumber="229">
|
||
<th id="76AD5D9B4F308DED21B7FC4C6CA7FC69" box="[159,650,929,951]" gridcol="0" gridrow="11" pageId="4" pageNumber="229">Distance between anus and anal-fin origin</th>
|
||
<td id="76AD5D9B4F308DED221DFC4C6DA3FC69" box="[821,910,929,951]" gridcol="1" gridrow="11" pageId="4" pageNumber="229">2.7</td>
|
||
<td id="76AD5D9B4F308DED252BFC4C6A77FC69" box="[1027,1114,929,951]" gridcol="2" gridrow="11" pageId="4" pageNumber="229">2.5</td>
|
||
<td id="76AD5D9B4F308DED25FDFC4C6B01FC69" box="[1237,1324,929,951]" gridcol="3" gridrow="11" pageId="4" pageNumber="229">2.5</td>
|
||
</tr>
|
||
<tr id="357C34E74F308DED21B7FC246B01FC01" box="[159,1324,969,991]" gridrow="12" pageId="4" pageNumber="229">
|
||
<th id="76AD5D9B4F308DED21B7FC246CA7FC01" box="[159,650,969,991]" gridcol="0" gridrow="12" pageId="4" pageNumber="229">Caudal-peduncle depth</th>
|
||
<td id="76AD5D9B4F308DED221DFC246DA3FC01" box="[821,910,969,991]" gridcol="1" gridrow="12" pageId="4" pageNumber="229">8.3</td>
|
||
<td id="76AD5D9B4F308DED252BFC246A77FC01" box="[1027,1114,969,991]" gridcol="2" gridrow="12" pageId="4" pageNumber="229">9.3</td>
|
||
<td id="76AD5D9B4F308DED25FDFC246B01FC01" box="[1237,1324,969,991]" gridcol="3" gridrow="12" pageId="4" pageNumber="229">9.4</td>
|
||
</tr>
|
||
<tr id="357C34E74F308DED21B7FC1C6B01FBD9" box="[159,1324,1009,1031]" gridrow="13" pageId="4" pageNumber="229">
|
||
<th id="76AD5D9B4F308DED21B7FC1C6CA7FBD9" box="[159,650,1009,1031]" gridcol="0" gridrow="13" pageId="4" pageNumber="229">Caudal-peduncle length</th>
|
||
<td id="76AD5D9B4F308DED221DFC1C6DA3FBD9" box="[821,910,1009,1031]" gridcol="1" gridrow="13" pageId="4" pageNumber="229">15.8</td>
|
||
<td id="76AD5D9B4F308DED252BFC1C6A77FBD9" box="[1027,1114,1009,1031]" gridcol="2" gridrow="13" pageId="4" pageNumber="229">18.0</td>
|
||
<td id="76AD5D9B4F308DED25FDFC1C6B01FBD9" box="[1237,1324,1009,1031]" gridcol="3" gridrow="13" pageId="4" pageNumber="229">17.2</td>
|
||
</tr>
|
||
<tr id="357C34E74F308DED21B7FBF76B01FBEE" box="[159,1324,1050,1072]" gridrow="14" pageId="4" pageNumber="229">
|
||
<th id="76AD5D9B4F308DED21B7FBF76CA7FBEE" box="[159,650,1050,1072]" gridcol="0" gridrow="14" pageId="4" pageNumber="229">Dorsal fin-base length</th>
|
||
<td id="76AD5D9B4F308DED221DFBF76DA3FBEE" box="[821,910,1050,1072]" gridcol="1" gridrow="14" pageId="4" pageNumber="229">10.3</td>
|
||
<td id="76AD5D9B4F308DED252BFBF76A77FBEE" box="[1027,1114,1050,1072]" gridcol="2" gridrow="14" pageId="4" pageNumber="229">9.0</td>
|
||
<td id="76AD5D9B4F308DED25FDFBF76B01FBEE" box="[1237,1324,1050,1072]" gridcol="3" gridrow="14" pageId="4" pageNumber="229">10.3</td>
|
||
</tr>
|
||
<tr id="357C34E74F308DED21B7FBAF6B01FB86" box="[159,1324,1090,1112]" gridrow="15" pageId="4" pageNumber="229">
|
||
<th id="76AD5D9B4F308DED21B7FBAF6CA7FB86" box="[159,650,1090,1112]" gridcol="0" gridrow="15" pageId="4" pageNumber="229">Anal fin-base length</th>
|
||
<td id="76AD5D9B4F308DED221DFBAF6DA3FB86" box="[821,910,1090,1112]" gridcol="1" gridrow="15" pageId="4" pageNumber="229">8.4</td>
|
||
<td id="76AD5D9B4F308DED252BFBAF6A77FB86" box="[1027,1114,1090,1112]" gridcol="2" gridrow="15" pageId="4" pageNumber="229">7.8</td>
|
||
<td id="76AD5D9B4F308DED25FDFBAF6B01FB86" box="[1237,1324,1090,1112]" gridcol="3" gridrow="15" pageId="4" pageNumber="229">8.4</td>
|
||
</tr>
|
||
<tr id="357C34E74F308DED21B7FB876B01FB5E" box="[159,1324,1130,1152]" gridrow="16" pageId="4" pageNumber="229">
|
||
<th id="76AD5D9B4F308DED21B7FB876CA7FB5E" box="[159,650,1130,1152]" gridcol="0" gridrow="16" pageId="4" pageNumber="229">Pelvic-fin length</th>
|
||
<td id="76AD5D9B4F308DED221DFB876DA3FB5E" box="[821,910,1130,1152]" gridcol="1" gridrow="16" pageId="4" pageNumber="229">12.1</td>
|
||
<td id="76AD5D9B4F308DED252BFB876A77FB5E" box="[1027,1114,1130,1152]" gridcol="2" gridrow="16" pageId="4" pageNumber="229">10.7</td>
|
||
<td id="76AD5D9B4F308DED25FDFB876B01FB5E" box="[1237,1324,1130,1152]" gridcol="3" gridrow="16" pageId="4" pageNumber="229">11.8</td>
|
||
</tr>
|
||
<tr id="357C34E74F308DED21B7FB7F6B01FB76" box="[159,1324,1170,1192]" gridrow="17" pageId="4" pageNumber="229">
|
||
<th id="76AD5D9B4F308DED21B7FB7F6CA7FB76" box="[159,650,1170,1192]" gridcol="0" gridrow="17" pageId="4" pageNumber="229">Pectoral-fin length</th>
|
||
<td id="76AD5D9B4F308DED221DFB7F6DA3FB76" box="[821,910,1170,1192]" gridcol="1" gridrow="17" pageId="4" pageNumber="229">18.4</td>
|
||
<td id="76AD5D9B4F308DED252BFB7F6A77FB76" box="[1027,1114,1170,1192]" gridcol="2" gridrow="17" pageId="4" pageNumber="229">18.3</td>
|
||
<td id="76AD5D9B4F308DED25FDFB7F6B01FB76" box="[1237,1324,1170,1192]" gridcol="3" gridrow="17" pageId="4" pageNumber="229">18.8</td>
|
||
</tr>
|
||
<tr id="357C34E74F308DED21B7FB566B01FB0F" box="[159,1324,1211,1233]" gridrow="18" pageId="4" pageNumber="229" rowspan-1="1" rowspan-2="1" rowspan-3="1">
|
||
<th id="76AD5D9B4F308DED21B7FB566CA7FB0F" box="[159,650,1211,1233]" gridcol="0" gridrow="18" pageId="4" pageNumber="229">In percent of head length</th>
|
||
</tr>
|
||
<tr id="357C34E74F308DED21B7FB0E6B01FB27" box="[159,1324,1251,1273]" gridrow="19" pageId="4" pageNumber="229">
|
||
<th id="76AD5D9B4F308DED21B7FB0E6CA7FB27" box="[159,650,1251,1273]" gridcol="0" gridrow="19" pageId="4" pageNumber="229">Head depth at nape</th>
|
||
<td id="76AD5D9B4F308DED221DFB0E6DA3FB27" box="[821,910,1251,1273]" gridcol="1" gridrow="19" pageId="4" pageNumber="229">14</td>
|
||
<td id="76AD5D9B4F308DED252BFB0E6A77FB27" box="[1027,1114,1251,1273]" gridcol="2" gridrow="19" pageId="4" pageNumber="229">14</td>
|
||
<td id="76AD5D9B4F308DED25FDFB0E6B01FB27" box="[1237,1324,1251,1273]" gridcol="3" gridrow="19" pageId="4" pageNumber="229">15</td>
|
||
</tr>
|
||
<tr id="357C34E74F308DED21B7FAE66B01FAFF" box="[159,1324,1291,1313]" gridrow="20" pageId="4" pageNumber="229">
|
||
<th id="76AD5D9B4F308DED21B7FAE66CA7FAFF" box="[159,650,1291,1313]" gridcol="0" gridrow="20" pageId="4" pageNumber="229">Maximum head width</th>
|
||
<td id="76AD5D9B4F308DED221DFAE66DA3FAFF" box="[821,910,1291,1313]" gridcol="1" gridrow="20" pageId="4" pageNumber="229">18</td>
|
||
<td id="76AD5D9B4F308DED252BFAE66A77FAFF" box="[1027,1114,1291,1313]" gridcol="2" gridrow="20" pageId="4" pageNumber="229">16</td>
|
||
<td id="76AD5D9B4F308DED25FDFAE66B01FAFF" box="[1237,1324,1291,1313]" gridcol="3" gridrow="20" pageId="4" pageNumber="229">18</td>
|
||
</tr>
|
||
<tr id="357C34E74F308DED21B7FAD96B01FA94" box="[159,1324,1332,1354]" gridrow="21" pageId="4" pageNumber="229">
|
||
<th id="76AD5D9B4F308DED21B7FAD96CA7FA94" box="[159,650,1332,1354]" gridcol="0" gridrow="21" pageId="4" pageNumber="229">Mouth width</th>
|
||
<td id="76AD5D9B4F308DED221DFAD96DA3FA94" box="[821,910,1332,1354]" gridcol="1" gridrow="21" pageId="4" pageNumber="229">8</td>
|
||
<td id="76AD5D9B4F308DED252BFAD96A77FA94" box="[1027,1114,1332,1354]" gridcol="2" gridrow="21" pageId="4" pageNumber="229">8</td>
|
||
<td id="76AD5D9B4F308DED25FDFAD96B01FA94" box="[1237,1324,1332,1354]" gridcol="3" gridrow="21" pageId="4" pageNumber="229">8</td>
|
||
</tr>
|
||
<tr id="357C34E74F308DED21B7FAB16B01FAAC" box="[159,1324,1372,1394]" gridrow="22" pageId="4" pageNumber="229">
|
||
<th id="76AD5D9B4F308DED21B7FAB16CA7FAAC" box="[159,650,1372,1394]" gridcol="0" gridrow="22" pageId="4" pageNumber="229">Inner rostral-barbel length</th>
|
||
<td id="76AD5D9B4F308DED221DFAB16DA3FAAC" box="[821,910,1372,1394]" gridcol="1" gridrow="22" pageId="4" pageNumber="229">4</td>
|
||
<td id="76AD5D9B4F308DED252BFAB16A77FAAC" box="[1027,1114,1372,1394]" gridcol="2" gridrow="22" pageId="4" pageNumber="229">4</td>
|
||
<td id="76AD5D9B4F308DED25FDFAB16B01FAAC" box="[1237,1324,1372,1394]" gridcol="3" gridrow="22" pageId="4" pageNumber="229">4</td>
|
||
</tr>
|
||
<tr id="357C34E74F308DED21B7FA696B01FA44" box="[159,1324,1412,1434]" gridrow="23" pageId="4" pageNumber="229">
|
||
<th id="76AD5D9B4F308DED21B7FA696CA7FA44" box="[159,650,1412,1434]" gridcol="0" gridrow="23" pageId="4" pageNumber="229">Outer rostral-barbel length</th>
|
||
<td id="76AD5D9B4F308DED221DFA696DA3FA44" box="[821,910,1412,1434]" gridcol="1" gridrow="23" pageId="4" pageNumber="229">7</td>
|
||
<td id="76AD5D9B4F308DED252BFA696A77FA44" box="[1027,1114,1412,1434]" gridcol="2" gridrow="23" pageId="4" pageNumber="229">6</td>
|
||
<td id="76AD5D9B4F308DED25FDFA696B01FA44" box="[1237,1324,1412,1434]" gridcol="3" gridrow="23" pageId="4" pageNumber="229">5</td>
|
||
</tr>
|
||
<tr id="357C34E74F308DED21B7FA416B01FA1C" box="[159,1324,1452,1474]" gridrow="24" pageId="4" pageNumber="229">
|
||
<th id="76AD5D9B4F308DED21B7FA416CA7FA1C" box="[159,650,1452,1474]" gridcol="0" gridrow="24" pageId="4" pageNumber="229">Maxillary-barbel length</th>
|
||
<td id="76AD5D9B4F308DED221DFA416DA3FA1C" box="[821,910,1452,1474]" gridcol="1" gridrow="24" pageId="4" pageNumber="229">5</td>
|
||
<td id="76AD5D9B4F308DED252BFA416A77FA1C" box="[1027,1114,1452,1474]" gridcol="2" gridrow="24" pageId="4" pageNumber="229">5</td>
|
||
<td id="76AD5D9B4F308DED25FDFA416B01FA1C" box="[1237,1324,1452,1474]" gridcol="3" gridrow="24" pageId="4" pageNumber="229">4</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BF336A54F30721621EFF9EB6FDEF82D" blockId="4.[151,1437,1542,2035]" pageId="4" pageNumber="229">
|
||
Dorsal fin with 7½ branched rays. Anal fin with 5½ branched rays. Caudal fin with 8+8 (n=1) or 8+7 (n=2) branched rays. Pectoral fin with 10 and pelvic fin with 5 branched rays. Many small unculi on dorsal surface of unbranched and branched pectoral-fin ray 1–3 (
|
||
<figureCitation id="13772A204F3072162381F9A36CC0F9B9" box="[681,749,1614,1639]" captionStart="FIGURE 5" captionStartId="6.[151,249,1254,1276]" captionTargetBox="[418,1176,176,1230]" captionTargetId="figure@6.[418,1176,175,1231]" captionTargetPageId="6" captionText="FIGURE 5. Eidinemacheilus proudlovei, FSJF 3542, paratype, 48.5 SL; right pectoral fin, dorsal view showing many small unculi; Iraqi Kurdistan: Ephemeral spring in Tabeen Gorge." httpUri="https://zenodo.org/record/159365/files/figure.png" pageId="4" pageNumber="229">Fig. 5</figureCitation>
|
||
). Scales absent. No central, but 1–2 lateral pores on each side in supratemporal canal, 6–7 pores in supraorbital, 7–9 pores in preoperculo-mandibular, 4+8–9 pores in infraorbital canal. Lateral line complete, Reaching to a point above origin. Anterior nostril opening on anterior side of a low, pointed and flap-like tube. Nostrils adjacent to each other, tip of anterior nostril overlaps posterior nostril when folded backwards. No suborbital flap, Groove or slit in fishes examined. Mouth very large, slightly arched (
|
||
<figureCitation id="13772A204F3072162462F9336BA3F929" box="[1354,1422,1758,1783]" captionStart="FIGURE 6" captionStartId="7.[151,249,1966,1988]" captionTargetBox="[222,1360,193,1930]" captionTargetId="figure@7.[222,1360,193,1932]" captionTargetPageId="7" captionText="FIGURE 6. Eidinemacheilus proudlovei, ZFMK-ICH 102318, holotype, 53.6 mm SL; lateral, and ventral view of head showing large mouth; Iraqi Kurdistan: Ephemeral spring in Tabeen Gorge." httpUri="https://zenodo.org/record/159366/files/figure.png" pageId="4" pageNumber="229">Fig. 6</figureCitation>
|
||
). Lips moderately thick, upper lip smooth or with fine ridges, lower lip with deep furrows. A median interruption in lower lip. Upper lip with median incision or a medial furrow. Lips do not meet medially, leaving mouth parts exposed. Processus dentiformis absent. No median notch in lower jaw. Both jaws large and wide, bordered by a cornified sharp edge. Barbels short, Inner rostral barbell reaching base of outer rostral barbel; outer rostral barbel reaching base of maxillary barbel. Maxillary barbel reaching to base of outer rostral barbel when folded to front. Bony swim bladder capsule (
|
||
<figureCitation id="13772A204F30721620F6F85B6C0CF811" box="[478,545,1974,1999]" captionStart="FIGURE 7" captionStartId="8.[151,249,1952,1974]" captionTargetBox="[176,1406,193,1918]" captionTargetId="figure@8.[176,1408,193,1920]" captionTargetPageId="8" captionText="FIGURE 7. Eidinemacheilus proudlovei, FSJF 3542, paratype, 48.5 SL; Iraqi Kurdistan: Ephemeral spring in Tabeen Gorge; E. smithi, FSJF 3541, 34 mm SL; Iran: Natural well-like opening being the origin of the stream Kay-e-Ru; bony swim bladder capsule, dorsal, ventral, posterior and anterior views." httpUri="https://zenodo.org/record/159367/files/figure.png" pageId="4" pageNumber="229">Fig. 7</figureCitation>
|
||
) with large posterior process and very narrow manubrium. A small and narrow epural bone in caudal skeleton.
|
||
</paragraph>
|
||
<caption id="DF33662D4F31721721BFFA266F01F9DE" httpUri="https://zenodo.org/record/159363/files/figure.png" pageId="5" pageNumber="230" targetBox="[151,1434,193,1461]" targetPageId="5">
|
||
<paragraph id="8BF336A54F31721721BFFA266F01F9DE" blockId="5.[151,1436,1483,1536]" pageId="5" pageNumber="230">
|
||
<emphasis id="B938EAB74F31721721BFFA266F3BFA3E" bold="true" box="[151,278,1483,1505]" pageId="5" pageNumber="230">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C4C4D264F3172172008FA266C6DFA3F" box="[288,576,1483,1505]" class="Actinopterygii" family="Nemacheilidae" genus="Eidinemacheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="230" phylum="Chordata" rank="species" species="proudlovei">
|
||
<emphasis id="B938EAB74F3172172008FA266C6DFA3F" box="[288,576,1483,1505]" italics="true" pageId="5" pageNumber="230">Eidinemacheilus proudlovei</emphasis>
|
||
</taxonomicName>
|
||
, paratypes, FSJF 3542, 48.5, 45.5 mm SL; Iraqi Kurdistan: Ephemeral spring in Tabeen Gorge.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF33662D4F31721721BFF8356BB0F830" box="[151,1437,2008,2030]" httpUri="https://zenodo.org/record/159364/files/figure.png" pageId="5" pageNumber="230" targetBox="[151,1436,1616,1986]" targetPageId="5">
|
||
<paragraph id="8BF336A54F31721721BFF8356BB0F830" blockId="5.[151,1437,2008,2030]" box="[151,1437,2008,2030]" pageId="5" pageNumber="230">
|
||
<emphasis id="B938EAB74F31721721BFF8356F3FF833" bold="true" box="[151,274,2008,2030]" pageId="5" pageNumber="230">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C4C4D264F3172172030F8356C19F830" box="[280,564,2008,2030]" class="Actinopterygii" family="Nemacheilidae" genus="Eidinemacheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="230" phylum="Chordata" rank="species" species="proudlovei">
|
||
<emphasis id="B938EAB74F3172172030F8356C19F830" box="[280,564,2008,2030]" italics="true" pageId="5" pageNumber="230">Eidinemacheilus proudlovei</emphasis>
|
||
</taxonomicName>
|
||
, alive, not preserved, 45 mm SL; Iraqi Kurdistan: Ephemeral spring in Tabeen Gorge.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF33662D4F32721421BFFB0B6CDEFAC5" httpUri="https://zenodo.org/record/159365/files/figure.png" pageId="6" pageNumber="231" targetBox="[418,1176,176,1230]" targetPageId="6">
|
||
<paragraph id="8BF336A54F32721421BFFB0B6CDEFAC5" blockId="6.[151,1436,1254,1307]" pageId="6" pageNumber="231">
|
||
<emphasis id="B938EAB74F32721421BFFB0B6F3EFB22" bold="true" box="[151,275,1254,1276]" pageId="6" pageNumber="231">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C4C4D264F3272142034FB0B6C14FB22" box="[284,569,1254,1276]" class="Actinopterygii" family="Nemacheilidae" genus="Eidinemacheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="proudlovei">
|
||
<emphasis id="B938EAB74F3272142034FB0B6C14FB22" box="[284,569,1254,1276]" italics="true" pageId="6" pageNumber="231">Eidinemacheilus proudlovei</emphasis>
|
||
</taxonomicName>
|
||
, FSJF 3542, paratype, 48.5 SL; right pectoral fin, dorsal view showing many small unculi; Iraqi Kurdistan: Ephemeral spring in Tabeen Gorge.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF336A54F32721421EFFAA56DD9FABF" blockId="6.[151,1437,1352,2025]" box="[199,1012,1352,1377]" pageId="6" pageNumber="231">
|
||
<emphasis id="B938EAB74F32721421EFFAA56F62FABF" bold="true" box="[199,335,1352,1377]" pageId="6" pageNumber="231">Coloration.</emphasis>
|
||
In live, body and fins pink; yellowish white in preservation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C356652E4F32721421EFFA816AA3F9CB" pageId="6" pageNumber="231" type="distribution">
|
||
<paragraph id="8BF336A54F32721421EFFA816AA3F9CB" blockId="6.[151,1437,1352,2025]" pageId="6" pageNumber="231">
|
||
<emphasis id="B938EAB74F32721421EFFA816F4DFA5B" bold="true" box="[199,352,1388,1413]" pageId="6" pageNumber="231">Distribution.</emphasis>
|
||
<taxonomicName id="4C4C4D264F3272142040FA816C89FA5B" box="[360,676,1388,1413]" class="Actinopterygii" family="Nemacheilidae" genus="Eidinemacheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="proudlovei">
|
||
<emphasis id="B938EAB74F3272142040FA816C89FA5B" box="[360,676,1388,1413]" italics="true" pageId="6" pageNumber="231">Eidinemacheilus proudlovei</emphasis>
|
||
</taxonomicName>
|
||
was washed out from an aquifer into an ephemeral spring close to the village Kanishok. The spring flowed into a small stream, which belongs to the Tabeen drainage. The Tabeen flows to the Little Zab River at the village of Surqawshan. The Little Zab is a tributary of the Tigris. The Tabeen catchment is well known to have many springs and Karezes (underground irrigation systems similar to qanāts in
|
||
<collectingCountry id="F35B76354F32721421BFFA106EE5F9CB" box="[151,200,1533,1557]" name="Iran" pageId="6" pageNumber="231">Iran</collectingCountry>
|
||
), which are fed by a relatively high winter precipitation in the surrounding mountains.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C356652E4F32721421EFF9CD6B6AF983" pageId="6" pageNumber="231" type="etymology">
|
||
<paragraph id="8BF336A54F32721421EFF9CD6B6AF983" blockId="6.[151,1437,1352,2025]" pageId="6" pageNumber="231">
|
||
<emphasis id="B938EAB74F32721421EFF9CD6F63F9E7" bold="true" box="[199,334,1568,1593]" pageId="6" pageNumber="231">Etymology.</emphasis>
|
||
Named for Graham Proudlove (Manchester Museum, the University of Manchester), a world expert on subterranean fishes, for his continuous efforts documenting the diversity of troglomorphic fishes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C356652E4F32721B21EFF9856CA5F9E4" lastPageId="9" lastPageNumber="234" pageId="6" pageNumber="231" type="discussion">
|
||
<paragraph id="8BF336A54F32721B21EFF9856AF0FB9F" blockId="6.[151,1437,1352,2025]" lastBlockId="9.[151,1437,921,1702]" lastPageId="9" lastPageNumber="234" pageId="6" pageNumber="231">
|
||
<emphasis id="B938EAB74F32721421EFF9856F16F95F" bold="true" box="[199,315,1640,1665]" pageId="6" pageNumber="231">Remarks.</emphasis>
|
||
The genus
|
||
<taxonomicName id="4C4C4D264F32721420E8F9856C56F95F" box="[448,635,1640,1665]" class="Actinopterygii" family="Nemacheilidae" genus="Eidinemacheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="231" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAB74F32721420E8F9856C56F95F" box="[448,635,1640,1665]" italics="true" pageId="6" pageNumber="231">Eidinemacheilus</emphasis>
|
||
</taxonomicName>
|
||
was recently introduced by Hashemzadeh Segherloo
|
||
<emphasis id="B938EAB74F32721425F4F9846B38F95F" box="[1244,1301,1640,1665]" italics="true" pageId="6" pageNumber="231">et al.</emphasis>
|
||
(2016) as a genus-group name for the troglomorphic
|
||
<emphasis id="B938EAB74F3272142351F9616D4DF97B" box="[633,864,1676,1701]" italics="true" pageId="6" pageNumber="231">
|
||
<taxonomicName id="4C4C4D264F3272142351F9616D71F97B" box="[633,860,1676,1701]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="231" phylum="Chordata" rank="species" species="smithi">Nemacheilus smithi</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
which is endemic to the Tigris drainage in
|
||
<collectingCountry id="F35B76354F3272142440F9606BB4F97A" box="[1384,1433,1677,1700]" name="Iran" pageId="6" pageNumber="231">Iran</collectingCountry>
|
||
.
|
||
<taxonomicName id="4C4C4D264F32721421BFF95D6F7FF917" box="[151,338,1712,1737]" class="Actinopterygii" family="Nemacheilidae" genus="Eidinemacheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="231" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAB74F32721421BFF95D6F7FF917" box="[151,338,1712,1737]" italics="true" pageId="6" pageNumber="231">Eidinemacheilus</emphasis>
|
||
</taxonomicName>
|
||
was distinguished from all genera of nemacheilid loaches in the Middle East by having 7+7 branched caudal-fin rays (vs. 9+8, rarely 8+8), the absence of eyes and pigmentation (vs. presence) and having a reduced set or no head canals (vs. all head canals complete or almost complete). Ideally, the diagnosis of a genus covers all included species and allows to distinguish it from all remaining genera. Nemacheilid loaches are a group of fishes where generic diagnoses are traditionally vague and unclear (
|
||
<bibRefCitation id="EFDD4B544F327214228EF8AD6AF7F887" author="Banarescu" box="[934,1242,1856,1881]" pageId="6" pageNumber="231" refString="Banarescu, P. M. & Nalbant, T. T. (1995) A generical classification of Nemacheilinae with description of two new genera (Teleostei: Cypriniformes: Cobitidae). Travaux du Museum d'Histoire Naturelle " Grigore Antipa ", 35, 429 - 495." type="journal article" year="1995">Bănărescu & Nalbant 1995</bibRefCitation>
|
||
, see also
|
||
<bibRefCitation id="EFDD4B544F327214246AF8AD6EFEF8A3" author="Kottelat" pageId="6" pageNumber="231" refString="Kottelat, M. (2012) Conspectus Cobitidum: An inventory of the loaches of the world (Teleostei: Cypriniformes: Cobitoidei). The Raffles Bulletin of Zoology Supplement, 26, 1 - 199." type="journal article" year="2012">Kottelat 2012</bibRefCitation>
|
||
), due to a limited set of useful characters as compared to other cypriniform families (
|
||
<bibRefCitation id="EFDD4B544F32721425C8F8896FB9F87E" author="Prokofiev" pageId="6" pageNumber="231" refString="Prokofiev, A. M. (2009) Problems of the classification and phylogeny of Nemacheiline loaches of the group lacking the preethmoid I (Cypriniformes: Balitoridae: Nemacheilinae). Journal of Ichthyology, 49, 874 - 898. http: // dx. doi. org / 10.1134 / s 0032945209100051" type="journal article" year="2009">Prokofiev 2009, Prokofiev pers. comm</bibRefCitation>
|
||
). In his detailed osteological study,
|
||
<bibRefCitation id="EFDD4B544F327214220CF8656DC9F87E" author="Prokofiev" box="[804,996,1928,1953]" pageId="6" pageNumber="231" refString="Prokofiev, A. M. (2009) Problems of the classification and phylogeny of Nemacheiline loaches of the group lacking the preethmoid I (Cypriniformes: Balitoridae: Nemacheilinae). Journal of Ichthyology, 49, 874 - 898. http: // dx. doi. org / 10.1134 / s 0032945209100051" type="journal article" year="2009">Prokofiev (2009)</bibRefCitation>
|
||
partly failed to recognise the genera of nemacheilid loaches from the Middle East as currently understood, and his results were only partly confirmed by molecular analyses (Freyhof
|
||
<emphasis id="B938EAB74F32721420F3F83C6C26F837" box="[475,523,2000,2025]" italics="true" pageId="6" pageNumber="231">et al</emphasis>
|
||
. 2014;
|
||
<bibRefCitation id="EFDD4B544F3272142377F83D6D14F837" author="Freyhof" box="[607,825,2000,2025]" pageId="6" pageNumber="231" refString="Freyhof, J., Sayyadzadeh, G., Esmaeili, H. R. & Geiger, M. (2015) Review of the genus Paraschistura from Iran with description of six new species (Teleostei: Nemacheilidae). Ichthyological Exploration of Freshwaters, 26, 1 - 48." type="journal article" year="2015">
|
||
Freyhof
|
||
<emphasis id="B938EAB74F3272142396F83C6CC2F837" box="[702,751,2000,2025]" italics="true" pageId="6" pageNumber="231">et al</emphasis>
|
||
. 2015
|
||
</bibRefCitation>
|
||
). Hashemzadeh Segherloo
|
||
<emphasis id="B938EAB74F327214254EF83C6ABBF837" box="[1126,1174,2000,2025]" italics="true" pageId="6" pageNumber="231">et al</emphasis>
|
||
. (2016) mentioned that all diagnostic characters for
|
||
<taxonomicName id="4C4C4D264F3D721B20F6FC746CB4FC6C" box="[478,665,921,946]" class="Actinopterygii" family="Nemacheilidae" genus="Eidinemacheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="234" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAB74F3D721B20F6FC746CB4FC6C" box="[478,665,921,946]" italics="true" pageId="9" pageNumber="234">Eidinemacheilus</emphasis>
|
||
</taxonomicName>
|
||
are, or could be somehow related to the adaptation of
|
||
<taxonomicName id="4C4C4D264F3D721B243EFC776B50FC6C" box="[1302,1405,921,946]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="234" phylum="Chordata" rank="species" species="smithi">
|
||
<emphasis id="B938EAB74F3D721B243EFC776B50FC6C" box="[1302,1405,921,946]" italics="true" pageId="9" pageNumber="234">E. smithi</emphasis>
|
||
</taxonomicName>
|
||
to subterranean life. Including
|
||
<taxonomicName id="4C4C4D264F3D721B20FDFC536C40FC0B" box="[469,621,956,981]" class="Actinopterygii" family="Nemacheilidae" genus="Eidinemacheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="234" phylum="Chordata" rank="species" species="proudlovei">
|
||
<emphasis id="B938EAB74F3D721B20FDFC536C40FC0B" box="[469,621,956,981]" italics="true" pageId="9" pageNumber="234">E. proudlovei</emphasis>
|
||
</taxonomicName>
|
||
in the same genus renders one character, the head canals (reduced in
|
||
<taxonomicName id="4C4C4D264F3D721B24ACFC536EF6FC24" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="234" phylum="Chordata" rank="species" species="smithi">
|
||
<emphasis id="B938EAB74F3D721B24ACFC536EF6FC24" italics="true" pageId="9" pageNumber="234">E. smithi</emphasis>
|
||
</taxonomicName>
|
||
, normally developed in
|
||
<taxonomicName id="4C4C4D264F3D721B232CFC0F6C8FFC24" box="[516,674,993,1018]" class="Actinopterygii" family="Nemacheilidae" genus="Eidinemacheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="234" phylum="Chordata" rank="species" species="proudlovei">
|
||
<emphasis id="B938EAB74F3D721B232CFC0F6C8FFC24" box="[516,674,993,1018]" italics="true" pageId="9" pageNumber="234">E. proudlovei</emphasis>
|
||
</taxonomicName>
|
||
) no longer diagnostic for
|
||
<taxonomicName id="4C4C4D264F3D721B22CCFC0C6AB2FC24" box="[996,1183,993,1018]" class="Actinopterygii" family="Nemacheilidae" genus="Eidinemacheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="234" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAB74F3D721B22CCFC0C6AB2FC24" box="[996,1183,993,1018]" italics="true" pageId="9" pageNumber="234">Eidinemacheilus</emphasis>
|
||
</taxonomicName>
|
||
. The low number of branched caudal-fin rays (7+
|
||
<quantity id="4CB49B404F3D721B20C9FBE86C23FBC0" box="[481,526,1029,1054]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="9" pageNumber="234" unit="in" value="7.0">7 in</quantity>
|
||
<taxonomicName id="4C4C4D264F3D721B233FFBEB6C53FBC0" box="[535,638,1029,1054]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="234" phylum="Chordata" rank="species" species="smithi">
|
||
<emphasis id="B938EAB74F3D721B233FFBEB6C53FBC0" box="[535,638,1029,1054]" italics="true" pageId="9" pageNumber="234">E. smithi</emphasis>
|
||
</taxonomicName>
|
||
, 8+7 or 8+
|
||
<quantity id="4CB49B404F3D721B222AFBE86D02FBC0" box="[770,815,1029,1054]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="9" pageNumber="234" unit="in" value="8.0">8 in</quantity>
|
||
<taxonomicName id="4C4C4D264F3D721B2210FBEB6DFFFBC0" box="[824,978,1029,1054]" class="Actinopterygii" family="Nemacheilidae" genus="Eidinemacheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="234" phylum="Chordata" rank="species" species="proudlovei">
|
||
<emphasis id="B938EAB74F3D721B2210FBEB6DFFFBC0" box="[824,978,1029,1054]" italics="true" pageId="9" pageNumber="234">E. proudlovei</emphasis>
|
||
</taxonomicName>
|
||
) and the lack of eyes and pigmentation remain diagnostic for
|
||
<taxonomicName id="4C4C4D264F3D721B20A5FBC46C65FB9C" box="[397,584,1065,1090]" class="Actinopterygii" family="Nemacheilidae" genus="Eidinemacheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="234" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAB74F3D721B20A5FBC46C65FB9C" box="[397,584,1065,1090]" italics="true" pageId="9" pageNumber="234">Eidinemacheilus</emphasis>
|
||
</taxonomicName>
|
||
, when compared to other nemacheilids in the Middle East.
|
||
</paragraph>
|
||
<caption id="DF33662D4F33721521BFF8436DA4F83D" httpUri="https://zenodo.org/record/159366/files/figure.png" pageId="7" pageNumber="232" targetBox="[222,1360,193,1930]" targetPageId="7">
|
||
<paragraph id="8BF336A54F33721521BFF8436DA4F83D" blockId="7.[151,1436,1966,2019]" pageId="7" pageNumber="232">
|
||
<emphasis id="B938EAB74F33721521BFF8436F3AF81A" bold="true" box="[151,279,1966,1988]" pageId="7" pageNumber="232">FIGURE 6.</emphasis>
|
||
<taxonomicName id="4C4C4D264F337215200AF8436C6EF81A" box="[290,579,1966,1988]" class="Actinopterygii" family="Nemacheilidae" genus="Eidinemacheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="232" phylum="Chordata" rank="species" species="proudlovei">
|
||
<emphasis id="B938EAB74F337215200AF8436C6EF81A" box="[290,579,1966,1988]" italics="true" pageId="7" pageNumber="232">Eidinemacheilus proudlovei</emphasis>
|
||
</taxonomicName>
|
||
, ZFMK-ICH 102318, holotype, 53.6 mm SL; lateral, and ventral view of head showing large mouth; Iraqi Kurdistan: Ephemeral spring in Tabeen Gorge.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF33662D4F3C721A21BFF84D6C82F82D" httpUri="https://zenodo.org/record/159367/files/figure.png" pageId="8" pageNumber="233" targetBox="[176,1406,193,1918]" targetPageId="8">
|
||
<paragraph id="8BF336A54F3C721A21BFF84D6C82F82D" blockId="8.[151,1436,1952,2035]" pageId="8" pageNumber="233">
|
||
<emphasis id="B938EAB74F3C721A21BFF84D6F3EF86B" bold="true" box="[151,275,1952,1974]" pageId="8" pageNumber="233">FIGURE 7.</emphasis>
|
||
<taxonomicName id="4C4C4D264F3C721A2033F84D6C15F868" box="[283,568,1952,1974]" class="Actinopterygii" family="Nemacheilidae" genus="Eidinemacheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="233" phylum="Chordata" rank="species" species="proudlovei">
|
||
<emphasis id="B938EAB74F3C721A2033F84D6C15F868" box="[283,568,1952,1974]" italics="true" pageId="8" pageNumber="233">Eidinemacheilus proudlovei</emphasis>
|
||
</taxonomicName>
|
||
, FSJF 3542, paratype, 48.5 SL; Iraqi Kurdistan: Ephemeral spring in Tabeen Gorge;
|
||
<taxonomicName id="4C4C4D264F3C721A21BFF82D6EDCF80A" box="[151,241,1982,2005]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="233" phylum="Chordata" rank="species" species="smithi">
|
||
<emphasis id="B938EAB74F3C721A21BFF82D6EDCF80A" box="[151,241,1982,2005]" italics="true" pageId="8" pageNumber="233">E. smithi</emphasis>
|
||
</taxonomicName>
|
||
, FSJF 3541, 34 mm SL; Iran: Natural well-like opening being the origin of the stream Kay-e-Ru; bony swim bladder capsule, dorsal, ventral, posterior and anterior views.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF33662D4F3D721B21BFFCD06F56FCAF" httpUri="https://zenodo.org/record/159368/files/figure.png" pageId="9" pageNumber="234" targetBox="[384,1203,193,807]" targetPageId="9">
|
||
<paragraph id="8BF336A54F3D721B21BFFCD06F56FCAF" blockId="9.[151,1435,829,881]" pageId="9" pageNumber="234">
|
||
<emphasis id="B938EAB74F3D721B21BFFCD06F3EFC8C" bold="true" box="[151,275,829,851]" pageId="9" pageNumber="234">FIGURE 8.</emphasis>
|
||
Ephemeral spring in Tabeen Gorge, type locality of
|
||
<taxonomicName id="4C4C4D264F3D721B221CFCD36D93FC8D" box="[820,958,829,851]" class="Actinopterygii" family="Nemacheilidae" genus="Eidinemacheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="234" phylum="Chordata" rank="species" species="proudlovei">
|
||
<emphasis id="B938EAB74F3D721B221CFCD36D93FC8D" box="[820,958,829,851]" italics="true" pageId="9" pageNumber="234">E. proudlovei</emphasis>
|
||
</taxonomicName>
|
||
. Korsh Ararat sitting at the head of the spring during a documentary.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF336A54F3D721B21EFFBA06D23FA13" blockId="9.[151,1437,921,1702]" pageId="9" pageNumber="234">
|
||
At the moment, we find it surprising that both subterranean nemacheilid loaches known so far apart within the Tigris drainage belong to the same genus. For now it can only be speculated that both species have evolved from a common (possibly subterranean) ancestor already a substantial time ago — several million years, if we assume a molecular clock-like behaviour of COI. Given that we only studied one mitochondrial marker (COI) we did not opt to perform an in-depth analysis of the absolute divergence time. In addition, there are no robust calibration points, such as geologic events, and rates of molecular evolution are expected to be heterogeneous among different lineages (e.g.
|
||
<bibRefCitation id="EFDD4B544F3D721B2015FAC86C09FAE0" author="Lavinia" box="[317,548,1317,1342]" pageId="9" pageNumber="234" refString="Lavinia, P. D., Kerr, K. C., Tubaro, P. L., Hebert, P. D. & Lijtmaer, D. A. (2016) Calibrating the molecular clock beyond cytochrome b: assessing the evolutionary rate of COI in birds. Journal of Avian Biology, 47, 84 - 91. http: // dx. doi. org / 10.1111 / jav. 00766" type="journal article" year="2016">
|
||
Lavinia
|
||
<emphasis id="B938EAB74F3D721B20B7FACB6FFBFAE0" box="[415,470,1317,1342]" italics="true" pageId="9" pageNumber="234">et al</emphasis>
|
||
. 2016
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFDD4B544F3D721B231DFAC86D27FAE3" author="Wilke" box="[565,778,1317,1342]" pageId="9" pageNumber="234" refString="Wilke, T., Schultheiß, R. & Albrecht, C. (2009) As time goes by: a simple fool's guide to molecular clock approaches in invertebrates. American Malacological Bulletin, 27, 25 - 45. http: // dx. doi. org / 10.4003 / 006.027.0203" type="journal article" year="2009">
|
||
Wilke
|
||
<emphasis id="B938EAB74F3D721B23ADFACB6C97FAE0" box="[645,698,1317,1342]" italics="true" pageId="9" pageNumber="234">et al</emphasis>
|
||
. 2009
|
||
</bibRefCitation>
|
||
), a problem especially given our heterogeneous sample composition. Yet (and despite the uncertainties of mitochondrial DNA molecular clock calibration) the highly divergent COI clusters (>8% K2P-distance) of the two
|
||
<taxonomicName id="4C4C4D264F3D721B220BFA806DF3FA58" box="[803,990,1389,1414]" class="Actinopterygii" family="Nemacheilidae" genus="Eidinemacheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="234" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EAB74F3D721B220BFA806DF3FA58" box="[803,990,1389,1414]" italics="true" pageId="9" pageNumber="234">Eidinemacheilus</emphasis>
|
||
</taxonomicName>
|
||
species are indicative of at least four million years of unique evolutionary history, using the rate proposed in several studies (
|
||
<bibRefCitation id="EFDD4B544F3D721B2544FA7C6B04FA77" author="April" box="[1132,1321,1425,1450]" pageId="9" pageNumber="234" refString="April, J., Mayden, R. L., Hanner, R. H. & Bernatchez, L. (2011) Genetic calibration of species diversity among North America's freshwater fishes. Proceedings of the National Academy of Sciences, 108, 10602 - 10607. http: // dx. doi. org / 10.1073 / pnas. 1016437108" type="journal article" year="2011">
|
||
April
|
||
<emphasis id="B938EAB74F3D721B2598FA7F6ACCFA74" box="[1200,1249,1425,1450]" italics="true" pageId="9" pageNumber="234">et al</emphasis>
|
||
. 2011
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFDD4B544F3D721B241DFA7F6EDBFA13" author="Brown" pageId="9" pageNumber="234" refString="Brown, W. M., George, M. & Wilson, A. C. (1979) Rapid evolution of animal mitochondrial DNA. Proceedings of the National Academy of Sciences, 76, 1967 - 1971. http: // dx. doi. org / 10.1073 / pnas. 76.4.1967" type="journal article" year="1979">
|
||
Brown
|
||
<emphasis id="B938EAB74F3D721B24A0FA7F6E80FA10" italics="true" pageId="9" pageNumber="234">et al</emphasis>
|
||
. 1979
|
||
</bibRefCitation>
|
||
; Durand
|
||
<emphasis id="B938EAB74F3D721B2074FA5B6FA0FA10" box="[348,397,1461,1486]" italics="true" pageId="9" pageNumber="234">et al</emphasis>
|
||
. 2003;
|
||
<bibRefCitation id="EFDD4B544F3D721B20CAFA586D2CFA13" author="Zardoya" box="[482,769,1461,1486]" pageId="9" pageNumber="234" refString="Zardoya, R. & Doadrio, I. (1999) Molecular evidence on the evolutionary and biogeographical patterns of European cyprinids. Journal of Molecular Evolution, 49, 227 - 237." type="journal article" year="1999">Zardoya & Doadrio 1999</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF336A54F3D721B21EFFA346CA5F9E4" blockId="9.[151,1437,921,1702]" pageId="9" pageNumber="234">
|
||
The circumstances of the discovery of
|
||
<taxonomicName id="4C4C4D264F3D721B23ADFA376D32FA2C" box="[645,799,1497,1522]" class="Actinopterygii" family="Nemacheilidae" genus="Eidinemacheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="234" phylum="Chordata" rank="species" species="proudlovei">
|
||
<emphasis id="B938EAB74F3D721B23ADFA376D32FA2C" box="[645,799,1497,1522]" italics="true" pageId="9" pageNumber="234">E. proudlovei</emphasis>
|
||
</taxonomicName>
|
||
, and the never before reported wash-out of fishes from this aquifer, raise the question how many other populations and species of subterranean fishes might still exist unearthed in the Middle East and elsewhere.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C356652E4F3D721B21EFF9A86CABF978" pageId="9" pageNumber="234" type="materials_examined">
|
||
<paragraph id="8BF336A54F3D721B21EFF9A86CABF978" blockId="9.[151,1437,921,1702]" pageId="9" pageNumber="234">
|
||
<materialsCitation id="3B243CF84F3D721B21EFF9A86CABF978" ID-GBIF-Occurrence="1316260496" collectionCode="FSJF" country="Iran" county="Natural" latitude="33.07722" location="Baq-e-Levan" longLatPrecision="20" longitude="48.593056" municipality="Sirom River" pageId="9" pageNumber="234" specimenCode="FSJF 3431, 1, FSJF 3541, 4" specimenCount="1" stateProvince="Lorestan">
|
||
<emphasis id="B938EAB74F3D721B21EFF9A86FFAF980" bold="true" box="[199,471,1605,1630]" pageId="9" pageNumber="234">Comparative material.</emphasis>
|
||
<taxonomicName id="4C4C4D264F3D721B20F5F9A86CCFF980" box="[477,738,1605,1630]" class="Actinopterygii" family="Nemacheilidae" genus="Eidinemacheilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="234" phylum="Chordata" rank="species" species="smithi">
|
||
<emphasis id="B938EAB74F3D721B20F5F9A86CCFF980" box="[477,738,1605,1630]" italics="true" pageId="9" pageNumber="234">Eidinemacheilus smithi</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<specimenCode id="DBEA9EDE4F3D721B23D9F9A86DA8F983" box="[753,901,1605,1629]" pageId="9" pageNumber="234">
|
||
<collectionCode id="ED5DAE604F3D721B23D9F9A86D07F983" box="[753,810,1605,1629]" pageId="9" pageNumber="234">FSJF</collectionCode>
|
||
3431, 1
|
||
</specimenCode>
|
||
,
|
||
<quantity id="4CB49B404F3D721B22BBF9A86DCFF983" box="[915,994,1605,1629]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="9" pageNumber="234" unit="mm" value="28.0">28 mm</quantity>
|
||
SL;
|
||
<specimenCode id="DBEA9EDE4F3D721B253FF9A86A80F983" box="[1047,1197,1605,1630]" pageId="9" pageNumber="234">
|
||
<collectionCode id="ED5DAE604F3D721B253FF9A86A7CF983" box="[1047,1105,1605,1629]" pageId="9" pageNumber="234">FSJF</collectionCode>
|
||
3541, 4
|
||
</specimenCode>
|
||
,
|
||
<quantity id="4CB49B404F3D721B2591F9A86B1FF983" box="[1209,1330,1605,1629]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" metricValueMax="3.8" metricValueMin="3.0" pageId="9" pageNumber="234" unit="mm" value="34.0" valueMax="38.0" valueMin="30.0">30–38 mm</quantity>
|
||
SL;
|
||
<collectingCountry id="F35B76354F3D721B244FF9AB6BB5F983" box="[1383,1432,1606,1629]" name="Iran" pageId="9" pageNumber="234">Iran</collectingCountry>
|
||
:
|
||
<collectingRegion id="4988F8474F3D721B21BFF9876ED4F95F" box="[151,249,1642,1665]" country="Iran" name="Lorestan" pageId="9" pageNumber="234">Lorestan</collectingRegion>
|
||
prov.:
|
||
<collectingCounty id="62924E294F3D721B2064F9846F8DF95C" box="[332,416,1641,1666]" pageId="9" pageNumber="234">Natural</collectingCounty>
|
||
well-like opening being the origin of the stream Kay-e-Ru flowing to
|
||
<collectingMunicipality id="6B97ACDF4F3D721B25F9F9846B72F95C" box="[1233,1375,1641,1666]" pageId="9" pageNumber="234">Sirom River</collectingMunicipality>
|
||
, near
|
||
<location id="8E93607E4F3D721B21BFF9636F04F97B" LSID="urn:lsid:plazi:treatment:03E587B34F37721B21BFFE5A6CABF978:8E93607E4F3D721B21BFF9636F04F97B" box="[151,297,1678,1701]" country="Iran" county="Natural" latitude="33.07722" longLatPrecision="20" longitude="48.593056" municipality="Sirom River" name="Baq-e-Levan" pageId="9" pageNumber="234" stateProvince="Lorestan">Baq-e-Levan</location>
|
||
oasis,
|
||
<geoCoordinate id="EE7850624F3D721B205EF9606FD5F97B" box="[374,504,1677,1701]" direction="north" orientation="latitude" pageId="9" pageNumber="234" precision="15" value="33.07722">33°04'38''N</geoCoordinate>
|
||
<geoCoordinate id="EE7850624F3D721B20D7F9606CAFF978" box="[511,642,1677,1702]" direction="east" orientation="longitude" pageId="9" pageNumber="234" precision="15" value="48.593056">48°35'35''E</geoCoordinate>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |