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<mods:title id="5250CC8AA9305CB56F4F7EE8432634EF">Taxonomy of the genus Diochus Erichson (Coleoptera: Staphylinidae, Staphylininae, Diochini) of Papua New Guinea, Vanuatu and Solomon Islands</mods:title>
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<mods:namePart id="C5180F628C905EA6C224E2D4433F8F39">Huang, Tian</mods:namePart>
<mods:affiliation id="32D6A7F9FBF05157B4CB424FBD73A69E">Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, 1 Beichen West Rd., Chaoyang District, Beijing 100101, P. R. China. &amp; University of the Chinese Academy of Sciences, 19 A Yuquan Rd., Shijingshan District, Beijing 100049, P. R. China.</mods:affiliation>
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<mods:namePart id="338A84ACFB58D53BA9756A6EC4A27FC3">Janák, Jiří</mods:namePart>
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<mods:namePart id="5762FFE8011C12E5D3221F9B58483AA0">Zhou, Hong-Zhang</mods:namePart>
<mods:affiliation id="C95C6DFFD26A3A158F1645E02D078601">Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, 1 Beichen West Rd., Chaoyang District, Beijing 100101, P. R. China. &amp; University of the Chinese Academy of Sciences, 19 A Yuquan Rd., Shijingshan District, Beijing 100049, P. R. China.</mods:affiliation>
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<paragraph id="8BF336B5FFF5FFD8B9C088A8FA25FAB2" blockId="1.[151,1431,1700,1727]" box="[151,1431,1700,1727]" pageId="1" pageNumber="104">
<heading id="D0BB81D9FFF5FFD8B9C088A8FA25FAB2" bold="true" box="[151,1431,1700,1727]" centered="true" fontSize="11" level="1" pageId="1" pageNumber="104" reason="1">
<emphasis id="B938EAA7FFF5FFD8B9C088A8FA25FAB2" bold="true" box="[151,1431,1700,1727]" pageId="1" pageNumber="104">
Key to species of the genus
<taxonomicName id="4C4C4D36FFF5FFD8B8BB88A8FDE3FAB3" ID-CoL="8VRMP" authority="Erichson, 1839" box="[492,593,1700,1727]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="104" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAA7FFF5FFD8B8BB88A8FDE3FAB3" bold="true" box="[492,593,1700,1727]" italics="true" pageId="1" pageNumber="104">Diochus</emphasis>
</taxonomicName>
from
<collectingCountry id="F35B7625FFF5FFD8BBCA88A8FC22FAB2" box="[669,912,1700,1726]" name="Papua New Guinea" pageId="1" pageNumber="104">Papua New Guinea</collectingCountry>
,
<collectingCountry id="F35B7625FFF5FFD8BACB88A8FBB5FAB2" box="[924,1031,1700,1726]" name="Vanuatu" pageId="1" pageNumber="104">Vanuatu</collectingCountry>
and
<collectingCountry id="F35B7625FFF5FFD8BD1288A8FAA1FAB2" box="[1093,1299,1700,1726]" name="Solomon Islands" pageId="1" pageNumber="104">Solomon Islands</collectingCountry>
(
<figureCitation id="13772A30FFF5FFD8BC7588A8FA3CFAB2" box="[1314,1422,1700,1726]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2-2" captionStart-2="FIGURE 5" captionStartId-0="2.[151,250,1874,1898]" captionStartId-1="4.[151,250,1943,1967]" captionStartId-2="13.[151,250,1874,1898]" captionTargetBox-0="[154,1432,744,1846]" captionTargetBox-1="[164,1423,184,1913]" captionTargetBox-2="[161,1426,535,1841]" captionTargetId-0="figure-232@2.[151,1435,741,1850]" captionTargetId-1="figure-23@4.[162,1425,181,1919]" captionTargetId-2="figure-147@13.[151,1436,525,1851]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="13" captionText-0="FIGURE 1. Habitus of four species from Papua New Guinea, Vanuatu and Solomon Islands in dorsal view:A. Diochus elongatus sp. nov.; B. D. chambrieri sp. nov.; C. D. papuanus Cameron, 1939, D. D. loebli sp. nov. (Scale bar: 0.5 mm)." captionText-1="FIGURE 2-2. Diochus elongatus sp. nov.: A. aedeagus, dorsal view; B. male sternite VIII; C. male tergite VIII; D. male tergites IXX; E. male sternite IX; F. female sternite VIII; G. female sternite X; H. female sternite IX; I. spermatheca.(Scale bars: A = 0.05 mm, BI = 0.1 mm)." captionText-2="FIGURE 5. Diochus papuanus Cameron, 1939:A. forebody, dorsal view; B. antenna; C. male sternite VIII; D. male tergite VIII; E. male sternite IX; F. male tergites IXX; G. aedeagus, dorsal view; H. labels. (Scale bars: A = 0.5 mm, BG = 0.1 mm)." figureDoi-0="http://doi.org/10.5281/zenodo.13915581" figureDoi-1="http://doi.org/10.5281/zenodo.13915585" figureDoi-2="http://doi.org/10.5281/zenodo.13915598" httpUri-0="https://zenodo.org/record/13915581/files/figure.png" httpUri-1="https://zenodo.org/record/13915585/files/figure.png" httpUri-2="https://zenodo.org/record/13915598/files/figure.png" pageId="1" pageNumber="104">Figs 15</figureCitation>
)
</emphasis>
</heading>
</paragraph>
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1 Elytra extremely short, one sixth shorter than pronotum (
<figureCitation id="13772A30FFF5FFD8BA5388EFFC85FAF4" box="[772,823,1762,1785]" captionStart="FIGURE 5" captionStartId="13.[151,250,1874,1898]" captionTargetBox="[161,1426,535,1841]" captionTargetId="figure-147@13.[151,1436,525,1851]" captionTargetPageId="13" captionText="FIGURE 5. Diochus papuanus Cameron, 1939:A. forebody, dorsal view; B. antenna; C. male sternite VIII; D. male tergite VIII; E. male sternite IX; F. male tergites IXX; G. aedeagus, dorsal view; H. labels. (Scale bars: A = 0.5 mm, BG = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.13915598" httpUri="https://zenodo.org/record/13915598/files/figure.png" pageId="1" pageNumber="104">Fig 5</figureCitation>
)...........................
<taxonomicName id="4C4C4D36FFF5FFD8BDDF88EFFA2EFAF4" ID-CoL="369ZD" authority="Cameron, 1939" authorityName="Cameron" authorityYear="1939" box="[1160,1436,1763,1784]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="104" phylum="Arthropoda" rank="species" species="papuanus">
<emphasis id="B938EAA7FFF5FFD8BDDF88EFFAB0FAF4" box="[1160,1282,1763,1784]" italics="true" pageId="1" pageNumber="104">D. papuanus</emphasis>
<bibRefCitation id="EFDD4B44FFF5FFD8BC5F88EFFA2EFAF4" author="Cameron, M." box="[1288,1436,1763,1784]" pageId="1" pageNumber="104" pagination="139 - 153" refId="ref8513" refString="Cameron, M. (1939) New Staphylinidae (Col.) from New Guinea. The Annals and Magazine of Natural History, Series 11, 3 (13), 139 - 153. https: // doi. org / 10.1080 / 03745481.1939.9723583" type="journal article" year="1939">Cameron, 1939</bibRefCitation>
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</paragraph>
<paragraph id="8BF336B5FFF5FFD8B9C0890CFA2EFB19" blockId="1.[151,1437,1762,2019]" box="[151,1436,1792,1814]" pageId="1" pageNumber="104">
<keyLead id="30BD9580FFF5FFD8B9C0890CFA2EFB19" box="[151,1436,1792,1814]" pageId="1" pageNumber="104">- Elytra of normal length, about as long as pronotum.......................................................... 2</keyLead>
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<paragraph id="8BF336B5FFF5FFD8B9C08911FA2EFB5C" blockId="1.[151,1437,1762,2019]" pageId="1" pageNumber="104">
<keyLead id="30BD9580FFF5FFD8B9C08911FA2EFB5C" pageId="1" pageNumber="104">
2 Antennomeres very long; antennomere 3 distinctly elongate, about 5 times longer than wide; body length
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(
<figureCitation id="13772A30FFF5FFD8BC0C8911FA22FB3E" box="[1371,1424,1821,1843]" captionStart="FIGURE 2-2" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[164,1423,184,1913]" captionTargetId="figure-23@4.[162,1425,181,1919]" captionTargetPageId="4" captionText="FIGURE 2-2. Diochus elongatus sp. nov.: A. aedeagus, dorsal view; B. male sternite VIII; C. male tergite VIII; D. male tergites IXX; E. male sternite IX; F. female sternite VIII; G. female sternite X; H. female sternite IX; I. spermatheca.(Scale bars: A = 0.05 mm, BI = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.13915585" httpUri="https://zenodo.org/record/13915585/files/figure.png" pageId="1" pageNumber="104">Fig 2</figureCitation>
)...................................................................................
<taxonomicName id="4C4C4D36FFF5FFD8BD988937FAFBFB5C" authorityName="Huang &amp; Janák &amp; Zhou" authorityYear="2024" box="[1231,1353,1851,1872]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="104" phylum="Arthropoda" rank="species" species="elongatus" status="sp. nov.">
<emphasis id="B938EAA7FFF5FFD8BD988937FAFBFB5C" box="[1231,1353,1851,1872]" italics="true" pageId="1" pageNumber="104">D. elongatus</emphasis>
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<taxonomicNameLabel id="A20B57DCFFF5FFD8BC078937FA2EFB5C" box="[1360,1436,1851,1872]" pageId="1" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BF336B5FFF5FFD8B9C08954FA2EFB61" blockId="1.[151,1437,1762,2019]" box="[151,1436,1880,1902]" pageId="1" pageNumber="104">
<keyLead id="30BD9580FFF5FFD8B9C08954FA2EFB61" box="[151,1436,1880,1902]" pageId="1" pageNumber="104">
- Antennomeres shorter; antennomere 3 slightly elongate, about 2 to 4 times longer than wide; body length less than
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.. 3
</keyLead>
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<keyStep id="30B82E10FFF5FFD8B9C08979FA2EFBEE" pageId="1" pageNumber="104">
<paragraph id="8BF336B5FFF5FFD8B9C08979FA2EFBA4" blockId="1.[151,1437,1762,2019]" pageId="1" pageNumber="104">
<keyLead id="30BD9580FFF5FFD8B9C08979FA2EFBA4" pageId="1" pageNumber="104">
3 Antennomere 3 more elongate, distinctly longer than antennomere 2; head narrower at posterior angle; tube of spermatheca shorter, slightly coiled (Fig 3)...............................................................
<taxonomicName id="4C4C4D36FFF5FFD8BDA7899FFAF6FBA4" authorityName="Huang &amp; Janák &amp; Zhou" authorityYear="2024" box="[1264,1348,1939,1960]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="104" phylum="Arthropoda" rank="species" species="loebli" status="sp. nov.">
<emphasis id="B938EAA7FFF5FFD8BDA7899FFAF6FBA4" box="[1264,1348,1939,1960]" italics="true" pageId="1" pageNumber="104">D. loebli</emphasis>
</taxonomicName>
,
<emphasis id="B938EAA7FFF5FFD8BC07899FFA2EFBA4" bold="true" box="[1360,1436,1939,1960]" pageId="1" pageNumber="104">
<taxonomicNameLabel id="A20B57DCFFF5FFD8BC07899FFA2EFBA4" box="[1360,1436,1939,1960]" pageId="1" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BF336B5FFF5FFD8B9C089BCFA2EFBEE" blockId="1.[151,1437,1762,2019]" pageId="1" pageNumber="104">
<keyLead id="30BD9580FFF5FFD8B9C089BCFA2EFBEE" pageId="1" pageNumber="104">
- Antennomere 3 less elongate, nearly equal to antennomere 2; head wider at posterior angle; tube of spermatheca longer, strongly coiled (Fig 4)................................................................
<taxonomicName id="4C4C4D36FFF5FFD8BD9589C2FAF8FBEE" authorityName="Huang &amp; Janák &amp; Zhou" authorityYear="2024" box="[1218,1354,1997,2019]" class="Insecta" family="Staphylinidae" genus="Diochus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="104" phylum="Arthropoda" rank="species" species="chambrieri" status="sp. nov.">
<emphasis id="B938EAA7FFF5FFD8BD9589C2FAF8FBEE" box="[1218,1354,1997,2019]" italics="true" pageId="1" pageNumber="104">D. chambrieri</emphasis>
</taxonomicName>
<emphasis id="B938EAA7FFF5FFD8BC0789C1FA2EFBEE" bold="true" box="[1360,1436,1997,2018]" pageId="1" pageNumber="104">
<taxonomicNameLabel id="A20B57DCFFF5FFD8BC0789C1FA2EFBEE" box="[1360,1436,1997,2018]" pageId="1" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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