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<document id="2FD642BC5889666C93AD8015E27169E2" ID-DOI="10.11646/zootaxa.2631.1.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="5304047" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629700474853" checkinUser="felipe" docAuthor="Mahunka, Sándor" docDate="2010" docId="03E587998C254B60FF32FAA6FCC3FE9C" docLanguage="en" docName="zt02631p068.pdf" docOrigin="Zootaxa 2631 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2631.1.5" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Hymenobelba exclamationis Mahunka 2010, sp. nov." docType="treatment" docVersion="4" lastPageNumber="65" masterDocId="FFDCFFE18C244B63FFA5FF9CFFF8FFDB" masterDocTitle="Two new species of Ameroidea (Acari: Oribatida) from Madagascar" masterLastPageNumber="68" masterPageNumber="62" pageNumber="63" updateTime="1699097852112" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="8340C4D8A76019BA4F7DCDA9365F9F7F">Two new species of Ameroidea (Acari: Oribatida) from Madagascar</mods:title>
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<mods:namePart id="32EB83641E1F59246D23CF708BA96114">Mahunka, Sándor</mods:namePart>
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<mods:date id="CECF36B495150C0A5BEE11219142B4FE">2010</mods:date>
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<treatment id="03E587998C254B60FF32FAA6FCC3FE9C" ID-DOI="http://doi.org/10.5281/zenodo.5311044" ID-Zenodo-Dep="5311044" LSID="urn:lsid:plazi:treatment:03E587998C254B60FF32FAA6FCC3FE9C" httpUri="http://treatment.plazi.org/id/03E587998C254B60FF32FAA6FCC3FE9C" lastPageId="3" lastPageNumber="65" pageId="1" pageNumber="63">
<subSubSection id="C35665048C254B62FF32FAA6FDAAFA8F" box="[151,594,1338,1364]" pageId="1" pageNumber="63" type="nomenclature">
<paragraph id="8BF3368F8C254B62FF32FAA6FDAAFA8F" blockId="1.[151,594,1338,1402]" box="[151,594,1338,1364]" pageId="1" pageNumber="63">
<heading id="D0BB81E38C254B62FF32FAA6FDAAFA8F" bold="true" box="[151,594,1338,1364]" fontSize="11" level="1" pageId="1" pageNumber="63" reason="1">
<taxonomicName id="4C4C4D0C8C254B62FF32FAA6FE15FA8F" ID-CoL="8TC8G" authority="Mahunka, 2010" authorityName="Mahunka" authorityYear="2010" box="[151,493,1338,1364]" class="Arachnida" family="Ameridae" genus="Hymenobelba" kingdom="Animalia" order="Sarcoptiformes" pageId="1" pageNumber="63" phylum="Arthropoda" rank="species" species="exclamationis" status="sp. nov.">Hymenobelba exclamationis</taxonomicName>
<taxonomicNameLabel id="A20B57E68C254B62FE50FAA6FDAAFA8F" box="[501,594,1338,1364]" pageId="1" pageNumber="63" rank="species">
<emphasis id="B938EA9D8C254B62FE50FAA6FDB3FA8F" bold="true" box="[501,587,1338,1364]" pageId="1" pageNumber="63">sp. nov</emphasis>
.
</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35665048C254B62FF32FAFCFEEFFAA1" box="[151,279,1376,1402]" pageId="1" pageNumber="63" type="description">
<paragraph id="8BF3368F8C254B62FF32FAFCFEEFFAA1" blockId="1.[151,594,1338,1402]" box="[151,279,1376,1402]" pageId="1" pageNumber="63">
(
<figureCitation id="13772A0A8C254B62FF3AFAFCFEF6FAA1" box="[159,270,1376,1402]" captionStart="FIGURES 16" captionStartId="2.[151,271,1760,1784]" captionTargetBox="[305,1278,681,1731]" captionTargetId="figure-326@2.[302,1284,662,1752]" captionTargetPageId="2" captionText="FIGURES 16. Hymenobelba exclamationis n. sp.: 1. body in dorsal view; 2. body in ventral view; 3. epimer 3 with enantiophysis S; 4. anteromedian part of prodorsum; 5. claw of leg I; 6. claw of leg IV." figureDoi="http://doi.org/10.5281/zenodo.5304049" httpUri="https://zenodo.org/record/5304049/files/figure.png" pageId="1" pageNumber="63">Figs. 16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35665048C254B62FF32FA36FE16F9A4" pageId="1" pageNumber="63" type="diagnosis">
<paragraph id="8BF3368F8C254B62FF32FA36FE16F9A4" blockId="1.[151,1436,1450,2036]" pageId="1" pageNumber="63">
<emphasis id="B938EA9D8C254B62FF32FA36FEEAFA1F" bold="true" box="[151,274,1450,1476]" pageId="1" pageNumber="63">Diagnosis</emphasis>
. Body and legs partly covered by cerotegument. Prodorsum with longitudinal unpaired costula, bearing lamellar setae. Interlamellar setae long, arising near to each other. Sensillus very long, finely ciliate. All ten pairs of notogastral setae thin, long. Apodemes and epimeral borders short, partly reduced. Enantiophysis
<emphasis id="B938EA9D8C254B62FEE2F987FEAEF9EF" box="[327,342,1563,1588]" italics="true" pageId="1" pageNumber="63">S</emphasis>
present. Epimeral setae varying in length, setae
<emphasis id="B938EA9D8C254B62FC3EF987FC40F9EF" box="[923,952,1563,1588]" italics="true" pageId="1" pageNumber="63">1b</emphasis>
and
<emphasis id="B938EA9D8C254B62FC56F987FBE8F9EF" box="[1011,1040,1563,1588]" italics="true" pageId="1" pageNumber="63">1c</emphasis>
longest of all, directed forwards. Strong ventral neotrichy, some setae in the aggenital region dilated, others long, setiform. Claw of leg I long, much thinner than the others.
</paragraph>
</subSubSection>
<subSubSection id="C35665048C254B61FF63F916FE6EFDAF" lastPageId="2" lastPageNumber="64" pageId="1" pageNumber="63" type="description">
<paragraph id="8BF3368F8C254B62FF63F916FBBBF97F" blockId="1.[151,1436,1450,2036]" box="[198,1091,1674,1700]" pageId="1" pageNumber="63">
<emphasis id="B938EA9D8C254B62FF63F916FE7DF97F" bold="true" box="[198,389,1674,1700]" pageId="1" pageNumber="63">Measurements.</emphasis>
Length of body: 818900 µm, width of body: 415510 µm.
</paragraph>
<paragraph id="8BF3368F8C254B62FF63F92CFCFEF871" blockId="1.[151,1436,1450,2036]" pageId="1" pageNumber="63">
<emphasis id="B938EA9D8C254B62FF63F92CFEA9F912" box="[198,337,1712,1737]" italics="true" pageId="1" pageNumber="63">Prodorsum.</emphasis>
Rostral part narrow, rostrum conical with a pair of weak longitudinal laths and a pair of well observable lateral tubercles bearing rostral setae. Prodosum with an unpaired, long, distally dilated, strong lath (
<figureCitation id="13772A0A8C254B62FF77F966FEE2F8CF" box="[210,282,1786,1812]" captionStart="FIGURES 16" captionStartId="2.[151,271,1760,1784]" captionTargetBox="[305,1278,681,1731]" captionTargetId="figure-326@2.[302,1284,662,1752]" captionTargetPageId="2" captionText="FIGURES 16. Hymenobelba exclamationis n. sp.: 1. body in dorsal view; 2. body in ventral view; 3. epimer 3 with enantiophysis S; 4. anteromedian part of prodorsum; 5. claw of leg I; 6. claw of leg IV." figureDoi="http://doi.org/10.5281/zenodo.5304049" httpUri="https://zenodo.org/record/5304049/files/figure.png" pageId="1" pageNumber="63">Fig. 4</figureCitation>
), which possesses a rounded tubercle behind as a longitudinal mark of exclamation-shape. All four pairs of prodorsal setae long, thin, setiform, lamellar setae arising on the distal end of costuliform crest. Interlamellar setae inserted on small tubercles, very near to one another. All setae long, their length:
<emphasis id="B938EA9D8C254B62FA92F8D9FF09F85F" italics="true" pageId="1" pageNumber="63">le =ro&gt; in&gt; ex.</emphasis>
A weak transversal line in interbothridial region present. Bothridium cup-shaped, sensillus very long, setiform, curving slightly distally, unilaterally ciliate.
</paragraph>
<paragraph id="8BF3368F8C254B61FF60F829FC81FF01" blockId="1.[151,1436,1450,2036]" lastBlockId="2.[151,1436,152,628]" lastPageId="2" lastPageNumber="64" pageId="1" pageNumber="63">
<emphasis id="B938EA9D8C254B62FF60F829FEB5F815" box="[197,333,1973,1998]" italics="true" pageId="1" pageNumber="63">Notogaster.</emphasis>
Anterior margin of notogaster slightly convex, with a pair of very small humeral processes. Notogastral surface without lateral cavities. Ten pairs of notogastral setae, all filiform, mostly with bent or slightly flagellate distal end (
<figureCitation id="13772A0A8C264B61FDABFF04FDA2FF69" box="[526,602,152,178]" captionStart="FIGURES 16" captionStartId="2.[151,271,1760,1784]" captionTargetBox="[305,1278,681,1731]" captionTargetId="figure-326@2.[302,1284,662,1752]" captionTargetPageId="2" captionText="FIGURES 16. Hymenobelba exclamationis n. sp.: 1. body in dorsal view; 2. body in ventral view; 3. epimer 3 with enantiophysis S; 4. anteromedian part of prodorsum; 5. claw of leg I; 6. claw of leg IV." figureDoi="http://doi.org/10.5281/zenodo.5304049" httpUri="https://zenodo.org/record/5304049/files/figure.png" pageId="2" pageNumber="64">Fig. 1</figureCitation>
). Setae
<emphasis id="B938EA9D8C264B61FD61FF04FD22FF6D" box="[708,730,152,182]" italics="true" pageId="2" pageNumber="64">
c
<subScript id="17C834CA8C264B61FD77FF34FD22FF6D" attach="left" box="[722,730,168,182]" fontSize="6" pageId="2" pageNumber="64">2</subScript>
</emphasis>
except setae
<emphasis id="B938EA9D8C264B61FC2CFF04FC4EFF6A" box="[905,950,152,177]" italics="true" pageId="2" pageNumber="64">p m</emphasis>
uch shorter than the notogastral setae. Lyrifissures
<emphasis id="B938EA9D8C264B61FE8DFF5CFEBDFF02" box="[296,325,192,217]" italics="true" pageId="2" pageNumber="64">im</emphasis>
located longitudinally, between setae
<emphasis id="B938EA9D8C264B61FCA0FF5CFCE4FF02" box="[773,796,192,217]" italics="true" pageId="2" pageNumber="64">la</emphasis>
and
<emphasis id="B938EA9D8C264B61FCF0FF5CFC8AFF02" box="[853,882,192,217]" italics="true" pageId="2" pageNumber="64">lm</emphasis>
.
</paragraph>
<paragraph id="8BF3368F8C264B61FF60FF7AFC04FE91" blockId="2.[151,1436,152,628]" pageId="2" pageNumber="64">
<emphasis id="B938EA9D8C264B61FF60FF7AFE06FF24" box="[197,510,230,255]" italics="true" pageId="2" pageNumber="64">Lateral part of podosoma.</emphasis>
Prodorsum characteristically excavate behind the rostrum and in middle part. Pedotecta I very large, rounded distally. Tutorium absent. Pedotecta II very small, triangular. Enatiophysis
<emphasis id="B938EA9D8C264B61FA28FE97FA64FEFF" box="[1421,1436,267,292]" italics="true" pageId="2" pageNumber="64">S</emphasis>
large, triangular, its distal part bent outwards. Setae
<emphasis id="B938EA9D8C264B61FD5DFEACFCECFE92" box="[760,788,304,329]" italics="true" pageId="2" pageNumber="64">3c</emphasis>
arising on its basis.
</paragraph>
<paragraph id="8BF3368F8C264B61FF60FECAFA97FDF1" blockId="2.[151,1436,152,628]" pageId="2" pageNumber="64">
<emphasis id="B938EA9D8C264B61FF60FECAFE92FEB4" box="[197,362,342,367]" italics="true" pageId="2" pageNumber="64">Ventral parts.</emphasis>
Epimeral region weakly sclerotised, without well developed structure. Only sejugal and apodemes IV visible, sejugal one strongest of all. Sternal apodeme absent. Length of setae highly varying, setae
<emphasis id="B938EA9D8C264B61FF7DFE3CFF0DFE62" box="[216,245,416,441]" italics="true" pageId="2" pageNumber="64">1b</emphasis>
and
<emphasis id="B938EA9D8C264B61FE8BFE3CFEB1FE62" box="[302,329,416,441]" italics="true" pageId="2" pageNumber="64">1c</emphasis>
longest of all, setae
<emphasis id="B938EA9D8C264B61FD9CFE3CFDAEFE62" box="[569,598,416,441]" italics="true" pageId="2" pageNumber="64">1a</emphasis>
and
<emphasis id="B938EA9D8C264B61FD2BFE3CFD54FE62" box="[654,684,416,441]" italics="true" pageId="2" pageNumber="64">2a</emphasis>
shortest of all. Setae
<emphasis id="B938EA9D8C264B61FC03FE3CFC3AFE62" box="[934,962,416,441]" italics="true" pageId="2" pageNumber="64">3c</emphasis>
arising on the enatiophysis (
<figureCitation id="13772A0A8C264B61FAB4FE3CFAA0FE61" box="[1297,1368,416,442]" captionStart="FIGURES 16" captionStartId="2.[151,271,1760,1784]" captionTargetBox="[305,1278,681,1731]" captionTargetId="figure-326@2.[302,1284,662,1752]" captionTargetPageId="2" captionText="FIGURES 16. Hymenobelba exclamationis n. sp.: 1. body in dorsal view; 2. body in ventral view; 3. epimer 3 with enantiophysis S; 4. anteromedian part of prodorsum; 5. claw of leg I; 6. claw of leg IV." figureDoi="http://doi.org/10.5281/zenodo.5304049" httpUri="https://zenodo.org/record/5304049/files/figure.png" pageId="2" pageNumber="64">Fig. 3</figureCitation>
), seta
<emphasis id="B938EA9D8C264B61FF32FE5AFF4CFE04" box="[151,180,454,479]" italics="true" pageId="2" pageNumber="64">4b</emphasis>
and
<emphasis id="B938EA9D8C264B61FF54FE5AFEF5FE04" box="[241,269,454,479]" italics="true" pageId="2" pageNumber="64">4c</emphasis>
located near to each other. Epimeral setal formula: 3-1-3-3. Genital setae arising in longitudinal rows. Aggenital and adanal neotrichy present. Some setae in median part of ventral plate dilated basally. Anal setae arising medially on two weak longitudinal crests. Lyrifissures
<emphasis id="B938EA9D8C264B61FC16FD8CFC20FDF2" box="[947,984,528,553]" italics="true" pageId="2" pageNumber="64">iad</emphasis>
located far from the anal aperture.
</paragraph>
<paragraph id="8BF3368F8C264B61FF60FDAAFE6EFDAF" blockId="2.[151,1436,152,628]" pageId="2" pageNumber="64">
<emphasis id="B938EA9D8C264B61FF60FDAAFEFCFD94" box="[197,260,566,591]" italics="true" pageId="2" pageNumber="64">Legs.</emphasis>
All legs conspicuously thin and long, monodactylous. Claws on legs I thinner and smaller than on legs IIIV (
<figureCitation id="13772A0A8C264B61FEBBFDC6FE72FDAF" box="[286,394,602,628]" captionStart="FIGURES 16" captionStartId="2.[151,271,1760,1784]" captionTargetBox="[305,1278,681,1731]" captionTargetId="figure-326@2.[302,1284,662,1752]" captionTargetPageId="2" captionText="FIGURES 16. Hymenobelba exclamationis n. sp.: 1. body in dorsal view; 2. body in ventral view; 3. epimer 3 with enantiophysis S; 4. anteromedian part of prodorsum; 5. claw of leg I; 6. claw of leg IV." figureDoi="http://doi.org/10.5281/zenodo.5304049" httpUri="https://zenodo.org/record/5304049/files/figure.png" pageId="2" pageNumber="64">Figs 56</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF3366078C264B61FF32F97CFBCBF8CC" ID-DOI="http://doi.org/10.5281/zenodo.5304049" ID-Zenodo-Dep="5304049" httpUri="https://zenodo.org/record/5304049/files/figure.png" pageId="2" pageNumber="64" startId="2.[151,271,1760,1784]" targetBox="[305,1278,681,1731]" targetPageId="2">
<paragraph id="8BF3368F8C264B61FF32F97CFBCBF8CC" blockId="2.[151,1436,1760,1816]" pageId="2" pageNumber="64">
<emphasis id="B938EA9D8C264B61FF32F97CFEBEF923" bold="true" box="[151,326,1760,1784]" pageId="2" pageNumber="64">FIGURES 16.</emphasis>
<taxonomicName id="4C4C4D0C8C264B61FEE8F97DFD7AF923" authority="Mahunka, 2010" authorityName="Mahunka" authorityYear="2010" box="[333,642,1761,1784]" class="Arachnida" family="Ameridae" genus="Hymenobelba" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="64" phylum="Arthropoda" rank="species" species="exclamationis" status="sp. nov.">
<emphasis id="B938EA9D8C264B61FEE8F97DFD7AF923" box="[333,642,1761,1784]" italics="true" pageId="2" pageNumber="64">Hymenobelba exclamationis</emphasis>
</taxonomicName>
<emphasis id="B938EA9D8C264B61FD2FF97CFD30F923" bold="true" box="[650,712,1760,1784]" pageId="2" pageNumber="64">
<taxonomicNameLabel id="A20B57E68C264B61FD2FF97CFD30F923" box="[650,712,1760,1784]" pageId="2" pageNumber="64" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: 1. body in dorsal view; 2. body in ventral view; 3. epimer 3 with enantiophysis
<emphasis id="B938EA9D8C264B61FE95F89DFEC5F8C3" box="[304,317,1793,1816]" italics="true" pageId="2" pageNumber="64">S</emphasis>
; 4. anteromedian part of prodorsum; 5. claw of leg I; 6. claw of leg IV.
</paragraph>
</caption>
<subSubSection id="C35665048C264B61FF60F8DAFD91F82E" pageId="2" pageNumber="64" type="discussion">
<paragraph id="8BF3368F8C264B61FF60F8DAFD91F82E" blockId="2.[151,1436,1862,2037]" pageId="2" pageNumber="64">
<emphasis id="B938EA9D8C264B61FF60F8DAFEC7F8BB" bold="true" box="[197,319,1862,1888]" pageId="2" pageNumber="64">Remarks.</emphasis>
The new species is well characterised by the presence of the prodorsal sculpture, by the form and length of the prodorsal and notogastral setae, by the distance of the lamellar and interlamellar ones and finally by the absence of the notogastral cavity. Considering most of these features
<taxonomicName id="4C4C4D0C8C264B61FB4EF80DFEC2F814" authority="Mahunka, 2010" authorityName="Mahunka" authorityYear="2010" class="Arachnida" family="Ameridae" genus="Hymenobelba" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="64" phylum="Arthropoda" rank="species" species="exclamationis" status="sp. nov.">
<emphasis id="B938EA9D8C264B61FB4EF80DFEC2F814" italics="true" pageId="2" pageNumber="64">Hymenobelba exclamationis</emphasis>
</taxonomicName>
<emphasis id="B938EA9D8C264B61FEE7F82AFE58F80B" bold="true" box="[322,416,1974,2000]" pageId="2" pageNumber="64">
<taxonomicNameLabel id="A20B57E68C264B61FEE7F82AFE58F80B" box="[322,416,1974,2000]" pageId="2" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
stand nearest to
<taxonomicName id="4C4C4D0C8C264B61FDCCF82AFC71F80B" authority="Balogh, 1962" authorityName="Balogh" authorityYear="1962" box="[617,905,1974,2000]" class="Arachnida" family="Ameridae" genus="Hymenobelba" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="64" phylum="Arthropoda" rank="species" species="ypsilon">
<emphasis id="B938EA9D8C264B61FDCCF82AFD1BF814" box="[617,739,1974,1999]" italics="true" pageId="2" pageNumber="64">H. ypsilon</emphasis>
Balogh, 1962
</taxonomicName>
, however, the prodorsal structure of the new species is much simpler than the others.
</paragraph>
</subSubSection>
<subSubSection id="C35665048C274B60FF60FF0BFC2EFF27" pageId="3" pageNumber="65" type="materials_examined">
<paragraph id="8BF3368F8C274B60FF60FF0BFC2EFF27" blockId="3.[151,1437,151,327]" pageId="3" pageNumber="65">
<emphasis id="B938EA9D8C274B60FF60FF0BFE46FF6A" bold="true" box="[197,446,151,177]" pageId="3" pageNumber="65">Material examined.</emphasis>
<materialsCitation id="3B243CD28C274B60FE62FF04FBDDFF0C" collectingDate="1998-08-26" collectionCode="T" collectorName="Leg. Dr. &amp; Pocs" country="Madagascar" location="Maromizaha forest" pageId="3" pageNumber="65" specimenCount="2" stateProvince="Toamasina" typeStatus="holotype">
<typeStatus id="54F7882D8C274B60FE62FF04FDC3FF69" box="[455,571,152,178]" pageId="3" pageNumber="65" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F35B761F8C274B60FDEDFF04FCCBFF69" box="[584,819,152,178]" name="Madagascar" pageId="3" pageNumber="65">Malagasy Republic</collectingCountry>
,
<collectingRegion id="4988F86D8C274B60FCE4FF04FBD1FF69" box="[833,1065,152,178]" country="Madagascar" name="Toamasina" pageId="3" pageNumber="65">Toamasina Province</collectingRegion>
.
<location id="8E9360548C274B60FB91FF04FAE8FF69" LSID="urn:lsid:plazi:treatment:03E587998C254B60FF32FAA6FCC3FE9C:8E9360548C274B60FB91FF04FAE8FF69" box="[1076,1296,152,178]" country="Madagascar" name="Maromizaha forest" pageId="3" pageNumber="65" stateProvince="Toamasina">Maromizaha forest</location>
.
<date id="FFF2104F8C274B60FABEFF04FF2DFF0C" pageId="3" pageNumber="65" value="1998-08-26">
<collectingDate id="EFB6E9A78C274B60FABEFF04FF2DFF0C" pageId="3" pageNumber="65" value="1998-08-26">26. August 1998</collectingDate>
</date>
.
<collectorName id="26B953598C274B60FF7AFF21FEBAFF0C" box="[223,322,189,215]" pageId="3" pageNumber="65">Leg. Dr.</collectorName>
<collectionCode id="ED5DAE4A8C274B60FEEFFF21FEA5FF0C" box="[330,349,189,215]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="65">T</collectionCode>
.
<collectorName id="26B953598C274B60FECCFF21FE59FF0C" box="[361,417,189,215]" pageId="3" pageNumber="65">Pócs</collectorName>
(9890) (Afr-923)).
<specimenCount id="9D4AFD068C274B60FD29FF21FCECFF0C" box="[652,788,189,215]" pageId="3" pageNumber="65" type="generic" typeStatus="paratypes">2 paratypes</specimenCount>
from the same sample
</materialsCitation>
.
<materialsCitation id="3B243CD28C274B60FB94FF21FC2AFF27" collectionCode="HNHM, MHNG" country="Madagascar" pageId="3" pageNumber="65" specimenCount="2" stateProvince="Toamasina" typeStatus="holotype">
<typeStatus id="54F7882D8C274B60FB94FF21FB67FF0C" box="[1073,1183,189,215]" pageId="3" pageNumber="65" type="holotype">Holotype</typeStatus>
(1784-HO-10) and
<specimenCount id="9D4AFD068C274B60FA28FF21FF04FF27" pageId="3" pageNumber="65" type="generic" typeStatus="paratype">1 paratype</specimenCount>
(1784-PO-10) deposited in the
<collectionCode id="ED5DAE4A8C274B60FDCAFF7EFD34FF27" box="[623,716,226,252]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="3" pageNumber="65">HNHM</collectionCode>
,
<specimenCount id="9D4AFD068C274B60FD7DFF7EFCAAFF27" box="[728,850,226,252]" pageId="3" pageNumber="65" type="generic" typeStatus="paratype">1 paratype</specimenCount>
in
<collectionCode id="ED5DAE4A8C274B60FCDDFF7EFC2AFF27" box="[888,978,226,252]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="3" pageNumber="65" type="Museum">MHNG</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35665048C274B60FF60FE9BFCC3FE9C" pageId="3" pageNumber="65" type="etymology">
<paragraph id="8BF3368F8C274B60FF60FE9BFCC3FE9C" blockId="3.[151,1437,151,327]" pageId="3" pageNumber="65">
<emphasis id="B938EA9D8C274B60FF60FE9BFEA1FEFA" bold="true" box="[197,345,263,289]" pageId="3" pageNumber="65">Etymology.</emphasis>
The name refers to the presence of a longitudinal mark of exclamation-shape (signum exclamationis) median costula in the interlamellar region.
</paragraph>
</subSubSection>
</treatment>
</document>