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<document id="2BBBFC69E57EB3CA6221ABDBA1EB5936" ID-CLB-Dataset="6630" ID-DOI="10.5281/zenodo.4648829" ID-GBIF-Dataset="ef7c103d-43ea-4929-8ceb-401a3053d901" ID-ISSN="1536-9307" ID-Zenodo-Dep="4648829" ID-ZooBank="634A674E-E08C-42DB-96E0-F6E5416D71CC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1617136517793" checkinUser="carolina" docAuthor="Kovařík, František, Lowe, Graeme &amp; Elmi, Hassan Sh Abdirahman" docDate="2020" docId="03C087E94510CD39FEBBFF63FBB8F852" docLanguage="en" docName="Euscorpius.322.1-23.pdf" docOrigin="Euscorpius 322" docStyle="DocumentStyle:5E96F6379CE20B0E8E4F9E7DF8E1C0DF.2:Euscorpius.2020-.journal_article" docStyleId="5E96F6379CE20B0E8E4F9E7DF8E1C0DF" docStyleName="Euscorpius.2020-.journal_article" docStyleVersion="2" docTitle="Pandinurus awalei Kovařík &amp; Lowe &amp; Elmi 2020, sp. n." docType="treatment" docVersion="7" lastPageNumber="15" masterDocId="FFF9FF914518CD29FF98FF89FF99FFA2" masterDocTitle="Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part XXV. Description of Pandinurus awalei sp. n. and the male of Pandiborellius somalilandus (Kovařík, 2012), with remarks on recent synonymies (Scorpionidae: Pandininae)" masterLastPageNumber="21" masterPageNumber="1" pageNumber="7" updateTime="1698935510268" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="48D65764D1B5ACBA78430F17B582B473">Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part XXV. Description of Pandinurus awalei sp. n. and the male of Pandiborellius somalilandus (Kovařík, 2012), with remarks on recent synonymies (Scorpionidae: Pandininae)</mods:title>
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<figureCitation id="13522A7A4510CD21FEA0FE85FE23FE86" box="[312,442,268,292]" captionStart-0="Figures 1224" captionStart-1="Figures 2528" captionStart-10="Figures 7576" captionStart-2="Figures 2930" captionStart-3="Figures 3139" captionStart-4="Figures 4047" captionStart-5="Figures 4852" captionStart-6="Figures 5466" captionStart-7="Figures 6769" captionStart-8="Figures 7072" captionStart-9="Figures 7374" captionStartId-0="7.[116,191,1952,1974]" captionStartId-1="9.[116,193,1994,2016]" captionStartId-10="18.[116,193,1992,2014]" captionStartId-2="10.[116,193,1642,1664]" captionStartId-3="11.[115,192,1985,2007]" captionStartId-4="12.[113,190,1958,1980]" captionStartId-5="13.[116,193,1982,2004]" captionStartId-6="14.[116,193,1955,1977]" captionStartId-7="15.[115,192,1990,2012]" captionStartId-8="16.[113,190,1385,1407]" captionStartId-9="17.[113,190,2018,2040]" captionTargetBox-0="[113,1474,224,1939]" captionTargetBox-1="[122,1474,251,1978]" captionTargetBox-10="[113,1474,219,1983]" captionTargetBox-2="[129,1446,240,1626]" captionTargetBox-3="[115,1476,231,1970]" captionTargetBox-4="[131,1464,270,1922]" captionTargetBox-5="[113,1474,252,1973]" captionTargetBox-6="[113,1454,238,1947]" captionTargetBox-7="[205,1387,241,1976]" captionTargetBox-8="[152,1437,259,1352]" captionTargetBox-9="[113,1474,234,2011]" captionTargetId-0="figure-106@7.[113,1474,221,1941]" captionTargetId-1="figure-43@9.[113,1474,234,1992]" captionTargetId-10="figure-47@18.[113,1474,216,1983]" captionTargetId-2="figure-237@10.[113,1474,234,1636]" captionTargetId-3="figure-76@11.[115,1476,231,1982]" captionTargetId-4="figure-82@12.[114,1475,244,1940]" captionTargetId-5="figure-52@13.[113,1474,234,1973]" captionTargetId-6="figure-107@14.[113,1474,227,1949]" captionTargetId-7="figure-35@15.[113,1474,229,1979]" captionTargetId-8="figure-354@16.[113,1474,244,1368]" captionTargetId-9="figure-31@17.[113,1474,234,2011]" captionTargetPageId-0="7" captionTargetPageId-1="9" captionTargetPageId-10="18" captionTargetPageId-2="10" captionTargetPageId-3="11" captionTargetPageId-4="12" captionTargetPageId-5="13" captionTargetPageId-6="14" captionTargetPageId-7="15" captionTargetPageId-8="16" captionTargetPageId-9="17" captionText-0="Figures 1224: Pedipalp segments. Figures 1222. Pandiborellius somalilandus, male from locality No. 19SL, pedipalp chela dorsal (12), external (13) and ventral (14) views, patella dorsal (15), external (16) and ventral (17) views, femur and trochanter dorsal (18), and ventral (19) views, movable finger dentition under white (20) and UV (21) light and fixed finger (22) dentition under UV light. Trichobothrial pattern indicated in Figures 1218. Figures 2324. Pandinurus awalei sp. n., male holotype, movable finger (23) and fixed finger (24) dentition under UV light." captionText-1="Figures 2528. Pandinurus awalei sp. n., male holotype, dorsal (25) and ventral (26) views, and right chelicera in dorsal (27) and ventral (28) aspects. Scale bar: 10 mm (2526)." captionText-10="Figures 7576. Pandinurus awalei sp. n., male paratype shortly after maturity ecdysis (75) and young female paratype two months after maturity ecdysis (76)." captionText-2="Figures 2930. Pandinurus awalei sp. n., female paratype in dorsal (29) and ventral (30) views. Scale bar: 10 mm." captionText-3="Figures 3139. Pandinurus awalei sp. n. Figures 31, 33, 3538. Male holotype, carapace and tergites IIV (31), coxosternal area and sternites (33), and left legs IIV, retrolateral aspect (3538). Figures 32, 34, and 39. Female paratype, carapace and tergites IIII (32), coxosternal area and sternites IIIIV (34), and left leg IV, retrolateral aspect (39)." captionText-4="Figures 4047: Pandinurus awalei sp. n. Figures 40, 4547. Male holotype, telson lateral (40), and metasoma and telson lateral (45), ventral (46), and dorsal (47) views. Figures 4144. Paratype female, telson lateral (41), and metasoma and telson lateral (42), ventral (43), and dorsal (44) views. Scale bars: 10 mm (4244, 4547)." captionText-5="Figures 4852: Pandinurus awalei sp. n. Figures 4852. Male holotype, pedipalp chela and patella dorsal (48), external (49) and ventral (50) views, femur and trochanter dorsal (51), and ventral (52) views. Figure 53. Type locality" captionText-6="Figures 5466: Pandinurus awalei sp. n. Figures 5461, 6466. Female paratype, pedipalp chela dorsal (54), external (55) and ventral (56) views, patella dorsal (57), external (58) and ventral (59) views, femur and trochanter dorsal (60), and ventral (61) views, movable finger dentition under white light (64); right chelicera dorsal (65) and ventral (66) aspects. Trichobothrial pattern indicated in Figures 5459. Figures 6263. Male holotype, movable finger dentition under UV (62) and white (63) light." captionText-7="Figures 6769. Pandinurus awalei sp. n., left whole hemispermatophore, anterior (67), posterior-concave (68) and convex (69) views. Scale bar: 4 mm." captionText-8="Figures 7072. Pandinurus awalei sp. n., left hemispermatophore, capsule, anterior (70), concave (71) and convex (72) views. Scale bar: 1 mm." captionText-9="Figures 7374. Pandinurus awalei sp. n., female paratype at the type locality (73) and with newborns (74)." figureDoi-0="http://doi.org/10.5281/zenodo.4648838" figureDoi-1="http://doi.org/10.5281/zenodo.4648842" figureDoi-10="http://doi.org/10.5281/zenodo.4648862" figureDoi-2="http://doi.org/10.5281/zenodo.4648844" figureDoi-3="http://doi.org/10.5281/zenodo.4648846" figureDoi-4="http://doi.org/10.5281/zenodo.4648850" figureDoi-5="http://doi.org/10.5281/zenodo.4648852" figureDoi-6="http://doi.org/10.5281/zenodo.4648854" figureDoi-7="http://doi.org/10.5281/zenodo.4648856" figureDoi-8="http://doi.org/10.5281/zenodo.4648858" figureDoi-9="http://doi.org/10.5281/zenodo.4648860" httpUri-0="https://zenodo.org/record/4648838/files/figure.png" httpUri-1="https://zenodo.org/record/4648842/files/figure.png" httpUri-10="https://zenodo.org/record/4648862/files/figure.png" httpUri-2="https://zenodo.org/record/4648844/files/figure.png" httpUri-3="https://zenodo.org/record/4648846/files/figure.png" httpUri-4="https://zenodo.org/record/4648850/files/figure.png" httpUri-5="https://zenodo.org/record/4648852/files/figure.png" httpUri-6="https://zenodo.org/record/4648854/files/figure.png" httpUri-7="https://zenodo.org/record/4648856/files/figure.png" httpUri-8="https://zenodo.org/record/4648858/files/figure.png" httpUri-9="https://zenodo.org/record/4648860/files/figure.png" pageId="8" pageNumber="7">Figs. 2376</figureCitation>
,
<figureCitation id="13522A7A4510CD21FE5DFE84FE7BFE87" box="[453,482,269,293]" captionStart="Figure 83" captionStartId="20.[113,181,2018,2040]" captionTargetBox="[113,1474,944,2005]" captionTargetId="figure-31@20.[113,1474,941,2005]" captionTargetPageId="20" captionText="Figure 83. Map showing confirmed distribution of Pandinurus spp." figureDoi="http://doi.org/10.5281/zenodo.4648868" httpUri="https://zenodo.org/record/4648868/files/figure.png" pageId="8" pageNumber="7">83</figureCitation>
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1845, 5
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<collectingDate id="EF93E9D74510CD21FFE9FDC4FF7AFDC7" box="[113,227,589,613]" pageId="8" pageNumber="7" value="2020-02-27">27.II.2020</collectingDate>
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<figureCitation id="13522A7A4510CD21FF75FDC5FEF4FDC6" box="[237,365,588,612]" captionStart="Figures 2528" captionStartId="9.[116,193,1994,2016]" captionTargetBox="[122,1474,251,1978]" captionTargetId="figure-43@9.[113,1474,234,1992]" captionTargetPageId="9" captionText="Figures 2528. Pandinurus awalei sp. n., male holotype, dorsal (25) and ventral (26) views, and right chelicera in dorsal (27) and ventral (28) aspects. Scale bar: 10 mm (2526)." figureDoi="http://doi.org/10.5281/zenodo.4648842" httpUri="https://zenodo.org/record/4648842/files/figure.png" pageId="8" pageNumber="7">Figs. 2528</figureCitation>
,
<figureCitation id="13522A7A4510CD21FEEFFDC4FD92FDC6" box="[375,523,588,613]" captionStart="Figures 3139" captionStartId="11.[115,192,1985,2007]" captionTargetBox="[115,1476,231,1970]" captionTargetId="figure-76@11.[115,1476,231,1982]" captionTargetPageId="11" captionText="Figures 3139. Pandinurus awalei sp. n. Figures 31, 33, 3538. Male holotype, carapace and tergites IIV (31), coxosternal area and sternites (33), and left legs IIV, retrolateral aspect (3538). Figures 32, 34, and 39. Female paratype, carapace and tergites IIII (32), coxosternal area and sternites IIIIV (34), and left leg IV, retrolateral aspect (39)." figureDoi="http://doi.org/10.5281/zenodo.4648846" httpUri="https://zenodo.org/record/4648846/files/figure.png" pageId="8" pageNumber="7">31, 33, 3538</figureCitation>
,
<figureCitation id="13522A7A4510CD21FD8DFDC4FD1BFDC6" box="[533,642,588,613]" captionStart-0="Figures 4047" captionStart-1="Figures 4852" captionStartId-0="12.[113,190,1958,1980]" captionStartId-1="13.[116,193,1982,2004]" captionTargetBox-0="[131,1464,270,1922]" captionTargetBox-1="[113,1474,252,1973]" captionTargetId-0="figure-82@12.[114,1475,244,1940]" captionTargetId-1="figure-52@13.[113,1474,234,1973]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionText-0="Figures 4047: Pandinurus awalei sp. n. Figures 40, 4547. Male holotype, telson lateral (40), and metasoma and telson lateral (45), ventral (46), and dorsal (47) views. Figures 4144. Paratype female, telson lateral (41), and metasoma and telson lateral (42), ventral (43), and dorsal (44) views. Scale bars: 10 mm (4244, 4547)." captionText-1="Figures 4852: Pandinurus awalei sp. n. Figures 4852. Male holotype, pedipalp chela and patella dorsal (48), external (49) and ventral (50) views, femur and trochanter dorsal (51), and ventral (52) views. Figure 53. Type locality" figureDoi-0="http://doi.org/10.5281/zenodo.4648850" figureDoi-1="http://doi.org/10.5281/zenodo.4648852" httpUri-0="https://zenodo.org/record/4648850/files/figure.png" httpUri-1="https://zenodo.org/record/4648852/files/figure.png" pageId="8" pageNumber="7">40, 4552</figureCitation>
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<figureCitation id="13522A7A4510CD21FD14FDC5FD48FDC6" box="[652,721,588,612]" captionStart="Figures 5466" captionStartId="14.[116,193,1955,1977]" captionTargetBox="[113,1454,238,1947]" captionTargetId="figure-107@14.[113,1474,227,1949]" captionTargetPageId="14" captionText="Figures 5466: Pandinurus awalei sp. n. Figures 5461, 6466. Female paratype, pedipalp chela dorsal (54), external (55) and ventral (56) views, patella dorsal (57), external (58) and ventral (59) views, femur and trochanter dorsal (60), and ventral (61) views, movable finger dentition under white light (64); right chelicera dorsal (65) and ventral (66) aspects. Trichobothrial pattern indicated in Figures 5459. Figures 6263. Male holotype, movable finger dentition under UV (62) and white (63) light." figureDoi="http://doi.org/10.5281/zenodo.4648854" httpUri="https://zenodo.org/record/4648854/files/figure.png" pageId="8" pageNumber="7">6263</figureCitation>
,
<figureCitation id="13522A7A4510CD21FD43FDC5FF16FD27" captionStart-0="Figures 6769" captionStart-1="Figures 7072" captionStartId-0="15.[115,192,1990,2012]" captionStartId-1="16.[113,190,1385,1407]" captionTargetBox-0="[205,1387,241,1976]" captionTargetBox-1="[152,1437,259,1352]" captionTargetId-0="figure-35@15.[113,1474,229,1979]" captionTargetId-1="figure-354@16.[113,1474,244,1368]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="Figures 6769. Pandinurus awalei sp. n., left whole hemispermatophore, anterior (67), posterior-concave (68) and convex (69) views. Scale bar: 4 mm." captionText-1="Figures 7072. Pandinurus awalei sp. n., left hemispermatophore, capsule, anterior (70), concave (71) and convex (72) views. Scale bar: 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4648856" figureDoi-1="http://doi.org/10.5281/zenodo.4648858" httpUri-0="https://zenodo.org/record/4648856/files/figure.png" httpUri-1="https://zenodo.org/record/4648858/files/figure.png" pageId="8" pageNumber="7">67 72</figureCitation>
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,
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(
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, scorpion born
<date id="FFD7103F4510CD21FE55FDE4FDC8FD27" box="[461,593,621,645]" pageId="8" pageNumber="7" value="2019-07-18">18.VII.2019</date>
,
<figureCitation id="13522A7A4510CD21FDC2FDE5FD40FD27" box="[602,729,620,645]" captionStart="Figures 2930" captionStartId="10.[116,193,1642,1664]" captionTargetBox="[129,1446,240,1626]" captionTargetId="figure-237@10.[113,1474,234,1636]" captionTargetPageId="10" captionText="Figures 2930. Pandinurus awalei sp. n., female paratype in dorsal (29) and ventral (30) views. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4648844" httpUri="https://zenodo.org/record/4648844/files/figure.png" pageId="8" pageNumber="7">Figs. 2930</figureCitation>
,
<figureCitation id="13522A7A4510CD21FD7AFDE4FF2FFD07" captionStart="Figures 3139" captionStartId="11.[115,192,1985,2007]" captionTargetBox="[115,1476,231,1970]" captionTargetId="figure-76@11.[115,1476,231,1982]" captionTargetPageId="11" captionText="Figures 3139. Pandinurus awalei sp. n. Figures 31, 33, 3538. Male holotype, carapace and tergites IIV (31), coxosternal area and sternites (33), and left legs IIV, retrolateral aspect (3538). Figures 32, 34, and 39. Female paratype, carapace and tergites IIII (32), coxosternal area and sternites IIIIV (34), and left leg IV, retrolateral aspect (39)." figureDoi="http://doi.org/10.5281/zenodo.4648846" httpUri="https://zenodo.org/record/4648846/files/figure.png" pageId="8" pageNumber="7">32, 34, 39</figureCitation>
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<figureCitation id="13522A7A4510CD21FF59FD04FE9CFD07" box="[193,261,653,677]" captionStart="Figures 4047" captionStartId="12.[113,190,1958,1980]" captionTargetBox="[131,1464,270,1922]" captionTargetId="figure-82@12.[114,1475,244,1940]" captionTargetPageId="12" captionText="Figures 4047: Pandinurus awalei sp. n. Figures 40, 4547. Male holotype, telson lateral (40), and metasoma and telson lateral (45), ventral (46), and dorsal (47) views. Figures 4144. Paratype female, telson lateral (41), and metasoma and telson lateral (42), ventral (43), and dorsal (44) views. Scale bars: 10 mm (4244, 4547)." figureDoi="http://doi.org/10.5281/zenodo.4648850" httpUri="https://zenodo.org/record/4648850/files/figure.png" pageId="8" pageNumber="7">4144</figureCitation>
,
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,
<figureCitation id="13522A7A4510CD21FE37FD04FE6CFD07" box="[431,501,653,677]" captionStart="Figures 7374" captionStartId="17.[113,190,2018,2040]" captionTargetBox="[113,1474,234,2011]" captionTargetId="figure-31@17.[113,1474,234,2011]" captionTargetPageId="17" captionText="Figures 7374. Pandinurus awalei sp. n., female paratype at the type locality (73) and with newborns (74)." figureDoi="http://doi.org/10.5281/zenodo.4648860" httpUri="https://zenodo.org/record/4648860/files/figure.png" pageId="8" pageNumber="7">7374</figureCitation>
),
<specimenCount id="9D6FFD764510CD21FD90FD04FDB0FD07" box="[520,553,653,677]" pageId="8" pageNumber="7" type="female">1♀</specimenCount>
(
<typeStatus id="54D2885D4510CD21FDAEFD04FD0DFD07" box="[566,660,653,677]" pageId="8" pageNumber="7" type="paratype">paratype</typeStatus>
still alive, ecdysis
<date id="FFD7103F4510CD21FF51FD24FEDDFD67" box="[201,324,685,709]" pageId="8" pageNumber="7" value="2019-09-14">14.IX.2019</date>
, maturity ecdysis
<date id="FFD7103F4510CD21FD93FD24FD00FD67" box="[523,665,685,709]" pageId="8" pageNumber="7" value="2020-08-22">22.VIII.2020</date>
,
<figureCitation id="13522A7A4510CD21FD3EFD25FD6EFD66" box="[678,759,684,708]" captionStart="Figures 7576" captionStartId="18.[116,193,1992,2014]" captionTargetBox="[113,1474,219,1983]" captionTargetId="figure-47@18.[113,1474,216,1983]" captionTargetPageId="18" captionText="Figures 7576. Pandinurus awalei sp. n., male paratype shortly after maturity ecdysis (75) and young female paratype two months after maturity ecdysis (76)." figureDoi="http://doi.org/10.5281/zenodo.4648862" httpUri="https://zenodo.org/record/4648862/files/figure.png" pageId="8" pageNumber="7">Fig. 76</figureCitation>
),
<specimenCount id="9D6FFD764510CD21FFE9FD45FF3FFD46" box="[113,166,716,740]" pageId="8" pageNumber="7" type="generic">1juv.</specimenCount>
(
<typeStatus id="54D2885D4510CD21FF2DFD44FE8FFD47" box="[181,278,717,741]" pageId="8" pageNumber="7" type="paratype">paratype</typeStatus>
), leg. F. KovařÍk et
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<collectionCode id="ED78AE3A4510CD21FE75FD44FE66FD46" box="[493,511,717,740]" name="Tavera, Department of Geology and Geophysics" pageId="8" pageNumber="7">T</collectionCode>
.
<collectorName id="269C53294510CD21FD92FD45FDFAFD46" box="[522,611,716,740]" pageId="8" pageNumber="7">Mazuch</collectorName>
</materialsCitation>
,
<specimenCount id="9D6FFD764510CD21FDF7FD44FD09FD47" box="[623,656,717,741]" pageId="8" pageNumber="7" type="male">1♂</specimenCount>
(
<typeStatus id="54D2885D4510CD21FD07FD44FD66FD47" box="[671,767,717,741]" pageId="8" pageNumber="7" type="paratype">paratype</typeStatus>
, born
<date id="FFD7103F4510CD21FF30FD64FEB5FCA7" box="[168,300,749,773]" pageId="8" pageNumber="7" value="2019-07-18">18.VII.2019</date>
, 2
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ecdysis
<date id="FFD7103F4510CD21FE34FD65FDCEFCA6" box="[428,599,748,772]" pageId="8" pageNumber="7" value="2019-09-19" valueMax="2019-09-25" valueMin="2019-09-19">19.25.IX.2019</date>
, 3
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ecdysis
<date id="FFD7103F4510CD21FD4CFD65FF74FC87" pageId="8" pageNumber="7" value="2019-11-15" valueMax="2019-11-22" valueMin="2019-11-15">15. 22.XI.2019</date>
, 4
<superScript id="7C1C9BB74510CD21FE9EFC82FE8BFCBB" attach="left" box="[262,274,779,793]" fontSize="6" pageId="8" pageNumber="7">th</superScript>
ecdysis
<date id="FFD7103F4510CD21FEE8FC84FD86FC87" box="[368,543,781,805]" pageId="8" pageNumber="7" value="2020-01-11" valueMax="2020-02-20" valueMin="2020-01-11">11.I.20.II.2020</date>
, 5
<superScript id="7C1C9BB74510CD21FDA0FC82FDDDFCBB" attach="none" box="[568,580,779,793]" fontSize="6" pageId="8" pageNumber="7">th</superScript>
, maturity ecdysis
<date id="FFD7103F4510CD21FFE9FCA4FF6CFCE7" box="[113,245,813,837]" pageId="8" pageNumber="7" value="2020-07-11">11.VII.2020</date>
,
<figureCitation id="13522A7A4510CD21FE98FCA5FED4FCE6" box="[256,333,812,836]" captionStart="Figures 7576" captionStartId="18.[116,193,1992,2014]" captionTargetBox="[113,1474,219,1983]" captionTargetId="figure-47@18.[113,1474,216,1983]" captionTargetPageId="18" captionText="Figures 7576. Pandinurus awalei sp. n., male paratype shortly after maturity ecdysis (75) and young female paratype two months after maturity ecdysis (76)." figureDoi="http://doi.org/10.5281/zenodo.4648862" httpUri="https://zenodo.org/record/4648862/files/figure.png" pageId="8" pageNumber="7">Fig. 75</figureCitation>
),
<specimenCount id="9D6FFD764510CD21FEF8FCA4FE4DFCE6" box="[352,468,812,836]" pageId="8" pageNumber="7" type="juvenile">7 juveniles</specimenCount>
or adult females (
<typeStatus id="54D2885D4510CD21FD0AFCA4FD62FCE7" box="[658,763,813,837]" pageId="8" pageNumber="7" type="paratype">paratypes</typeStatus>
) after 5
<superScript id="7C1C9BB74510CD21FF2FFCC2FF5AFCFB" attach="left" box="[183,195,843,857]" fontSize="6" pageId="8" pageNumber="7">th</superScript>
ecdysis still alive (born
<date id="FFD7103F4510CD21FE52FCC4FDD6FCC7" box="[458,591,845,869]" pageId="8" pageNumber="7" value="2019-07-18">18.VII.2019</date>
, 2
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ecdysis
<date id="FFD7103F4510CD21FD4CFCC4FF77FC26" pageId="8" pageNumber="7" value="2019-09-19" valueMax="2019-09-25" valueMin="2019-09-19">19. 25.IX.2019</date>
, 3
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ecdysis
<date id="FFD7103F4510CD21FEECFCE5FD87FC26" box="[372,542,876,900]" pageId="8" pageNumber="7" value="2019-11-15" valueMax="2019-11-22" valueMin="2019-11-15">15.22.XI.2019</date>
, 4
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ecdysis
<date id="FFD7103F4510CD21FD3CFCE4FF58FC07" pageId="8" pageNumber="7" value="2020-01-11" valueMax="2020-02-20" valueMin="2020-01-11">11.I.20. II.2020</date>
, 5
<superScript id="7C1C9BB74510CD21FF42FC02FF7FFC3B" attach="left" box="[218,230,907,921]" fontSize="6" pageId="8" pageNumber="7">th</superScript>
ecdysis
<date id="FFD7103F4510CD21FEDAFC05FDBAFC06" box="[322,547,908,932]" pageId="8" pageNumber="7" value="2020-06-10" valueMax="2020-08-25" valueMin="2020-06-10">
10.
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<collectionCode id="ED78AE3A4510CD21FEF8FC05FEE6FC06" box="[352,383,908,932]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="8" pageNumber="7">VI</collectionCode>
.25.VIII.2020
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</date>
).
</paragraph>
</subSubSection>
<subSubSection id="C37365744510CD21FFE9FC44FD9CFB67" pageId="8" pageNumber="7" type="etymology">
<paragraph id="8BD636FF4510CD21FFE9FC44FD9CFB67" blockId="8.[113,771,972,1221]" pageId="8" pageNumber="7">
ETYMOLOGY. Named after Ahmed Ibrahim Awale, an ecologist from Somaliland, a university lecturer, chairman of Candlelight NGO, and co-founder and chairman of Somaliland Biodiversity Foundation. He is also the author of several scientific papers and books including
<emphasis id="B91DEAED4510CD21FDE2FBC4FD31FB26" italics="true" pageId="8" pageNumber="7">
Environment in Crisis: Selected Essays on
<collectingRegion id="49ADF81D4510CD21FE51FBE5FD8AFB26" box="[457,531,1132,1156]" country="Ethiopia" name="Sumale" pageId="8" pageNumber="7">Somali</collectingRegion>
Environment
</emphasis>
(2016); and
<emphasis id="B91DEAED4510CD21FF3AFB05FD16FB06" box="[162,655,1164,1188]" italics="true" pageId="8" pageNumber="7">Introduction to Plants in Central Somaliland</emphasis>
(the latter together with Helen
<bibRefCitation id="EFF84B0E4510CD21FED4FB25FE60FB67" author="PICKERING" box="[332,505,1196,1221]" pageId="8" pageNumber="7" refId="ref6527" refString="PICKERING, H. &amp; A. I. AWALE. 2018. Introduction to Plants in Central Somaliland. Poynte Invisibile (Redsea Cultural Foundation), Hargeysa, Somaliland, 143 pp." type="book" year="2018">Pickering, 2018</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37365744510CD21FFE9FB64FE3EF827" pageId="8" pageNumber="7" type="diagnosis">
<paragraph id="8BD636FF4510CD21FFE9FB64FE3EF827" blockId="8.[113,772,1260,1925]" pageId="8" pageNumber="7">
DIAGNOSIS (
<specimenCount id="9D6FFD764510CD21FF6AFB64FE93FAA7" box="[242,266,1261,1285]" pageId="8" pageNumber="7" type="male"></specimenCount>
<specimenCount id="9D6FFD764510CD21FE92FB64FE82FAA7" box="[266,283,1261,1285]" pageId="8" pageNumber="7" type="female"></specimenCount>
). Total length
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. Color uniformly reddish brown to black; legs yellow; chela orange to brown but yellow in young specimens; telson yellowish brown to black. Chelicerae yellowish brown, reticulate, with black fingers and anterior margin. Carapace lacking carinae but with sparse granules, anterior part smooth. External trichobothria on patella number 1719 (5
<emphasis id="B91DEAED4510CD21FEE3FA25FE0DFA66" box="[379,404,1452,1476]" italics="true" pageId="8" pageNumber="7">eb</emphasis>
, 46
<emphasis id="B91DEAED4510CD21FE57FA25FE6AFA66" box="[463,499,1452,1476]" italics="true" pageId="8" pageNumber="7">esb</emphasis>
, 2
<emphasis id="B91DEAED4510CD21FD8BFA24FDABFA66" box="[531,562,1453,1476]" italics="true" pageId="8" pageNumber="7">em</emphasis>
, 3
<emphasis id="B91DEAED4510CD21FDCAFA24FDE9FA66" box="[594,624,1453,1476]" italics="true" pageId="8" pageNumber="7">est</emphasis>
, 3
<emphasis id="B91DEAED4510CD21FD17FA24FD3BFA66" box="[655,674,1453,1476]" italics="true" pageId="8" pageNumber="7">et</emphasis>
); ventral trichobothria on patella number 3543; internal trichobothria on chela number 3, accessory external trichobothrium
<emphasis id="B91DEAED4510CD21FD72FA64FC9AF9A6" box="[746,771,1517,1540]" italics="true" pageId="8" pageNumber="7">ea</emphasis>
on chela absent, ventral trichobothria on chela number 11 13. Pedipalp densely hirsute, mainly on chela. Granules on dorsal surface of chela of pedipalp conical and rounded. Lobe of chela granulated with the same intensity as whole dorsal surface of chela. External surface of chela with granules and without carinae which could be indicated in females. Chela of male length/ width ratio 1.87. Pectine teeth
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both sexes. Dorsal carinae on second through fourth metasomal segments granulate by large pointed tooth. Spiniform formula of tarsomere II = 6-7/4: 6-7/4: 7/5: 78/5. Tarsomere II with 2 spines on inclined anteroventral surface. Length to width ratio of metasoma V of male 2.79.
</paragraph>
</subSubSection>
<subSubSection id="C37365744510CD39FFE9F824FBB8F852" lastPageId="16" lastPageNumber="15" pageId="8" pageNumber="7" type="description">
<subSubSection id="C37365744510CD23FFE9F824FBF3F85B" lastPageId="10" lastPageNumber="9" pageId="8" pageNumber="7" type="description">
<paragraph id="8BD636FF4510CD21FFE9F824FEDBF846" blockId="8.[113,771,1964,2020]" pageId="8" pageNumber="7">
DESCRIPTION. Habitus as shown in
<figureCitation id="13522A7A4510CD21FE76F825FDE9F866" box="[494,624,1964,1988]" captionStart="Figures 2528" captionStartId="9.[116,193,1994,2016]" captionTargetBox="[122,1474,251,1978]" captionTargetId="figure-43@9.[113,1474,234,1992]" captionTargetPageId="9" captionText="Figures 2528. Pandinurus awalei sp. n., male holotype, dorsal (25) and ventral (26) views, and right chelicera in dorsal (27) and ventral (28) aspects. Scale bar: 10 mm (2526)." figureDoi="http://doi.org/10.5281/zenodo.4648842" httpUri="https://zenodo.org/record/4648842/files/figure.png" pageId="8" pageNumber="7">Figs. 2526</figureCitation>
,
<figureCitation id="13522A7A4510CD21FDE4F824FD5BF867" box="[636,706,1965,1989]" captionStart="Figures 2930" captionStartId="10.[116,193,1642,1664]" captionTargetBox="[129,1446,240,1626]" captionTargetId="figure-237@10.[113,1474,234,1636]" captionTargetPageId="10" captionText="Figures 2930. Pandinurus awalei sp. n., female paratype in dorsal (29) and ventral (30) views. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4648844" httpUri="https://zenodo.org/record/4648844/files/figure.png" pageId="8" pageNumber="7">2930</figureCitation>
. Total length
<quantity id="4C919B1A4510CD21FF23F845FEA7F846" box="[187,318,1996,2020]" metricMagnitude="-2" metricUnit="m" metricValue="9.75" metricValueMax="10.5" metricValueMin="9.0" pageId="8" pageNumber="7" unit="mm" value="97.5" valueMax="105.0" valueMin="90.0">90105 mm</quantity>
.
</paragraph>
<paragraph id="8BD636FF4510CD21FCA8FF63FBC0FE01" blockId="8.[816,1475,234,2019]" pageId="8" pageNumber="7">
<emphasis id="B91DEAED4510CD21FCA8FF63FC32FEA0" bold="true" box="[816,939,234,258]" pageId="8" pageNumber="7">Coloration</emphasis>
(
<figureCitation id="13522A7A4510CD21FC24FF62FBD8FEA0" box="[956,1089,234,259]" captionStart="Figures 2528" captionStartId="9.[116,193,1994,2016]" captionTargetBox="[122,1474,251,1978]" captionTargetId="figure-43@9.[113,1474,234,1992]" captionTargetPageId="9" captionText="Figures 2528. Pandinurus awalei sp. n., male holotype, dorsal (25) and ventral (26) views, and right chelicera in dorsal (27) and ventral (28) aspects. Scale bar: 10 mm (2526)." figureDoi="http://doi.org/10.5281/zenodo.4648842" httpUri="https://zenodo.org/record/4648842/files/figure.png" pageId="8" pageNumber="7">Figs. 2526</figureCitation>
,
<figureCitation id="13522A7A4510CD21FBD6FF62FB0AFEA1" box="[1102,1171,235,259]" captionStart="Figures 2930" captionStartId="10.[116,193,1642,1664]" captionTargetBox="[129,1446,240,1626]" captionTargetId="figure-237@10.[113,1474,234,1636]" captionTargetPageId="10" captionText="Figures 2930. Pandinurus awalei sp. n., female paratype in dorsal (29) and ventral (30) views. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4648844" httpUri="https://zenodo.org/record/4648844/files/figure.png" pageId="8" pageNumber="7">2930</figureCitation>
,
<figureCitation id="13522A7A4510CD21FB38FF62FB7FFEA1" box="[1184,1254,235,259]" captionStart="Figures 4047" captionStartId="12.[113,190,1958,1980]" captionTargetBox="[131,1464,270,1922]" captionTargetId="figure-82@12.[114,1475,244,1940]" captionTargetPageId="12" captionText="Figures 4047: Pandinurus awalei sp. n. Figures 40, 4547. Male holotype, telson lateral (40), and metasoma and telson lateral (45), ventral (46), and dorsal (47) views. Figures 4144. Paratype female, telson lateral (41), and metasoma and telson lateral (42), ventral (43), and dorsal (44) views. Scale bars: 10 mm (4244, 4547)." figureDoi="http://doi.org/10.5281/zenodo.4648850" httpUri="https://zenodo.org/record/4648850/files/figure.png" pageId="8" pageNumber="7">4346</figureCitation>
,
<figureCitation id="13522A7A4510CD21FB6BFF62FAA3FEA1" box="[1267,1338,235,259]" captionStart-0="Figures 7374" captionStart-1="Figures 7576" captionStartId-0="17.[113,190,2018,2040]" captionStartId-1="18.[116,193,1992,2014]" captionTargetBox-0="[113,1474,234,2011]" captionTargetBox-1="[113,1474,219,1983]" captionTargetId-0="figure-31@17.[113,1474,234,2011]" captionTargetId-1="figure-47@18.[113,1474,216,1983]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionText-0="Figures 7374. Pandinurus awalei sp. n., female paratype at the type locality (73) and with newborns (74)." captionText-1="Figures 7576. Pandinurus awalei sp. n., male paratype shortly after maturity ecdysis (75) and young female paratype two months after maturity ecdysis (76)." figureDoi-0="http://doi.org/10.5281/zenodo.4648860" figureDoi-1="http://doi.org/10.5281/zenodo.4648862" httpUri-0="https://zenodo.org/record/4648860/files/figure.png" httpUri-1="https://zenodo.org/record/4648862/files/figure.png" pageId="8" pageNumber="7">7376</figureCitation>
). Base color uniform reddish brown to black, young specimens more pale (
<figureCitation id="13522A7A4510CD21FCA0FEA2FC15FEE0" box="[824,908,298,323]" captionStart="Figures 7576" captionStartId="18.[116,193,1992,2014]" captionTargetBox="[113,1474,219,1983]" captionTargetId="figure-47@18.[113,1474,216,1983]" captionTargetPageId="18" captionText="Figures 7576. Pandinurus awalei sp. n., male paratype shortly after maturity ecdysis (75) and young female paratype two months after maturity ecdysis (76)." figureDoi="http://doi.org/10.5281/zenodo.4648862" httpUri="https://zenodo.org/record/4648862/files/figure.png" pageId="8" pageNumber="7">Fig. 75</figureCitation>
). Legs yellow, pedipalp chela orange to brown but yellow in young specimens, and telson yellowish brown to black. Chelicerae yellowish brown, reticulate, with black fingers and anterior margin.
</paragraph>
<paragraph id="8BD636FF4510CD21FCA8FE23FB20FC61" blockId="8.[816,1475,234,2019]" pageId="8" pageNumber="7">
<emphasis id="B91DEAED4510CD21FCA8FE23FC04FE60" bold="true" box="[816,925,426,450]" pageId="8" pageNumber="7">Pedipalps</emphasis>
(
<figureCitation id="13522A7A4510CD21FC2BFE22FBA0FE61" box="[947,1081,427,451]" captionStart="Figures 1224" captionStartId="7.[116,191,1952,1974]" captionTargetBox="[113,1474,224,1939]" captionTargetId="figure-106@7.[113,1474,221,1941]" captionTargetPageId="7" captionText="Figures 1224: Pedipalp segments. Figures 1222. Pandiborellius somalilandus, male from locality No. 19SL, pedipalp chela dorsal (12), external (13) and ventral (14) views, patella dorsal (15), external (16) and ventral (17) views, femur and trochanter dorsal (18), and ventral (19) views, movable finger dentition under white (20) and UV (21) light and fixed finger (22) dentition under UV light. Trichobothrial pattern indicated in Figures 1218. Figures 2324. Pandinurus awalei sp. n., male holotype, movable finger (23) and fixed finger (24) dentition under UV light." figureDoi="http://doi.org/10.5281/zenodo.4648838" httpUri="https://zenodo.org/record/4648838/files/figure.png" pageId="8" pageNumber="7">Figs. 2324</figureCitation>
,
<figureCitation id="13522A7A4510CD21FBD3FE23FB16FE60" box="[1099,1167,426,450]" captionStart="Figures 4852" captionStartId="13.[116,193,1982,2004]" captionTargetBox="[113,1474,252,1973]" captionTargetId="figure-52@13.[113,1474,234,1973]" captionTargetPageId="13" captionText="Figures 4852: Pandinurus awalei sp. n. Figures 4852. Male holotype, pedipalp chela and patella dorsal (48), external (49) and ventral (50) views, femur and trochanter dorsal (51), and ventral (52) views. Figure 53. Type locality" figureDoi="http://doi.org/10.5281/zenodo.4648852" httpUri="https://zenodo.org/record/4648852/files/figure.png" pageId="8" pageNumber="7">4852</figureCitation>
,
<figureCitation id="13522A7A4510CD21FB38FE23FB7EFE60" box="[1184,1255,426,450]" captionStart="Figures 5466" captionStartId="14.[116,193,1955,1977]" captionTargetBox="[113,1454,238,1947]" captionTargetId="figure-107@14.[113,1474,227,1949]" captionTargetPageId="14" captionText="Figures 5466: Pandinurus awalei sp. n. Figures 5461, 6466. Female paratype, pedipalp chela dorsal (54), external (55) and ventral (56) views, patella dorsal (57), external (58) and ventral (59) views, femur and trochanter dorsal (60), and ventral (61) views, movable finger dentition under white light (64); right chelicera dorsal (65) and ventral (66) aspects. Trichobothrial pattern indicated in Figures 5459. Figures 6263. Male holotype, movable finger dentition under UV (62) and white (63) light." figureDoi="http://doi.org/10.5281/zenodo.4648854" httpUri="https://zenodo.org/record/4648854/files/figure.png" pageId="8" pageNumber="7">5464</figureCitation>
). Pedipalps densely hirsute, mainly on the chela. Femur smooth with several large granules dorsally, and four carinae composed of several strong granules. Patella smooth and rugose externally, with four carinae. Granules on dorsoexternal surface of chela conical and rounded. Lobe of chela granulated with same intensity as whole dorsal surface of chela. Internal surface of chela granulated by conical usually pointed granules mainly in anterior part. Dentate margins of fixed and movable fingers of pedipalp with distinct granules in two parallel rows, present in anterior half of fingers. Posterior half of fingers almost without granules in male, with distinct granules in a row in female.
<emphasis id="B91DEAED4510CD21FB58FC82FAE5FC81" box="[1216,1404,779,803]" italics="true" pageId="8" pageNumber="7">Trichobothriotaxy</emphasis>
(
<figureCitation id="13522A7A4510CD21FA16FC82FCEFFCE0" captionStart="Figures 5466" captionStartId="14.[116,193,1955,1977]" captionTargetBox="[113,1454,238,1947]" captionTargetId="figure-107@14.[113,1474,227,1949]" captionTargetPageId="14" captionText="Figures 5466: Pandinurus awalei sp. n. Figures 5461, 6466. Female paratype, pedipalp chela dorsal (54), external (55) and ventral (56) views, patella dorsal (57), external (58) and ventral (59) views, femur and trochanter dorsal (60), and ventral (61) views, movable finger dentition under white light (64); right chelicera dorsal (65) and ventral (66) aspects. Trichobothrial pattern indicated in Figures 5459. Figures 6263. Male holotype, movable finger dentition under UV (62) and white (63) light." figureDoi="http://doi.org/10.5281/zenodo.4648854" httpUri="https://zenodo.org/record/4648854/files/figure.png" pageId="8" pageNumber="7">Figs. 5459</figureCitation>
). External trichobothria on the patella number 1719 (5
<emphasis id="B91DEAED4510CD21FCA8FCC2FCD0FCC1" box="[816,841,843,867]" italics="true" pageId="8" pageNumber="7">eb</emphasis>
, 46
<emphasis id="B91DEAED4510CD21FC19FCC2FC3DFCC1" box="[897,932,843,867]" italics="true" pageId="8" pageNumber="7">esb</emphasis>
, 2
<emphasis id="B91DEAED4510CD21FC5AFCC2FC78FCC0" box="[962,993,843,866]" italics="true" pageId="8" pageNumber="7">em</emphasis>
, 3
<emphasis id="B91DEAED4510CD21FC67FCC2FB85FCC0" box="[1023,1052,843,866]" italics="true" pageId="8" pageNumber="7">est</emphasis>
, 3
<emphasis id="B91DEAED4510CD21FBA2FCC2FBD4FCC0" box="[1082,1101,843,866]" italics="true" pageId="8" pageNumber="7">et</emphasis>
); accessory external trichobothrium
<emphasis id="B91DEAED4510CD21FCA8FCE2FCD0FC20" box="[816,841,875,898]" italics="true" pageId="8" pageNumber="7">ea</emphasis>
on chela absent, ventral trichobothria on patella number 3543; internal trichobothria on chela number 3, ventral trichobothria on chela number 1113.
</paragraph>
<paragraph id="8BD636FF4510CD21FCA8FC42FB9EFAA1" blockId="8.[816,1475,234,2019]" pageId="8" pageNumber="7">
<emphasis id="B91DEAED4510CD21FCA8FC42FBB5FC41" bold="true" box="[816,1068,971,995]" pageId="8" pageNumber="7">Metasoma and telson</emphasis>
(
<figureCitation id="13522A7A4510CD21FBA6FC42FB5EFC41" box="[1086,1223,971,995]" captionStart="Figures 4047" captionStartId="12.[113,190,1958,1980]" captionTargetBox="[131,1464,270,1922]" captionTargetId="figure-82@12.[114,1475,244,1940]" captionTargetPageId="12" captionText="Figures 4047: Pandinurus awalei sp. n. Figures 40, 4547. Male holotype, telson lateral (40), and metasoma and telson lateral (45), ventral (46), and dorsal (47) views. Figures 4144. Paratype female, telson lateral (41), and metasoma and telson lateral (42), ventral (43), and dorsal (44) views. Scale bars: 10 mm (4244, 4547)." figureDoi="http://doi.org/10.5281/zenodo.4648850" httpUri="https://zenodo.org/record/4648850/files/figure.png" pageId="8" pageNumber="7">Figs. 4047</figureCitation>
). Metasomal segments IIV each with a total of 8 complete carinae of which the ventral on segments IIII are smooth. Other carinae sparsely granulated. Segment V with five carinae developed and granulated. Dorsal and lateral surfaces of the segments rugose with several granules, segments IVV more granulated. Dorsal carinae on segments IIIV granulate with large pointed tooth. Entire surfaces of metasoma and telson hirsute with long setae. Telson smooth to rugose, bulbous, with aculeus shorter than vesicle.
</paragraph>
<paragraph id="8BD636FF4510CD21FCA8FA83FA06F9C1" blockId="8.[816,1475,234,2019]" pageId="8" pageNumber="7">
<emphasis id="B91DEAED4510CD21FCA8FA83FBCAFA81" bold="true" box="[816,1107,1290,1315]" pageId="8" pageNumber="7">Carapace and mesosoma</emphasis>
(
<figureCitation id="13522A7A4510CD21FBF0FA82FB77FA81" box="[1128,1262,1291,1315]" captionStart="Figures 3139" captionStartId="11.[115,192,1985,2007]" captionTargetBox="[115,1476,231,1970]" captionTargetId="figure-76@11.[115,1476,231,1982]" captionTargetPageId="11" captionText="Figures 3139. Pandinurus awalei sp. n. Figures 31, 33, 3538. Male holotype, carapace and tergites IIV (31), coxosternal area and sternites (33), and left legs IIV, retrolateral aspect (3538). Figures 32, 34, and 39. Female paratype, carapace and tergites IIII (32), coxosternal area and sternites IIIIV (34), and left leg IV, retrolateral aspect (39)." figureDoi="http://doi.org/10.5281/zenodo.4648846" httpUri="https://zenodo.org/record/4648846/files/figure.png" pageId="8" pageNumber="7">Figs. 3134</figureCitation>
). Carapace smooth without carinae, sparsely covered by large granules medially and laterally, and finely granulated with small granules posteriorly mainly in the male. Anterior margin of carapace bilobate, strongly emarginate medially, bearing several macrosetae. Three lateral eyes present on each side. Tergites finely granulated in the male, almost smooth in the female. Pectinal tooth count
<quantity id="4C919B1A4510CD21FB90FA62FBFDF9A1" box="[1032,1124,1515,1539]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" metricValueMax="4.826" metricValueMin="4.318" pageId="8" pageNumber="7" unit="in" value="18.0" valueMax="19.0" valueMin="17.0">1719 in</quantity>
both sexes. Pectine marginal tips extend to the anterior first quarter of sternite IV in the male, and anterior third quarter of sternite III in the female. Sternites smooth, without carinae, but with two longitudinal furrows.
</paragraph>
<paragraph id="8BD636FF4510CD21FCA8F9E3FB9CF881" blockId="8.[816,1475,234,2019]" pageId="8" pageNumber="7">
<emphasis id="B91DEAED4510CD21FCA8F9E3FC30F920" bold="true" box="[816,937,1642,1666]" pageId="8" pageNumber="7">Chelicerae</emphasis>
(
<figureCitation id="13522A7A4510CD21FC5AF9E2FBD4F921" box="[962,1101,1643,1667]" captionStart="Figures 2528" captionStartId="9.[116,193,1994,2016]" captionTargetBox="[122,1474,251,1978]" captionTargetId="figure-43@9.[113,1474,234,1992]" captionTargetPageId="9" captionText="Figures 2528. Pandinurus awalei sp. n., male holotype, dorsal (25) and ventral (26) views, and right chelicera in dorsal (27) and ventral (28) aspects. Scale bar: 10 mm (2526)." figureDoi="http://doi.org/10.5281/zenodo.4648842" httpUri="https://zenodo.org/record/4648842/files/figure.png" pageId="8" pageNumber="7">Figs. 2728</figureCitation>
,
<figureCitation id="13522A7A4510CD21FBF9F9E3FB31F920" box="[1121,1192,1642,1666]" captionStart="Figures 5466" captionStartId="14.[116,193,1955,1977]" captionTargetBox="[113,1454,238,1947]" captionTargetId="figure-107@14.[113,1474,227,1949]" captionTargetPageId="14" captionText="Figures 5466: Pandinurus awalei sp. n. Figures 5461, 6466. Female paratype, pedipalp chela dorsal (54), external (55) and ventral (56) views, patella dorsal (57), external (58) and ventral (59) views, femur and trochanter dorsal (60), and ventral (61) views, movable finger dentition under white light (64); right chelicera dorsal (65) and ventral (66) aspects. Trichobothrial pattern indicated in Figures 5459. Figures 6263. Male holotype, movable finger dentition under UV (62) and white (63) light." figureDoi="http://doi.org/10.5281/zenodo.4648854" httpUri="https://zenodo.org/record/4648854/files/figure.png" pageId="8" pageNumber="7">6566</figureCitation>
). Movable finger dorsal margin with one large subdistal (
<emphasis id="B91DEAED4510CD21FB00F902FB29F901" box="[1176,1200,1675,1699]" italics="true" pageId="8" pageNumber="7">sd</emphasis>
) denticle; ventral margin smooth; ventral distal (
<emphasis id="B91DEAED4510CD21FBA1F922FBCBF961" box="[1081,1106,1707,1731]" italics="true" pageId="8" pageNumber="7">vd</emphasis>
) denticle longer than prominent dorsal (
<emphasis id="B91DEAED4510CD21FC1AF942FC04F941" box="[898,925,1739,1763]" italics="true" pageId="8" pageNumber="7">dd</emphasis>
) denticle. Fixed finger with four denticles, median (
<emphasis id="B91DEAED4510CD21FCA1F962FCD5F8A0" box="[825,844,1771,1794]" italics="true" pageId="8" pageNumber="7">m</emphasis>
) and basal (
<emphasis id="B91DEAED4510CD21FC7DF962FC6BF8A1" box="[997,1010,1771,1795]" italics="true" pageId="8" pageNumber="7">b</emphasis>
) denticles fused into bicusp; no ventral accessory denticles.
</paragraph>
<paragraph id="8BD636FF4510CD21FCA8F8A2FABBF841" blockId="8.[816,1475,234,2019]" pageId="8" pageNumber="7">
<emphasis id="B91DEAED4510CD21FCA8F8A2FCFCF8E1" bold="true" box="[816,869,1835,1859]" pageId="8" pageNumber="7">Legs</emphasis>
(
<figureCitation id="13522A7A4510CD21FCEDF8A2FC61F8E0" box="[885,1016,1834,1859]" captionStart="Figures 3139" captionStartId="11.[115,192,1985,2007]" captionTargetBox="[115,1476,231,1970]" captionTargetId="figure-76@11.[115,1476,231,1982]" captionTargetPageId="11" captionText="Figures 3139. Pandinurus awalei sp. n. Figures 31, 33, 3538. Male holotype, carapace and tergites IIV (31), coxosternal area and sternites (33), and left legs IIV, retrolateral aspect (3538). Figures 32, 34, and 39. Female paratype, carapace and tergites IIII (32), coxosternal area and sternites IIIIV (34), and left leg IV, retrolateral aspect (39)." figureDoi="http://doi.org/10.5281/zenodo.4648846" httpUri="https://zenodo.org/record/4648846/files/figure.png" pageId="8" pageNumber="7">Figs. 3539</figureCitation>
). All legs without distinct carinae, smooth. Tarsomeres hirsute, with setae and macrosetae. Spiniform formula of tarsomere II = 6-7/4: 6-7/4: 7/5: 78/5. Tarsomere II with 2 spines on inclined anteroventral surface but there could be an additional spiniform seta on external margin indicated by another not well developed spina.
</paragraph>
<caption id="DF1666774511CD20FFECF843FE5EF85F" ID-DOI="http://doi.org/10.5281/zenodo.4648842" ID-Zenodo-Dep="4648842" httpUri="https://zenodo.org/record/4648842/files/figure.png" pageId="9" pageNumber="8" startId="9.[116,193,1994,2016]" targetBox="[122,1474,251,1978]" targetPageId="9">
<paragraph id="8BD636FF4511CD20FFECF843FE5EF85F" blockId="9.[116,1474,1994,2045]" pageId="9" pageNumber="8">
<emphasis id="B91DEAED4511CD20FFECF843FE9BF87D" bold="true" box="[116,258,1994,2016]" pageId="9" pageNumber="8">Figures 2528</emphasis>
.
<taxonomicName id="4C694D7C4511CD20FE96F843FE5AF87D" authority="Kovařík &amp; Lowe &amp; Elmi, 2020" authorityName="Kovařík &amp; Lowe &amp; Elmi" authorityYear="2020" box="[270,451,1994,2015]" class="Arachnida" family="Scorpionidae" genus="Pandinurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="awalei" status="sp. nov.">
<emphasis id="B91DEAED4511CD20FE96F843FE5AF87D" box="[270,451,1994,2015]" italics="true" pageId="9" pageNumber="8">Pandinurus awalei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57964511CD20FE51F843FE64F87D" box="[457,509,1994,2015]" pageId="9" pageNumber="8" rank="species">
<emphasis id="B91DEAED4511CD20FE51F843FE79F87D" bold="true" box="[457,480,1994,2015]" pageId="9" pageNumber="8">sp</emphasis>
.
<emphasis id="B91DEAED4511CD20FE73F843FE61F87D" bold="true" box="[491,504,1994,2015]" pageId="9" pageNumber="8">n</emphasis>
.
</taxonomicNameLabel>
, male holotype, dorsal (25) and ventral (26) views, and right chelicera in dorsal (27) and ventral (28) aspects. Scale bar: 10 mm (2526).
</paragraph>
</caption>
<caption id="DF1666774512CD23FFECF9E3FB5EF922" ID-DOI="http://doi.org/10.5281/zenodo.4648844" ID-Zenodo-Dep="4648844" box="[116,1223,1642,1665]" httpUri="https://zenodo.org/record/4648844/files/figure.png" pageId="10" pageNumber="9" startId="10.[116,193,1642,1664]" targetBox="[129,1446,240,1626]" targetPageId="10">
<paragraph id="8BD636FF4512CD23FFECF9E3FB5EF922" blockId="10.[116,1223,1642,1665]" box="[116,1223,1642,1665]" pageId="10" pageNumber="9">
<emphasis id="B91DEAED4512CD23FFECF9E3FE9AF9DD" bold="true" box="[116,259,1642,1664]" pageId="10" pageNumber="9">Figures 2930</emphasis>
.
<taxonomicName id="4C694D7C4512CD23FE97F9E2FE5CF922" authority="Kovařík &amp; Lowe &amp; Elmi, 2020" authorityName="Kovařík &amp; Lowe &amp; Elmi" authorityYear="2020" box="[271,453,1643,1664]" class="Arachnida" family="Scorpionidae" genus="Pandinurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="awalei" status="sp. nov.">
<emphasis id="B91DEAED4512CD23FE97F9E2FE5CF922" box="[271,453,1643,1664]" italics="true" pageId="10" pageNumber="9">Pandinurus awalei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57964512CD23FE53F9E2FD99F922" box="[459,512,1643,1664]" pageId="10" pageNumber="9" rank="species">
<emphasis id="B91DEAED4512CD23FE53F9E2FE7BF922" bold="true" box="[459,482,1643,1664]" pageId="10" pageNumber="9">sp</emphasis>
.
<emphasis id="B91DEAED4512CD23FE75F9E2FE63F922" bold="true" box="[493,506,1643,1664]" pageId="10" pageNumber="9">n</emphasis>
.
</taxonomicNameLabel>
, female paratype in dorsal (29) and ventral (30) views. Scale bar: 10 mm.
</paragraph>
</caption>
<paragraph id="8BD636FF4512CD23FFECF928FBDAF87B" blockId="10.[116,775,1697,2041]" lastBlockId="10.[819,1478,1697,2041]" pageId="10" pageNumber="9">
<emphasis id="B91DEAED4512CD23FFECF928FEC2F91B" bold="true" box="[116,347,1697,1721]" pageId="10" pageNumber="9">Hemispermatophore</emphasis>
. (
<figureCitation id="13522A7A4512CD23FEE5F928FD93F91B" box="[381,522,1697,1721]" captionStart-0="Figures 6769" captionStart-1="Figures 7072" captionStartId-0="15.[115,192,1990,2012]" captionStartId-1="16.[113,190,1385,1407]" captionTargetBox-0="[205,1387,241,1976]" captionTargetBox-1="[152,1437,259,1352]" captionTargetId-0="figure-35@15.[113,1474,229,1979]" captionTargetId-1="figure-354@16.[113,1474,244,1368]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="Figures 6769. Pandinurus awalei sp. n., left whole hemispermatophore, anterior (67), posterior-concave (68) and convex (69) views. Scale bar: 4 mm." captionText-1="Figures 7072. Pandinurus awalei sp. n., left hemispermatophore, capsule, anterior (70), concave (71) and convex (72) views. Scale bar: 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4648856" figureDoi-1="http://doi.org/10.5281/zenodo.4648858" httpUri-0="https://zenodo.org/record/4648856/files/figure.png" httpUri-1="https://zenodo.org/record/4648858/files/figure.png" pageId="10" pageNumber="9">Figs. 6772</figureCitation>
). Lamelliform. Distal lamina long, with robust hook on anterior margin near base. Lamina constricted immediately distal to hook, but gradually widening more distally. Apex of lamina with sharp posterior deflection at ca. 70° angle relative to proximal axis, tapering to a narrow, slightly bent tip. Portion of distal lamina proximal to hook with broad, shallow trough on convex side.Truncal flexure, tectum, hemisolenos and clasper strongly developed. Trunk relatively short, ca. half of length of distal lamina, gradually tapered towards base, with weakly sclerotized diagonal axial rib. Proximal 2/3 of axial rib running along posterior margin of trunk. Both left and right hemispermatophores displayed similar morphology. Measurements of left hemispermatophore (mm): distal lamina: total length from truncal flexure 9.90; length from truncal flexure to base of hook 2.39; length from base of hook to apical deflection 6.37; width immediately proximal to hook 0.97, immediately distal to hook 0.69; width at apical deflection 1.03. Trunk: length 3.47; distal width 1.47, basal width 0.77. Pedicel: length 1.91. Morphometric ratios: distal lamina total length/ trunk length 2.85; distal lamina proximal to hook length/ distal to hook length 3.08.
</paragraph>
<paragraph id="8BD636FF4512CD23FCABF868FBF3F85B" blockId="10.[819,1478,1697,2041]" box="[819,1130,2017,2041]" pageId="10" pageNumber="9">
<emphasis id="B91DEAED4512CD23FCABF868FC43F85B" bold="true" box="[819,986,2017,2041]" pageId="10" pageNumber="9">Measurements</emphasis>
. See
<tableCitation id="C6EB03444512CD23FB8CF868FBFAF85B" box="[1044,1123,2017,2041]" captionStart="Table 1" captionStartId="5.[113,169,996,1017]" captionTargetId="graphics-572@5.[112,1477,237,957]" captionTargetPageId="5" captionText="Table 1. Comparative measurements of adults of Pandiborellius somalilandus and Pandinurus awalei sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." httpUri="http://table.plazi.org/id/DF166677451DCD2CFFE9FC6DFD25FBB5" pageId="10" pageNumber="9" tableUuid="DF166677451DCD2CFFE9FC6DFD25FBB5">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<caption id="DF1666774513CD22FFEBF848FD45F7B3" ID-DOI="http://doi.org/10.5281/zenodo.4648846" ID-Zenodo-Dep="4648846" httpUri="https://zenodo.org/record/4648846/files/figure.png" pageId="11" pageNumber="10" startId="11.[115,192,1985,2007]" targetBox="[115,1476,231,1970]" targetPageId="11">
<paragraph id="8BD636FF4513CD22FFEBF848FD45F7B3" blockId="11.[115,1473,1985,2065]" pageId="11" pageNumber="10">
<emphasis id="B91DEAED4513CD22FFEBF848FE98F874" bold="true" box="[115,257,1985,2007]" pageId="11" pageNumber="10">Figures 3139</emphasis>
.
<taxonomicName id="4C694D7C4513CD22FE94F848FE58F874" authority="Kovařík &amp; Lowe &amp; Elmi, 2020" authorityName="Kovařík &amp; Lowe &amp; Elmi" authorityYear="2020" box="[268,449,1985,2006]" class="Arachnida" family="Scorpionidae" genus="Pandinurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="awalei" status="sp. nov.">
<emphasis id="B91DEAED4513CD22FE94F848FE58F874" box="[268,449,1985,2006]" italics="true" pageId="11" pageNumber="10">Pandinurus awalei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57964513CD22FE5EF848FE65F875" box="[454,508,1985,2007]" pageId="11" pageNumber="10" rank="species">
<emphasis id="B91DEAED4513CD22FE5EF848FE44F874" bold="true" box="[454,477,1985,2006]" pageId="11" pageNumber="10">sp</emphasis>
.
<emphasis id="B91DEAED4513CD22FE70F848FE6CF874" bold="true" box="[488,501,1985,2006]" pageId="11" pageNumber="10">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="B91DEAED4513CD22FD99F848FDF2F874" bold="true" box="[513,619,1985,2007]" pageId="11" pageNumber="10">Figures 31</emphasis>
,
<emphasis id="B91DEAED4513CD22FDEEF848FD17F874" bold="true" box="[630,654,1985,2006]" pageId="11" pageNumber="10">33</emphasis>
,
<emphasis id="B91DEAED4513CD22FD01F848FD4CF874" bold="true" box="[665,725,1985,2007]" pageId="11" pageNumber="10">3538</emphasis>
. Male holotype, carapace and tergites IIV (31), coxosternal area and sternites (33), and left legs IIV, retrolateral aspect (3538).
<emphasis id="B91DEAED4513CD22FDF8F857FD53F851" bold="true" box="[608,714,2014,2036]" pageId="11" pageNumber="10">Figures 32</emphasis>
,
<emphasis id="B91DEAED4513CD22FD4EF857FD77F851" bold="true" box="[726,750,2014,2035]" pageId="11" pageNumber="10">34</emphasis>
, and
<emphasis id="B91DEAED4513CD22FCBAF857FCA3F851" bold="true" box="[802,826,2014,2035]" pageId="11" pageNumber="10">39</emphasis>
. Female paratype, carapace and tergites IIII (32), coxosternal area and sternites IIIIV (34), and left leg IV, retrolateral aspect (39).
</paragraph>
</caption>
<caption id="DF1666774514CD25FFE9F82FFDAFF854" ID-DOI="http://doi.org/10.5281/zenodo.4648850" ID-Zenodo-Dep="4648850" httpUri="https://zenodo.org/record/4648850/files/figure.png" pageId="12" pageNumber="11" startId="12.[113,190,1958,1980]" targetBox="[131,1464,270,1922]" targetPageId="12">
<paragraph id="8BD636FF4514CD25FFE9F82FFDAFF854" blockId="12.[113,1472,1958,2038]" pageId="12" pageNumber="11">
<emphasis id="B91DEAED4514CD25FFE9F82FFE99F819" bold="true" box="[113,256,1958,1980]" pageId="12" pageNumber="11">Figures 4047</emphasis>
:
<taxonomicName id="4C694D7C4514CD25FE95F82FFE5AF819" authority="Kovařík &amp; Lowe &amp; Elmi, 2020" authorityName="Kovařík &amp; Lowe &amp; Elmi" authorityYear="2020" box="[269,451,1958,1979]" class="Arachnida" family="Scorpionidae" genus="Pandinurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="awalei" status="sp. nov.">
<emphasis id="B91DEAED4514CD25FE95F82FFE5AF819" box="[269,451,1958,1979]" italics="true" pageId="12" pageNumber="11">Pandinurus awalei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57964514CD25FE51F82FFE66F81E" box="[457,511,1958,1980]" pageId="12" pageNumber="11" rank="species">
<emphasis id="B91DEAED4514CD25FE51F82FFE79F819" bold="true" box="[457,480,1958,1979]" pageId="12" pageNumber="11">sp</emphasis>
.
<emphasis id="B91DEAED4514CD25FE73F82FFE61F819" bold="true" box="[491,504,1958,1979]" pageId="12" pageNumber="11">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="B91DEAED4514CD25FD9DF82FFDF6F819" bold="true" box="[517,623,1958,1980]" pageId="12" pageNumber="11">Figures 40</emphasis>
,
<emphasis id="B91DEAED4514CD25FDE3F82FFD2EF819" bold="true" box="[635,695,1958,1980]" pageId="12" pageNumber="11">4547</emphasis>
. Male holotype, telson lateral (40), and metasoma and telson lateral (45), ventral (46), and dorsal (47) views.
<emphasis id="B91DEAED4514CD25FEE7F84AFD97F87A" bold="true" box="[383,526,1987,2009]" pageId="12" pageNumber="11">Figures 4144</emphasis>
. Paratype female, telson lateral (41), and metasoma and telson lateral (42), ventral (43), and dorsal (44) views. Scale bars: 10 mm (4244, 4547).
</paragraph>
</caption>
<caption id="DF1666774515CD24FFECF837FC53F853" ID-DOI="http://doi.org/10.5281/zenodo.4648852" ID-Zenodo-Dep="4648852" httpUri="https://zenodo.org/record/4648852/files/figure.png" pageId="13" pageNumber="12" startId="13.[116,193,1982,2004]" targetBox="[113,1474,252,1973]" targetPageId="13">
<paragraph id="8BD636FF4515CD24FFECF837FC53F853" blockId="13.[116,1474,1982,2033]" pageId="13" pageNumber="12">
<emphasis id="B91DEAED4515CD24FFECF837FE9BF876" bold="true" box="[116,258,1982,2004]" pageId="13" pageNumber="12">Figures 4852</emphasis>
:
<taxonomicName id="4C694D7C4515CD24FE97F836FE5DF876" authority="Kovařík &amp; Lowe &amp; Elmi, 2020" authorityName="Kovařík &amp; Lowe &amp; Elmi" authorityYear="2020" box="[271,452,1983,2004]" class="Arachnida" family="Scorpionidae" genus="Pandinurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="awalei" status="sp. nov.">
<emphasis id="B91DEAED4515CD24FE97F836FE5DF876" box="[271,452,1983,2004]" italics="true" pageId="13" pageNumber="12">Pandinurus awalei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57964515CD24FE52F836FD99F876" box="[458,512,1983,2004]" pageId="13" pageNumber="12" rank="species">
<emphasis id="B91DEAED4515CD24FE52F836FE78F876" bold="true" box="[458,481,1983,2004]" pageId="13" pageNumber="12">sp</emphasis>
.
<emphasis id="B91DEAED4515CD24FE74F836FE60F876" bold="true" box="[492,505,1983,2004]" pageId="13" pageNumber="12">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="B91DEAED4515CD24FD9DF837FD0DF876" bold="true" box="[517,660,1982,2004]" pageId="13" pageNumber="12">Figures 4852</emphasis>
. Male holotype, pedipalp chela and patella dorsal (48), external (49) and ventral (50) views, femur and trochanter dorsal (51), and ventral (52) views.
<emphasis id="B91DEAED4515CD24FD47F852FCD8F852" bold="true" box="[735,833,2011,2033]" pageId="13" pageNumber="12">Figure 53</emphasis>
. Type locality
</paragraph>
</caption>
<caption id="DF1666774516CD27FFECF82AFC04F7AD" ID-DOI="http://doi.org/10.5281/zenodo.4648854" ID-Zenodo-Dep="4648854" httpUri="https://zenodo.org/record/4648854/files/figure.png" pageId="14" pageNumber="13" startId="14.[116,193,1955,1977]" targetBox="[113,1454,238,1947]" targetPageId="14">
<paragraph id="8BD636FF4516CD27FFECF82AFC04F7AD" blockId="14.[116,1474,1955,2063]" pageId="14" pageNumber="13">
<emphasis id="B91DEAED4516CD27FFECF82AFE9AF81A" bold="true" box="[116,259,1955,1977]" pageId="14" pageNumber="13">Figures 5466</emphasis>
:
<taxonomicName id="4C694D7C4516CD27FE89F82AFE5EF81A" authority="Kovařík &amp; Lowe &amp; Elmi, 2020" authorityName="Kovařík &amp; Lowe &amp; Elmi" authorityYear="2020" box="[273,455,1955,1976]" class="Arachnida" family="Scorpionidae" genus="Pandinurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="awalei" status="sp. nov.">
<emphasis id="B91DEAED4516CD27FE89F82AFE5EF81A" box="[273,455,1955,1976]" italics="true" pageId="14" pageNumber="13">Pandinurus awalei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57964516CD27FE56F82AFD9CF81A" box="[462,517,1955,1976]" pageId="14" pageNumber="13" rank="species">
<emphasis id="B91DEAED4516CD27FE56F82AFE7CF81A" bold="true" box="[462,485,1955,1976]" pageId="14" pageNumber="13">sp</emphasis>
.
<emphasis id="B91DEAED4516CD27FE69F82AFE67F81A" bold="true" box="[497,510,1955,1976]" pageId="14" pageNumber="13">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="B91DEAED4516CD27FD93F82AFD02F81A" bold="true" box="[523,667,1955,1977]" pageId="14" pageNumber="13">Figures 5461</emphasis>
,
<emphasis id="B91DEAED4516CD27FD3FF82AFD7AF81A" bold="true" box="[679,739,1955,1976]" pageId="14" pageNumber="13">6466</emphasis>
. Female paratype, pedipalp chela dorsal (54), external (55) and ventral (56) views, patella dorsal (57), external (58) and ventral (59) views, femur and trochanter dorsal (60), and ventral (61) views, movable finger dentition under white light (64); right chelicera dorsal (65) and ventral (66) aspects. Trichobothrial pattern indicated in Figures 5459.
<emphasis id="B91DEAED4516CD27FAEDF855FF29F7AC" bold="true" pageId="14" pageNumber="13">Figures 6263</emphasis>
. Male holotype, movable finger dentition under UV (62) and white (63) light.
</paragraph>
</caption>
<caption id="DF1666774517CD26FFEBF84FFF45F85B" ID-DOI="http://doi.org/10.5281/zenodo.4648856" ID-Zenodo-Dep="4648856" httpUri="https://zenodo.org/record/4648856/files/figure.png" pageId="15" pageNumber="14" startId="15.[115,192,1990,2012]" targetBox="[205,1387,241,1976]" targetPageId="15">
<paragraph id="8BD636FF4517CD26FFEBF84FFF45F85B" blockId="15.[115,1473,1990,2041]" pageId="15" pageNumber="14">
<emphasis id="B91DEAED4517CD26FFEBF84FFE9BF879" bold="true" box="[115,258,1990,2012]" pageId="15" pageNumber="14">Figures 6769</emphasis>
.
<taxonomicName id="4C694D7C4517CD26FE96F84FFE5CF879" authority="Kovařík &amp; Lowe &amp; Elmi, 2020" authorityName="Kovařík &amp; Lowe &amp; Elmi" authorityYear="2020" box="[270,453,1990,2011]" class="Arachnida" family="Scorpionidae" genus="Pandinurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="awalei" status="sp. nov.">
<emphasis id="B91DEAED4517CD26FE96F84FFE5CF879" box="[270,453,1990,2011]" italics="true" pageId="15" pageNumber="14">Pandinurus awalei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57964517CD26FE53F84FFD99F87E" box="[459,512,1990,2012]" pageId="15" pageNumber="14" rank="species">
<emphasis id="B91DEAED4517CD26FE53F84FFE7BF879" bold="true" box="[459,482,1990,2011]" pageId="15" pageNumber="14">sp</emphasis>
.
<emphasis id="B91DEAED4517CD26FE76F84FFE62F879" bold="true" box="[494,507,1990,2011]" pageId="15" pageNumber="14">n</emphasis>
.
</taxonomicNameLabel>
, left whole hemispermatophore, anterior (67), posterior-concave (68) and convex (69) views. Scale bar: 4 mm.
</paragraph>
</caption>
<caption id="DF1666774508CD39FFE9FAE0FF07FA3E" ID-DOI="http://doi.org/10.5281/zenodo.4648858" ID-Zenodo-Dep="4648858" httpUri="https://zenodo.org/record/4648858/files/figure.png" pageId="16" pageNumber="15" startId="16.[113,190,1385,1407]" targetBox="[152,1437,259,1352]" targetPageId="16">
<paragraph id="8BD636FF4508CD39FFE9FAE0FF07FA3E" blockId="16.[113,1471,1385,1436]" pageId="16" pageNumber="15">
<emphasis id="B91DEAED4508CD39FFE9FAE0FE98FADC" bold="true" box="[113,257,1385,1407]" pageId="16" pageNumber="15">Figures 7072</emphasis>
.
<taxonomicName id="4C694D7C4508CD39FE96FAE0FE5CFADC" authority="Kovařík &amp; Lowe &amp; Elmi, 2020" authorityName="Kovařík &amp; Lowe &amp; Elmi" authorityYear="2020" box="[270,453,1385,1406]" class="Arachnida" family="Scorpionidae" genus="Pandinurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="awalei" status="sp. nov.">
<emphasis id="B91DEAED4508CD39FE96FAE0FE5CFADC" box="[270,453,1385,1406]" italics="true" pageId="16" pageNumber="15">Pandinurus awalei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57964508CD39FE54FAE3FD9BFADD" box="[460,514,1386,1407]" pageId="16" pageNumber="15" rank="species">
<emphasis id="B91DEAED4508CD39FE54FAE3FE7AFADD" bold="true" box="[460,483,1386,1407]" pageId="16" pageNumber="15">sp</emphasis>
.
<emphasis id="B91DEAED4508CD39FE68FAE3FE64FADD" bold="true" box="[496,509,1386,1407]" pageId="16" pageNumber="15">n</emphasis>
.
</taxonomicNameLabel>
, left hemispermatophore, capsule, anterior (70), concave (71) and convex (72) views. Scale bar: 1 mm.
</paragraph>
</caption>
<subSubSection id="C37365744508CD39FFE9FA30FBB8F852" pageId="16" pageNumber="15" type="discussion">
<paragraph id="8BD636FF4508CD39FFE9FA30FE3FF9F3" blockId="16.[113,771,1465,1617]" pageId="16" pageNumber="15">
REMARKS. Overall structure of the hemispermatophore of
<taxonomicName id="4C694D7C4508CD39FFE9FA50FEA2FA53" authority="Kovařík &amp; Lowe &amp; Elmi, 2020" authorityName="Kovařík &amp; Lowe &amp; Elmi" authorityYear="2020" box="[113,315,1497,1521]" class="Arachnida" family="Scorpionidae" genus="Pandinurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="awalei" status="sp. nov.">
<emphasis id="B91DEAED4508CD39FFE9FA50FEA2FA53" box="[113,315,1497,1521]" italics="true" pageId="16" pageNumber="15">Pandinurus awalei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57964508CD39FED9FA50FEE4FA52" box="[321,381,1497,1521]" pageId="16" pageNumber="15" rank="species">
<emphasis id="B91DEAED4508CD39FED9FA50FEC3FA53" bold="true" box="[321,346,1497,1521]" pageId="16" pageNumber="15">sp</emphasis>
.
<emphasis id="B91DEAED4508CD39FEFFFA50FEEFFA53" bold="true" box="[359,374,1497,1521]" pageId="16" pageNumber="15">n</emphasis>
.
</taxonomicNameLabel>
is quite similar to those recorded for seven other
<taxonomicName id="4C694D7C4508CD39FF61FA70FEECF9B3" authorityName="Fet" authorityYear="1997" box="[249,373,1529,1553]" class="Arachnida" family="Scorpionidae" genus="Pandinurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEAED4508CD39FF61FA70FEECF9B3" box="[249,373,1529,1553]" italics="true" pageId="16" pageNumber="15">Pandinurus</emphasis>
</taxonomicName>
spp. (KovařÍk et al., 2017c), which also have a long, narrow distal lamina that is sharply deflected at its apex, and a short trunk.
</paragraph>
<paragraph id="8BD636FF4508CD39FFE9F9F0FEEBF8D3" blockId="16.[113,771,1657,2033]" pageId="16" pageNumber="15">
AFFINITIES.
<taxonomicName id="4C694D7C4508CD39FF68F9F0FE23F933" authority="Kovařík &amp; Lowe &amp; Elmi, 2020" authorityName="Kovařík &amp; Lowe &amp; Elmi" authorityYear="2020" box="[240,442,1657,1681]" class="Arachnida" family="Scorpionidae" genus="Pandinurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="awalei" status="sp. nov.">
<emphasis id="B91DEAED4508CD39FF68F9F0FE23F933" box="[240,442,1657,1681]" italics="true" pageId="16" pageNumber="15">Pandinurus awalei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57964508CD39FE59F9F0FE64F932" box="[449,509,1657,1681]" pageId="16" pageNumber="15" rank="species">
<emphasis id="B91DEAED4508CD39FE59F9F0FE43F933" bold="true" box="[449,474,1657,1681]" pageId="16" pageNumber="15">sp</emphasis>
.
<emphasis id="B91DEAED4508CD39FE70F9F0FE6EF933" bold="true" box="[488,503,1657,1681]" pageId="16" pageNumber="15">n</emphasis>
.
</taxonomicNameLabel>
is reliably distinguished from all other
<taxonomicName id="4C694D7C4508CD39FE8FF910FE0AF913" authorityName="Fet" authorityYear="1997" box="[279,403,1689,1713]" class="Arachnida" family="Scorpionidae" genus="Pandinurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEAED4508CD39FE8FF910FE0AF913" box="[279,403,1689,1713]" italics="true" pageId="16" pageNumber="15">Pandinurus</emphasis>
</taxonomicName>
species by the following unique combination of characters: accessory external trichobothrium
<emphasis id="B91DEAED4508CD39FFE9F953FF13F953" box="[113,138,1754,1777]" italics="true" pageId="16" pageNumber="15">ea</emphasis>
on pedipalp chela absent; granules on dorsal surface of chela of pedipalp conical and rounded, not pointed; legs yellow; pedipalp chela densely hirsute; pedipalp chela of male length/ width ratio 1.87; spiniform formula of tarsomere II = 6-7/4: 6-7/4: 7/5: 78/5.
</paragraph>
<paragraph id="8BD636FF4508CD39FF39F8F0FB9DF973" blockId="16.[113,771,1657,2033]" lastBlockId="16.[816,1474,1465,1745]" pageId="16" pageNumber="15">
The
<typeStatus id="54D2885D4508CD39FF46F8F0FE94F833" box="[222,269,1913,1937]" pageId="16" pageNumber="15">type</typeStatus>
locality of
<taxonomicName id="4C694D7C4508CD39FE33F8F0FD1DF833" authority="Kovařík &amp; Lowe &amp; Elmi, 2020" authorityName="Kovařík &amp; Lowe &amp; Elmi" authorityYear="2020" box="[427,644,1913,1937]" class="Arachnida" family="Scorpionidae" genus="Pandinurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="awalei" status="sp. nov.">
<emphasis id="B91DEAED4508CD39FE33F8F0FD1DF833" box="[427,644,1913,1937]" italics="true" pageId="16" pageNumber="15">Pandinurus awalei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57964508CD39FD0FF8F0FD46F832" box="[663,735,1913,1937]" pageId="16" pageNumber="15" rank="species">
<emphasis id="B91DEAED4508CD39FD0FF8F0FD29F833" bold="true" box="[663,688,1913,1937]" pageId="16" pageNumber="15">sp</emphasis>
.
<emphasis id="B91DEAED4508CD39FD51F8F0FD41F833" bold="true" box="[713,728,1913,1937]" pageId="16" pageNumber="15">n</emphasis>
.
</taxonomicNameLabel>
is relatively close to known areas of distribution of
<taxonomicName id="4C694D7C4508CD39FDE2F813FEC8F873" authority="KovarIk et al., 2017" authorityName="KovarIk" authorityYear="2017" class="Arachnida" family="Scorpionidae" genus="Pandinurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="hangarale">
<emphasis id="B91DEAED4508CD39FDE2F813FC9AF813" box="[634,771,1945,1969]" italics="true" pageId="16" pageNumber="15">P. hangarale</emphasis>
KovařÍk et al., 2017
</taxonomicName>
,
<taxonomicName id="4C694D7C4508CD39FEC5F833FD49F873" authority="KovarIk et al., 2017" authorityName="KovarIk" authorityYear="2017" box="[349,720,1977,2003]" class="Arachnida" family="Scorpionidae" genus="Pandinurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="kmoniceki">
<emphasis id="B91DEAED4508CD39FEC5F833FE7EF873" box="[349,487,1977,2001]" italics="true" pageId="16" pageNumber="15">P. kmoniceki</emphasis>
KovařÍk et al., 2017
</taxonomicName>
, and
<taxonomicName id="4C694D7C4508CD39FFE9F853FE3EF853" authority="(Pocock, 1896)" baseAuthorityName="Pocock" baseAuthorityYear="1896" box="[113,423,2009,2033]" class="Arachnida" family="Scorpionidae" genus="Pandinurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="phillipsi">
<emphasis id="B91DEAED4508CD39FFE9F853FF75F853" box="[113,236,2009,2033]" italics="true" pageId="16" pageNumber="15">P. phillipsi</emphasis>
(Pocock, 1896)
</taxonomicName>
. These other species differ as follows:
<taxonomicName id="4C694D7C4508CD39FC0EFA33FBBEFA73" box="[918,1063,1465,1489]" class="Arachnida" family="Scorpionidae" genus="Pandinurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="hangarale">
<emphasis id="B91DEAED4508CD39FC0EFA33FBBEFA73" box="[918,1063,1465,1489]" italics="true" pageId="16" pageNumber="15">P. hangarale</emphasis>
</taxonomicName>
has the whole body including legs dark (figs.
<quantity id="4C919B1A4508CD39FC32FA50FB95FA53" box="[938,1036,1497,1521]" metricMagnitude="-1" metricUnit="m" metricValue="5.715" metricValueMax="5.842" metricValueMin="5.588" pageId="16" pageNumber="15" unit="in" value="22.5" valueMax="23.0" valueMin="22.0">2223 in</quantity>
KovařÍk et al., 2017a: 7) and has male pedipalp chela length/ width ratio of 1.59;
<taxonomicName id="4C694D7C4508CD39FA91FA73FA0AF9B3" box="[1289,1427,1529,1553]" class="Arachnida" family="Scorpionidae" genus="Pandinurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="kmoniceki">
<emphasis id="B91DEAED4508CD39FA91FA73FA0AF9B3" box="[1289,1427,1529,1553]" italics="true" pageId="16" pageNumber="15">P. kmoniceki</emphasis>
</taxonomicName>
and
<taxonomicName id="4C694D7C4508CD39FCA8F993FC3CF993" baseAuthorityName="Pocock" baseAuthorityYear="1896" box="[816,933,1561,1585]" class="Arachnida" family="Scorpionidae" genus="Pandinurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="phillipsi">
<emphasis id="B91DEAED4508CD39FCA8F993FC3CF993" box="[816,933,1561,1585]" italics="true" pageId="16" pageNumber="15">P. phillipsi</emphasis>
</taxonomicName>
have accessory external trichobothrium
<emphasis id="B91DEAED4508CD39FAC7F993FAE1F993" box="[1375,1400,1562,1585]" italics="true" pageId="16" pageNumber="15">ea</emphasis>
on the chela present and located between trichobothria
<emphasis id="B91DEAED4508CD39FAD3F9B0FAF6F9F3" box="[1355,1391,1593,1617]" italics="true" pageId="16" pageNumber="15">esb</emphasis>
and
<emphasis id="B91DEAED4508CD39FA31F9B0FA5BF9F3" box="[1449,1474,1593,1617]" italics="true" pageId="16" pageNumber="15">eb</emphasis>
on the base of the fixed finger (fig.
<quantity id="4C919B1A4508CD39FB30F9D0FB49F9D3" box="[1192,1232,1625,1649]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="16" pageNumber="15" unit="in" value="6.0">6 in</quantity>
KovařÍk et al., 2017b: 4), and have a pedipalp chela that is not as densely hirsute as in
<taxonomicName id="4C694D7C4508CD39FCE8F913FC4EF913" authority="Kovařík &amp; Lowe &amp; Elmi, 2020" authorityName="Kovařík &amp; Lowe &amp; Elmi" authorityYear="2020" box="[880,983,1689,1713]" class="Arachnida" family="Scorpionidae" genus="Pandinurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="awalei" status="sp. nov.">
<emphasis id="B91DEAED4508CD39FCE8F913FC4EF913" box="[880,983,1689,1713]" italics="true" pageId="16" pageNumber="15">P. awalei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57964508CD39FC79F910FBBBF912" box="[993,1058,1689,1713]" pageId="16" pageNumber="15" rank="species">
<emphasis id="B91DEAED4508CD39FC79F910FC63F913" bold="true" box="[993,1018,1689,1713]" pageId="16" pageNumber="15">sp</emphasis>
.
<emphasis id="B91DEAED4508CD39FB94F910FB82F913" bold="true" box="[1036,1051,1689,1713]" pageId="16" pageNumber="15">n</emphasis>
.
</taxonomicNameLabel>
These species also differ genetically (unpublished data).
</paragraph>
<paragraph id="8BD636FF4508CD39FCA8F970FBB8F852" blockId="16.[816,1474,1785,2033]" pageId="16" pageNumber="15">
COMMENTS ON LOCALITIES AND LIFE STRATEGY.
<taxonomicName id="4C694D7C4508CD39FAD9F970FCE3F893" authority="Kovařík &amp; Lowe &amp; Elmi, 2020" authorityName="Kovařík &amp; Lowe &amp; Elmi" authorityYear="2020" class="Arachnida" family="Scorpionidae" genus="Pandinurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="awalei" status="sp. nov.">
<emphasis id="B91DEAED4508CD39FAD9F970FCE3F893" italics="true" pageId="16" pageNumber="15">Pandinurus awalei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57964508CD39FC11F890FC56F892" box="[905,975,1817,1841]" pageId="16" pageNumber="15" rank="species">
<emphasis id="B91DEAED4508CD39FC11F890FC3AF893" bold="true" box="[905,931,1817,1841]" pageId="16" pageNumber="15">sp</emphasis>
.
<emphasis id="B91DEAED4508CD39FC21F890FC51F893" bold="true" box="[953,968,1817,1841]" pageId="16" pageNumber="15">n</emphasis>
.
</taxonomicNameLabel>
inhabits rocky mountain areas in central Somaliland. The
<typeStatus id="54D2885D4508CD39FC75F8B0FBBEF8F3" box="[1005,1063,1849,1873]" pageId="16" pageNumber="15">types</typeStatus>
were collected by day under rocks in open terrain. At this locality, the first author also recorded
<taxonomicName id="4C694D7C4508CD39FCA8F8F0FB2BF833" authority="Pocock, 1900" authorityName="Pocock" authorityYear="1900" box="[816,1202,1913,1937]" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="berberensis">
<emphasis id="B91DEAED4508CD39FCA8F8F0FB90F833" box="[816,1033,1913,1937]" italics="true" pageId="16" pageNumber="15">Buthus berberensis</emphasis>
Pocock, 1900
</taxonomicName>
,
<taxonomicName id="4C694D7C4508CD39FB59F8F3FC46F813" authority="(Pocock, 1896)" baseAuthorityName="Pocock" baseAuthorityYear="1896" class="Arachnida" family="Buthidae" genus="Hottentotta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="polystictus">
<emphasis id="B91DEAED4508CD39FB59F8F3FA5BF833" box="[1217,1474,1913,1937]" italics="true" pageId="16" pageNumber="15">Hottentotta polystictus</emphasis>
(Pocock, 1896)
</taxonomicName>
,
<taxonomicName id="4C694D7C4508CD39FC75F810FAE6F813" authority="KovarIk, 2019" authorityName="KovarIk" authorityYear="2019" box="[1005,1407,1945,1971]" class="Arachnida" family="Buthidae" genus="Neobuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="solegladi">
<emphasis id="B91DEAED4508CD39FC75F810FB4AF813" box="[1005,1235,1945,1969]" italics="true" pageId="16" pageNumber="15">Neobuthus solegladi</emphasis>
KovařÍk, 2019
</taxonomicName>
(
<typeStatus id="54D2885D4508CD39FA09F810FA5BF813" box="[1425,1474,1945,1969]" pageId="16" pageNumber="15">type</typeStatus>
locality),
<taxonomicName id="4C694D7C4508CD39FC3AF830FA24F873" authority="KovarIk et al. 2019" authorityName="KovarIk" authorityYear="2019" box="[930,1469,1977,2003]" class="Arachnida" family="Buthidae" genus="Parabuthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="somalilandus">
<emphasis id="B91DEAED4508CD39FC3AF830FB5CF873" box="[930,1221,1977,2001]" italics="true" pageId="16" pageNumber="15">Parabuthus somalilandus</emphasis>
KovařÍk et al. 2019
</taxonomicName>
, and
<taxonomicName id="4C694D7C4508CD39FCF8F853FC63F853" authorityName="Peters" authorityYear="1861" box="[864,1018,2010,2033]" class="Arachnida" family="Hemiscorpiidae" genus="Hemiscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEAED4508CD39FCF8F853FC63F853" box="[864,1018,2010,2033]" italics="true" pageId="16" pageNumber="15">Hemiscorpius</emphasis>
</taxonomicName>
sp.
</paragraph>
</subSubSection>
</subSubSection>
</treatment>
</document>