treatments-xml/data/03/C0/87/03C087E74553FF9DFF3CFDA4FE424A71.xml
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<document id="60A0B67E918723B38DB72D39A02400CA" ID-CLB-Dataset="39188" ID-DOI="10.11646/zootaxa.4105.6.1" ID-GBIF-Dataset="9f3191f8-cd86-4a17-ad5b-0c454e0acb8d" ID-ISSN="1175-5326" ID-Zenodo-Dep="271319" ID-ZooBank="DCB6A5BB-46C9-4D05-8B4A-C6E4CBABB6F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461796872515" checkinUser="plazi" docAuthor="Taylor, Robert W. &amp; Alpert, Gary D." docDate="2016" docId="03C087E74553FF9DFF3CFDA4FE424A71" docLanguage="en" docName="zootaxa.4105.6.1.pdf.imf" docOrigin="Zootaxa 4105 (6)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Metapone tillyardi Wheeler 1919" docType="treatment" docVersion="17" lastPageNumber="539" masterDocId="FFF9FF9F4577FFBBFFABFFA4FF9D4811" masterDocTitle="The myrmicine ant genus Metapone Forel (Hymenoptera: Formicidae): a global taxonomic review with descriptions of twelve new species" masterLastPageNumber="545" masterPageNumber="501" pageNumber="537" updateTime="1698662027110" updateUser="ExternalLinkService">
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<mods:title id="DBA3C074A1FC0A7270A516A733DB6D13">The myrmicine ant genus Metapone Forel (Hymenoptera: Formicidae): a global taxonomic review with descriptions of twelve new species</mods:title>
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<treatment id="03C087E74553FF9DFF3CFDA4FE424A71" ID-DOI="http://doi.org/10.5281/zenodo.5641482" ID-GBIF-Taxon="119651738" ID-Zenodo-Dep="5641482" LSID="urn:lsid:plazi:treatment:03C087E74553FF9DFF3CFDA4FE424A71" httpUri="http://treatment.plazi.org/id/03C087E74553FF9DFF3CFDA4FE424A71" lastPageId="38" lastPageNumber="539" pageId="36" pageNumber="537">
<subSubSection id="C373657A4553FF9FFF3CFDA4FDC54A0B" box="[151,600,511,538]" pageId="36" pageNumber="537" type="nomenclature">
<paragraph id="8BD636F14553FF9FFF3CFDA4FDC54A0B" blockId="36.[151,600,511,571]" box="[151,600,511,538]" pageId="36" pageNumber="537">
<heading id="D09E819D4553FF9FFF3CFDA4FDC54A0B" bold="true" box="[151,600,511,538]" fontSize="11" level="1" pageId="36" pageNumber="537" reason="1">
<emphasis id="B91DEAE34553FF9FFF3CFDA4FDC54A0B" bold="true" box="[151,600,511,538]" pageId="36" pageNumber="537">
24.
<taxonomicName id="4C694D724553FF9FFF68FE5BFDC54A0B" ID-CoL="42CPS" authority="Wheeler, 1919" authorityName="Wheeler" authorityYear="1919" box="[195,600,511,538]" class="Insecta" family="Formicidae" genus="Metapone" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="537" phylum="Arthropoda" rank="species" species="tillyardi">
<emphasis id="B91DEAE34553FF9FFF68FE5BFE3D4A08" bold="true" box="[195,416,511,537]" italics="true" pageId="36" pageNumber="537">Metapone tillyardi</emphasis>
Wheeler, 1919
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C373657A4553FF9FFF3CFD86FEDE4A2A" box="[151,323,546,571]" pageId="36" pageNumber="537" type="description">
<paragraph id="8BD636F14553FF9FFF3CFD86FEDE4A2A" blockId="36.[151,600,511,571]" box="[151,323,546,571]" pageId="36" pageNumber="537">
(
<figureCitation id="13522A744553FF9FFF34FD86FF624A2A" box="[159,255,546,571]" captionStart="FIGURES 131 136. M" captionStartId="37.[151,264,1596,1619]" captionTargetBox="[152,1436,303,1575]" captionTargetId="figure@37.[151,1436,301,1575]" captionTargetPageId="37" captionText="FIGURES 131 136. M. tillyardi, worker, Brooklana, NSW, HW 1.01, PMW 0.91, MSL 1.83; FIGURES 137 140. gyne, HW 1.15, MSL 2.38." httpUri="https://zenodo.org/record/271344/files/figure.png" pageId="36" pageNumber="537">Figs 131</figureCitation>
140)
</paragraph>
</subSubSection>
<subSubSection id="C373657A4553FF9FFF3CFDCFFBF14A91" box="[151,1132,618,640]" pageId="36" pageNumber="537" type="reference_group">
<paragraph id="8BD636F14553FF9FFF3CFDCFFBF14A91" blockId="36.[151,1132,618,640]" box="[151,1132,618,640]" pageId="36" pageNumber="537">
<treatmentCitationGroup id="AB7911DF4553FF9FFF3CFDCFFBF14A91" box="[151,1132,618,640]" pageId="36" pageNumber="537">
<taxonomicName id="4C694D724553FF9FFF3CFDCFFE784A91" ID-CoL="42CPS" authority="Wheeler, 1919b: 187" authorityName="Wheeler" authorityPageNumber="187" authorityYear="1919" box="[151,485,618,640]" class="Insecta" family="Formicidae" genus="Metapone" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="537" phylum="Arthropoda" rank="species" species="tillyardi">
<emphasis id="B91DEAE34553FF9FFF3CFDCFFE974A91" box="[151,266,618,640]" italics="true" pageId="36" pageNumber="537">M. tillyardi</emphasis>
<treatmentCitation id="0AC810E04553FF9FFEA4FDCEFE784A91" author="Wheeler" box="[271,485,618,640]" page="187" pageId="36" pageNumber="537" year="1919">
<bibRefCitation id="EFF84B004553FF9FFEA4FDCEFE2E4A91" author="Wheeler" box="[271,435,618,640]" pageId="36" pageNumber="537" refString="Wheeler, W. M. (1919 b) The ants of the genus Metapone Forel. Annals of the Entomological Society of America, 12, 173 - 191. http: // dx. doi. org / 10.1093 / aesa / 12.3.173" type="journal article" year="1919" yearSuffix="b">Wheeler, 1919b</bibRefCitation>
: 187
</treatmentCitation>
</taxonomicName>
,
<figureCitation id="13522A744553FF9FFE5BFDCEFDB94A91" box="[496,548,618,640]" captionStart="FIGURES 5 7" captionStartId="8.[151,264,1951,1974]" captionTargetBox="[151,1436,903,1929]" captionTargetId="figure@8.[151,1436,903,1930]" captionTargetPageId="8" captionText="FIGURES 5 7. Metapone emersoni, worker, Perinet, Madagascar, HW 0.75, PMW 0.56, MSL 1.20." httpUri="https://zenodo.org/record/271321/files/figure.png" pageId="36" pageNumber="537">Fig 6</figureCitation>
a, b—Worker;
<typeStatus id="54D288534553FF9FFD6AFDCFFD6E4A91" box="[705,755,619,640]" pageId="36" pageNumber="537">Type</typeStatus>
locality:
<taxonomicName id="4C694D724553FF9FFCF8FDCEFC3B4A91" ID-CoL="8XKZJ" box="[851,934,618,640]" class="Insecta" family="Staphylinidae" genus="Dorrigo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="537" phylum="Arthropoda" rank="genus">Dorrigo</taxonomicName>
, New South
<collectingCountry id="F37E76614553FF9FFB83FDCEFBF44A91" box="[1064,1129,618,640]" name="United Kingdom" pageId="36" pageNumber="537">Wales</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C373657A4553FF9FFF3CFD0AFC454D2F" pageId="36" pageNumber="537" type="distribution">
<paragraph id="8BD636F14553FF9FFF3CFD0AFA924CC1" blockId="36.[151,1437,686,2026]" pageId="36" pageNumber="537">
<emphasis id="B91DEAE34553FF9FFF3CFD0AFDA64AD6" bold="true" box="[151,571,686,711]" pageId="36" pageNumber="537">Distribution, material examined.</emphasis>
<collectingCountry id="F37E76614553FF9FFDFBFD0BFD6E4AD6" box="[592,755,687,711]" name="Australia" pageId="36" pageNumber="537">AUSTRALIA</collectingCountry>
: QUEENSLAND:
<emphasis id="B91DEAE34553FF9FFC4DFD0AFA954AD6" box="[998,1288,686,711]" italics="true" pageId="36" pageNumber="537">Lords Table, W slopes</emphasis>
,
<geoCoordinate id="EE5D50364553FF9FFA8EFD0BFA0A4AD6" box="[1317,1431,686,711]" direction="south" orientation="latitude" pageId="36" pageNumber="537" precision="15" value="-22.651388">22°39'5''S</geoCoordinate>
,
<geoCoordinate id="EE5D50364553FF9FFF3CFD77FE854AFA" box="[151,280,722,747]" direction="east" orientation="longitude" pageId="36" pageNumber="537" precision="15" value="148.00194">148°00'7''E</geoCoordinate>
, nest in rotting log, vine scrub,
<quantity id="4C919B144553FF9FFDD1FD77FD364AFD" box="[634,683,723,748]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="36" pageNumber="537" unit="m" value="60.0">60m</quantity>
(C. Burwell,
<date id="FFD710314553FF9FFCEEFD70FC384AFD" box="[837,933,723,748]" pageId="36" pageNumber="537">7 March</date>
, 2006, QMBA) dealate gyne,
<specimenCount id="9D6FFD784553FF9FFB5EFD77FAF94AFD" box="[1269,1380,723,748]" pageId="36" pageNumber="537" type="worker">8 workers</specimenCount>
on 3 pins;
<emphasis id="B91DEAE34553FF9FFF73FD5CFD9B4B01" box="[216,518,759,784]" italics="true" pageId="36" pageNumber="537">Stony Creek, via Samford,</emphasis>
<geoCoordinate id="EE5D50364553FF9FFDBBFD5CFDF74B01" box="[528,618,759,784]" direction="south" orientation="latitude" pageId="36" pageNumber="537" precision="925" value="-27.333334">27°20'S</geoCoordinate>
,
<geoCoordinate id="EE5D50364553FF9FFDD1FD53FD7E4B01" box="[634,739,759,784]" direction="east" orientation="longitude" pageId="36" pageNumber="537" precision="925" value="153.8">153°48'E</geoCoordinate>
,
<quantity id="4C919B144553FF9FFD58FD53FCAF4B01" box="[755,818,759,784]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="36" pageNumber="537" unit="m" value="850.0">850m</quantity>
(HJ &amp;GBM,
<date id="FFD710314553FF9FFC73FD5CFB204B01" box="[984,1213,759,784]" pageId="36" pageNumber="537" value="1995-10-22" valueMax="1995-02-02" valueMin="1995-10-22">22 Oct2 Feb 1995</date>
, open forest, flight intercept trap), Gyne.
<emphasis id="B91DEAE34553FF9FFE3FFCBBFE414B27" box="[404,476,799,822]" italics="true" pageId="36" pageNumber="537">Banyo</emphasis>
[
<geoCoordinate id="EE5D50364553FF9FFE47FCBBFDDA4B26" box="[492,583,795,823]" direction="south" orientation="latitude" pageId="36" pageNumber="537" precision="925" value="-27.366667">27°22S</geoCoordinate>
,
<geoCoordinate id="EE5D50364553FF9FFDFDFCBAFD2F4B26" box="[598,690,795,823]" direction="east" orientation="longitude" pageId="36" pageNumber="537" precision="925" value="53.066666">53°04E</geoCoordinate>
], branch of pine tree (
<date id="FFD710314553FF9FFC6EFCBAFBFE4B26" box="[965,1123,798,823]" pageId="36" pageNumber="537" value="2005-10">October 2005</date>
J. Gray acc 12394) 7 alate queens (QMBA).
<emphasis id="B91DEAE34553FF9FFEF4FCE2FDE04B4F" box="[351,637,837,862]" italics="true" pageId="36" pageNumber="537">Lamington National Park</emphasis>
: 11 alate gynes with various data, as follows:
<geoCoordinate id="EE5D50364553FF9FFBD4FCE2FB7E4B4F" box="[1151,1251,834,862]" direction="south" orientation="latitude" pageId="36" pageNumber="537" precision="55" value="-28.142">28.142°S</geoCoordinate>
,
<geoCoordinate id="EE5D50364553FF9FFB44FCE1FAF94B4C" box="[1263,1380,834,862]" direction="east" orientation="longitude" pageId="36" pageNumber="537" precision="55" value="153.133">153.133°E</geoCoordinate>
, Plot #IQ-300-D,
<quantity id="4C919B144553FF9FFE8FFCC9FEFB4B94" box="[292,358,877,901]" metricMagnitude="2" metricUnit="m" metricValue="2.48" pageId="36" pageNumber="537" unit="m" value="248.0">248m</quantity>
,
<date id="FFD710314553FF9FFEDEFCC9FE744B95" box="[373,489,877,901]" pageId="36" pageNumber="537" value="2013-01-23">1323 Jan</date>
207 (GBM, 22162, n=1);
<geoCoordinate id="EE5D50364553FF9FFCB5FCC8FC1F4B94" box="[798,898,873,901]" direction="south" orientation="latitude" pageId="36" pageNumber="537" precision="55" value="-28.151">28.151°S</geoCoordinate>
,
<geoCoordinate id="EE5D50364553FF9FFC3AFCC8FB984B95" box="[913,1029,873,901]" direction="east" orientation="longitude" pageId="36" pageNumber="537" precision="55" value="153.138">153.138°E</geoCoordinate>
, Plot #IQ-300-C,
<quantity id="4C919B144553FF9FFB7DFCC8FA844B94" box="[1238,1305,876,901]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="36" pageNumber="537" unit="m" value="260.0">260m</quantity>
,
<date id="FFD710314553FF9FFA8CFCC9FA064B95" box="[1319,1435,877,901]" pageId="36" pageNumber="537" value="2013-01-23">1323 Jan</date>
207 (GBM, 22161, n=2);
<geoCoordinate id="EE5D50364553FF9FFE1EFC37FD844BBA" box="[437,537,911,940]" direction="south" orientation="latitude" pageId="36" pageNumber="537" precision="55" value="-28.155">28.155°S</geoCoordinate>
,
<geoCoordinate id="EE5D50364553FF9FFD8EFC37FD074BBA" box="[549,666,911,940]" direction="east" orientation="longitude" pageId="36" pageNumber="537" precision="55" value="153.139">153.139°E</geoCoordinate>
, Plot #IQ-300-B,
<quantity id="4C919B144553FF9FFCC9FC37FC394BBA" box="[866,932,915,939]" metricMagnitude="2" metricUnit="m" metricValue="2.82" pageId="36" pageNumber="537" unit="m" value="282.0">282m</quantity>
,
<date id="FFD710314553FF9FFC04FC37FBC24BBA" box="[943,1119,915,939]" pageId="36" pageNumber="537" value="2007-01-13" valueMax="2007-01-23" valueMin="2007-01-13">1323 Jan 2007</date>
(GBM &amp; RM, 22183, n=1); same data (IBSCA 22139, GL &amp; NS, n=4); same data (Plot #IQ-300-C,
<quantity id="4C919B144553FF9FFC15FC1DFB9C4BC3" box="[958,1025,953,978]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="36" pageNumber="537" unit="m" value="260.0">260m</quantity>
, n=1);
<geoCoordinate id="EE5D50364553FF9FFBFBFC1EFB294BC3" box="[1104,1204,950,978]" direction="south" orientation="latitude" pageId="36" pageNumber="537" precision="55" value="-28.193">28.193°S</geoCoordinate>
,
<geoCoordinate id="EE5D50364553FF9FFB6BFC1DFAA84BC0" box="[1216,1333,950,978]" direction="east" orientation="longitude" pageId="36" pageNumber="537" precision="55" value="153.128">153.128°E</geoCoordinate>
, IBISCA 700c,
<date id="FFD710314553FF9FFF77FC45FEEA4BE9" box="[220,375,993,1017]" pageId="36" pageNumber="537">2 Nov9 Dec</date>
, 2008 (GBM, n=1);
<geoCoordinate id="EE5D50364553FF9FFDCBFC45FD594BE8" box="[608,708,989,1017]" direction="south" orientation="latitude" pageId="36" pageNumber="537" precision="55" value="-28.227">28.227°S</geoCoordinate>
,
<geoCoordinate id="EE5D50364553FF9FFD79FC44FCDB4BE9" box="[722,838,989,1017]" direction="east" orientation="longitude" pageId="36" pageNumber="537" precision="55" value="153.131">153.131°E</geoCoordinate>
, Plot #IQ-900-D,
<quantity id="4C919B144553FF9FFBBFFC45FBCA4BE8" box="[1044,1111,993,1017]" metricMagnitude="2" metricUnit="m" metricValue="9.2" pageId="36" pageNumber="537" unit="m" value="920.0">920m</quantity>
,
<date id="FFD710314553FF9FFBC8FC45FA8B4BE8" box="[1123,1302,993,1017]" pageId="36" pageNumber="537" value="2007-01-14" valueMax="2007-01-24" valueMin="2007-01-14">1424 Jan 2007</date>
(CL &amp; NS, 221s, n=1) (QMBA). NEW SOUTH
<collectingCountry id="F37E76614553FF9FFD96FBA1FD024C0C" box="[573,671,1029,1053]" name="United Kingdom" pageId="36" pageNumber="537">WALES</collectingCountry>
:
<taxonomicName id="4C694D724553FF9FFD01FBA1FCC54C0C" authority="Plateau" authorityName="Plateau" box="[682,856,1028,1053]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="36" pageNumber="537" phylum="Tracheophyta" rank="genus">
<emphasis id="B91DEAE34553FF9FFD01FBA1FCC54C0C" box="[682,856,1028,1053]" italics="true" pageId="36" pageNumber="537">Acacia Plateau</emphasis>
</taxonomicName>
(28;152) (J. Armstrong,
<date id="FFD710314553FF9FFBDEFBA1FB234C0C" box="[1141,1214,1029,1053]" pageId="36" pageNumber="537" value="48-01-03">3.1.48</date>
, ANIC) alate gyne.
<emphasis id="B91DEAE34553FF9FFF3CFB8CFE914C50" box="[151,268,1064,1089]" italics="true" pageId="36" pageNumber="537">Brooklana</emphasis>
(30;152) (W.W. Froggatt, ANIC) dealate gyne.
<taxonomicName id="4C694D724553FF9FFC8CFB8DFC1E4C51" box="[807,899,1065,1088]" class="Insecta" family="Staphylinidae" genus="Dorrigo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="537" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEAE34553FF9FFC8CFB8DFC1E4C51" box="[807,899,1065,1088]" italics="true" pageId="36" pageNumber="537">Dorrigo</emphasis>
</taxonomicName>
(30;152) 5
<typeStatus id="54D288534553FF9FFBACFB8DFBC34C50" box="[1031,1118,1065,1089]" pageId="36" pageNumber="537" type="syntype">syntype</typeStatus>
workers from the following collections: ANIC (
<specimenCount id="9D6FFD784553FF9FFED8FBE9FE604C74" box="[371,509,1100,1125]" pageId="36" pageNumber="537" type="generic">3 specimens</specimenCount>
on 2 pins), MCZC (2-1), QMBA (1).
<emphasis id="B91DEAE34553FF9FFC0DFBE8FBA34C74" box="[934,1086,1100,1125]" italics="true" pageId="36" pageNumber="537">Sunshine Bay</emphasis>
<geoCoordinate id="EE5D50364553FF9FFBEEFBE8FB354C74" box="[1093,1192,1100,1125]" direction="south" orientation="latitude" pageId="36" pageNumber="537" precision="925" value="-35.75">35°45 S</geoCoordinate>
<geoCoordinate id="EE5D50364553FF9FFB05FBE8FABC4C75" box="[1198,1313,1100,1125]" direction="east" orientation="longitude" pageId="36" pageNumber="537" precision="925" value="150.21666">150°13 E</geoCoordinate>
ex Willow (
<taxonomicName id="4C694D724553FF9FFF0BFBD4FF454C98" box="[160,216,1136,1161]" class="Ascidiacea" family="Polycitoridae" genus="Salix" kingdom="Animalia" order="Enterogona" pageId="36" pageNumber="537" phylum="Chordata" rank="genus">
<emphasis id="B91DEAE34553FF9FFF0BFBD4FF454C98" box="[160,216,1136,1161]" italics="true" pageId="36" pageNumber="537">Salix</emphasis>
</taxonomicName>
sp.) branch in caravan park J.A.L Watson &amp; RWT
<date id="FFD710314553FF9FFC86FBD4FC094C98" box="[813,916,1136,1161]" pageId="36" pageNumber="537" value="1990-05-05">5 v 1990</date>
ANIC. All Queensland labels from localities except Lords Table indicate collection in rainforest by Malaise or flight intercept trap. Note that the word “pine” on the Banyo labels is given in parentheses. It presumably does not refer to an exotic tree of genus
<taxonomicName id="4C694D724553FF9FFB62FB1DFA944CC1" box="[1225,1289,1209,1232]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="36" pageNumber="537" phylum="Tracheophyta" rank="genus">
<emphasis id="B91DEAE34553FF9FFB62FB1DFA944CC1" box="[1225,1289,1209,1232]" italics="true" pageId="36" pageNumber="537">Pinus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD636F14553FF9FFF6CFB78FC454D2F" blockId="36.[151,1437,686,2026]" pageId="36" pageNumber="537">
Apparently widespread in S.E. Queensland and northeastern NSW, at least between S latitudes 22 and 35. The
<typeStatus id="54D288534553FF9FFF3CFAA6FF4F4D0B" box="[151,210,1282,1306]" pageId="36" pageNumber="537">types</typeStatus>
were almost certainly collected in rainforest at
<taxonomicName id="4C694D724553FF9FFCA3FAA5FCFC4D0B" box="[776,865,1281,1306]" class="Insecta" family="Staphylinidae" genus="Dorrigo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="537" phylum="Arthropoda" rank="genus">Dorrigo</taxonomicName>
National Park (
<geoCoordinate id="EE5D50364553FF9FFBB6FAA6FBE54D0B" box="[1053,1144,1281,1306]" direction="south" orientation="latitude" pageId="36" pageNumber="537" precision="925" value="-30.366667">30°22'S</geoCoordinate>
,
<geoCoordinate id="EE5D50364553FF9FFB23FAA5FB6E4D0B" box="[1160,1267,1281,1306]" direction="east" orientation="longitude" pageId="36" pageNumber="537" precision="925" value="152.75">152°45'E</geoCoordinate>
). Worker/gyne identity of unassociated specimens is confirmed by the Lords Table series.
</paragraph>
</subSubSection>
<subSubSection id="C373657A4553FF9EFF6CFAEDFAB448E9" lastPageId="37" lastPageNumber="538" pageId="36" pageNumber="537" type="description">
<paragraph id="8BD636F14553FF9FFF6CFAEDFEE24EB7" blockId="36.[151,1437,686,2026]" pageId="36" pageNumber="537">
<emphasis id="B91DEAE34553FF9FFF6CFAEDFE3E4D73" bold="true" box="[199,419,1353,1378]" pageId="36" pageNumber="537">Worker diagnosis.</emphasis>
General and diagnostic features as illustrated and keyed here, and in Wheelers detailed (1919b) description and figures. Note however that the original description (
<bibRefCitation id="EFF84B004553FF9FFC58FAC9FB354D97" author="Wheeler" box="[1011,1192,1389,1414]" pageId="36" pageNumber="537" refString="Wheeler, W. M. (1919 b) The ants of the genus Metapone Forel. Annals of the Entomological Society of America, 12, 173 - 191. http: // dx. doi. org / 10.1093 / aesa / 12.3.173" type="journal article" year="1919" yearSuffix="b">Wheeler, 1919b</bibRefCitation>
) did not mention that some of the worker
<typeStatus id="54D288534553FF9FFEDCFA36FE2F4DBB" box="[375,434,1426,1450]" pageId="36" pageNumber="537">types</typeStatus>
have reduced eyes, and less massive propodeal and waist node configuration than that of the
<typeStatus id="54D288534553FF9FFF6DFA11FEB14DDF" box="[198,300,1461,1486]" pageId="36" pageNumber="537" type="lectotype">lectotype</typeStatus>
(see above). Clypeus projecting anteriorly less than in
<taxonomicName id="4C694D724553FF9FFC67FA13FBC64DDF" box="[972,1115,1461,1486]" class="Insecta" family="Formicidae" genus="Metapone" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="537" phylum="Arthropoda" rank="species" species="mjobergi">
<emphasis id="B91DEAE34553FF9FFC67FA13FBC64DDF" box="[972,1115,1461,1486]" italics="true" pageId="36" pageNumber="537">M. mjobergi</emphasis>
</taxonomicName>
; anterior border somewhat obscurely bicuspid, with a pair of blunt projections at approximately its outer quarters, the space between them shallowly excavated, concave and raised almost to their apices. Eyes and waist nodes as discussed above. Subpetiolar process differing from
<taxonomicName id="4C694D724553FF9FFD83F986FD294E2B" box="[552,692,1569,1594]" class="Insecta" family="Formicidae" genus="Metapone" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="537" phylum="Arthropoda" rank="species" species="mjobergi">
<emphasis id="B91DEAE34553FF9FFD83F986FD294E2B" box="[552,692,1569,1594]" italics="true" pageId="36" pageNumber="537">M. mjobergi</emphasis>
</taxonomicName>
as in couplet 6 of the key to Australian species above. Posterior subpetiolar face as in
<taxonomicName id="4C694D724553FF9FFE27F9E3FD8B4E4F" box="[396,534,1605,1630]" class="Insecta" family="Formicidae" genus="Metapone" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="537" phylum="Arthropoda" rank="species" species="mjobergi">
<emphasis id="B91DEAE34553FF9FFE27F9E3FD8B4E4F" box="[396,534,1605,1630]" italics="true" pageId="36" pageNumber="537">M. mjobergi</emphasis>
</taxonomicName>
. The Sunshine Bay specimens are relatively small and gracile and have reduced petioles. They were taken from a colony which might not have been large enough ergonomically to produce large, less-gracile workers.
</paragraph>
<paragraph id="8BD636F14553FF9EFF6CF916FAB448E9" blockId="36.[151,1437,686,2026]" lastBlockId="37.[151,1437,151,248]" lastPageId="37" lastPageNumber="538" pageId="36" pageNumber="537">
<emphasis id="B91DEAE34553FF9FFF6CF916FED64ED8" box="[199,331,1714,1737]" italics="true" pageId="36" pageNumber="537">Dimensions</emphasis>
: (MCZC
<typeStatus id="54D288534553FF9FFE12F916FD8A4EDB" box="[441,535,1714,1738]" pageId="36" pageNumber="537" type="syntype">Syntype</typeStatus>
, the 2 intact ANIC
<typeStatus id="54D288534553FF9FFD44F916FCC94EDB" box="[751,852,1714,1738]" pageId="36" pageNumber="537" type="syntype">syntypes</typeStatus>
, other specimens not
<typeStatus id="54D288534553FF9FFBEFF916FB344EDB" box="[1092,1193,1714,1738]" pageId="36" pageNumber="537" type="syntype">syntypes</typeStatus>
, Sunshine Bay series: TL ca: 6.3, 6.4, 6.7, 6.9, 5.6, 6.6; HL: 1.30, 1.32, 1.42, 1.47, 1.24, 1.45; HW: 1.01, 1.04, 1.11, 1.17, 0.99, 1.14; CI: 78, 79, 78, 79, 80, 79; CpL: 0.42, 0.40, -, 0.48, 0.40, 0.48; CpI: 32, 30, -, 33, -, -; MSL: 1.83, 1.86, 2.05, 2.00, 1.56, 1.85; PML: 1.02, 1.08, 1.15, 1.27, 1.00, 1.00; PMW: 0.91, 0.93, 0.98, 0.96, 0.75, 0.85; PMI: 56, 58, 56, 63, 64, 54; PDW: 0.84, 0.88, 0.94, -, 0.65, 0.72; PetL: 0.56, -, 0.61, 0.53, 0 42, 0.49; PetW: 0.83, 0.91, 0.89, 0.78, 0.57, 0.68; PetH: -, -, -, -, 0.70, 0.84 PpetL: 0.51, -, 0.55, 0.52, 0.37, 0.42; PpetW: 0.78, 0.81; 0.86, 0.77, 0.58, 0.67; PpetH: -, -; -, -; 0.59, 0.70; GW: 1.20, 1.21, 1.33, 1.45, 1.01, 1.1. Two damaged, otherwise unmeasured ANIC
<typeStatus id="54D288534553FF9FFF3CF80AFF644FD7" box="[151,249,1966,1990]" pageId="36" pageNumber="537" type="syntype">syntypes</typeStatus>
lacking waist nodes and gasters have HW 1.01 and
<quantity id="4C919B144553FF9FFCCCF80AFC4C4FD4" box="[871,977,1966,1990]" metricMagnitude="-3" metricUnit="m" metricValue="1.11" pageId="36" pageNumber="537" unit="mm" value="1.11">1.11 mm</quantity>
. Some
<typeStatus id="54D288534553FF9FFB81F80AFB1C4FD7" box="[1066,1153,1966,1990]" pageId="36" pageNumber="537" type="syntype">syntype</typeStatus>
measurements were not possible due to glue or anatomical obstruction. Note also the differences in petiole and postpetiole dimensions in the two ANIC
<typeStatus id="54D288534552FF9EFEFEFF3CFE2648A1" box="[341,443,152,176]" pageId="37" pageNumber="538" type="syntype">syntypes</typeStatus>
. The larger specimen has a relatively small petiole and postpetiole and virtually imperceptible, vestigial eyes, while the smaller specimen has large eyes, much as illustrated in
<figureCitation id="13522A744552FF9EFB7BFF18FA0A48C5" box="[1232,1431,188,213]" captionStart="FIGURES 131 136. M" captionStartId="37.[151,264,1596,1619]" captionTargetBox="[152,1436,303,1575]" captionTargetId="figure@37.[151,1436,301,1575]" captionTargetPageId="37" captionText="FIGURES 131 136. M. tillyardi, worker, Brooklana, NSW, HW 1.01, PMW 0.91, MSL 1.83; FIGURES 137 140. gyne, HW 1.15, MSL 2.38." httpUri="https://zenodo.org/record/271344/files/figure.png" pageId="37" pageNumber="538">Figs 131 and 132</figureCitation>
, and more massive propodeal and waist node structure. Possible significance of these details is unknown.
</paragraph>
</subSubSection>
<caption id="DF1666794552FF9EFF3CF998FEA04E60" httpUri="https://zenodo.org/record/271344/files/figure.png" pageId="37" pageNumber="538" targetBox="[152,1436,303,1575]" targetPageId="37">
<paragraph id="8BD636F14552FF9EFF3CF998FEA04E60" blockId="37.[151,1436,1596,1649]" pageId="37" pageNumber="538">
<emphasis id="B91DEAE34552FF9EFF3CF998FEF04E43" bold="true" box="[151,365,1596,1619]" pageId="37" pageNumber="538">FIGURES 131136.</emphasis>
<taxonomicName id="4C694D724552FF9EFED8F99AFE7B4E43" box="[371,486,1596,1619]" class="Insecta" family="Formicidae" genus="Metapone" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="538" phylum="Arthropoda" rank="species" species="tillyardi">
<emphasis id="B91DEAE34552FF9EFED8F99AFE7B4E43" box="[371,486,1596,1619]" italics="true" pageId="37" pageNumber="538">M. tillyardi</emphasis>
</taxonomicName>
, worker, Brooklana, NSW, HW 1.01, PMW 0.91, MSL 1.83;
<emphasis id="B91DEAE34552FF9EFBF3F998FAB34E43" bold="true" box="[1112,1326,1596,1619]" pageId="37" pageNumber="538">FIGURES 137140.</emphasis>
gyne, HW 1.15, MSL 2.38.
</paragraph>
</caption>
<subSubSection id="C373657A4552FF9EFF6CF93CFC954F50" pageId="37" pageNumber="538" type="diagnosis">
<paragraph id="8BD636F14552FF9EFF6CF93CFD6F4EC4" blockId="37.[151,1437,1688,2038]" pageId="37" pageNumber="538">
<emphasis id="B91DEAE34552FF9EFF6CF93CFE184EA0" bold="true" box="[199,389,1688,1713]" pageId="37" pageNumber="538">Gyne diagnosis.</emphasis>
General features as illustrated. Clypeus and subpetiolar process as in worker. Mesosoma in dorsal view more bulky than in gynes of
<taxonomicName id="4C694D724552FF9EFDC9F91AFD764EC4" box="[610,747,1724,1749]" class="Insecta" family="Formicidae" genus="Metapone" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="538" phylum="Arthropoda" rank="species" species="mjobergi">
<emphasis id="B91DEAE34552FF9EFDC9F91AFD764EC4" box="[610,747,1724,1749]" italics="true" pageId="37" pageNumber="538">M. mjobergi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD636F14552FF9EFF6CF946FC954F50" blockId="37.[151,1437,1688,2038]" pageId="37" pageNumber="538">
<emphasis id="B91DEAE34552FF9EFF6CF946FED64EE8" box="[199,331,1762,1785]" italics="true" pageId="37" pageNumber="538">Dimensions</emphasis>
: The available gynes vary little in size (HW
<quantity id="4C919B144552FF9EFCE3F945FC704EE9" box="[840,1005,1761,1785]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.19" metricValueMin="1.11" pageId="37" pageNumber="538" unit="mm" value="1.15" valueMax="1.19" valueMin="1.11">1.111.19 mm</quantity>
, N = 17). One from Banyo with mean HW has: TL: ca 8.4, HL: 1.47, HW: 1.13, CI: 77, CpL: 0.42, CpI: 29, MSL: 2.38, PetL: 0.58, PetW: 0.64, PetH: 0.72, PpetL: 0.48, PpetW: 0.66, PpetH: 0.67, GW: 1.21.
</paragraph>
</subSubSection>
<subSubSection id="C373657A4552FF9DFF6CF8E9FE424A71" lastPageId="38" lastPageNumber="539" pageId="37" pageNumber="538" type="discussion">
<paragraph id="8BD636F14552FF9DFF6CF8E9FD2549BC" blockId="37.[151,1437,1688,2038]" lastBlockId="38.[151,1437,151,608]" lastPageId="38" lastPageNumber="539" pageId="37" pageNumber="538">
<emphasis id="B91DEAE34552FF9EFF6CF8E9FED34F77" bold="true" box="[199,334,1869,1894]" pageId="37" pageNumber="538">Discussion:</emphasis>
<taxonomicName id="4C694D724552FF9EFEF5F8EAFE784F77" box="[350,485,1869,1894]" class="Insecta" family="Formicidae" genus="Metapone" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="538" phylum="Arthropoda" rank="species" species="tillyardi">
<emphasis id="B91DEAE34552FF9EFEF5F8EAFE784F77" box="[350,485,1869,1894]" italics="true" pageId="37" pageNumber="538">M. tillyardi</emphasis>
</taxonomicName>
is apparently a relatively widespread species ranging at least from southeast Queensland to the vicinity of Sunshine Bay on the south shore of Batemans Bay, NSW. Its worker caste is significantly morphologically variable, possibly evidencing incipient caste polymorphism, on the following grounds: (1) The available
<taxonomicName id="4C694D724552FF9EFE4CF81CFDDD4FC0" box="[487,576,1976,2001]" class="Insecta" family="Staphylinidae" genus="Dorrigo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="537" phylum="Arthropoda" rank="genus">Dorrigo</taxonomicName>
worker
<typeStatus id="54D288534552FF9EFD06F81DFD754FC0" box="[685,744,1977,2001]" pageId="37" pageNumber="538">types</typeStatus>
include specimens with relatively large eyes and robust propodeal, petiolar and postpetiolar structure (as illustrated in
<figureCitation id="13522A744552FF9EFCC8F879FB9F4FE7" box="[867,1026,2013,2038]" captionStart="FIGURES 131 136. M" captionStartId="37.[151,264,1596,1619]" captionTargetBox="[152,1436,303,1575]" captionTargetId="figure@37.[151,1436,301,1575]" captionTargetPageId="37" captionText="FIGURES 131 136. M. tillyardi, worker, Brooklana, NSW, HW 1.01, PMW 0.91, MSL 1.83; FIGURES 137 140. gyne, HW 1.15, MSL 2.38." httpUri="https://zenodo.org/record/271344/files/figure.png" pageId="37" pageNumber="538">Figs 131136</figureCitation>
), along with others which are more gracile, with less robust propodea and petiolar nodes, and with eyes very reduced to absent. Wheelers original description and illustrations (
<bibRefCitation id="EFF84B004551FF9DFE4EFF18FD0148C4" author="Wheeler" box="[485,668,188,213]" pageId="38" pageNumber="539" refString="Wheeler, W. M. (1919 b) The ants of the genus Metapone Forel. Annals of the Entomological Society of America, 12, 173 - 191. http: // dx. doi. org / 10.1093 / aesa / 12.3.173" type="journal article" year="1919" yearSuffix="b">Wheeler, 1919b</bibRefCitation>
) featured only the former, overlooking this variability in the typeseries. (2) Ten workers from Sunshine Bay satisfy the
<taxonomicName id="4C694D724551FF9DFD58FF45FCEC48E9" box="[755,881,223,248]" class="Insecta" family="Formicidae" genus="Metapone" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="539" phylum="Arthropoda" rank="species" species="tillyardi">
<emphasis id="B91DEAE34551FF9DFD58FF45FCEC48E9" box="[755,881,223,248]" italics="true" pageId="38" pageNumber="539">M. tillyardi</emphasis>
</taxonomicName>
diagnosis but the specimens are mostly relatively small. They have even less-robust propodeal and waist structure than the gracile, optically-reduced
<taxonomicName id="4C694D724551FF9DFAE8FEA0FA01490C" box="[1347,1436,260,285]" class="Insecta" family="Staphylinidae" genus="Dorrigo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="537" phylum="Arthropoda" rank="genus">Dorrigo</taxonomicName>
<typeStatus id="54D288534551FF9DFF3CFE8CFF604951" box="[151,253,296,320]" pageId="38" pageNumber="539" type="syntype">syntypes</typeStatus>
. They also have small, essentially vestigial eyes similar to those of the more gracile
<taxonomicName id="4C694D724551FF9DFB14FE83FA854951" box="[1215,1304,295,320]" class="Insecta" family="Staphylinidae" genus="Dorrigo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="537" phylum="Arthropoda" rank="genus">Dorrigo</taxonomicName>
<typeStatus id="54D288534551FF9DFA8BFE8CFAC24951" box="[1312,1375,296,320]" pageId="38" pageNumber="539">types</typeStatus>
. This interpretation depends in part on a reasonable assumption that the
<taxonomicName id="4C694D724551FF9DFC29FEE8FC464974" box="[898,987,332,357]" class="Insecta" family="Staphylinidae" genus="Dorrigo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="537" phylum="Arthropoda" rank="genus">Dorrigo</taxonomicName>
and Batemans Bay series are genuinely conspecific. The alternative possibility that two species are represented in Wheelers
<typeStatus id="54D288534551FF9DFBFDFED4FB1B4999" box="[1110,1158,368,392]" pageId="38" pageNumber="539">type</typeStatus>
series, with one of them present also at Sunshine Bay, seems improbable.
</paragraph>
<paragraph id="8BD636F14551FF9DFF6CFE13FEB849E4" blockId="38.[151,1437,151,608]" pageId="38" pageNumber="539">
<emphasis id="B91DEAE34551FF9DFF6CFE13FEFD49C1" bold="true" box="[199,352,439,464]" pageId="38" pageNumber="539">Illustrations.</emphasis>
The illustrated gyne (
<figureCitation id="13522A744551FF9DFDCFFE13FD5749C1" box="[612,714,439,464]" captionStart="FIGURES 131 136. M" captionStartId="37.[151,264,1596,1619]" captionTargetBox="[152,1436,303,1575]" captionTargetId="figure@37.[151,1436,301,1575]" captionTargetPageId="37" captionText="FIGURES 131 136. M. tillyardi, worker, Brooklana, NSW, HW 1.01, PMW 0.91, MSL 1.83; FIGURES 137 140. gyne, HW 1.15, MSL 2.38." httpUri="https://zenodo.org/record/271344/files/figure.png" pageId="38" pageNumber="539">Figs 136</figureCitation>
140) is the confidently identified Brooklana, NSW, specimen listed above.
</paragraph>
<paragraph id="8BD636F14551FF9DFF6CFE5BFB864A09" blockId="38.[151,1437,151,608]" box="[199,1051,511,536]" pageId="38" pageNumber="539">
<emphasis id="B91DEAE34551FF9DFF6CFE5BFEA64A09" bold="true" box="[199,315,511,536]" pageId="38" pageNumber="539">Remarks.</emphasis>
<taxonomicName id="4C694D724551FF9DFEE9FDA5FE5D4A09" box="[322,448,511,536]" class="Insecta" family="Formicidae" genus="Metapone" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="539" phylum="Arthropoda" rank="species" species="tillyardi">
<emphasis id="B91DEAE34551FF9DFEE9FDA5FE5D4A09" box="[322,448,511,536]" italics="true" pageId="38" pageNumber="539">M. tillyardi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C694D724551FF9DFE5DFDA5FD1D4A09" box="[502,640,511,536]" class="Insecta" family="Formicidae" genus="Metapone" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="539" phylum="Arthropoda" rank="species" species="mjobergi">
<emphasis id="B91DEAE34551FF9DFE5DFDA5FD1D4A09" box="[502,640,511,536]" italics="true" pageId="38" pageNumber="539">M. mjobergi</emphasis>
</taxonomicName>
are similar, possibly related species.
</paragraph>
<paragraph id="8BD636F14551FF9DFF6CFD80FE424A71" blockId="38.[151,1437,151,608]" pageId="38" pageNumber="539">
<emphasis id="B91DEAE34551FF9DFF6CFD80FDE44A2C" bold="true" box="[199,633,548,573]" pageId="38" pageNumber="539">Congeneric sympatric associations.</emphasis>
The Lamington National Park Record implies that
<taxonomicName id="4C694D724551FF9DFB5FFD81FAE64A2C" box="[1268,1403,548,573]" class="Insecta" family="Formicidae" genus="Metapone" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="539" phylum="Arthropoda" rank="species" species="tillyardi">
<emphasis id="B91DEAE34551FF9DFB5FFD81FAE64A2C" box="[1268,1403,548,573]" italics="true" pageId="38" pageNumber="539">M. tillyardi</emphasis>
</taxonomicName>
is sympatric there with
<emphasis id="B91DEAE34551FF9DFE2FFDEDFE424A71" box="[388,479,583,608]" italics="true" pageId="38" pageNumber="539">
<taxonomicName id="4C694D724551FF9DFE2FFDEDFE464A71" box="[388,475,583,608]" class="Insecta" family="Formicidae" genus="Metapone" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="539" phylum="Arthropoda" rank="species" species="leae">M. leae</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>