622 lines
70 KiB
XML
622 lines
70 KiB
XML
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<mods:title id="1659CB9323623DE52DE34C5BC9D1F837">Studies on Adriaphaenops Noesske with the description of five new species from the Dinarides (Coleoptera: Carabidae: Trechini)</mods:title>
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Genus
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locality: Čatol jama des Bjelasica-Gebirges (weitere Umgebung von Gacko) im nord-ostherzegowinischen Karstlande.
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(
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sg.
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:79
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,
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:159
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,
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:166
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,
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(sg.
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<taxonomicName id="4C694D04FFA5FF9315370CFBFE24FD46" ID-CoL="8KTZK" box="[313,469,738,760]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="genus">Adriaphaenops</taxonomicName>
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:34
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,
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:33
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,
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||
<bibRefCitation id="EFF84B76FFA5FF9316BC0D18FCAAFCA9" author="Monguzzi" box="[690,859,769,791]" pageId="2" pageNumber="503" refString="Monguzzi, R. (1993) Dalmataphaenops (n. gen.) chiarae (n. sp.) nuovo eccezionale Trechino troglobio della regione Dinarica e considerazioni sul genere Aphaenopsis G. Muller, 1913. Natura Bresciana, Annali del Museo civico di Scienze naturali di Brescia, 28 (1992), 231 - 242." type="journal article" year="1993">Monguzzi, 1993</bibRefCitation>
|
||
:238
|
||
</treatmentCitation>
|
||
,
|
||
<treatmentCitation id="0AC81096FFA5FF93179C0D18FB2DFCA9" author="Casale" box="[914,1244,769,791]" page="421" pageId="2" pageNumber="503" year="1994">
|
||
<bibRefCitation id="EFF84B76FFA5FF93179C0D18FB41FCA9" author="Casale" box="[914,1200,769,791]" pageId="2" pageNumber="503" refString="Casale, A. & Gueorguiev, V. B. (1994) Albanotrechus beroni, nuovo genere, nuova specie di Trechini cavernicoli di Albania (Coleoptera, Carabidae). Bolletino del Museo regionale di Scienze naturali di Torino, 12 (2), 413 - 423." type="journal article" year="1994">Casale & Guéorguiev, 1994</bibRefCitation>
|
||
:421
|
||
</treatmentCitation>
|
||
.
|
||
<treatmentCitation id="0AC81096FFA5FF9310EB0D18FEEDFC88" author="Drovenik" page="43" pageId="2" pageNumber="503" year="1994">
|
||
<bibRefCitation id="EFF84B76FFA5FF9310EB0D18FF0FFC88" author="Drovenik" pageId="2" pageNumber="503" refString="Drovenik, B. & Peks, H. (1994) Catalogus faunae Carabiden den Balkanlander, Coleoptera Carabidae. Schwanfelder Coleopterologische Mitteilungen, Schwanfeld, 103 pp." type="book" year="1994">Drovenik & Peks, 1994</bibRefCitation>
|
||
:43
|
||
</treatmentCitation>
|
||
,
|
||
<treatmentCitation id="0AC81096FFA5FF9315280D38FDD9FC88" author="Moravec" box="[294,552,800,822]" page="289" pageId="2" pageNumber="503" year="2003">
|
||
<bibRefCitation id="EFF84B76FFA5FF9315280D38FE0DFC88" author="Moravec" box="[294,508,800,822]" pageId="2" pageNumber="503" refString="Moravec, P., Ueno, S. - I. & Belousov, I. A. (2003) Trechini, In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palearctic Coleoptera, Uol. 1. Apollo Books, Stenstrup, pp. 288 - 346." type="journal article" year="2003">
|
||
Moravec
|
||
<emphasis id="B91DEA95FFA5FF9315890D38FE4BFC88" box="[391,442,800,822]" italics="true" pageId="2" pageNumber="503">et al.</emphasis>
|
||
, 2003
|
||
</bibRefCitation>
|
||
:289
|
||
</treatmentCitation>
|
||
, Quéinnec, 2008:157, Quéinnec & Pavićević, 2008:144.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BD63687FFA5FF9314990D27FBBFFCEA" blockId="2.[151,1436,616,853]" box="[151,1102,830,853]" pageId="2" pageNumber="503">
|
||
<bibRefCitation id="EFF84B76FFA5FF9314990D27FDE9FCEA" author="Queinnec" box="[151,536,830,852]" pageId="2" pageNumber="503" refString="Queinnec, E., Pavicevic, D. & Ollivier, E. (2008) Preliminary description of a new Adriaphaenops from Lebrsnik Nountain (Hercegovina, BiH) (Coleoptera, Carabidae, Trechinae). In: Pavicevic, D. & Perreau, M. (Eds.), Advances in the studies of the fauna of the Balkan Peninsula. Papers dedicated to the memory of Guido Nonveiller. Institute for Nature Conservation of Serbia, Belgrade, Monographs, 22, pp. 154 - 156." type="journal article" year="2008">Quéinnec, Pavićević & Ollivier, 2008</bibRefCitation>
|
||
:154, Lakota
|
||
<emphasis id="B91DEA95FFA5FF9316950D59FD3FFCEA" box="[667,718,830,853]" italics="true" pageId="2" pageNumber="503">et al.</emphasis>
|
||
, 2010:100, Lohaj & Lakota, 2010:78.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C373650CFFA5FF9214990D98FD7BFD10" lastPageId="3" lastPageNumber="504" pageId="2" pageNumber="503" type="diagnosis">
|
||
<paragraph id="8BD63687FFA5FF9314990D98FE9AFBB8" blockId="2.[151,1437,897,2038]" pageId="2" pageNumber="503">
|
||
<emphasis id="B91DEA95FFA5FF9314990D98FEE0FC24" bold="true" box="[151,273,897,922]" pageId="2" pageNumber="503">Diagnosis.</emphasis>
|
||
Small to medium-sized trechine genus, L:
|
||
<quantity id="4C919B62FFA5FF93172B0D98FC38FC27" box="[805,969,897,922]" metricMagnitude="-3" metricUnit="m" metricValue="4.625" metricValueMax="5.75" metricValueMin="3.5" pageId="2" pageNumber="503" unit="mm" value="4.625" valueMax="5.75" valueMin="3.5">3.5–5.75 mm</quantity>
|
||
, TL:
|
||
<quantity id="4C919B62FFA5FF9310080D98FB5BFC27" box="[1030,1194,897,922]" metricMagnitude="-3" metricUnit="m" metricValue="4.375" metricValueMax="5.35" metricValueMin="3.4" pageId="2" pageNumber="503" unit="mm" value="4.375" valueMax="5.35" valueMin="3.4">3.4–5.35 mm</quantity>
|
||
, with aphaenopsoid characters: elongate head and pronotum, elytra ovoid with base strongly narrowed, distinctly wider than head and pronotum, body apterous, depigmented, strongly flattened, more or less pubescent. Antennae and legs elongate, densely pubescent.
|
||
</paragraph>
|
||
<paragraph id="8BD63687FFA5FF9314C90A08FE86FAD0" blockId="2.[151,1437,897,2038]" pageId="2" pageNumber="503">
|
||
Head varying in shape, from almost rounded (
|
||
<taxonomicName id="4C694D04FFA5FF9316DB0A0BFCDFFB97" box="[725,814,1042,1065]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="pretneri">
|
||
<emphasis id="B91DEA95FFA5FF9316DB0A0BFCDFFB97" box="[725,814,1042,1065]" italics="true" pageId="2" pageNumber="503">pretneri</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C694D04FFA5FF9317330A0BFC6DFB97" box="[829,924,1042,1065]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="perreaui">
|
||
<emphasis id="B91DEA95FFA5FF9317330A0BFC6DFB97" box="[829,924,1042,1065]" italics="true" pageId="2" pageNumber="503">perreaui</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C694D04FFA5FF9317A20A08FBE8FB94" box="[940,1049,1041,1066]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="jasminkoi" status="sp. nov.">
|
||
<emphasis id="B91DEA95FFA5FF9317A20A08FBE8FB94" box="[940,1049,1041,1066]" italics="true" pageId="2" pageNumber="503">jasminkoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22E57EEFFA5FF93102F0A0BFB8BFB94" box="[1057,1146,1042,1066]" pageId="2" pageNumber="503" rank="species">sp. nov.</taxonomicNameLabel>
|
||
) to more or less elongate (other species), as long as wide (
|
||
<taxonomicName id="4C694D04FFA5FF9316050A2FFD81FBF3" box="[523,624,1078,1101]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="zupcense">
|
||
<emphasis id="B91DEA95FFA5FF9316050A2FFD81FBF3" box="[523,624,1078,1101]" italics="true" pageId="2" pageNumber="503">zupcense</emphasis>
|
||
</taxonomicName>
|
||
) or distinctly longer than wide (other species), as wide as pronotum, or slightly narrower. Eyes totally absent. Frontal furrows moderately impressed, incomplete, usually shorter than half of head length, or reaching middle of head (
|
||
<taxonomicName id="4C694D04FFA5FF9316810A64FD1FFB28" box="[655,750,1149,1174]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="mlejneki" status="sp. nov.">
|
||
<emphasis id="B91DEA95FFA5FF9316810A64FD1FFB28" box="[655,750,1149,1174]" italics="true" pageId="2" pageNumber="503">mlejneki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22E57EEFFA5FF9316F80A67FCA3FB28" box="[758,850,1150,1174]" pageId="2" pageNumber="503" rank="species">sp. nov.</taxonomicNameLabel>
|
||
). Usually two pairs of supraorbital setae present, in
|
||
<taxonomicName id="4C694D04FFA5FF9314990AB8FEF5FB04" box="[151,260,1185,1210]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="albanicus" status="sp. nov.">
|
||
<emphasis id="B91DEA95FFA5FF9314990AB8FEF5FB04" box="[151,260,1185,1210]" italics="true" pageId="2" pageNumber="503">albanicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22E57EEFFA5FF9315020ABBFE95FB07" box="[268,356,1186,1209]" pageId="2" pageNumber="503" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C694D04FFA5FF9315920AB8FE0AFB04" box="[412,507,1185,1210]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="mlejneki" status="sp. nov.">
|
||
<emphasis id="B91DEA95FFA5FF9315920AB8FE0AFB04" box="[412,507,1185,1210]" italics="true" pageId="2" pageNumber="503">mlejneki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22E57EEFFA5FF93160A0ABBFDADFB07" box="[516,604,1186,1209]" pageId="2" pageNumber="503" rank="species">sp. nov.</taxonomicNameLabel>
|
||
anterior pair is absent, in
|
||
<taxonomicName id="4C694D04FFA5FF9317890AB8FBFAFB04" box="[903,1035,1185,1210]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="staudacheri">
|
||
<emphasis id="B91DEA95FFA5FF9317890AB8FBFAFB04" box="[903,1035,1185,1210]" italics="true" pageId="2" pageNumber="503">staudacheri</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C694D04FFA5FF9310150ABBFB8BFB07" box="[1051,1146,1186,1209]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="perreaui">
|
||
<emphasis id="B91DEA95FFA5FF9310150ABBFB8BFB07" box="[1051,1146,1186,1209]" italics="true" pageId="2" pageNumber="503">perreaui</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C694D04FFA5FF9310BD0ABBFACDFB07" box="[1203,1340,1186,1209]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="njegosiensis" status="sp. nov.">
|
||
<emphasis id="B91DEA95FFA5FF9310BD0ABBFACDFB07" box="[1203,1340,1186,1209]" italics="true" pageId="2" pageNumber="503">njegosiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22E57EEFFA5FF93114A0ABBFA6DFB07" box="[1348,1436,1186,1209]" pageId="2" pageNumber="503" rank="species">sp. nov.</taxonomicNameLabel>
|
||
both pairs are missing or reduced (indistinguishable from pubescence). Mandibles very long and slender, at apex pointed, right mandible without tooth. Clypeus with four (
|
||
<taxonomicName id="4C694D04FFA5FF9317330AF0FBEEFABC" box="[829,1055,1257,1282]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="antroherponomimus">
|
||
<emphasis id="B91DEA95FFA5FF9317330AF0FBEEFABC" box="[829,1055,1257,1282]" italics="true" pageId="2" pageNumber="503">antroherponomimus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C694D04FFA5FF93103E0AF3FB78FABF" box="[1072,1161,1258,1281]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="pretneri">
|
||
<emphasis id="B91DEA95FFA5FF93103E0AF3FB78FABF" box="[1072,1161,1258,1281]" italics="true" pageId="2" pageNumber="503">pretneri</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C694D04FFA5FF9310950AF3FB27FABF" box="[1179,1238,1258,1281]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="stirni">
|
||
<emphasis id="B91DEA95FFA5FF9310950AF3FB27FABF" box="[1179,1238,1258,1281]" italics="true" pageId="2" pageNumber="503">stirni</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C694D04FFA5FF93111D0AF0FA83FABC" box="[1299,1394,1257,1282]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="mlejneki" status="sp. nov.">
|
||
<emphasis id="B91DEA95FFA5FF93111D0AF0FA83FABC" box="[1299,1394,1257,1282]" italics="true" pageId="2" pageNumber="503">mlejneki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22E57EEFFA5FF9311720AF3FF39FA98" pageId="2" pageNumber="503" rank="species">sp. nov.</taxonomicNameLabel>
|
||
) or three (other species) pairs of setae, labrum with three pairs. Mentum imperfectly fused with submentum. Antennae long and slender, covered with dense pubescence, scape as long as pedicel, antennomeres
|
||
<date id="FFD71047FFA5FF9310F30B2BFA6DFAF4" box="[1277,1436,1329,1354]" pageId="2" pageNumber="503" value="2011-07-03" valueMax="2011-08-03" valueMin="2011-07-03">3–7 and 8 –11</date>
|
||
nearly equally long.
|
||
</paragraph>
|
||
<paragraph id="8BD63687FFA5FF9314C90B63FE83FA40" blockId="2.[151,1437,897,2038]" pageId="2" pageNumber="503">
|
||
Pronotum elongate, distinctly longer than wide, variously shaped. Basal part as wide as anterior (
|
||
<taxonomicName id="4C694D04FFA5FF9311140B63FA82FA2F" box="[1306,1395,1402,1425]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="pretneri">
|
||
<emphasis id="B91DEA95FFA5FF9311140B63FA82FA2F" box="[1306,1395,1402,1425]" italics="true" pageId="2" pageNumber="503">pretneri</emphasis>
|
||
</taxonomicName>
|
||
) or narrower (other species), anterior angles distinct, at apex pointed, or more or less rounded, posterior angles acute, sharp (
|
||
<taxonomicName id="4C694D04FFA5FF9314E80BDBFEBAFA67" box="[230,331,1474,1497]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="zupcense">
|
||
<emphasis id="B91DEA95FFA5FF9314E80BDBFEBAFA67" box="[230,331,1474,1497]" italics="true" pageId="2" pageNumber="503">zupcense</emphasis>
|
||
</taxonomicName>
|
||
), or obtuse, more or less rounded (other species). One pair of antero-lateral setae present; basolateral setae absent.
|
||
</paragraph>
|
||
<paragraph id="8BD63687FFA5FF9314C90810FA62F96B" blockId="2.[151,1437,897,2038]" pageId="2" pageNumber="503">
|
||
Elytra subovate elongate, with maximum width at middle or apical third, distinctly wider than head and pronotum. Basal part strongly narrowed, pedunculate, shoulders totally effaced. Scutellum small, single pair of basal scutellar setiferous pores present. Elytral striae absent. Pubescence of elytra sparse (
|
||
<taxonomicName id="4C694D04FFA5FF9310970848FB2EF9D4" box="[1177,1247,1617,1642]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="kevser">
|
||
<emphasis id="B91DEA95FFA5FF9310970848FB2EF9D4" box="[1177,1247,1617,1642]" italics="true" pageId="2" pageNumber="503">kevser</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C694D04FFA5FF9310E1084BFA83F9D7" box="[1263,1394,1618,1641]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="rumijaensis" status="sp. nov.">
|
||
<emphasis id="B91DEA95FFA5FF9310E1084BFA83F9D7" box="[1263,1394,1618,1641]" italics="true" pageId="2" pageNumber="503">rumijaensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91DEA95FFA5FF9311740848FF39F930" bold="true" pageId="2" pageNumber="503">
|
||
<taxonomicNameLabel id="A22E57EEFFA5FF9311740848FF39F930" pageId="2" pageNumber="503" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
) or more or less dense (other species), site of stria 3 with four (three discal and one preapical) (
|
||
<taxonomicName id="4C694D04FFA5FF931140086CFA65F930" box="[1358,1428,1653,1678]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="kevser">
|
||
<emphasis id="B91DEA95FFA5FF931140086CFA65F930" box="[1358,1428,1653,1678]" italics="true" pageId="2" pageNumber="503">kevser</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C694D04FFA5FF9314990883FEEBF90F" box="[151,282,1690,1713]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="rumijaensis" status="sp. nov.">
|
||
<emphasis id="B91DEA95FFA5FF9314990883FEEBF90F" box="[151,282,1690,1713]" italics="true" pageId="2" pageNumber="503">rumijaensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91DEA95FFA5FF93152E0880FE88F90C" bold="true" box="[288,377,1689,1714]" pageId="2" pageNumber="503">
|
||
<taxonomicNameLabel id="A22E57EEFFA5FF93152E0880FE88F90C" box="[288,377,1689,1714]" pageId="2" pageNumber="503" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
) or three (two discal and one preapical) dorsal pairs of setae (other species). Humeral group of umbilicate pores not aggregated, first pore of humeral group isolated and situated before level of first discal setae.
|
||
</paragraph>
|
||
<paragraph id="8BD63687FFA5FF9314C908FBFC14F8A0" blockId="2.[151,1437,897,2038]" pageId="2" pageNumber="503">Legs long and slender, pro- and mesotarsi short, claws very long and acute, without denticulation, densely pubescent. First two tarsomeres of male protarsi dilated at internal margins.</paragraph>
|
||
<paragraph id="8BD63687FFA5FF9314C90930FA98F837" blockId="2.[151,1437,897,2038]" pageId="2" pageNumber="503">
|
||
Aedeagus (
|
||
<figureCitation id="13522A02FFA5FF93154A0930FE23F8FC" box="[324,466,1833,1858]" captionStart="FIGURES 23 – 30" captionStartId="11.[151,264,1951,1974]" captionTargetBox="[160,1426,209,1922]" captionTargetId="figure@11.[151,1436,193,1930]" captionTargetPageId="11" captionText="FIGURES 23 – 30. Adriaphaenops spp., aedeagus, left lateral aspect, 23. A. antroherponomimus, 24. A. stirni, 25. A. zupcense zupcense, 26. A. zupcense tartariensis, 27. A. kevser, 28. A. perreaui, 29. A. mlejneki sp. nov., 30. A. njegosiensis sp. nov." httpUri="https://zenodo.org/record/272640/files/figure.png" pageId="2" pageNumber="503">Figs. 23–30</figureCitation>
|
||
) from short and wide (
|
||
<emphasis id="B91DEA95FFA5FF9316DF0933FBD1F8FC" box="[721,1056,1833,1858]" italics="true" pageId="2" pageNumber="503">
|
||
<taxonomicName id="4C694D04FFA5FF9316DF0933FCC2F8FF" box="[721,819,1834,1857]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="perreaui">perreaui</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C694D04FFA5FF9317300930FBD1F8FC" box="[830,1056,1833,1858]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="antroherponomimus">antroherponomimus</taxonomicName>
|
||
</emphasis>
|
||
) to elongate and slender in lateral aspect, apex variously shaped. Endophallus generally without distinct sclerotised copulatory piece or this piece is only very slightly defined. Parameres from short and wide to long, each with two long and thick setae at apex.
|
||
</paragraph>
|
||
<paragraph id="8BD63687FFA5FF9214C9098CFB68FEDB" blockId="2.[151,1437,897,2038]" lastBlockId="3.[151,1437,151,1406]" lastPageId="3" lastPageNumber="504" pageId="2" pageNumber="503">
|
||
Female genitalia (
|
||
<figureCitation id="13522A02FFA5FF93159F098CFDD2F810" box="[401,547,1941,1966]" captionStart="FIGURES 31 – 36" captionStartId="12.[151,264,1943,1966]" captionTargetBox="[189,1396,207,1898]" captionTargetId="figure@12.[169,1410,193,1907]" captionTargetPageId="12" captionText="FIGURES 31 – 36. Adriaphaenops spp., female genitalia, gonocoxite 1 and 2 (basal and apical segments of gonostylus), 31. A. zupcense zupcense, 32. A. kevser, 33. A. albanicus sp. nov., 34. A. jasminkoi sp. nov., 35. A. rumijaensis sp. nov., 36. A. mlejneki sp. nov.." httpUri="https://zenodo.org/record/200277/files/figure.png" pageId="2" pageNumber="503">Figs. 31–36</figureCitation>
|
||
): examination was possible only in six species:
|
||
<emphasis id="B91DEA95FFA5FF931031098FFA82F810" box="[1087,1395,1941,1966]" italics="true" pageId="2" pageNumber="503">
|
||
<taxonomicName id="4C694D04FFA5FF931031098FFB56F813" box="[1087,1191,1942,1965]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="zupcense">zupcense</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C694D04FFA5FF9310BD098CFB0BF810" box="[1203,1274,1941,1966]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="kevser">kevser</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C694D04FFA5FF931108098CFA82F810" box="[1286,1395,1941,1966]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="albanicus" status="sp. nov.">albanicus</taxonomicName>
|
||
</emphasis>
|
||
<emphasis id="B91DEA95FFA5FF931174098CFF39F86C" bold="true" pageId="2" pageNumber="503">
|
||
<taxonomicNameLabel id="A22E57EEFFA5FF931174098CFF39F86C" pageId="2" pageNumber="503" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C694D04FFA5FF9314D609A0FEB4F86C" box="[216,325,1977,2002]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="jasminkoi" status="sp. nov.">
|
||
<emphasis id="B91DEA95FFA5FF9314D609A0FEB4F86C" box="[216,325,1977,2002]" italics="true" pageId="2" pageNumber="503">jasminkoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91DEA95FFA5FF93154109A0FE5AF86C" bold="true" box="[335,427,1977,2002]" pageId="2" pageNumber="503">
|
||
<taxonomicNameLabel id="A22E57EEFFA5FF93154109A0FE5AF86C" box="[335,427,1977,2002]" pageId="2" pageNumber="503" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C694D04FFA5FF9315B509A3FDCFF86F" box="[443,574,1978,2001]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="rumijaensis" status="sp. nov.">
|
||
<emphasis id="B91DEA95FFA5FF9315B509A3FDCFF86F" box="[443,574,1978,2001]" italics="true" pageId="2" pageNumber="503">rumijaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22E57EEFFA5FF93164909A3FD50F86F" box="[583,673,1978,2001]" pageId="2" pageNumber="503" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C694D04FFA5FF9316D209A0FCCAF86C" box="[732,827,1977,2002]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="mlejneki" status="sp. nov.">
|
||
<emphasis id="B91DEA95FFA5FF9316D209A0FCCAF86C" box="[732,827,1977,2002]" italics="true" pageId="2" pageNumber="503">mlejneki</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91DEA95FFA5FF93174A09A0FC51F86C" bold="true" box="[836,928,1977,2002]" pageId="2" pageNumber="503">
|
||
<taxonomicNameLabel id="A22E57EEFFA5FF93174A09A0FC51F86C" box="[836,928,1977,2002]" pageId="2" pageNumber="503" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
. Gonocoxite 1 with one to three short setae close to the distal margin (
|
||
<emphasis id="B91DEA95FFA5FF9315CB09C4FD77F848" box="[453,646,2013,2038]" italics="true" pageId="2" pageNumber="503">
|
||
<taxonomicName id="4C694D04FFA5FF9315CB09C4FDFDF848" box="[453,524,2013,2038]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="kevser">kevser</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C694D04FFA5FF93161709C4FD77F848" box="[537,646,2013,2038]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="jasminkoi" status="sp. nov.">jasminkoi</taxonomicName>
|
||
</emphasis>
|
||
<emphasis id="B91DEA95FFA5FF93168309C4FD19F848" bold="true" box="[653,744,2013,2038]" pageId="2" pageNumber="503">
|
||
<taxonomicNameLabel id="A22E57EEFFA5FF93168309C4FD19F848" box="[653,744,2013,2038]" pageId="2" pageNumber="503" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C694D04FFA5FF9316F909C7FC8BF84B" box="[759,890,2014,2037]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="rumijaensis" status="sp. nov.">
|
||
<emphasis id="B91DEA95FFA5FF9316F909C7FC8BF84B" box="[759,890,2014,2037]" italics="true" pageId="2" pageNumber="503">rumijaensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91DEA95FFA5FF93178C09C4FC2DF848" bold="true" box="[898,988,2013,2038]" pageId="2" pageNumber="503">
|
||
<taxonomicNameLabel id="A22E57EEFFA5FF93178C09C4FC2DF848" box="[898,988,2013,2038]" pageId="2" pageNumber="503" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
), or without them (
|
||
<emphasis id="B91DEA95FFA5FF9310B509C7FA6DF848" box="[1211,1436,2013,2038]" italics="true" pageId="2" pageNumber="503">
|
||
<taxonomicName id="4C694D04FFA5FF9310B509C7FAD2F84B" box="[1211,1315,2014,2037]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="zupcense">zupcense</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C694D04FFA5FF93112109C4FA6DF848" box="[1327,1436,2013,2038]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="503" phylum="Arthropoda" rank="species" species="albanicus" status="sp. nov.">albanicus</taxonomicName>
|
||
</emphasis>
|
||
<emphasis id="B91DEA95FFA4FF9214990E8EFF03FF0E" bold="true" box="[151,242,151,176]" pageId="3" pageNumber="504">
|
||
<taxonomicNameLabel id="A22E57EEFFA4FF9214990E8EFF03FF0E" box="[151,242,151,176]" pageId="3" pageNumber="504" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C694D04FFA4FF92150F0E8EFE91FF0E" box="[257,352,151,176]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="mlejneki" status="sp. nov.">
|
||
<emphasis id="B91DEA95FFA4FF92150F0E8EFE91FF0E" box="[257,352,151,176]" italics="true" pageId="3" pageNumber="504">mlejneki</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91DEA95FFA4FF9215660E8EFE32FF0E" bold="true" box="[360,451,151,176]" pageId="3" pageNumber="504">
|
||
<taxonomicNameLabel id="A22E57EEFFA4FF9215660E8EFE32FF0E" box="[360,451,151,176]" pageId="3" pageNumber="504" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
), gonocoxite 2 short, at apex rounded (
|
||
<taxonomicName id="4C694D04FFA4FF9217880E8EFC3CFF0E" box="[902,973,151,176]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="kevser">
|
||
<emphasis id="B91DEA95FFA4FF9217880E8EFC3CFF0E" box="[902,973,151,176]" italics="true" pageId="3" pageNumber="504">kevser</emphasis>
|
||
</taxonomicName>
|
||
) or more or less pointed (other species), with two–ten very short setae (
|
||
<emphasis id="B91DEA95FFA4FF9215F50EA5FCBEFF6A" box="[507,847,188,213]" italics="true" pageId="3" pageNumber="504">
|
||
<taxonomicName id="4C694D04FFA4FF9215F50EA5FD9AFF6B" box="[507,619,188,213]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="jasminkoi">jasminkoi</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C694D04FFA4FF9216760EA5FD4EFF6B" box="[632,703,188,213]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="kevser">kevser</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C694D04FFA4FF9216C20EA4FCBEFF6A" box="[716,847,189,212]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="rumijaensis" status="sp. nov.">rumijaensis</taxonomicName>
|
||
</emphasis>
|
||
<emphasis id="B91DEA95FFA4FF9217560EA5FC45FF6B" bold="true" box="[856,948,188,213]" pageId="3" pageNumber="504">
|
||
<taxonomicNameLabel id="A22E57EEFFA4FF9217560EA5FC45FF6B" box="[856,948,188,213]" pageId="3" pageNumber="504" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
) or without them (
|
||
<taxonomicName id="4C694D04FFA4FF9210810EA4FB05FF6A" box="[1167,1268,189,212]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="zupcense">
|
||
<emphasis id="B91DEA95FFA4FF9210810EA4FB05FF6A" box="[1167,1268,189,212]" italics="true" pageId="3" pageNumber="504">zupcense</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C694D04FFA4FF92110A0EA5FA80FF6B" box="[1284,1393,188,213]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="albanicus" status="sp. nov.">
|
||
<emphasis id="B91DEA95FFA4FF92110A0EA5FA80FF6B" box="[1284,1393,188,213]" italics="true" pageId="3" pageNumber="504">albanicus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91DEA95FFA4FF9211740EA5FF39FF46" bold="true" pageId="3" pageNumber="504">
|
||
<taxonomicNameLabel id="A22E57EEFFA4FF9211740EA5FF39FF46" pageId="3" pageNumber="504" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C694D04FFA4FF9214D90EC6FEC7FF46" box="[215,310,223,248]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="mlejneki" status="sp. nov.">
|
||
<emphasis id="B91DEA95FFA4FF9214D90EC6FEC7FF46" box="[215,310,223,248]" italics="true" pageId="3" pageNumber="504">mlejneki</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91DEA95FFA4FF9215300EC6FE68FF46" bold="true" box="[318,409,223,248]" pageId="3" pageNumber="504">
|
||
<taxonomicNameLabel id="A22E57EEFFA4FF9215300EC6FE68FF46" box="[318,409,223,248]" pageId="3" pageNumber="504" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
); sternal sensorial fovea small, with one long seta (
|
||
<emphasis id="B91DEA95FFA4FF9217E90EF8FAC3FF46" box="[999,1330,223,248]" italics="true" pageId="3" pageNumber="504">
|
||
<taxonomicName id="4C694D04FFA4FF9217E90EF8FBBEFF46" box="[999,1103,225,248]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="zupcense">zupcense</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C694D04FFA4FF9210550EC6FB53FF46" box="[1115,1186,223,248]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="kevser">kevser</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C694D04FFA4FF9210A10EF8FAC3FF46" box="[1199,1330,225,248]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="rumijaensis" status="sp. nov.">rumijaensis</taxonomicName>
|
||
</emphasis>
|
||
<emphasis id="B91DEA95FFA4FF9211340EC6FA64FF46" bold="true" box="[1338,1429,223,248]" pageId="3" pageNumber="504">
|
||
<taxonomicNameLabel id="A22E57EEFFA4FF9211340EC6FA64FF46" box="[1338,1429,223,248]" pageId="3" pageNumber="504" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C694D04FFA4FF9214990F1DFF07FEA3" box="[151,246,260,285]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="mlejneki" status="sp. nov.">
|
||
<emphasis id="B91DEA95FFA4FF9214990F1DFF07FEA3" box="[151,246,260,285]" italics="true" pageId="3" pageNumber="504">mlejneki</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91DEA95FFA4FF92150C0F1DFE90FEA3" bold="true" box="[258,353,260,285]" pageId="3" pageNumber="504">
|
||
<taxonomicNameLabel id="A22E57EEFFA4FF92150C0F1DFE90FEA3" box="[258,353,260,285]" pageId="3" pageNumber="504" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
) or this seta is missing (
|
||
<taxonomicName id="4C694D04FFA4FF92169D0F1DFCF1FEA3" box="[659,768,260,285]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="albanicus" status="sp. nov.">
|
||
<emphasis id="B91DEA95FFA4FF92169D0F1DFCF1FEA3" box="[659,768,260,285]" italics="true" pageId="3" pageNumber="504">albanicus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91DEA95FFA4FF9217020F1DFC9AFEA3" bold="true" box="[780,875,260,285]" pageId="3" pageNumber="504">
|
||
<taxonomicNameLabel id="A22E57EEFFA4FF9217020F1DFC9AFEA3" box="[780,875,260,285]" pageId="3" pageNumber="504" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C694D04FFA4FF9217700F1DFC1AFEA3" box="[894,1003,260,285]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="jasminkoi" status="sp. nov.">
|
||
<emphasis id="B91DEA95FFA4FF9217700F1DFC1AFEA3" box="[894,1003,260,285]" italics="true" pageId="3" pageNumber="504">jasminkoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91DEA95FFA4FF9217F90F1DFBA7FEA3" bold="true" box="[1015,1110,260,285]" pageId="3" pageNumber="504">
|
||
<taxonomicNameLabel id="A22E57EEFFA4FF9217F90F1DFBA7FEA3" box="[1015,1110,260,285]" pageId="3" pageNumber="504" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
). We suppose that in older specimen examined (old dry preparation, specimen from long exposed traps) some especially short setae could be broken. This opinion should be confirmed by the examination of additional fresh specimen.
|
||
</paragraph>
|
||
<paragraph id="8BD63687FFA4FF9214C90F76FD7BFD10" blockId="3.[151,1437,151,1406]" pageId="3" pageNumber="504">
|
||
<emphasis id="B91DEA95FFA4FF9214C90F76FDF7FE36" bold="true" box="[199,518,367,392]" pageId="3" pageNumber="504">Geographical distribution.</emphasis>
|
||
So far known from underground habitats (caves, pits) in Bosnia & Hercegovina,
|
||
<materialsCitation id="3B013CDAFFA4FF9214990F8CFD7BFD10" ID-GBIF-Occurrence="1416605845" country="Albania" elevation="220" location="Lebrsnik Mts." municipality="Durmitor Mts." pageId="3" pageNumber="504" specimenCount="1">
|
||
Montenegro and northern
|
||
<collectingCountry id="F37E7617FFA4FF9215CA0F8DFDECFE13" box="[452,541,404,429]" name="Albania" pageId="3" pageNumber="504">Albania</collectingCountry>
|
||
(see Distribution map,
|
||
<figureCitation id="13522A02FFA4FF92173E0F8DFC7EFE12" box="[816,911,404,429]" captionStart="FIGURE 37" captionStartId="13.[151,250,1179,1201]" captionTargetBox="[154,1432,197,1153]" captionTargetId="figure@13.[150,1436,193,1157]" captionTargetPageId="13" captionText="FIGURE 37. Distribution map of Adriaphaenops species: 1. A. antroherponomimus, 2. A. pretneri, 3. A. staudacheri, 4. A. stirni, 5. A. zupcense, 6. A. kevser, 7. A. perreaui, 8. A. albanicus sp. nov., 9. A. jasminkoi sp. nov., 10. A. rumijaensis sp. nov., 11. A. mlejneki sp. nov., 12. A. njegosiensis sp. nov.." httpUri="https://zenodo.org/record/200278/files/figure.png" pageId="3" pageNumber="504">Fig. 37</figureCitation>
|
||
). Eight species are only known from a single limestone karst caves, four species (
|
||
<emphasis id="B91DEA95FFA4FF92163F0FA0FC9DFE6E" box="[561,876,439,464]" italics="true" pageId="3" pageNumber="504">
|
||
<taxonomicName id="4C694D04FFA4FF92163F0FA0FD7CFE6E" box="[561,653,441,464]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="pretneri">pretneri</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C694D04FFA4FF9216970FA0FCF0FE6E" box="[665,769,441,464]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="zupcense">zupcense</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C694D04FFA4FF9217030FAEFC9DFE6E" box="[781,876,439,464]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="mlejneki" status="sp. nov.">mlejneki</taxonomicName>
|
||
</emphasis>
|
||
<emphasis id="B91DEA95FFA4FF92177A0FAEFC3EFE6E" bold="true" box="[884,975,439,464]" pageId="3" pageNumber="504">
|
||
<taxonomicNameLabel id="A22E57EEFFA4FF92177A0FAEFC3EFE6E" box="[884,975,439,464]" pageId="3" pageNumber="504" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C694D04FFA4FF9210090FA0FB7BFE6E" box="[1031,1162,441,464]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="rumijaensis" status="sp. nov.">
|
||
<emphasis id="B91DEA95FFA4FF9210090FA0FB7BFE6E" box="[1031,1162,441,464]" italics="true" pageId="3" pageNumber="504">rumijaensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91DEA95FFA4FF92109C0FAEFB1CFE6E" bold="true" box="[1170,1261,439,464]" pageId="3" pageNumber="504">
|
||
<taxonomicNameLabel id="A22E57EEFFA4FF92109C0FAEFB1CFE6E" box="[1170,1261,439,464]" pageId="3" pageNumber="504" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
<emphasis id="B91DEA95FFA4FF9210E30FA1FB07FE71" box="[1261,1270,440,463]" italics="true" pageId="3" pageNumber="504">)</emphasis>
|
||
were found in more (two or three) underground localities.
|
||
<taxonomicName id="4C694D04FFA4FF9216A70FC5FCA5FE4B" box="[681,852,476,501]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91DEA95FFA4FF9216A70FC5FCA5FE4B" box="[681,852,476,501]" italics="true" pageId="3" pageNumber="504">Adriaphaenops</emphasis>
|
||
</taxonomicName>
|
||
inhabits subterranean habitats situated from low altitudes (
|
||
<taxonomicName id="4C694D04FFA4FF9215060C18FE90FDA6" box="[264,353,513,536]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="pretneri">
|
||
<emphasis id="B91DEA95FFA4FF9215060C18FE90FDA6" box="[264,353,513,536]" italics="true" pageId="3" pageNumber="504">pretneri</emphasis>
|
||
</taxonomicName>
|
||
from Vjetrenica cave on Popovo polje, ca
|
||
<elevation id="0044D1B4FFA4FF92175B0C19FC2BFDA6" box="[853,986,511,536]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="504" unit="m" value="220.0">
|
||
<quantity id="4C919B62FFA4FF92175B0C19FC6FFDA9" box="[853,926,512,536]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="504" unit="m" value="220.0">220 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
) thorough middle regions up to highly situated caves (
|
||
<taxonomicName id="4C694D04FFA4FF9215460C3DFE61FD83" box="[328,400,548,573]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="kevser">
|
||
<emphasis id="B91DEA95FFA4FF9215460C3DFE61FD83" box="[328,400,548,573]" italics="true" pageId="3" pageNumber="504">kevser</emphasis>
|
||
</taxonomicName>
|
||
from Vilina pećina cave on
|
||
<location id="8EB6605CFFA4FF9216D70C3DFC8AFD82" LSID="urn:lsid:plazi:treatment:03C08791FFA5FF9214990C38FAFFFAC0:8EB6605CFFA4FF9216D70C3DFC8AFD82" box="[729,891,548,573]" country="Albania" municipality="Durmitor Mts." name="Lebrsnik Mts." pageId="3" pageNumber="504">Lebršnik Mts.</location>
|
||
, ca
|
||
<elevation id="0044D1B4FFA4FF9217A50C3DFBCEFD83" box="[939,1087,548,573]" metricMagnitude="3" metricUnit="m" metricValue="1.8199999999999998" pageId="3" pageNumber="504" unit="m" value="1820.0">
|
||
<quantity id="4C919B62FFA4FF9217A50C3DFBF3FD82" box="[939,1026,548,572]" metricMagnitude="3" metricUnit="m" metricValue="1.8199999999999998" pageId="3" pageNumber="504" unit="m" value="1820.0">1820 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
and
|
||
<taxonomicName id="4C694D04FFA4FF9210740C3CFB2EFD82" box="[1146,1247,549,572]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" sensu="lato" species="zupcense">
|
||
<emphasis id="B91DEA95FFA4FF9210740C3CFB2EFD82" box="[1146,1247,549,572]" italics="true" pageId="3" pageNumber="504">zupcense</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22E57EEFFA4FF9210E60C3DFAF8FD83" box="[1256,1289,548,573]" pageId="3" pageNumber="504" sensu="lato">s.l.</taxonomicNameLabel>
|
||
from Pećina u Zupcima cave, ca
|
||
<elevation id="0044D1B4FFA4FF9215790C51FDF6FDDE" box="[375,519,583,608]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="3" pageNumber="504" unit="m" value="2100.0">
|
||
<quantity id="4C919B62FFA4FF9215790C51FE3DFDE1" box="[375,460,584,608]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="3" pageNumber="504" unit="m" value="2100.0">2100 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
and Jama na Vjetrenim Brdima, ca
|
||
<elevation id="0044D1B4FFA4FF9217970C51FBD8FDDE" box="[921,1065,583,608]" metricMagnitude="3" metricUnit="m" metricValue="2.17" pageId="3" pageNumber="504" unit="m" value="2170.0">
|
||
<quantity id="4C919B62FFA4FF9217970C51FC1FFDE1" box="[921,1006,584,608]" metricMagnitude="3" metricUnit="m" metricValue="2.17" pageId="3" pageNumber="504" unit="m" value="2170.0">2170 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
on
|
||
<collectingMunicipality id="6BB2ACFDFFA4FF92105C0C5EFB09FDDE" box="[1106,1272,583,608]" pageId="3" pageNumber="504">Durmitor Mts.</collectingMunicipality>
|
||
). Low situated caves are warmer and mostly drier, with air temperature about 8–9 o C, mountain caves are significantly colder and wetter, with air temperature between 2–4 o C.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C373650CFFA4FF9214C90CA0FAFFFAC0" pageId="3" pageNumber="504" type="description">
|
||
<paragraph id="8BD63687FFA4FF9214C90CA0FC84FBA8" blockId="3.[151,1437,151,1406]" pageId="3" pageNumber="504">
|
||
<emphasis id="B91DEA95FFA4FF9214C90CA0FEC9FD6C" bold="true" box="[199,312,697,722]" pageId="3" pageNumber="504">Bionomy.</emphasis>
|
||
Virtually nothing is known about the bionomy of
|
||
<taxonomicName id="4C694D04FFA4FF9217600CA0FBE8FD6C" box="[878,1049,697,722]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91DEA95FFA4FF9217600CA0FBE8FD6C" box="[878,1049,697,722]" italics="true" pageId="3" pageNumber="504">Adriaphaenops</emphasis>
|
||
</taxonomicName>
|
||
as well as other Dinaric genera of aphaenopsoid trechine beetles. Most of the taxa known so far, with the exception of
|
||
<taxonomicName id="4C694D04FFA4FF9210310CC4FEDAFCA4" authority="Lohaj & Lakota, 2010" authorityName="Lohaj & Lakota" authorityYear="2010" class="Insecta" family="Carabidae" genus="Acheroniotes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="mlejneki">
|
||
<emphasis id="B91DEA95FFA4FF9210310CC4FAC6FD48" box="[1087,1335,733,758]" italics="true" pageId="3" pageNumber="504">Acheroniotes mlejneki</emphasis>
|
||
Lohaj & Lakota, 2010
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C694D04FFA4FF92156C0D18FCA1FCA4" authority="Ganglbauer, 1891" authorityName="Ganglbauer" authorityYear="1891" box="[354,848,769,794]" class="Insecta" family="Carabidae" genus="Aphaenopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="apfelbecki">
|
||
<emphasis id="B91DEA95FFA4FF92156C0D18FD9AFCA4" box="[354,619,769,794]" italics="true" pageId="3" pageNumber="504">Aphaenopsis apfelbecki</emphasis>
|
||
(Ganglbauer, 1891)
|
||
</taxonomicName>
|
||
, were collected only individually or are known only from very few individuals. Representatives of these taxa were mostly found free walking or running on wet walls, speleothems and stones in the caves and pits, exceptionally collected also by pitfall traps baited with meat and/or cheese. Bionomy of
|
||
<taxonomicName id="4C694D04FFA4FF9215720D74FCFDFC38" authority="Lohaj & Lakota, 2010" authorityName="Lohaj & Lakota" authorityYear="2010" box="[380,780,877,902]" class="Insecta" family="Carabidae" genus="Acheroniotes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91DEA95FFA4FF9215720D74FDFEFC38" box="[380,527,877,902]" italics="true" pageId="3" pageNumber="504">Acheroniotes</emphasis>
|
||
Lohaj & Lakota, 2010
|
||
</taxonomicName>
|
||
is identical to other endogenous species of trechine genera like
|
||
<taxonomicName id="4C694D04FFA4FF9214C60D88FE46FC14" authority="Delarouzee" authorityName="Delarouzee" box="[200,439,913,938]" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91DEA95FFA4FF9214C60D88FEDCFC14" box="[200,301,913,938]" italics="true" pageId="3" pageNumber="504">Duvalius</emphasis>
|
||
Delarouzée
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C694D04FFA4FF9215CF0D88FD53FC14" authority="Jeannel" authorityName="Jeannel" box="[449,674,913,938]" class="Insecta" family="Carabidae" genus="Duvaliopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91DEA95FFA4FF9215CF0D88FDB6FC14" box="[449,583,913,938]" italics="true" pageId="3" pageNumber="504">Duvaliopsis</emphasis>
|
||
Jeannel
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="4C694D04FFA4FF9216C90D88FC31FC17" authority="Clairville. No" authorityName="Clairville. No" box="[711,960,913,938]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91DEA95FFA4FF9216C90D88FCEEFC14" box="[711,799,913,938]" italics="true" pageId="3" pageNumber="504">Trechus</emphasis>
|
||
Clairville. No
|
||
</taxonomicName>
|
||
specimens were found outside of the clay, on the walls or stones. Special behaviour was observed in
|
||
<taxonomicName id="4C694D04FFA4FF92173D0DACFB8EFC73" authority="Absolon. Most" authorityName="Absolon. Most" box="[819,1151,949,974]" class="Insecta" family="Carabidae" genus="Scotoplanetes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91DEA95FFA4FF92173D0DACFC3EFC70" box="[819,975,949,974]" italics="true" pageId="3" pageNumber="504">Scotoplanetes</emphasis>
|
||
Absolon. Most
|
||
</taxonomicName>
|
||
of the known specimens were found in cave hygropetric environment (
|
||
<bibRefCitation id="EFF84B76FFA4FF9216C80DC0FCBAFC4C" author="Sket" box="[710,843,985,1010]" pageId="3" pageNumber="504" refString="Sket, B. (2004) The cave hygropetric - a little known habitat and its inhabitants. Archiv fur Hydrobiologie, 160 (3), 413 - 425. https: // doi. org / 10.1127 / 0003 - 9136 / 2004 / 0160 - 0413" type="journal article" year="2004">Sket, 2004</bibRefCitation>
|
||
), in Vjetrenica cave (B&H) and Vodna jama pit (
|
||
<collectingCountry id="F37E7617FFA4FF9214900DE7FEDBFBA8" box="[158,298,1022,1046]" name="Serbia" pageId="3" pageNumber="504">Montenegro</collectingCountry>
|
||
) together with leptodirine beetle
|
||
<taxonomicName id="4C694D04FFA4FF9216910DE4FC84FBA8" authority="G. Muller." authorityName="G. Muller." box="[671,885,1021,1046]" class="Insecta" family="Leiodidae" genus="Hadesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91DEA95FFA4FF9216910DE4FD0DFBA8" box="[671,764,1021,1046]" italics="true" pageId="3" pageNumber="504">Hadesia</emphasis>
|
||
G. Müller.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BD63687FFA4FF9214C90A38FAFFFAC0" blockId="3.[151,1437,151,1406]" pageId="3" pageNumber="504">
|
||
<emphasis id="B91DEA95FFA4FF9214C90A38FDCCFB84" bold="true" box="[199,573,1057,1082]" pageId="3" pageNumber="504">Common external morphology.</emphasis>
|
||
Strongly flattened body with very long sensorial setae on elytra and/or head and pronotum can indicate that these taxa are inhabitants of narrow spaces like fissures, slots, rock debris and MSS (Mesovoid Shallow Substratum or originally
|
||
<emphasis id="B91DEA95FFA4FF9216980A70FC11FB3C" box="[662,992,1129,1154]" italics="true" pageId="3" pageNumber="504">Milieu Souterrain Superficiel</emphasis>
|
||
sensu
|
||
<bibRefCitation id="EFF84B76FFA4FF9210200A70FADAFB3C" author="Juberthie" box="[1070,1323,1129,1154]" pageId="3" pageNumber="504" refString="Juberthie, C., Delay, B. & Bouillon, M. (1980 a) Sur l´existence d´un milieu souterrain superficiel en zone non calcaire. Comptes rendus de l´Academie des Sciences, France, 290, 49 - 52." type="journal article" year="1980" yearSuffix="a">
|
||
Juberthie
|
||
<emphasis id="B91DEA95FFA4FF9210920A73FB24FB3C" box="[1180,1237,1129,1154]" italics="true" pageId="3" pageNumber="504">et al.</emphasis>
|
||
, 1980a
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFF84B76FFA4FF9211390A70FAB6FB3C" author="Juberthie" box="[1335,1351,1129,1154]" pageId="3" pageNumber="504" refString="Juberthie, C., Delay, B. & Bouillon, M. (1980 b) Extension du milieu souterrain en zone non-calcaire: description d´un nouveau milieu et son peuplement par les Coleopteres troglobies. Memoires de Biospeologie, 7, 19 - 52." type="journal article" year="1980" yearSuffix="b">b</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFF84B76FFA4FF92115A0A73FA61FB3C" author="Juberthie" box="[1364,1424,1130,1154]" pageId="3" pageNumber="504" refString="Juberthie, C., Bouillon, M. & Delay, B. (1981) Sur l´existence du milieu souterrain superfciel en zone calcaire. Memoires de Biospeologie, 8, 77 - 93." type="journal article" year="1981">1981</bibRefCitation>
|
||
). We suppose that conditions (temperature, humidity) and availability of food here is more stable than in large cave spaces, which they visited only occasionally. Morphology of mandibles, which are very long and slender, apically pointed and usually with robust teeth at the base of the right mandible, indicates that they are very active predators (
|
||
<bibRefCitation id="EFF84B76FFA4FF9214900AE0FE86FAAC" author="Casale" box="[158,375,1273,1298]" pageId="3" pageNumber="504" refString="Casale, A., Jalzic, B., Lohaj, R. & Mlejnek, R. (2012) Two new highly specialised subterranean beetles from the Velebit Massif (Croatia): Uelebitaphaenops (new genus) giganteus Casale & Jalzic, new species and Uelebitodromus ozrenlukici Lohaj, Mlejnek & Jalzic, new species (Coleoptera: Cholevidae: Leptodirini). Natura Croatica, 21 (1), 129 - 153." type="journal article" year="2012">
|
||
Casale
|
||
<emphasis id="B91DEA95FFA4FF9214FD0AE3FED7FAAC" box="[243,294,1273,1298]" italics="true" pageId="3" pageNumber="504">et al</emphasis>
|
||
., 2012
|
||
</bibRefCitation>
|
||
; Lohaj & Lakota, 2010; Lakota
|
||
<emphasis id="B91DEA95FFA4FF9216E70AE3FCEAFAAC" box="[745,795,1273,1298]" italics="true" pageId="3" pageNumber="504">et al</emphasis>
|
||
., 2010;
|
||
<bibRefCitation id="EFF84B76FFA4FF9217740AE0FB6CFAAC" author="Mlejnek" box="[890,1181,1273,1298]" pageId="3" pageNumber="504" refString="Mlejnek, R. & Zajicek, P. (2006) Reportaz ze speleologickych a biospeleologickych pruzkumu Cerne Hory v letech 2004 - 2005. A report from speleological and biospeleological survey of Monte Negro during 2004 - 2005. In czech, english abstract. Speleoforum, 25, 21 - 24." type="journal article" year="2006">Mlejnek & Zajíček, 2006</bibRefCitation>
|
||
; Mlejnek, Jalžić, pers. comm.). Interestingly, ecomorphological convergent evolution between Dinaric
|
||
<taxonomicName id="4C694D04FFA4FF92104B0B04FB01FA88" box="[1093,1264,1309,1334]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91DEA95FFA4FF92104B0B04FB01FA88" box="[1093,1264,1309,1334]" italics="true" pageId="3" pageNumber="504">Adriaphaenops</emphasis>
|
||
</taxonomicName>
|
||
and Pyrenean
|
||
<taxonomicName id="4C694D04FFA4FF9214990B58FEBBFAE4" box="[151,330,1345,1370]" class="Insecta" family="Carabidae" genus="Hydraphaenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91DEA95FFA4FF9214990B58FEBBFAE4" box="[151,330,1345,1370]" italics="true" pageId="3" pageNumber="504">Hydraphaenops</emphasis>
|
||
</taxonomicName>
|
||
species (eg.
|
||
<emphasis id="B91DEA95FFA4FF9215D70B5BFC5FFAE4" box="[473,942,1345,1370]" italics="true" pageId="3" pageNumber="504">
|
||
<taxonomicName id="4C694D04FFA4FF9215D70B5BFD88FAE7" box="[473,633,1346,1369]" class="Insecta" family="Carabidae" genus="Hydraphaenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="vasconicus">H. vasconicus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C694D04FFA4FF92168D0B5BFCE7FAE7" box="[643,790,1346,1369]" class="Insecta" family="Carabidae" genus="Hydraphaenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="navaricus">H. navaricus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C694D04FFA4FF92172F0B5BFC63FAE4" box="[801,914,1345,1370]" class="Insecta" family="Carabidae" genus="Hydraphaenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="elhersi">H. elhersi</taxonomicName>
|
||
...)
|
||
</emphasis>
|
||
provides evidences of adaptive processes as important drivers of phenotypic conservatism. Further research could develop these interesting topics.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |