197 lines
27 KiB
XML
197 lines
27 KiB
XML
<document id="61B553FA30D0D23DAB5D3E89CDCBCE84" ID-CLB-Dataset="8419" ID-DOI="10.11646/zootaxa.4852.4.2" ID-GBIF-Dataset="1430f8df-baac-425a-990d-e9db0b875e5d" ID-ISSN="1175-5326" ID-Zenodo-Dep="4410041" ID-ZooBank="61DA8090-7DC0-4968-9503-C7FB3A113ED2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609596614450" checkinUser="plazi" docAuthor="Hutchinson, Paul M. & Allsopp, Peter G." docDate="2020" docId="039F87D1FFA7CF14F5FFFAEC89A75ECD" docLanguage="en" docName="zootaxa.4852.4.2.pdf" docOrigin="Zootaxa 4852 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Epironastes Carne 1957" docType="treatment" docVersion="7" lastPageNumber="430" masterDocId="FFA6FFA9FFA6CF16F568FFD78B415D4C" masterDocTitle="Revision of Epironastes Carne, 1957 (Coleoptera: Scarabaeidae: Dynastinae Pentodontini: Pseudoryctina) with description of the female, behaviours, and habitats" masterLastPageNumber="448" masterPageNumber="428" pageNumber="429" updateTime="1698851051965" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="9F53F46392796464B511C495BCD7A1B2">Revision of Epironastes Carne, 1957 (Coleoptera: Scarabaeidae: Dynastinae Pentodontini: Pseudoryctina) with description of the female, behaviours, and habitats</mods:title>
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<mods:name id="21BB3847F6A9CA7E76FF110EEB2DB7B5" type="personal">
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<mods:namePart id="43DE3AB59D0E6A0DB0E670B625CAD329">Hutchinson, Paul M.</mods:namePart>
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<mods:affiliation id="00EE4860691BA55390C91DEF4D1204DC">Department of Agriculture and Food WA (Quarantine WA), Locked Bag 69, Welshpool DC, Western Australia, 6986, Australia. paul. hutchinson @ dpird. wa. gov. au; https: // orcid. org / 0000 - 0001 - 8747 - 1320</mods:affiliation>
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<mods:nameIdentifier id="F241B622B055DCD813F98312AE064ECB" type="email">paul.hutchinson@dpird.wa.gov.au</mods:nameIdentifier>
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<mods:namePart id="BBD14EDA88199A015C33A8FDE3B1EDBF">Allsopp, Peter G.</mods:namePart>
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<mods:affiliation id="BA02AAA5D5A6ED132E546CB78610284A">Queensland Museum, PO Box 3300, South Brisbane, Queensland, 4101, Australia.</mods:affiliation>
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<mods:title id="C8BAD8CC49AEBC1BB4746F48742703DE">Zootaxa</mods:title>
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<mods:date id="9C22C713842668D1FFF5F6ED3455F34B">2020</mods:date>
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<mods:number id="5F6BCCF8E19ED011973BDD086055CE5A">4852</mods:number>
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<treatment id="039F87D1FFA7CF14F5FFFAEC89A75ECD" ID-DOI="http://doi.org/10.5281/zenodo.4410027" ID-GBIF-Taxon="177217056" ID-Zenodo-Dep="4410027" LSID="urn:lsid:plazi:treatment:039F87D1FFA7CF14F5FFFAEC89A75ECD" httpUri="http://treatment.plazi.org/id/039F87D1FFA7CF14F5FFFAEC89A75ECD" lastPageId="2" lastPageNumber="430" pageId="1" pageNumber="429">
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<paragraph id="8B8936C7FFA7CF17F5FFFAEC8A875818" blockId="1.[151,454,1338,1365]" box="[151,454,1338,1365]" pageId="1" pageNumber="429">
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<heading id="D0C181ABFFA7CF17F5FFFAEC8A875818" bold="true" box="[151,454,1338,1365]" fontSize="11" level="1" pageId="1" pageNumber="429" reason="1">
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<taxonomicName id="4C364D44FFA7CF17F5FFFAEC8A875818" authority="Carne, 1957" authorityName="Carne" authorityYear="1957" box="[151,454,1338,1365]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="429" phylum="Arthropoda" rank="genus">
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<emphasis id="B942EAD5FFA7CF17F5FFFAEC8A875818" bold="true" box="[151,454,1338,1365]" pageId="1" pageNumber="429">
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<emphasis id="B942EAD5FFA7CF17F5FFFAEC8A675819" bold="true" box="[151,294,1339,1365]" italics="true" pageId="1" pageNumber="429">Epironastes</emphasis>
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<bibRefCitation id="EFA74B36FFA7CF17F446FAED8A875818" author="Carne, P. B." box="[302,454,1338,1364]" pageId="1" pageNumber="429" refId="ref9996" refString="Carne, P. B. (1957) A Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Commonwealth Scientific and Industrial Research Organisation, Melbourne, 284 pp." type="book" year="1957">Carne, 1957</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="C32C654CFFA7CF17F5FFFA5789A75898" pageId="1" pageNumber="429" type="reference_group">
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<paragraph id="8B8936C7FFA7CF17F5FFFA5788BA58FA" blockId="1.[151,1436,1407,1493]" pageId="1" pageNumber="429">
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<treatmentCitation id="0A9710D6FFA7CF17F5FFFA578A8E58DB" author="Carne, P. B." box="[151,463,1407,1431]" page="124" pageId="1" pageNumber="429" year="1957">
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<taxonomicName id="4C364D44FFA7CF17F5FFFA578A8E58DB" authority="Carne, 1957: 124" authorityName="Carne" authorityPageNumber="124" authorityYear="1957" box="[151,463,1407,1431]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="429" phylum="Arthropoda" rank="genus">
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<emphasis id="B942EAD5FFA7CF17F5FFFA578A4E58DB" box="[151,271,1408,1431]" italics="true" pageId="1" pageNumber="429">Epironastes</emphasis>
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<bibRefCitation id="EFA74B36FFA7CF17F470FAA88A8E58DB" author="Carne, P. B." box="[280,463,1407,1431]" pageId="1" pageNumber="429" refId="ref9996" refString="Carne, P. B. (1957) A Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Commonwealth Scientific and Industrial Research Organisation, Melbourne, 284 pp." type="book" year="1957">Carne, 1957: 124</bibRefCitation>
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</taxonomicName>
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, Figs. 322–333;
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<treatmentCitation id="0A9710D6FFA7CF17F7E8FAA8880458DB" author="Endrodi, S." box="[640,837,1407,1431]" page="382" pageId="1" pageNumber="429" year="1985">
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<bibRefCitation id="EFA74B36FFA7CF17F7E8FAA8880458DB" author="Endrodi, S." box="[640,837,1407,1431]" pageId="1" pageNumber="429" refId="ref10200" refString="Endrodi, S. (1985) The Dynastinae of the World. Junk, Dordrecht, 800 pp." type="book" year="1985">Endrődi 1985: 382</bibRefCitation>
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</treatmentCitation>
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, Figs. 1395–1397;
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<treatmentCitation id="0A9710D6FFA7CF17F167FAA88E5B58DB" author="Cassis, G. & Weir, T. A." box="[1039,1306,1407,1431]" page="400" pageId="1" pageNumber="429" year="1992">
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<bibRefCitation id="EFA74B36FFA7CF17F167FAA88E5B58DB" author="Cassis, G. & Weir, T. A." box="[1039,1306,1407,1431]" pageId="1" pageNumber="429" refId="ref10030" refString="Cassis, G. & Weir, T. A. (1992) Dynastinae. In: Houston, W. W. K. (ed), Zoological Catalogue of Australia. Coleoptera: Scarabaeoidea. Volume 9. Australian Government Publishing Service, Canberra, pp. 383 - 425." type="book" year="1992">Cassis & Weir 1992: 400</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0A9710D6FFA7CF17F04FFAA88A6258FA" author="Dechambre, R-P." page="67" pageId="1" pageNumber="429" year="2005">
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<bibRefCitation id="EFA74B36FFA7CF17F04FFAA88A6258FA" author="Dechambre, R-P." pageId="1" pageNumber="429" refId="ref10085" refString="Dechambre, R-P. (2005) Les Coleopteres du Monde. Vol. 30. Dynastidae Australiens et Oceaniens. Hillside Books, Canterbury, 132 pp." type="book" year="2005">Dechambre 2005: 67</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0A9710D6FFA7CF17F446FA498A9D58FA" author="Krajcik, M." box="[302,476,1438,1462]" page="42" pageId="1" pageNumber="429" year="2005">
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<bibRefCitation id="EFA74B36FFA7CF17F446FA498A9D58FA" author="Krajcik, M." box="[302,476,1438,1462]" pageId="1" pageNumber="429" pagination="1 - 122" refId="ref10336" refString="Krajcik, M. (2005) Dynastinae of the World, checklist. Animma. X Supplement, 2, 1 - 122." type="journal article" year="2005">Krajcik 2005: 42</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0A9710D6FFA7CF17F48FFA4989E358FA" author="Krajcik, M." box="[487,674,1438,1462]" page="103" pageId="1" pageNumber="429" year="2012">
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<bibRefCitation id="EFA74B36FFA7CF17F48FFA4989E358FA" author="Krajcik, M." box="[487,674,1438,1462]" pageId="1" pageNumber="429" pagination="1 - 278" refId="ref10361" refString="Krajcik, M. (2012) Checklist of the World Scarabaeoidea. Animma. X Supplement, 5, 1 - 278." type="journal article" year="2012">Krajcik 2012: 103</bibRefCitation>
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</treatmentCitation>
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; Weir
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||
<emphasis id="B942EAD5FFA7CF17F78BFA49885758FA" box="[739,790,1438,1462]" italics="true" pageId="1" pageNumber="429">et.al.</emphasis>
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2019: 524, Plate 54O.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8B8936C7FFA7CF17F5FFFA6A89A75898" blockId="1.[151,1436,1407,1493]" box="[151,742,1468,1493]" pageId="1" pageNumber="429">
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<emphasis id="B942EAD5FFA7CF17F5FFFA6A8A4E5898" box="[151,271,1469,1492]" italics="true" pageId="1" pageNumber="429">Apironastes</emphasis>
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:
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<bibRefCitation id="EFA74B36FFA7CF17F475FA6B8A915899" author="Endrodi, S." box="[285,464,1468,1493]" pageId="1" pageNumber="429" pagination="17 - 70" refId="ref10164" refString="Endrodi, S. (1974) Monographie der Dynastinae 4. Tribus: Pentodontini von Australien und Neu-Zeeland (Coleoptera). Folia Entomologica Hungarica, Series Nova, 27, 17 - 70." type="journal article" year="1974">Endrődi 1974: 38</bibRefCitation>
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,
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<figureCitation id="130D2A42FFA7CF17F4B3FA6B89135898" box="[475,594,1468,1492]" captionStart="FIGURES 43–50" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[172,1415,217,1919]" captionTargetId="figure-25@15.[172,1415,181,1919]" captionTargetPageId="15" captionText="FIGURES 43–50. Epironastes nigrisetosus Carne, 1957, male, Dempster Inlet specimen. 43, dorsum; 44, venter; 45, lateral; 46, head, lateral; 47, head, dorsal; 48, pygidium; 49, parameres; 50, aedeagus, lateral." figureDoi="http://doi.org/10.5281/zenodo.4410065" httpUri="https://zenodo.org/record/4410065/files/figure.png" pageId="1" pageNumber="429">Figs. 48–50</figureCitation>
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(misspelling).
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C654CFFA7CF17F5FFF9D488B55B51" box="[151,1012,1538,1565]" pageId="1" pageNumber="429" type="type_taxon">
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<paragraph id="8B8936C7FFA7CF17F5FFF9D488B55B51" blockId="1.[151,1437,1538,2033]" box="[151,1012,1538,1565]" pageId="1" pageNumber="429">
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<emphasis id="B942EAD5FFA7CF17F5FFF9D48A735B51" bold="true" box="[151,306,1538,1565]" pageId="1" pageNumber="429">
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<typeStatus id="548D8865FFA7CF17F5FFF9D48B935B51" box="[151,210,1539,1565]" pageId="1" pageNumber="429">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C364D44FFA7CF17F451F9D4899E5B51" authority="Carne, 1957" authorityName="Carne" authorityYear="1957" box="[313,735,1538,1565]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="429" phylum="Arthropoda" rank="species" species="nigrisetosus">
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<emphasis id="B942EAD5FFA7CF17F451F9D4890D5B50" box="[313,588,1539,1564]" italics="true" pageId="1" pageNumber="429">Epironastes nigrisetosus</emphasis>
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<bibRefCitation id="EFA74B36FFA7CF17F73BF9D4899E5B51" author="Carne, P. B." box="[595,735,1538,1565]" pageId="1" pageNumber="429" refId="ref9996" refString="Carne, P. B. (1957) A Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Commonwealth Scientific and Industrial Research Organisation, Melbourne, 284 pp." type="book" year="1957">Carne, 1957</bibRefCitation>
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</taxonomicName>
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, by original designation.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C32C654CFFA7CF14F5AFF9F18A825F71" lastPageId="2" lastPageNumber="430" pageId="1" pageNumber="429" type="description">
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<paragraph id="8B8936C7FFA7CF14F5AFF9F18F0B5C51" blockId="1.[151,1437,1538,2033]" lastBlockId="2.[151,1437,151,897]" lastPageId="2" lastPageNumber="430" pageId="1" pageNumber="429">
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<emphasis id="B942EAD5FFA7CF17F5AFF9F18A345B0C" bold="true" box="[199,373,1574,1600]" pageId="1" pageNumber="429">Redescription.</emphasis>
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Body dorsally dark brown-black, ventral surface brown or black,
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<quantity id="4CCE9B22FFA7CF17F10EF9F08FBD5B0D" box="[1126,1276,1575,1601]" metricMagnitude="-2" metricUnit="m" metricValue="1.5699999999999998" metricValueMax="2.17" metricValueMin="0.97" pageId="1" pageNumber="429" unit="mm" value="15.7" valueMax="21.7" valueMin="9.7">9.7–21.7 mm</quantity>
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long (apex of clypeus to tip of pygidium), 6.0–
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<quantity id="4CCE9B22FFA7CF17F77EF99C893D5B29" box="[534,636,1611,1637]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="1" pageNumber="429" unit="mm" value="12.6">12.6 mm</quantity>
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wide (widest at midlength of elytra). Mentum broad at base, slightly narrowed to ligula; ligula slightly or strongly reflexed, rounded or weakly bilobed; maxillae reduced to small subtriangular pieces with irregular inner margins; palps short, palpomere 4 cylindrical; mandibles exposed, evenly rounded; labrum concealed. Antenna with 10 antennomeres, club longer than shaft, scape and pedicel setose. Clypeus strongly narrowed, upturned at apex, weakly emarginate, deflexed across ridge joining antennal angles anterior to clypeofrontal suture; clypeofrontal suture usually obliterated, transverse, in middle weakly posteriorly arcuate. Frons rugulose; ocular canthi broad, coarsely punctate, surpassing outline of eye. Pronotum with anterior margin membranous, disc conspicuously punctate, basal ridge continuous, widest in apical half, basolateral angles well defined. Each elytron with sutural stria punctate or linear-punctate, punctures of disc simple or annulate, surface often microreticulate, epipleurae poorly developed, lateral setae extending posteriorly as far as first abdominal sternite. Postcoxal prosternal process apically tapering posteriorly or truncate, densely setose. Pygidium weakly or strongly convex, mostly setose across base, apical ridge mostly glabrous. Abdominal sternites setose, sternite 5 glabrous across middle; coxae with abundant yellow, orange or black vestiture. Protibiae tridentate, tarsomeres 1 and 5 subequal in length; metafemora moderately stout bearing a single sulcus; metatibiae dilated at apex, bicarinate, distal cilia short, stout and closely set, spurs usually broad, subparallel or subspatulate, not or only weakly curved; metatarsomeres slender, tarsomere 1 not dilated, equal to or longer than tarsomere 5.
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</paragraph>
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<paragraph id="8B8936C7FFA4CF14F5AFFEF08B8C5C9D" blockId="2.[151,1437,151,897]" pageId="2" pageNumber="430">
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<emphasis id="B942EAD5FFA4CF14F5AFFEF08A4A5C0D" bold="true" box="[199,267,295,321]" pageId="2" pageNumber="430">Male.</emphasis>
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Pronotum with anteromedian tubercle in front of an impression that is either confined to anterior half or continued over basal convexity by shallow impression. Antennal club subequal to or slightly longer than shaft, straight, elongate oval, setose on inner surface. Genitalia with phallobase short, broad, parameres narrow basally, usually laterally abruptly dilated in apical half, either with arcuate margin to apex or with pre-apical lateral process.
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</paragraph>
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<paragraph id="8B8936C7FFA4CF14F5AFFE0C8A825F71" blockId="2.[151,1437,151,897]" pageId="2" pageNumber="430">
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<emphasis id="B942EAD5FFA4CF14F5AFFE0C8A645CB9" bold="true" box="[199,293,475,501]" pageId="2" pageNumber="430">Female.</emphasis>
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Pronotum with anteromedian tubercle obsolete, impression as small, flattened region. Antennal club shorter than shaft. Abdominal sternites setose across middle. Metatarsomeres stout, shorter than tibial length, tarsomere 1 slightly broadened.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C654CFFA4CF14F5AFFD9189A75ECD" pageId="2" pageNumber="430" type="diagnosis">
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<paragraph id="8B8936C7FFA4CF14F5AFFD9189A75ECD" blockId="2.[151,1437,151,897]" pageId="2" pageNumber="430">
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<emphasis id="B942EAD5FFA4CF14F5AFFD918A905F2C" bold="true" box="[199,465,582,608]" pageId="2" pageNumber="430">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="4C364D44FFA4CF14F4B3FD9089215F2C" authorityName="Carne" authorityYear="1957" box="[475,608,583,608]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B942EAD5FFA4CF14F4B3FD9089215F2C" box="[475,608,583,608]" italics="true" pageId="2" pageNumber="430">Epironastes</emphasis>
|
||
</taxonomicName>
|
||
shares with
|
||
<taxonomicName id="4C364D44FFA4CF14F79EFD908F5C5F2D" authority="Sharp, 1873 a" authorityName="Sharp" authorityYear="1873" box="[758,1053,583,609]" class="Insecta" family="Dynastidae" genus="Teinogenys" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B942EAD5FFA4CF14F79EFD9088325F2C" box="[758,883,583,608]" italics="true" pageId="2" pageNumber="430">Teinogenys</emphasis>
|
||
Sharp, 1873 a
|
||
</taxonomicName>
|
||
broad, strongly compressed, and reflexed mentum; large exposed mandibles; and small maxillary palps. The two genera are at once distinguished by the upturned clypeus in
|
||
<taxonomicName id="4C364D44FFA4CF14F4C9FD5889675FE4" authorityName="Carne" authorityYear="1957" box="[417,550,655,680]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B942EAD5FFA4CF14F4C9FD5889675FE4" box="[417,550,655,680]" italics="true" pageId="2" pageNumber="430">Epironastes</emphasis>
|
||
</taxonomicName>
|
||
, and the presence in this genus of an anteromedian pronotal tubercle. The shape of the clypeus (
|
||
<figureCitation id="130D2A42FFA4CF14F420FD648AD95F81" box="[328,408,691,717]" captionStart="FIGURES 3–10" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[181,1404,193,1919]" captionTargetId="figure-19@4.[181,1407,181,1920]" captionTargetPageId="4" captionText="FIGURES 3–10. Epironastes abruptus Carne, 1957, male specimen #0465. 3, dorsum; 4, venter; 5, lateral; 6, head, lateral; 7, head, dorsal; 8, pygidium; 9, parameres; 10, aedeagus, lateral." figureDoi="http://doi.org/10.5281/zenodo.4410045" httpUri="https://zenodo.org/record/4410045/files/figure.png" pageId="2" pageNumber="430">Figs. 7</figureCitation>
|
||
,
|
||
<figureCitation id="130D2A42FFA4CF14F4CDFD658A825F81" box="[421,451,690,717]" captionStart="FIGURES 11–16" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[180,1401,181,1919]" captionTargetId="figure-25@5.[180,1408,181,1919]" captionTargetPageId="5" captionText="FIGURES 11–16. Epironastes abruptus Carne, 1957, female specimen #0464. 11, dorsum; 12, venter; 13, lateral; 14, head, lateral; 15, head, dorsal; 16, head and pronotum, frontal." figureDoi="http://doi.org/10.5281/zenodo.4410047" httpUri="https://zenodo.org/record/4410047/files/figure.png" pageId="2" pageNumber="430">15</figureCitation>
|
||
,
|
||
<figureCitation id="130D2A42FFA4CF14F4B9FD658AAE5F81" box="[465,495,690,717]" captionStart="FIGURES 21–28" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[188,1385,187,1920]" captionTargetId="figure-25@9.[188,1399,181,1920]" captionTargetPageId="9" captionText="FIGURES 21–28. Epironastes demarzi (Endrődi 1971), male specimen #1713. 21, dorsum; 22, venter; 23, lateral; 24, head, lateral; 25, head, dorsal; 26, pygidium; 27, parameres; 28, aedeagus, lateral." figureDoi="http://doi.org/10.5281/zenodo.4410053" httpUri="https://zenodo.org/record/4410053/files/figure.png" pageId="2" pageNumber="430">25</figureCitation>
|
||
,
|
||
<figureCitation id="130D2A42FFA4CF14F494FD64895B5F81" box="[508,538,691,717]" captionStart="FIGURES 32–39" captionStartId="12.[151,264,1943,1967]" captionTargetBox="[161,1405,189,1920]" captionTargetId="figure-19@12.[161,1427,181,1920]" captionTargetPageId="12" captionText="FIGURES 32–39. Epironastes limbatus Carne, 1957, male specimen #1703; 32, dorsum; 33, venter; 34, lateral; 35, head, lateral; 36, head, dorsal; 37, pygidium; 38, parameres; 39, aedeagus, lateral." figureDoi="http://doi.org/10.5281/zenodo.4410059" httpUri="https://zenodo.org/record/4410059/files/figure.png" pageId="2" pageNumber="430">36</figureCitation>
|
||
,
|
||
<figureCitation id="130D2A42FFA4CF14F740FD6489045F81" box="[552,581,691,717]" captionStart="FIGURES 43–50" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[172,1415,217,1919]" captionTargetId="figure-25@15.[172,1415,181,1919]" captionTargetPageId="15" captionText="FIGURES 43–50. Epironastes nigrisetosus Carne, 1957, male, Dempster Inlet specimen. 43, dorsum; 44, venter; 45, lateral; 46, head, lateral; 47, head, dorsal; 48, pygidium; 49, parameres; 50, aedeagus, lateral." figureDoi="http://doi.org/10.5281/zenodo.4410065" httpUri="https://zenodo.org/record/4410065/files/figure.png" pageId="2" pageNumber="430">47</figureCitation>
|
||
) separates
|
||
<taxonomicName id="4C364D44FFA4CF14F7ACFD6488085F80" authorityName="Carne" authorityYear="1957" box="[708,841,691,716]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B942EAD5FFA4CF14F7ACFD6488085F80" box="[708,841,691,716]" italics="true" pageId="2" pageNumber="430">Epironastes</emphasis>
|
||
</taxonomicName>
|
||
from all other
|
||
<taxonomicName id="4C364D44FFA4CF14F69EFD648FD95F81" authorityName="Carne" authorityYear="1957" box="[1014,1176,691,717]" class="Insecta" family="Dynastidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="subTribe" subTribe="Pseudoryctina">Pseudoryctina</taxonomicName>
|
||
. It is deflexed ventrad, either evenly curved or abruptly declivous, not from the clypeofrontal suture, but from a ridge joining the antennal angles, and is narrowed distally, the sides sinuate to a very narrow anterior face that is distinctly upturned. In this, and in some other characters,
|
||
<taxonomicName id="4C364D44FFA4CF14F486FCC889325E74" authorityName="Carne" authorityYear="1957" box="[494,627,799,824]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B942EAD5FFA4CF14F486FCC889325E74" box="[494,627,799,824]" italics="true" pageId="2" pageNumber="430">Epironastes</emphasis>
|
||
</taxonomicName>
|
||
looks similar to species of
|
||
<taxonomicName id="4C364D44FFA4CF14F6D8FCC88F9F5E75" authority="Hope, 1847" authorityName="Hope" authorityYear="1847" box="[944,1246,799,825]" class="Insecta" family="Dynastidae" genus="Semanopterus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B942EAD5FFA4CF14F6D8FCC88F0E5E75" box="[944,1103,799,825]" italics="true" pageId="2" pageNumber="430">Semanopterus</emphasis>
|
||
Hope, 1847
|
||
</taxonomicName>
|
||
, but
|
||
<taxonomicName id="4C364D44FFA4CF14F07FFCC88EDD5E74" authorityName="Carne" authorityYear="1957" box="[1303,1436,799,824]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B942EAD5FFA4CF14F07FFCC88EDD5E74" box="[1303,1436,799,824]" italics="true" pageId="2" pageNumber="430">Epironastes</emphasis>
|
||
</taxonomicName>
|
||
species have sexually dimorphic antennae and an upturned clypeus. The keys of
|
||
<bibRefCitation id="EFA74B36FFA4CF14F179FC948FE45E11" author="Carne, P. B." box="[1041,1189,834,861]" pageId="2" pageNumber="430" refId="ref9996" refString="Carne, P. B. (1957) A Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Commonwealth Scientific and Industrial Research Organisation, Melbourne, 284 pp." type="book" year="1957">Carne (1957)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFA74B36FFA4CF14F1B1FC948EDD5E11" author="Weir, T. A. & Lawrence, J. F. & Lemann, C. & Gunter, N. L." box="[1241,1436,835,861]" pageId="2" pageNumber="430" pagination="1919" refId="ref10501" refString="Weir, T. A., Lawrence, J. F., Lemann, C. & Gunter, N. L. (2019) Scarabaeidae: Dynastinae Macleay 1919 [sic]. In: Slipinski, A. & Lawrence, J. F. (Eds.), Australian Beetles. Volume 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton, pp. 516 - 530." type="book chapter" year="2019">
|
||
Weir
|
||
<emphasis id="B942EAD5FFA4CF14F07DFC948E0D5E11" box="[1301,1356,835,861]" italics="true" pageId="2" pageNumber="430">et al.</emphasis>
|
||
(2019)
|
||
</bibRefCitation>
|
||
will separate the genera of Australian
|
||
<taxonomicName id="4C364D44FFA4CF14F728FCB089A35ECD" authorityName="Carne" authorityYear="1957" box="[576,738,871,897]" class="Insecta" family="Dynastidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="430" phylum="Arthropoda" rank="subTribe" subTribe="Pseudoryctina">Pseudoryctina</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |