253 lines
28 KiB
XML
253 lines
28 KiB
XML
<document id="A6A026099D87182C0547E63F7817A362" ID-DOI="10.11646/zootaxa.4090.1.1" ID-GBIF-Dataset="e6b35f5a-435e-4473-b29e-1e4d842f84b0" ID-ISSN="1175-5326" ID-Zenodo-Dep="263299" ID-ZooBank="203BAC25-B542-48FE-B5AD-EBA8C0285833" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1457964633549" checkinUser="plazi" docAuthor="Fallon, Phillip J." docDate="2016" docId="039F87C4FAE5FF28CBAFB983FB6DF8EF" docLanguage="en" docName="15612-5136-1-PB.pdf.imf" docOrigin="Zootaxa 4090 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Douglassia Bartsch 1934" docType="treatment" docVersion="13" lastPageNumber="129" masterDocId="FFA6FFBCFA65FFA8CB38B91BFFE8FFAA" masterDocTitle="Taxonomic review of tropical western Atlantic shallow water Drilliidae (Mollusca: Gastropoda: Conoidea) including descriptions of 100 new species" masterLastPageNumber="363" masterPageNumber="1" pageNumber="129" updateTime="1698184378282" updateUser="plazi">
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<mods:titleInfo id="D2D09D3D020710F8F94900C357EBA6D8">
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<mods:title id="F74D93B95B54038967B181E524E580DD">Taxonomic review of tropical western Atlantic shallow water Drilliidae (Mollusca: Gastropoda: Conoidea) including descriptions of 100 new species</mods:title>
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<mods:name id="283B5F2C8DC920920F5741FF451E4F3B" type="personal">
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<mods:namePart id="2FFF0A1EBD675AB41F82AC77520C9208">Fallon, Phillip J.</mods:namePart>
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<mods:title id="E08D7612D54BA087899C8CC0E75B4730">Zootaxa</mods:title>
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<mods:part id="BC038DF82EA58D7CAF6ECB4F9CD0C33D">
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<mods:date id="E1468BA373EDF429AF8CC158AE757F91">2016</mods:date>
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<mods:number id="C06CFFAB86CD1DEFED7C4ADE8ED3D8E3">4090</mods:number>
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<mods:classification id="A9F71825475F785EBC5955A7352874CE">journal article</mods:classification>
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<mods:identifier id="11CB3EAF906CAA645B647317F5543767" type="DOI">10.11646/zootaxa.4090.1.1</mods:identifier>
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<treatment id="039F87C4FAE5FF28CBAFB983FB6DF8EF" ID-DOI="http://doi.org/10.5281/zenodo.6076380" ID-GBIF-Taxon="117911643" ID-Zenodo-Dep="6076380" LSID="urn:lsid:plazi:treatment:039F87C4FAE5FF28CBAFB983FB6DF8EF" httpUri="http://treatment.plazi.org/id/039F87C4FAE5FF28CBAFB983FB6DF8EF" lastPageNumber="129" pageId="128" pageNumber="129">
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<subSubSection id="C32C6559FAE5FF28CBAFB983FAFCFF59" pageId="128" pageNumber="129" type="nomenclature">
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<paragraph id="8B8936D2FAE5FF28CBAFB983FDCFFF18" blockId="128.[151,551,151,178]" box="[151,551,151,178]" pageId="128" pageNumber="129">
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<heading id="D0C181BEFAE5FF28CBAFB983FDCFFF18" bold="true" box="[151,551,151,178]" fontSize="11" level="1" pageId="128" pageNumber="129" reason="1">
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<emphasis id="B942EAC0FAE5FF28CBAFB983FDCFFF18" bold="true" box="[151,551,151,178]" pageId="128" pageNumber="129">
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Genus
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<taxonomicName id="4C364D51FAE5FF28CBD6B98CFDCFFF18" ID-CoL="7P36G" authority="Bartsch, 1934" authorityName="Bartsch" authorityYear="1934" box="[238,551,151,178]" class="Gastropoda" family="Drilliidae" genus="Douglassia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="128" pageNumber="129" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FAE5FF28CBD6B98CFE9EFF1B" bold="true" box="[238,374,151,177]" italics="true" pageId="128" pageNumber="129">Douglassia</emphasis>
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Bartsch, 1934
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8936D2FAE5FF28CBAFB9C5FAFCFF59" blockId="128.[151,1300,221,243]" box="[151,1300,221,243]" pageId="128" pageNumber="129">
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<typeStatus id="548D8870FAE5FF28CBAFB9C5FF21FF59" box="[151,201,222,243]" pageId="128" pageNumber="129">Type</typeStatus>
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species:
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<taxonomicName id="4C364D51FAE5FF28CA1EB9C6FD80FF59" authority="Bartsch, 1934" authorityName="Bartsch" authorityYear="1934" box="[294,616,221,243]" class="Gastropoda" family="Drilliidae" genus="Douglassia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="128" pageNumber="129" phylum="Mollusca" rank="species" species="enae">
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<emphasis id="B942EAC0FAE5FF28CA1EB9C6FE27FF59" box="[294,463,221,243]" italics="true" pageId="128" pageNumber="129">Douglassia enae</emphasis>
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, Bartsch, 1934
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</taxonomicName>
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by original designation (Recent, off the north coast of
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<collectingCountry id="F3217642FAE5FF28CFACB9C5FAE5FF59" box="[1172,1293,221,243]" name="Puerto Rico" pageId="128" pageNumber="129">Puerto Rico</collectingCountry>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C6559FAE5FF28CBAFB804FAFBFE47" pageId="128" pageNumber="129" type="diagnosis">
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<paragraph id="8B8936D2FAE5FF28CBAFB804FAFBFE47" blockId="128.[151,1437,287,565]" pageId="128" pageNumber="129">
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<emphasis id="B942EAC0FAE5FF28CBAFB804FEF9FE92" bold="true" box="[151,273,287,312]" pageId="128" pageNumber="129">Diagnosis.</emphasis>
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According to Bartsch (1934: 5–6), a medium-sized shell with smooth, round protoconch whorls; teleoconch whorls with a smooth concave sulcus devoid of ribs, remaining portion of whorls convex. Shell base short with a short columella and weak fasciole; aperture large with a deep anterior canal and anal sinus, the latter adjoining the suture. Inner lip recumbent with a parietal lobe posteriorly; outer lip with a strong stromboid notch. Axial ribs extend from sulcus, grow strongest on periphery, and evanesce on base near anterior fasciole; fine growth striae present overall. Whorls marked with fine spiral lines; anterior fasciole with spiral ridges.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C6559FAE5FF28CBFFB8E3FC61FDD7" pageId="128" pageNumber="129" type="key">
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<paragraph id="8B8936D2FAE5FF28CBFFB8E3FAF9FD9E" blockId="128.[151,1437,287,565]" pageId="128" pageNumber="129">
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<emphasis id="B942EAC0FAE5FF28CBFFB8E3FE5FFDBA" bold="true" box="[199,439,503,528]" pageId="128" pageNumber="129">Key characteristics.</emphasis>
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The presence of all the following characteristics, in addition to those that define the family, is diagnostic of species of
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<taxonomicName id="4C364D51FAE5FF28C92CBB07FD7CFD9F" box="[532,660,540,565]" class="Gastropoda" family="Drilliidae" genus="Douglassia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="128" pageNumber="129" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FAE5FF28C92CBB07FD7CFD9F" box="[532,660,540,565]" italics="true" pageId="128" pageNumber="129">Douglassia</emphasis>
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</taxonomicName>
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and separates the genus from other TWA drilliid genera:
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</paragraph>
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<paragraph id="8B8936D2FAE5FF28CBAFBB7FFC61FDD7" blockId="128.[151,1436,612,888]" box="[151,905,612,637]" pageId="128" pageNumber="129">1. Shell fusiform, stout, (W/L> 0.40), with a short anterior canal.</paragraph>
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</subSubSection>
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<subSubSection id="C32C6559FAE5FF28CBAFBB93FEAEF972" pageId="128" pageNumber="129" type="description">
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<paragraph id="8B8936D2FAE5FF28CBAFBB93FB34FD0A" blockId="128.[151,1436,612,888]" box="[151,1244,647,672]" pageId="128" pageNumber="129">2. Whorls with a concave sulcus lacking ribs in later whorls; ribs strongly convex below sulcus;</paragraph>
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<paragraph id="8B8936D2FAE5FF28CBAFBBB7FD4FFD6F" blockId="128.[151,1436,612,888]" box="[151,679,684,709]" pageId="128" pageNumber="129">3. Protoconch of 2½–3 smooth round whorls;</paragraph>
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<paragraph id="8B8936D2FAE5FF28CBAFBBCBFE2DFC9A" blockId="128.[151,1436,612,888]" pageId="128" pageNumber="129">4. Spiral microsculpture primarily on shell base and includes incised grooves, threads or ridges that strengthen anteriorly, strongest on anterior fasciole; remainder of shell surface may have microscopic spiral lines, but usually is smooth; and</paragraph>
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<paragraph id="8B8936D2FAE5FF28CBAFBA26FB18FCD2" blockId="128.[151,1436,612,888]" pageId="128" pageNumber="129">5. Cup-handle-like varix behind anal sinus (within ¼-turn of outer lip edge, usually less) such that the base of the fully developed shell appears lopsided (skewed to the right) when the shell is viewed ventrally.</paragraph>
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<paragraph id="8B8936D2FAE5FF28CBFFBAB3FD6BFBFA" blockId="128.[151,1437,935,1861]" pageId="128" pageNumber="129">
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<emphasis id="B942EAC0FAE5FF28CBFFBAB3FE22FC15" bold="true" box="[199,458,936,959]" pageId="128" pageNumber="129">Nomenclatural notes.</emphasis>
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<taxonomicName id="4C364D51FAE5FF28CAECBABCFDBCFC6A" box="[468,596,935,960]" class="Gastropoda" family="Drilliidae" genus="Douglassia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="128" pageNumber="129" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FAE5FF28CAECBABCFDBCFC6A" box="[468,596,935,960]" italics="true" pageId="128" pageNumber="129">Douglassia</emphasis>
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</taxonomicName>
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is difficult to distinguish from
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<taxonomicName id="4C364D51FAE5FF28C8FFBABCFA81FC6A" authority="Bartsch & Rehder, 1939" authorityName="Bartsch & Rehder" authorityYear="1939" box="[967,1385,935,960]" class="Gastropoda" family="Drilliidae" genus="Cerodrillia" kingdom="Animalia" order="Neogastropoda" pageId="128" pageNumber="129" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FAE5FF28C8FFBABCFBACFC6A" box="[967,1092,935,960]" italics="true" pageId="128" pageNumber="129">Cerodrillia</emphasis>
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Bartsch & Rehder, 1939
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</taxonomicName>
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and these two genera may prove to be congeneric. Some members of both genera have axials that are greatly reduced on the shoulder or, in later whorls of older individual especially, obsolete. The two genera differ principally in the number of protoconch whorls (2½–3 whorls in
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<taxonomicName id="4C364D51FAE5FF28C98EBD0FFCDEFB87" box="[694,822,1044,1069]" class="Gastropoda" family="Drilliidae" genus="Douglassia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="128" pageNumber="129" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FAE5FF28C98EBD0FFCDEFB87" box="[694,822,1044,1069]" italics="true" pageId="128" pageNumber="129">Douglassia</emphasis>
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</taxonomicName>
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versus 1¾–
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<quantity id="4CCE9B37FAE5FF28C8F8BD0FFC05FB87" box="[960,1005,1044,1069]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="128" pageNumber="129" unit="in" value="2.0">2 in</quantity>
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<taxonomicName id="4C364D51FAE5FF28C8CEBD0FFB9BFB87" box="[1014,1139,1044,1069]" class="Gastropoda" family="Drilliidae" genus="Cerodrillia" kingdom="Animalia" order="Neogastropoda" pageId="128" pageNumber="129" phylum="Mollusca" rank="genus">
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||
<emphasis id="B942EAC0FAE5FF28C8CEBD0FFB9BFB87" box="[1014,1139,1044,1069]" italics="true" pageId="128" pageNumber="129">Cerodrillia</emphasis>
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</taxonomicName>
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), and
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<taxonomicName id="4C364D51FAE5FF28CF85BD0FFAD5FB87" box="[1213,1341,1044,1069]" class="Gastropoda" family="Drilliidae" genus="Douglassia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="128" pageNumber="129" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FAE5FF28CF85BD0FFAD5FB87" box="[1213,1341,1044,1069]" italics="true" pageId="128" pageNumber="129">Douglassia</emphasis>
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</taxonomicName>
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usually, but not always, has more angular shoulders.
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</paragraph>
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<paragraph id="8B8936D2FAE5FF28CBFFBD47FB66FA82" blockId="128.[151,1437,935,1861]" pageId="128" pageNumber="129">
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Gibson (1962: 235) suggests that
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<taxonomicName id="4C364D51FAE5FF28C906BD47FD56FBDF" box="[574,702,1116,1141]" class="Gastropoda" family="Drilliidae" genus="Douglassia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="128" pageNumber="129" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FAE5FF28C906BD47FD56FBDF" box="[574,702,1116,1141]" italics="true" pageId="128" pageNumber="129">Douglassia</emphasis>
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</taxonomicName>
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should be synonymized with
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<taxonomicName id="4C364D51FAE5FF28CF32BD47FB66FBDF" box="[1034,1166,1116,1141]" class="Gastropoda" family="Drilliidae" genus="Splendrillia" kingdom="Animalia" order="Neogastropoda" pageId="128" pageNumber="129" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FAE5FF28CF32BD47FB66FBDF" box="[1034,1166,1116,1141]" italics="true" pageId="128" pageNumber="129">Splendrillia</emphasis>
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</taxonomicName>
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, presumably because of the absence of ribs in the sulcus and smooth shell, but the two genera have different varices—that of
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<taxonomicName id="4C364D51FAE5FF28CE20BD64FA75FB32" box="[1304,1437,1151,1176]" class="Gastropoda" family="Drilliidae" genus="Splendrillia" kingdom="Animalia" order="Neogastropoda" pageId="128" pageNumber="129" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FAE5FF28CE20BD64FA75FB32" box="[1304,1437,1151,1176]" italics="true" pageId="128" pageNumber="129">Splendrillia</emphasis>
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</taxonomicName>
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being hump-like about ⅓–turn from the edge of the outer lip. The varix of
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<taxonomicName id="4C364D51FAE5FF28CF37BDBFFB67FB17" box="[1039,1167,1188,1213]" class="Gastropoda" family="Drilliidae" genus="Douglassia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="128" pageNumber="129" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FAE5FF28CF37BDBFFB67FB17" box="[1039,1167,1188,1213]" italics="true" pageId="128" pageNumber="129">Douglassia</emphasis>
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</taxonomicName>
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(and of
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<taxonomicName id="4C364D51FAE5FF28CFC0BDBFFA9DFB17" box="[1272,1397,1188,1213]" class="Gastropoda" family="Drilliidae" genus="Cerodrillia" kingdom="Animalia" order="Neogastropoda" pageId="128" pageNumber="129" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FAE5FF28CFC0BDBFFA9DFB17" box="[1272,1397,1188,1213]" italics="true" pageId="128" pageNumber="129">Cerodrillia</emphasis>
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</taxonomicName>
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) is immediately behind the anal sinus (about ¼-turn or less) and cup handle-like, not hump-like. The
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<typeStatus id="548D8870FAE5FF28CFEFBDD3FAEFFB75" box="[1239,1287,1224,1247]" pageId="128" pageNumber="129">type</typeStatus>
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of the genus,
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<taxonomicName id="4C364D51FAE5FF28CBAFBDF6FF07FAAE" box="[151,239,1261,1284]" class="Gastropoda" family="Drilliidae" genus="Douglassia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="128" pageNumber="129" phylum="Mollusca" rank="species" species="enae">
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<emphasis id="B942EAC0FAE5FF28CBAFBDF6FF07FAAE" box="[151,239,1261,1284]" italics="true" pageId="128" pageNumber="129">D. enae</emphasis>
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</taxonomicName>
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, is morphologically closer to
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<taxonomicName id="4C364D51FAE5FF28C978BDF7FD55FAAF" box="[576,701,1260,1285]" class="Gastropoda" family="Drilliidae" genus="Cerodrillia" kingdom="Animalia" order="Neogastropoda" pageId="128" pageNumber="129" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FAE5FF28C978BDF7FD55FAAF" box="[576,701,1260,1285]" italics="true" pageId="128" pageNumber="129">Cerodrillia</emphasis>
|
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</taxonomicName>
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than to
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<taxonomicName id="4C364D51FAE5FF28C821BDF7FC76FAAF" box="[793,926,1260,1285]" class="Gastropoda" family="Drilliidae" genus="Splendrillia" kingdom="Animalia" order="Neogastropoda" pageId="128" pageNumber="129" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FAE5FF28C821BDF7FC76FAAF" box="[793,926,1260,1285]" italics="true" pageId="128" pageNumber="129">Splendrillia</emphasis>
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</taxonomicName>
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on the basis of the structure of its anal sinus, shell surface microsculpture, and cup-handle-like varix immediately behind the anal sinus.
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</paragraph>
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<paragraph id="8B8936D2FAE5FF28CBFFBC2FFEAEF972" blockId="128.[151,1437,935,1861]" pageId="128" pageNumber="129">
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<emphasis id="B942EAC0FAE5FF28CBFFBC2FFE6CFAE6" bold="true" box="[199,388,1332,1357]" pageId="128" pageNumber="129">Similar genera.</emphasis>
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<taxonomicName id="4C364D51FAE5FF28CAA8BC2FFCD1FAE6" authority="Bartsch & Rehder, 1939" authorityName="Bartsch & Rehder" authorityYear="1939" box="[400,825,1332,1357]" class="Gastropoda" family="Drilliidae" genus="Cerodrillia" kingdom="Animalia" order="Neogastropoda" pageId="128" pageNumber="129" phylum="Mollusca" rank="genus">
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||
<emphasis id="B942EAC0FAE5FF28CAA8BC2FFDE5FAE7" box="[400,525,1332,1357]" italics="true" pageId="128" pageNumber="129">Cerodrillia</emphasis>
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Bartsch & Rehder, 1939
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</taxonomicName>
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is similar and some specimens may be difficult to separate from
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<taxonomicName id="4C364D51FAE5FF28CA07BC4CFE57FADA" box="[319,447,1367,1392]" class="Gastropoda" family="Drilliidae" genus="Douglassia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="128" pageNumber="129" phylum="Mollusca" rank="genus">
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<emphasis id="B942EAC0FAE5FF28CA07BC4CFE57FADA" box="[319,447,1367,1392]" italics="true" pageId="128" pageNumber="129">Douglassia</emphasis>
|
||
</taxonomicName>
|
||
. Differences are discussed under nomenclatural notes above. Some members of two other genera,
|
||
<taxonomicName id="4C364D51FAE5FF28CA0CBC67FDA2FA3E" authority="Bartsch, 1943" authorityName="Bartsch" authorityYear="1943" box="[308,586,1404,1429]" class="Gastropoda" family="Pseudomelatomidae" genus="Viridrillia" kingdom="Animalia" order="Neogastropoda" pageId="128" pageNumber="129" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FAE5FF28CA0CBC67FE4CFA3F" box="[308,420,1404,1429]" italics="true" pageId="128" pageNumber="129">Viridrillia</emphasis>
|
||
Bartsch, 1943
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C364D51FAE5FF28C9BCBC67FC7EFA3E" authority="Bartsch, 1943" authorityName="Bartsch" authorityYear="1943" box="[644,918,1404,1429]" class="Gastropoda" family="Horaiclavidae" genus="Inodrillia" kingdom="Animalia" order="Neogastropoda" pageId="128" pageNumber="129" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FAE5FF28C9BCBC67FD18FA3F" box="[644,752,1404,1429]" italics="true" pageId="128" pageNumber="129">Inodrillia</emphasis>
|
||
Bartsch, 1943
|
||
</taxonomicName>
|
||
have an uncanny resemblance to
|
||
<taxonomicName id="4C364D51FAE5FF28CE24BC67FA73FA3F" box="[1308,1435,1404,1429]" class="Gastropoda" family="Drilliidae" genus="Douglassia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="128" pageNumber="129" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FAE5FF28CE24BC67FA73FA3F" box="[1308,1435,1404,1429]" italics="true" pageId="128" pageNumber="129">Douglassia</emphasis>
|
||
</taxonomicName>
|
||
(and to
|
||
<taxonomicName id="4C364D51FAE5FF28CBC0BC84FE9DFA12" box="[248,373,1439,1464]" class="Gastropoda" family="Drilliidae" genus="Cerodrillia" kingdom="Animalia" order="Neogastropoda" pageId="128" pageNumber="129" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FAE5FF28CBC0BC84FE9DFA12" box="[248,373,1439,1464]" italics="true" pageId="128" pageNumber="129">Cerodrillia</emphasis>
|
||
</taxonomicName>
|
||
). Tippett (1995: 134–135) illustrates the radula of
|
||
<taxonomicName id="4C364D51FAE5FF28C8E7BC84FA8EFA12" authority="Bartsch, 1943" authorityName="Bartsch" authorityYear="1943" box="[991,1382,1439,1464]" class="Gastropoda" family="Pseudomelatomidae" genus="Viridrillia" kingdom="Animalia" order="Neogastropoda" pageId="128" pageNumber="129" phylum="Mollusca" rank="species" species="williami">
|
||
<emphasis id="B942EAC0FAE5FF28C8E7BC84FB50FA12" box="[991,1208,1439,1464]" italics="true" pageId="128" pageNumber="129">Viridrillia williami</emphasis>
|
||
Bartsch, 1943
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C364D51FAE5FF28CBAFBCDFFDC4FA76" authority="Bartsch, 1943" authorityName="Bartsch" authorityYear="1943" box="[151,556,1476,1501]" class="Gastropoda" family="Pseudomelatomidae" genus="Viridrillia" kingdom="Animalia" order="Neogastropoda" pageId="128" pageNumber="129" phylum="Mollusca" rank="species" species="hendersoni">
|
||
<emphasis id="B942EAC0FAE5FF28CBAFBCDFFE62FA77" box="[151,394,1476,1501]" italics="true" pageId="128" pageNumber="129">Viridrillia hendersoni</emphasis>
|
||
Bartsch, 1943
|
||
</taxonomicName>
|
||
to demonstrate the more appropriate placement of the genus in
|
||
<taxonomicName id="4C364D51FAE5FF28CFCFBCDFFA70FA77" box="[1271,1432,1476,1501]" class="Gastropoda" family="Turridae" kingdom="Animalia" order="Neogastropoda" pageId="128" pageNumber="129" phylum="Mollusca" rank="subFamily" subFamily="Crassispirinae">Crassispirinae</taxonomicName>
|
||
, now
|
||
<taxonomicName id="4C364D51FAE5FF28CBF6BCFCFC8EF9AA" authority="Morrison, 1965 (Bouchet et al., 2011)" authorityName="Morrison, 1965 (Bouchet et al." authorityYear="2011" box="[206,870,1511,1536]" class="Gastropoda" family="Pseudomelatomidae" kingdom="Animalia" order="Neogastropoda" pageId="128" pageNumber="129" phylum="Mollusca" rank="family">
|
||
Pseudomelatomidae Morrison, 1965 (Bouchet
|
||
<emphasis id="B942EAC0FAE5FF28C9E3BCF3FCFCF9AA" box="[731,788,1511,1536]" italics="true" pageId="128" pageNumber="129">et al.</emphasis>
|
||
, 2011)
|
||
</taxonomicName>
|
||
. Earlier, Sysoev, in Taylor
|
||
<emphasis id="B942EAC0FAE5FF28CFACBCF3FB26F9AA" box="[1172,1230,1511,1536]" italics="true" pageId="128" pageNumber="129">et al.</emphasis>
|
||
(1993: 164) listed
|
||
<taxonomicName id="4C364D51FAE5FF28CBAFBF17FEEBF98F" box="[151,259,1548,1573]" class="Gastropoda" family="Horaiclavidae" genus="Inodrillia" kingdom="Animalia" order="Neogastropoda" pageId="128" pageNumber="129" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FAE5FF28CBAFBF17FEEBF98F" box="[151,259,1548,1573]" italics="true" pageId="128" pageNumber="129">Inodrillia</emphasis>
|
||
</taxonomicName>
|
||
as a crassispirine genus on the basis of its radular teeth.
|
||
<taxonomicName id="4C364D51FAE5FF28C8B1BF17FC1DF98F" box="[905,1013,1548,1573]" class="Gastropoda" family="Horaiclavidae" genus="Inodrillia" kingdom="Animalia" order="Neogastropoda" pageId="128" pageNumber="129" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FAE5FF28C8B1BF17FC1DF98F" box="[905,1013,1548,1573]" italics="true" pageId="128" pageNumber="129">Inodrillia</emphasis>
|
||
</taxonomicName>
|
||
has most recently been provisionally placed in
|
||
<taxonomicName id="4C364D51FAE5FF28CA2ABF34FD23F9E2" authority="Bouchet et al., 2011" authorityName="Bouchet et al." authorityYear="2011" box="[274,715,1583,1608]" class="Gastropoda" family="Horaiclavidae" kingdom="Animalia" order="Neogastropoda" pageId="128" pageNumber="129" phylum="Mollusca" rank="family">
|
||
Horaiclavidae (Bouchet
|
||
<emphasis id="B942EAC0FAE5FF28C90ABF2BFD9AF9E2" box="[562,626,1583,1608]" italics="true" pageId="128" pageNumber="129">et al.</emphasis>
|
||
, 2011)
|
||
</taxonomicName>
|
||
. Although very similar,
|
||
<taxonomicName id="4C364D51FAE5FF28C8CEBF34FB8EF9E2" box="[1014,1126,1583,1608]" class="Gastropoda" family="Pseudomelatomidae" genus="Viridrillia" kingdom="Animalia" order="Neogastropoda" pageId="128" pageNumber="129" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FAE5FF28C8CEBF34FB8EF9E2" box="[1014,1126,1583,1608]" italics="true" pageId="128" pageNumber="129">Viridrillia</emphasis>
|
||
</taxonomicName>
|
||
has subtle morphological characteristics not seen in any of the known
|
||
<taxonomicName id="4C364D51FAE5FF28C992BF4FFCC2F9C7" box="[682,810,1620,1645]" class="Gastropoda" family="Drilliidae" genus="Douglassia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="128" pageNumber="129" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FAE5FF28C992BF4FFCC2F9C7" box="[682,810,1620,1645]" italics="true" pageId="128" pageNumber="129">Douglassia</emphasis>
|
||
</taxonomicName>
|
||
(or
|
||
<taxonomicName id="4C364D51FAE5FF28C858BF4FFC35F9C7" box="[864,989,1620,1645]" class="Gastropoda" family="Drilliidae" genus="Cerodrillia" kingdom="Animalia" order="Neogastropoda" pageId="128" pageNumber="129" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FAE5FF28C858BF4FFC35F9C7" box="[864,989,1620,1645]" italics="true" pageId="128" pageNumber="129">Cerodrillia</emphasis>
|
||
</taxonomicName>
|
||
): an anterior fasciole that lacks spiral ridges, the presence of spiral threads or grooves over the entire shell, not just on the shell base, and a protoconch with obsolete riblets that nonetheless leave a wrinkled look. The protoconch in
|
||
<taxonomicName id="4C364D51FAE5FF28CF7BBF87FB2BF91F" box="[1091,1219,1692,1717]" class="Gastropoda" family="Drilliidae" genus="Douglassia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="128" pageNumber="129" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FAE5FF28CF7BBF87FB2BF91F" box="[1091,1219,1692,1717]" italics="true" pageId="128" pageNumber="129">Douglassia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C364D51FAE5FF28CE39BF87FA96F91F" box="[1281,1406,1692,1717]" class="Gastropoda" family="Drilliidae" genus="Cerodrillia" kingdom="Animalia" order="Neogastropoda" pageId="128" pageNumber="129" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FAE5FF28CE39BF87FA96F91F" box="[1281,1406,1692,1717]" italics="true" pageId="128" pageNumber="129">Cerodrillia</emphasis>
|
||
</taxonomicName>
|
||
is always smooth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32C6559FAE5FF28CBFFBFFFFB6DF8EF" pageId="128" pageNumber="129" type="distribution">
|
||
<paragraph id="8B8936D2FAE5FF28CBFFBFFFFB6DF8EF" blockId="128.[151,1437,935,1861]" pageId="128" pageNumber="129">
|
||
<emphasis id="B942EAC0FAE5FF28CBFFBFFFFE88F957" bold="true" box="[199,352,1764,1789]" pageId="128" pageNumber="129">Distribution.</emphasis>
|
||
The genus
|
||
<taxonomicName id="4C364D51FAE5FF28CAC8BFFFFD98F957" box="[496,624,1764,1789]" class="Gastropoda" family="Drilliidae" genus="Douglassia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="128" pageNumber="129" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B942EAC0FAE5FF28CAC8BFFFFD98F957" box="[496,624,1764,1789]" italics="true" pageId="128" pageNumber="129">Douglassia</emphasis>
|
||
</taxonomicName>
|
||
is distributed throughout the TWA, from North Carolina to northern
|
||
<collectingCountry id="F3217642FAE5FF28CBAFBE1CFF37F88A" box="[151,223,1799,1824]" name="Brazil" pageId="128" pageNumber="129">Brazil</collectingCountry>
|
||
. Most species are from water depths of
|
||
<quantity id="4CCE9B37FAE5FF28C98EBE13FD1BF8B5" box="[694,755,1800,1824]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="128" pageNumber="129" unit="m" value="30.0">30 m</quantity>
|
||
or greater, an exception being
|
||
<taxonomicName id="4C364D51FAE5FF28CF59BE13FB12F88A" box="[1121,1274,1799,1824]" class="Gastropoda" family="Drilliidae" genus="Douglassia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="128" pageNumber="129" phylum="Mollusca" rank="species" species="antillensis" status="sp. nov.">
|
||
<emphasis id="B942EAC0FAE5FF28CF59BE13FB12F88A" box="[1121,1274,1799,1824]" italics="true" pageId="128" pageNumber="129">D. antillensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A27157BBFAE5FF28CE33BE13FA7FF88A" box="[1291,1431,1799,1824]" pageId="128" pageNumber="129" rank="species">new species</taxonomicNameLabel>
|
||
, which is distributed throughout the eastern Caribbean and usually found above this depth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |