234 lines
28 KiB
XML
234 lines
28 KiB
XML
<document id="53C5B69555BF4725D961CB96B97F3308" ID-CLB-Dataset="22144" ID-DOI="10.5852/ejt.2017.309" ID-GBIF-Dataset="4e7bcd4d-1cdb-45cf-b2c3-54da2466f171" ID-ISSN="2118-9773" ID-Zenodo-Dep="3827189" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589468946001" checkinUser="carolina" docAuthor="Herbert, David G." docDate="2017" docId="039F87A56758FFBC4D0BFCADFAACFD12" docLanguage="en" docName="ejt-2017-309.pdf.imf" docOrigin="European Journal of Taxonomy 309" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Selatodryas Herbert 2017, gen. nov." docType="treatment" docUuid="72AF5C89-20EB-44EE-B2F5-6390BD6E0183" docUuidSource="ZooBank" docVersion="6" lastPageNumber="33" masterDocId="FFA6FFDD6747FF9C4F79FFC5FFA2FFF9" masterDocTitle="A new genus and eight new species of tail-wagger snails from eastern South Africa, with a key to genera within Sheldonia s. l. (Gastropoda: Urocyclidae)" masterLastPageNumber="50" masterPageNumber="1" pageNumber="32" updateTime="1698839690494" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:mods id="5762C7EA115738265AEABEBFA084DE76" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="70374C0CBD5B2B3F906323399551FD2D">
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<mods:title id="F605284D5EB9713A41E8C1B4EB12C29F">A new genus and eight new species of tail-wagger snails from eastern South Africa, with a key to genera within Sheldonia s. l. (Gastropoda: Urocyclidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="680047B7CC8FDBAAEF41432F592724EF" type="personal">
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<mods:roleTerm id="6136C0F25DE6E39A7152CFE933AB619D">Author</mods:roleTerm>
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<mods:namePart id="2886B6F66D94B494A7487E879854CC1C">Herbert, David G.</mods:namePart>
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<mods:typeOfResource id="ABBED95130D396C9FFBBE481AE2A9E0C">text</mods:typeOfResource>
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<mods:title id="D1CD41B53E54E091A02F3483799E9381">European Journal of Taxonomy</mods:title>
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</mods:titleInfo>
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<mods:part id="3796763E018AB06A13B764C3E8B36D1A">
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<mods:date id="4E9EA2E4E6C23C6B8654CE15D6AFA4EE">2017</mods:date>
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<mods:detail id="0C4B37091F8F3E2951F7826626D95BF7" type="pubDate">
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<mods:number id="BE56AAADC29737F72F39FB3ED95FF66D">2017-04-10</mods:number>
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<mods:detail id="1623CBD99E6C47594EDCAEE31EF7F120" type="issue">
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<mods:number id="DC807899B460A153BC2AB557C9C98D0F">309</mods:number>
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<mods:classification id="EB38D687E604168737F61DCAB6750A07">journal article</mods:classification>
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<mods:identifier id="7EC2EB42827747D28575CE5A78364321" type="CLB-Dataset">22144</mods:identifier>
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<mods:identifier id="7C74C3CF989BD6DA5B1986005DBD8D5D" type="DOI">10.5852/ejt.2017.309</mods:identifier>
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<mods:identifier id="B49C60B57F970FAF131CA8B0B003FC46" type="ISSN">2118-9773</mods:identifier>
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<mods:identifier id="9CD792C822C62EAC67534F633AC234FD" type="Zenodo-Dep">3827189</mods:identifier>
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<treatment id="039F87A56758FFBC4D0BFCADFAACFD12" ID-DOI="http://doi.org/10.5281/zenodo.3846858" ID-GBIF-Taxon="164151029" ID-Zenodo-Dep="3846858" LSID="urn:lsid:zoobank.org:act:72AF5C89-20EB-44EE-B2F5-6390BD6E0183" httpUri="http://treatment.plazi.org/id/039F87A56758FFBC4D0BFCADFAACFD12" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
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<subSubSection id="C32C65386758FF834D0BFCADFC60FC7B" box="[626,962,871,898]" pageId="31" pageNumber="32" type="nomenclature">
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<paragraph id="8B8936B36758FF834D0BFCADFC60FC7B" blockId="31.[368,1219,871,934]" box="[626,962,871,898]" pageId="31" pageNumber="32">
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<heading id="D0C181DF6758FF834D0BFCADFC60FC7B" box="[626,962,871,898]" centered="true" fontSize="11" level="2" pageId="31" pageNumber="32" reason="2">
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Genus
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<taxonomicName id="4C364D306758FF834DBDFCA2FCF2FC7B" ID-CoL="7PGHK" authority="Herbert, 2017" authorityName="Herbert" authorityYear="2017" box="[708,848,871,898]" class="Gastropoda" family="Urocyclidae" genus="Selatodryas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="genus" status="gen. nov.">
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<emphasis id="B942EAA16758FF834DBDFCA2FCF2FC7B" bold="true" box="[708,848,871,898]" italics="true" pageId="31" pageNumber="32">Selatodryas</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27157DA6758FF834C2EFCADFC60FC7B" box="[855,962,872,898]" pageId="31" pageNumber="32" rank="genus">gen. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C65386758FF834E09FC4EFB61FC5F" box="[368,1219,907,934]" pageId="31" pageNumber="32" type="description">
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<paragraph id="8B8936B36758FF834E09FC4EFB61FC5F" blockId="31.[368,1219,871,934]" box="[368,1219,907,934]" pageId="31" pageNumber="32">
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<uri id="FFA73AB16758FF834E09FC4EFB61FC5F" box="[368,1219,907,934]" pageId="31" pageNumber="32">
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urn:lsid:zoobank.org:act:
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<uuid id="FF900C666758FF834DF7FC4EFB61FC5F" box="[654,1219,907,934]" pageId="31" pageNumber="32">72AF5C89-20EB-44EE-B2F5-6390BD6E0183</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C65386758FF834FC4FC0BFD26FBEE" pageId="31" pageNumber="32" type="type_taxon">
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<paragraph id="8B8936B36758FF834FC4FC0BFEFAFC1E" blockId="31.[189,344,973,1000]" box="[189,344,973,1000]" pageId="31" pageNumber="32">
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<heading id="D0C181DF6758FF834FC4FC0BFEFAFC1E" bold="true" box="[189,344,973,1000]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
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<emphasis id="B942EAA16758FF834FC4FC0BFEFAFC1E" bold="true" box="[189,344,973,1000]" pageId="31" pageNumber="32">
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<typeStatus id="548D88116758FF834FC4FC0BFF58FC11" box="[189,250,974,1000]" pageId="31" pageNumber="32">Type</typeStatus>
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species
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8936B36758FF834FC4FC38FD26FBEE" blockId="31.[189,644,1021,1047]" box="[189,644,1021,1047]" pageId="31" pageNumber="32">
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<taxonomicName id="4C364D306758FF834FC4FC38FE6FFBEE" ID-CoL="4WD6Y" authority="Herbert, 2017" authorityName="Herbert" authorityYear="2017" box="[189,461,1021,1047]" class="Gastropoda" family="Urocyclidae" genus="Selatodryas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="roseosoma" status="gen. et sp. nov.">
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<emphasis id="B942EAA16758FF834FC4FC38FE6FFBEE" box="[189,461,1021,1047]" italics="true" pageId="31" pageNumber="32">Selatodryas roseosoma</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27157DA6758FF834EADFC38FD26FBEE" box="[468,644,1021,1047]" pageId="31" pageNumber="32" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C65386758FF834FC4FBFAFD45FAA6" pageId="31" pageNumber="32" type="diagnosis">
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<paragraph id="8B8936B36758FF834FC4FBFAFE94FBA0" blockId="31.[189,310,1087,1113]" box="[189,310,1087,1113]" pageId="31" pageNumber="32">
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<heading id="D0C181DF6758FF834FC4FBFAFE94FBA0" bold="true" box="[189,310,1087,1113]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
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<emphasis id="B942EAA16758FF834FC4FBFAFE94FBA0" bold="true" box="[189,310,1087,1113]" pageId="31" pageNumber="32">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8936B36758FF834FC4FBABFD45FAA6" blockId="31.[189,1399,1134,1375]" pageId="31" pageNumber="32">Shell small, delicate and glossy, periphery at or just below mid-whorl; umbilicus absent, edge of columella lip reflected and slightly thickened with a thin, whitish callus; columella running into axis of coiling rather than fusing with the parietal region; protoconch essentially smooth. Optic tentacles conspicuously dark. Penis long, coiled inside sheath; epiphallus long; caecum elongate, situated just proximal to mid-point of epiphallus, well separated from insertion of penial retractor muscle; penial verge absent; vagina long. Terminal branches of spermatophore spines bluntly bifid; distal portion of spermatophore tail smooth and relatively short.</paragraph>
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</subSubSection>
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<subSubSection id="C32C65386758FF834FC4FA4DFC7FFA0D" pageId="31" pageNumber="32" type="etymology">
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<paragraph id="8B8936B36758FF834FC4FA4DFEE6FA5B" blockId="31.[189,324,1416,1442]" box="[189,324,1416,1442]" pageId="31" pageNumber="32">
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<heading id="D0C181DF6758FF834FC4FA4DFEE6FA5B" bold="true" box="[189,324,1416,1442]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
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<emphasis id="B942EAA16758FF834FC4FA4DFEE6FA5B" bold="true" box="[189,324,1416,1442]" pageId="31" pageNumber="32">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8936B36758FF834FC4FA72FC7FFA0D" blockId="31.[189,1399,1461,1524]" pageId="31" pageNumber="32">From the Greek ‘σελάτης’, m., a snail, and ‘Δρυάς’, f., a Dryad or forest nymph; in reference to the delicate shell and preferred habitat of these snails. Gender feminine.</paragraph>
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</subSubSection>
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<subSubSection id="C32C65386758FFBC4FC4F9D8FAACFD12" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32" type="discussion">
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<paragraph id="8B8936B36758FF834FC4F9D8FE8DF9CE" blockId="31.[189,303,1565,1591]" box="[189,303,1565,1591]" pageId="31" pageNumber="32">
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<heading id="D0C181DF6758FF834FC4F9D8FE8DF9CE" bold="true" box="[189,303,1565,1591]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
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<emphasis id="B942EAA16758FF834FC4F9D8FE8DF9CE" bold="true" box="[189,303,1565,1591]" pageId="31" pageNumber="32">Remarks</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8936B36758FF834FC4F989FD95F8C4" blockId="31.[189,1399,1612,1854]" pageId="31" pageNumber="32">
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The morphology of the penis and epiphallus, and the bluntly bifid branch tips of the spermatophore spines of
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<taxonomicName id="4C364D306758FF834E48F9B5FE1EF973" authority="Herbert, 2017" authorityName="Herbert" authorityYear="2017" box="[305,444,1648,1674]" class="Gastropoda" family="Urocyclidae" genus="Selatodryas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="genus" status="gen. nov.">
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<emphasis id="B942EAA16758FF834E48F9B5FE1EF973" box="[305,444,1648,1674]" italics="true" pageId="31" pageNumber="32">Selatodryas</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27157DA6758FF834EBDF9B5FD93F973" box="[452,561,1648,1674]" pageId="31" pageNumber="32" rank="genus">gen. nov.</taxonomicNameLabel>
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suggest a close relationship with
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<emphasis id="B942EAA16758FF834CBEF9B5FBC9F970" box="[967,1131,1648,1674]" italics="true" pageId="31" pageNumber="32">
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<taxonomicName id="4C364D306758FF834CBEF9B5FB9EF973" authorityName="Ancey" authorityYear="1887" box="[967,1084,1648,1674]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="genus" sensu="stricto">Sheldonia</taxonomicName>
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<taxonomicNameLabel id="A27157DA6758FF834B3CF9B5FBC9F970" box="[1093,1131,1648,1673]" pageId="31" pageNumber="32" sensu="stricto">s.s.</taxonomicNameLabel>
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</emphasis>
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Similarly, the form of the columella region (complete absence of an umbilicus) is a character shared with some species of
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<taxonomicName id="4C364D306758FF834FC4F972FE90F928" authorityName="Ancey" authorityYear="1887" box="[189,306,1719,1745]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="genus" sensu="stricto">
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<emphasis id="B942EAA16758FF834FC4F972FE90F928" box="[189,306,1719,1745]" italics="true" pageId="31" pageNumber="32">Sheldonia</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27157DA6758FF834E42F97DFEC2F92B" box="[315,352,1720,1746]" pageId="31" pageNumber="32" sensu="stricto">
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<emphasis id="B942EAA16758FF834E42F97DFEFBF928" box="[315,345,1720,1745]" italics="true" pageId="31" pageNumber="32">s.s</emphasis>
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.
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</taxonomicNameLabel>
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(such as
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<taxonomicName id="4C364D306758FF834EADF97DFC92F928" authority="(Benson, 1848)" baseAuthorityName="Benson" baseAuthorityYear="1848" box="[468,816,1719,1746]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="trotteriana">
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<emphasis id="B942EAA16758FF834EADF97DFDD3F928" box="[468,625,1720,1746]" italics="true" pageId="31" pageNumber="32">S. trotteriana</emphasis>
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(Benson, 1848)
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</taxonomicName>
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,
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<taxonomicName id="4C364D306758FF834C39F97DFB37F92B" authority="(Pfeiffer, 1846)" baseAuthorityName="Pfeiffer" baseAuthorityYear="1846" box="[832,1173,1719,1746]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="natalensis">
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<emphasis id="B942EAA16758FF834C39F97DFC74F928" box="[832,982,1719,1746]" italics="true" pageId="31" pageNumber="32">S. natalensis</emphasis>
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(Pfeiffer, 1846)
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</taxonomicName>
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and
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<taxonomicName id="4C364D306758FF834BABF97DFE22F90F" authority="(Benson, 1850))" authorityName="" baseAuthorityName="Benson" baseAuthorityYear="1850" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="cotyledonis">
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<emphasis id="B942EAA16758FF834BABF97DFAD5F928" box="[1234,1399,1719,1746]" italics="true" pageId="31" pageNumber="32">S. cotyledonis</emphasis>
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(Benson, 1850))
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</taxonomicName>
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. However, two characters of
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<taxonomicName id="4C364D306758FF834DA1F91EFCC1F90C" authority="Herbert, 2017" authorityName="Herbert" authorityYear="2017" box="[728,867,1755,1781]" class="Gastropoda" family="Urocyclidae" genus="Selatodryas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="genus" status="gen. nov.">
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<emphasis id="B942EAA16758FF834DA1F91EFCC1F90C" box="[728,867,1755,1781]" italics="true" pageId="31" pageNumber="32">Selatodryas</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27157DA6758FF834C13F919FC74F90F" box="[874,982,1756,1782]" pageId="31" pageNumber="32" rank="genus">gen. nov.</taxonomicNameLabel>
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set the genus apart from
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<emphasis id="B942EAA16758FF834A78F91EFF41F8E0" italics="true" pageId="31" pageNumber="32">
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<taxonomicName id="4C364D306758FF834A78F91EFAD4F90C" authorityName="Ancey" authorityYear="1887" box="[1281,1398,1755,1781]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="genus" sensu="stricto">Sheldonia</taxonomicName>
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<taxonomicNameLabel id="A27157DA6758FF834FC4F8C5FF41F8E0" box="[189,227,1792,1817]" pageId="31" pageNumber="32" sensu="stricto">s.s.</taxonomicNameLabel>
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</emphasis>
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, namely the smooth protoconch and the short distal portion of the spermatophore tail that lacks a double row of blunt projections.
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</paragraph>
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<paragraph id="8B8936B36758FFBC4FC4F8A0FC3FFE69" blockId="31.[189,1399,1893,2027]" lastBlockId="32.[189,1398,267,400]" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
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Preliminary mtDNA data confirm this close relationship, placing the two species of
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<taxonomicName id="4C364D306758FF834B95F8A0FAD5F886" authority="Herbert, 2017" authorityName="Herbert" authorityYear="2017" box="[1260,1399,1893,1919]" class="Gastropoda" family="Urocyclidae" genus="Selatodryas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="genus" status="gen. nov.">
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<emphasis id="B942EAA16758FF834B95F8A0FAD5F886" box="[1260,1399,1893,1919]" italics="true" pageId="31" pageNumber="32">Selatodryas</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27157DA6758FF834FC4F84CFE8EF85A" box="[189,300,1929,1955]" pageId="31" pageNumber="32" rank="genus">gen. nov.</taxonomicNameLabel>
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in a well-supported clade sister to, but distinct from
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<taxonomicName id="4C364D306758FF834CB9F84CFB97F85A" authorityName="Ancey" authorityYear="1887" box="[960,1077,1929,1955]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="genus" sensu="stricto">
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<emphasis id="B942EAA16758FF834CB9F84CFB97F85A" box="[960,1077,1929,1955]" italics="true" pageId="31" pageNumber="32">Sheldonia</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27157DA6758FF834B38F84FFBC4F85A" box="[1089,1126,1929,1955]" pageId="31" pageNumber="32" sensu="stricto">
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<emphasis id="B942EAA16758FF834B38F84FFBFDF85A" box="[1089,1119,1930,1955]" italics="true" pageId="31" pageNumber="32">s.s</emphasis>
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.
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</taxonomicNameLabel>
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(
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<bibRefCitation id="EFA74B426758FF834B00F84CFF5FF83E" author="Herbert D. G. & Moussalli A." pageId="31" pageNumber="32" pagination="20 - 21" refId="ref24016" refString="Herbert D. G. & Moussalli A. 2009. Tail-wagger snails: molecules and morphology - systematics and biogeography (Pulmonata: Urocyclidae). Tenth Anniversary Conference of the Southern African Society for Systematic Biology, Illovo Beach, KwaZulu-Natal, 25 - 27 July, 2009: 20 - 21." type="proceedings paper" year="2009">Herbert & Moussalli 2009</bibRefCitation>
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).
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<taxonomicName id="4C364D306758FF834E6CF868FE02F83E" authority="Herbert, 2017" authorityName="Herbert" authorityYear="2017" box="[277,416,1965,1991]" class="Gastropoda" family="Urocyclidae" genus="Selatodryas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="genus" status="gen. nov.">
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<emphasis id="B942EAA16758FF834E6CF868FE02F83E" box="[277,416,1965,1991]" italics="true" pageId="31" pageNumber="32">Selatodryas</emphasis>
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</taxonomicName>
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|
<taxonomicNameLabel id="A27157DA6758FF834ED5F868FDBEF83E" box="[428,540,1965,1991]" pageId="31" pageNumber="32" rank="genus">gen. nov.</taxonomicNameLabel>
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is known only from high altitude forest (Southern Mistbelt). From a biogeographical perspective the ranges of the two genera do not overlap.
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<emphasis id="B942EAA16758FF834B45F814FB43F812" box="[1084,1249,2001,2027]" italics="true" pageId="31" pageNumber="32">
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|
<taxonomicName id="4C364D306758FF834B45F814FB13F812" authorityName="Ancey" authorityYear="1887" box="[1084,1201,2001,2027]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="genus" sensu="stricto">Sheldonia</taxonomicName>
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<taxonomicNameLabel id="A27157DA6758FF834BC2F817FB43F812" box="[1211,1249,2002,2027]" pageId="31" pageNumber="32" sensu="stricto">s.s.</taxonomicNameLabel>
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</emphasis>
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occurs over a large part of the Eastern and
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<collectingRegion id="49F2F8516767FFBC4D49FEC9FD7AFEDC" box="[560,728,267,294]" country="South Africa" name="Western Cape" pageId="32" pageNumber="33">Western Cape</collectingRegion>
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, from the Mbhashe River valley west to St Helena Bay, whereas
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<taxonomicName id="4C364D306767FFBC4E5CFEEAFE0DFEB0" authority="Herbert, 2017" authorityName="Herbert" authorityYear="2017" box="[293,431,303,329]" class="Gastropoda" family="Urocyclidae" genus="Selatodryas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="32" pageNumber="33" phylum="Mollusca" rank="genus" status="gen. nov.">
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<emphasis id="B942EAA16767FFBC4E5CFEEAFE0DFEB0" box="[293,431,303,329]" italics="true" pageId="32" pageNumber="33">Selatodryas</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27157DA6767FFBC4ECEFEEAFD80FEB0" box="[439,546,303,329]" pageId="32" pageNumber="33" rank="genus">gen. nov.</taxonomicNameLabel>
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|
occurs as an almost parapatric isolate to the north-east of
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|
<emphasis id="B942EAA16767FFBC4BB4FEEAFAD2FEB1" box="[1229,1392,303,329]" italics="true" pageId="32" pageNumber="33">
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|
<taxonomicName id="4C364D306767FFBC4BB4FEEAFAE0FEB0" authorityName="Ancey" authorityYear="1887" box="[1229,1346,303,329]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="32" pageNumber="33" phylum="Mollusca" rank="genus" sensu="stricto">Sheldonia</taxonomicName>
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<taxonomicNameLabel id="A27157DA6767FFBC4A33FEEAFAD2FEB1" box="[1354,1392,303,328]" pageId="32" pageNumber="33" sensu="stricto">s.s.</taxonomicNameLabel>
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</emphasis>
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, suggesting that
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<taxonomicName id="4C364D306767FFBC4E0CFE97FDA2FE95" authority="Herbert, 2017" authorityName="Herbert" authorityYear="2017" box="[373,512,338,364]" class="Gastropoda" family="Urocyclidae" genus="Selatodryas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="32" pageNumber="33" phylum="Mollusca" rank="genus" status="gen. nov.">
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|
<emphasis id="B942EAA16767FFBC4E0CFE97FDA2FE95" box="[373,512,338,364]" italics="true" pageId="32" pageNumber="33">Selatodryas</emphasis>
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|
</taxonomicName>
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|
<taxonomicNameLabel id="A27157DA6767FFBC4D7EFE97FDD0FE95" box="[519,626,338,364]" pageId="32" pageNumber="33" rank="genus">gen. nov.</taxonomicNameLabel>
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|
diverged from the ancestral
|
|
<taxonomicName id="4C364D306767FFBC4CB9FE97FB97FE95" authorityName="Ancey" authorityYear="1887" box="[960,1077,338,364]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="32" pageNumber="33" phylum="Mollusca" rank="genus">
|
|
<emphasis id="B942EAA16767FFBC4CB9FE97FB97FE95" box="[960,1077,338,364]" italics="true" pageId="32" pageNumber="33">Sheldonia</emphasis>
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|
</taxonomicName>
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|
stock before
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|
<taxonomicName id="4C364D306767FFBC4BADFE97FAEBFE95" authorityName="Ancey" authorityYear="1887" box="[1236,1353,338,364]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="32" pageNumber="33" phylum="Mollusca" rank="genus" sensu="stricto">
|
|
<emphasis id="B942EAA16767FFBC4BADFE97FAEBFE95" box="[1236,1353,338,364]" italics="true" pageId="32" pageNumber="33">Sheldonia</emphasis>
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|
</taxonomicName>
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|
<taxonomicNameLabel id="A27157DA6767FFBC4A28FE96FAD4FE95" box="[1361,1398,338,364]" pageId="32" pageNumber="33" sensu="stricto">
|
|
<emphasis id="B942EAA16767FFBC4A28FE96FACDFE95" box="[1361,1391,339,364]" italics="true" pageId="32" pageNumber="33">s.s</emphasis>
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|
.
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|
</taxonomicNameLabel>
|
|
radiated into the drier regions of the southern
|
|
<collectingRegion id="49F2F8516767FFBC4DAEFEB0FCB1FE76" box="[727,787,373,399]" country="South Africa" name="Western Cape" pageId="32" pageNumber="33">Cape</collectingRegion>
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and Karoo.
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</paragraph>
|
|
<paragraph id="8B8936B36767FFBC4FC4FE72FAACFD12" blockId="32.[189,1399,439,748]" pageId="32" pageNumber="33">
|
|
The two species of
|
|
<taxonomicName id="4C364D306767FFBC4EDDFE72FD8DFE28" authority="Herbert, 2017" authorityName="Herbert" authorityYear="2017" box="[420,559,439,465]" class="Gastropoda" family="Urocyclidae" genus="Selatodryas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="32" pageNumber="33" phylum="Mollusca" rank="genus" status="gen. nov.">
|
|
<emphasis id="B942EAA16767FFBC4EDDFE72FD8DFE28" box="[420,559,439,465]" italics="true" pageId="32" pageNumber="33">Selatodryas</emphasis>
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|
</taxonomicName>
|
|
<taxonomicNameLabel id="A27157DA6767FFBC4D4EFE72FD01FE28" box="[567,675,439,465]" pageId="32" pageNumber="33" rank="genus">gen. nov.</taxonomicNameLabel>
|
|
described below differ conspicuously in body coloration and this is reflected in their respective species names. However, the taxonomic value of body colour as a diagnostic character in southern African urocyclid snails must be viewed with caution. A number of species show considerable intra-specific variation in this regard, including
|
|
<taxonomicName id="4C364D306767FFBC4B11FDE4FAD4FDC2" authority="Herbert, 2017" authorityName="Herbert" authorityYear="2017" box="[1128,1398,545,571]" class="Gastropoda" family="Urocyclidae" genus="Kerkophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="terrestris" status="sp. nov.">
|
|
<emphasis id="B942EAA16767FFBC4B11FDE4FAD4FDC2" box="[1128,1398,545,571]" italics="true" pageId="32" pageNumber="33">Kerkophorus terrestris</emphasis>
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|
</taxonomicName>
|
|
<taxonomicNameLabel id="A27157DA6767FFBC4FC4FD81FEB8FDA7" box="[189,282,580,606]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
|
|
described above (
|
|
<figureCitation id="130D2A366767FFBC4E8FFD81FDEDFDA7" box="[502,591,580,606]" captionStart="Fig" captionStartId="24.[189,232,1930,1956]" captionTargetBox="[191,1398,1486,1888]" captionTargetId="figure@24.[189,1399,1485,1889]" captionTargetPageId="24" captionText="Fig. 17. Kerkophorus terrestris sp. nov., living animals. A. Collywobbles, E Cape, shell diameter 13.1 mm (holotype, NMSA W3979/T3862). B. Mount Thesiger, Port St Johns, E Cape, shell diameter 10.7 mm (paratype, NMSA V6515/T3859)." figureDoi="http://doi.org/10.5281/zenodo.3827227" httpUri="https://zenodo.org/record/3827227/files/figure.png" pageId="32" pageNumber="33">Fig. 17</figureCitation>
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|
), as well as
|
|
<taxonomicName id="4C364D306767FFBC4D91FD81FB00FDA7" authority="Herbert, 2016" authorityName="Herbert" authorityYear="2016" box="[744,1186,580,606]" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="monsmaripi">
|
|
<emphasis id="B942EAA16767FFBC4D91FD81FC51FDA7" box="[744,1011,580,606]" italics="true" pageId="32" pageNumber="33">Sheldonia monsmaripi</emphasis>
|
|
Herbert, 2016
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C364D306767FFBC4B99FD81FED9FD7B" authority="(Pfeiffer, 1846)" baseAuthorityName="Pfeiffer" baseAuthorityYear="1846" class="Gastropoda" family="Urocyclidae" genus="Sheldonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="natalensis">
|
|
<emphasis id="B942EAA16767FFBC4B99FD81FAD4FDA7" box="[1248,1398,580,606]" italics="true" pageId="32" pageNumber="33">S. natalensis</emphasis>
|
|
(Pfeiffer, 1846)
|
|
</taxonomicName>
|
|
. Such colour differences may relate to ecological rather than genetic factors. The above notwithstanding, in the two species of
|
|
<taxonomicName id="4C364D306767FFBC4DA9FD4EFCF9FD5C" authority="Herbert, 2017" authorityName="Herbert" authorityYear="2017" box="[720,859,651,677]" class="Gastropoda" family="Urocyclidae" genus="Selatodryas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="32" pageNumber="33" phylum="Mollusca" rank="genus" status="gen. nov.">
|
|
<emphasis id="B942EAA16767FFBC4DA9FD4EFCF9FD5C" box="[720,859,651,677]" italics="true" pageId="32" pageNumber="33">Selatodryas</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="A27157DA6767FFBC4C1AFD4EFC6CFD5C" box="[867,974,651,677]" pageId="32" pageNumber="33" rank="genus">gen. nov.</taxonomicNameLabel>
|
|
the colour difference is pronounced and geographically consistent, and there are further correlated differences in genital anatomy and spermatophore morphology, providing confirmatory evidence that the two are distinct species.
|
|
</paragraph>
|
|
</subSubSection>
|
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</treatment>
|
|
</document> |