278 lines
28 KiB
XML
278 lines
28 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.295.4905" ID-GBIF-Dataset="59cbf2a9-41ca-4f5a-a350-ef034991798b" ID-PMC="PMC3677376" ID-Pensoft-Pub="1313-2970-295-1" ID-PubMed="23794874" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-295-1" ModsDocOrigin="ZooKeys 295" ModsDocTitle="Revision of the Middle American clade of the ant genus Stenamma Westwood (Hymenoptera, Formicidae, Myrmicinae)" checkinTime="1451247354359" checkinUser="pensoft" docAuthor="Branstetter, Michael G." docDate="2013" docId="8419C0EA8859D32D1E938FCCFB795F7A" docLanguage="en" docName="ZooKeys 295: 1-277" docOrigin="ZooKeys 295" docSource="http://dx.doi.org/10.3897/zookeys.295.4905" docTitle="Stenamma vexator Branstetter, 2013, sp. n." docType="treatment" docVersion="4" lastPageNumber="223" masterDocId="FFB3D3328201C177FF99FFC5FFA8A72C" masterDocTitle="Revision of the Middle American clade of the ant genus Stenamma Westwood (Hymenoptera, Formicidae, Myrmicinae)" masterLastPageNumber="277" masterPageNumber="1" pageNumber="219" updateTime="1668155749461" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the Middle American clade of the ant genus Stenamma Westwood (Hymenoptera, Formicidae, Myrmicinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Branstetter, Michael G.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>295</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>277</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.295.4905</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.295.4905</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-295-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152043632" LSID="urn:lsid:zoobank.org:act:918C9F3B-9C8E-4B71-924A-060CA2CD8D3A" httpUri="http://treatment.plazi.org/id/8419C0EA8859D32D1E938FCCFB795F7A" lastPageId="222" lastPageNumber="223" pageId="218" pageNumber="219">
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<subSubSection pageId="218" pageNumber="219" type="nomenclature">
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<paragraph pageId="218" pageNumber="219">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:918C9F3B-9C8E-4B71-924A-060CA2CD8D3A" class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma vexator" order="Hymenoptera" pageId="218" pageNumber="219" phylum="Arthropoda" rank="species" species="vexator">Stenamma vexator</taxonomicName>
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<taxonomicNameLabel pageId="218" pageNumber="219">sp. n.</taxonomicNameLabel>
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Worker: Figures 165, 166; Queen: Figure 167; Map: Figure 168
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</paragraph>
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</subSubSection>
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<subSubSection pageId="218" pageNumber="219" type="reference_group">
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<paragraph pageId="218" pageNumber="219">
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma" order="Hymenoptera" pageId="218" pageNumber="219" phylum="Arthropoda" rank="genus">Stenamma</taxonomicName>
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mgb39 [variant 2 below] Branstetter, 2012: phylogeny.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="218" pageNumber="219" type="type material">
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<paragraph pageId="218" pageNumber="219">Type material.</paragraph>
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<paragraph pageId="218" pageNumber="219">
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Holotype worker.
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<normalizedToken originalValue="MÉXICO">MEXICO</normalizedToken>
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, Veracruz: 1.2mi [1.9km] S Huatusco, [ca.
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<geoCoordinate direction="north" orientation="latitude" precision="55" value="19.193">19.193°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="55" value="-96.956">96.956°W</geoCoordinate>
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], 1344m, 2-5 Aug 1969, cloud forest, [Berlese sample of sifted leaf litter] (S. & J. Peck, collection ANTC18239) [USNM, specimen CASENT0126485]. Paratypes: same data as holotype [1dq, 1w, CAS, CASENT0126500, CASENT0126495], [1dq, 1w, FMNH, CASENT126502, CASENT0126499], [1dq, 1w, INBio, CASENT0126503, CASENT0126506], [1w, JTLC, CASENT0126507], [1dq, 1w, LACM, CASENT0126487, CASENT0126486], [1dq, 1w, MGBPC, CASENT0126511, CASENT0126512], [1dq, 1w, MCZ, CASENT0126523, CASENT0126496], [1w, MZSP, CASENT0126517], [1dq, 1w, UCD, CASENT0126520, CASENT0126501], [1dq, 1w, UNAM, CASENT0126513, CASENT0126498], [1dq, 1w, USNM, CASENT0126490, CASENT0126492].
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</paragraph>
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</subSubSection>
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<subSubSection pageId="218" pageNumber="219" type="worker diagnosis">
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<paragraph pageId="218" pageNumber="219">Worker diagnosis.</paragraph>
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<paragraph pageId="218" pageNumber="219">
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Integument mostly black or dark brown to red-brown; small- to medium-sized species (see HL, ML, PrW below); anterior clypeal margin undulating, forming 2-4 blunt teeth; basal margin of mandible usually sinuous, with a slight basal depression, but without a tooth; face completely sculptured, mostly rugoreticulate; pilosity on gastral dorsum predominately suberect and relatively sparse; eye of moderate size (EL 0.10-0.15, REL 18-22), oval-shaped, with 5-8 ommatidia at greatest diameter; propodeal spines tuberculate to short (PSL 0.08-0.16, PSI 1.1-1.8), often robust; petiole in profile often distinctive, appearing somewhat elongate, with node broadly rounded and pointed posteriad, venter under node with a small concavity. Similar species:
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma crypticum" order="Hymenoptera" pageId="218" pageNumber="219" phylum="Arthropoda" rank="species" species="crypticum">Stenamma crypticum</taxonomicName>
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,
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma huachucanum" order="Hymenoptera" pageId="218" pageNumber="219" phylum="Arthropoda" rank="species" species="huachucanum">Stenamma huachucanum</taxonomicName>
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,
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma stictosomum" order="Hymenoptera" pageId="218" pageNumber="219" phylum="Arthropoda" rank="species" species="stictosomum">Stenamma stictosomum</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="218" pageNumber="219" type="geographic range">
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<paragraph pageId="218" pageNumber="219">Geographic range.</paragraph>
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<paragraph pageId="218" pageNumber="219">Mexico (Atlantic slope).</paragraph>
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</subSubSection>
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<subSubSection lastPageId="220" lastPageNumber="221" pageId="218" pageNumber="219" type="worker description">
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<paragraph pageId="218" pageNumber="219">Worker description.</paragraph>
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<paragraph pageId="218" pageNumber="219">(29 measured) HL 0.62-0.78 (0.65), HW 0.55-0.71 (0.58), FLD 0.15-0.20 (0.16), PCW 0.04-0.06 (0.04), SL 0.50-0.67 (0.55), EL 0.10-0.15 (0.12), ACL 0.48-0.60 (0.50), ML 0.77-1.02 (0.85), PrW 0.39-0.49 (0.43), PSL 0.08-0.16, SDL 0.07-0.10 (0.09), PL 0.31-0.40 (0.34), PH 0.17-0.23 (0.19), PW 0.14-0.19 (0.15), PPL 0.15-0.22 (0.20), PPH 0.16-0.21 (0.17), PPW 0.17-0.22 (0.19), MFL 0.53-0.76 (0.60), MTL 0.45-0.62 (0.48), CI 89-96 (89), SI 85-98 (94), REL 18-22 (21), FLI 25-29 (28), PSI 1.1-1.8 (1.6), PI 53-59 (59), MFI 93-107 (97), ACI1 65-71 (68), ACI2 87-98 (92)</paragraph>
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<paragraph lastPageId="220" lastPageNumber="221" pageId="219" pageNumber="220">
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<pageBreakToken pageId="219" pageNumber="220" start="start">Small-</pageBreakToken>
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to medium-sized species; general body color black, or dark brown to red-brown (type population), with appendages brown to orange-brown or yellow-brown, lighter in joints and toward extremities; setae golden brown; mandible with 6 teeth, 2 teeth nearest basal tooth sometimes more worn and less distinct, basal tooth often well-defined, projecting; basal margin of mandible usually sinuous, with a slight basal depression (type population), but without a tooth, some populations with basal margin almost straight; mandible mostly smooth, with scattered piligerous punctae, and some striations near base and along lateral surface; median lobe of clypeus with a pair of faint longitudinal carinulae that diverge anteriorly, apex of lobe with a short transverse carinula, remainder of clypeus mostly smooth; area in between median lobe and anterior clypeal margin forming a distinct concavity where mandibles insert; posterior extension of clypeus between frontal lobes of moderate width (PCW 0.04-0.06), with sides subparallel; frontal lobes of moderate width (FLD 0.15-0.20, FLI 25-29), not greatly covering torular lobes in full-face view; head roughly oval-shaped (CI 89-96), with posterior margin flat to slightly depressed medially; eye of moderate size (EL 0.10-0.15, REL 18-22), oval-shaped, with 5-8 ommatidia at greatest diameter; face usually strongly sculptured, mostly rugoreticulate, with some longitudinal rugae along midline (type population), some populations with sculpture somewhat polished and reticulate less distinct; scape of moderate length (SI 85-98), just reaching posterior margin of head when laid back; surface of scape mostly smooth, with scattered piligerous punctae; flagellum with distinct 4-segmented antennal club; mesosoma usually completely sculptured (type population), but some populations shinier overall and with pronotum mostly smooth and shiny; pronotal dorsum longitudinally rugose, with rugae dense to somewhat sparse and irregular, rugae usually becoming punctae and reticulae on dorsum of mesonotum; side of pronotum mostly punctate or rugose, often with a small to large patch of smooth cuticle on ventral half; mesopleuron and side of propodeum rugulose-punctate; propodeal dorsum with rugoreticulae near anterior margin, changing to transverse carinulae near posterior margin; promesonotum in profile low-domed, slightly asymmetrical, with apex posterior of midpoint, anterior declivity of pronotum sometimes forming a sharp transition with dorsum, sculpture on declivity punctate, contrasting with rugose dorsum; metanotal groove usually well-demarcated, of moderate depth and width; propodeal spines tuberculate to short (PSL 0.08-0.16, PSI 1.1-1.8), often somewhat robust (type population); petiole in profile often distinctive, appearing somewhat elongate (PL/HW 0.53-0.56), with node broadly rounded and pointed posteriad, venter under node often with a small concavity; peduncle sometimes noticeably slender, thickening toward node; postpetiole in profile subspherical, similar in size to petiolar node (PPH/PH 0.85-0.91); anterior face of petiolar and postpetiolar nodes smooth, remainder of waist mostly punctate; gaster mostly smooth and shiny, with scattered piligerous punctae; pilosity on head, short and bilayered, with some longer suberect setae, and many shorter decumbent setae, remainder of body dorsum with longer suberect setae; pilosity on gastral dorsum predominately suberect and relatively sparse; setae on scape mostly subdecumbent;
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<pageBreakToken pageId="220" pageNumber="221" start="start">setae</pageBreakToken>
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on legs mostly subdecumbent to appressed, with some longer suberect setae on femoral venters and coxae.
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</paragraph>
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<caption pageId="220" pageNumber="221">
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<paragraph pageId="220" pageNumber="221">
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Figure 165.
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma vexator" order="Hymenoptera" pageId="220" pageNumber="221" phylum="Arthropoda" rank="species" species="vexator">Stenamma vexator</taxonomicName>
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holotype worker (CASENT0126485) A Profile B Face C Dorsum D Anterior clypeal margin in anterodorsal view E Gaster.
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</paragraph>
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</caption>
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<caption pageId="220" pageNumber="221">
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<paragraph pageId="220" pageNumber="221">
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Figure 166.
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma vexator" order="Hymenoptera" pageId="220" pageNumber="221" phylum="Arthropoda" rank="species" species="vexator">Stenamma vexator</taxonomicName>
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worker variants. Face, profile, and dorsal views
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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Variant 1 (CASENT0604610)
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<normalizedToken originalValue="D–F">D-F</normalizedToken>
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Variant 2 (CASENT0605506)
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<normalizedToken originalValue="G–I">G-I</normalizedToken>
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Variant 3 (CASENT060472).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="220" pageNumber="221" type="queen description">
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<paragraph pageId="220" pageNumber="221">Queen description.</paragraph>
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<paragraph pageId="220" pageNumber="221">(11 measured) HL 0.66-0.77 (0.66), HW 0.60-0.70 (0.60), FLD 0.17-0.20 (0.18), PCW 0.05-0.07 (0.05), SL 0.53-0.64 (0.53), EL 0.18-0.20 (0.18), ACL 0.50-0.60 (0.51), ML 0.94-1.13 (0.94), PrW 0.53-0.64 (0.54), PSL 0.14-0.17 (0.15), SDL 0.09-0.12 (0.09), PL 0.37-0.44 (0.38), PH 0.19-0.23 (0.20), PW 0.16-0.19 (0.17), PPL 0.18-0.22 (0.20), PPH 0.18-0.23 (0.19), PPW 0.20-0.24 (0.20), MFL 0.59-0.75 (0.59), MTL 0.49-0.61 (0.51), CI 90-95 (92), SI 84-95 (87), REL 28-30 (30), FLI 26-30 (29), PSI 1.3-1.8 (1.6), MFI 90-104 (102), ACI1 65-68 (68), ACI2 88-98 (96).</paragraph>
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<paragraph pageId="220" pageNumber="221">Same as worker except for standard queen modifications and as follows: pronotum with transverse rugae and rugoreticulae on lateral surface, changing to carinulae and punctae mesad; mesoscutum densely longitudinally rugose; scutellum rugose to rugoreticulate, sometimes with a smooth patch mesad; side of propodeum rugulose, with faint punctae, dorsum with transverse carinulae; wing venation as in Figure 167D.</paragraph>
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<caption pageId="220" pageNumber="221">
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<paragraph pageId="220" pageNumber="221">
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Figure 167.
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma vexator" order="Hymenoptera" pageId="220" pageNumber="221" phylum="Arthropoda" rank="species" species="vexator">Stenamma vexator</taxonomicName>
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paratype queen (CASENT0126490) A Profile B Face C Dorsum D Wings.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="220" pageNumber="221" type="male">
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<paragraph pageId="220" pageNumber="221">Male.</paragraph>
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<paragraph pageId="220" pageNumber="221">Unknown.</paragraph>
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</subSubSection>
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<subSubSection pageId="220" pageNumber="221" type="biology">
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<paragraph pageId="220" pageNumber="221">Biology.</paragraph>
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<paragraph pageId="220" pageNumber="221">
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This species is known only from Winkler and Berlese samples of sifted leaf litter. It occurs from approximately 1000-2000 m elevation and inhabits a variety of montane mesic forest environments (e.g. cloud forest, wet oak forest, oak-pine forest, tropical subevergreen forest,
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<taxonomicName class="Magnoliopsida" family="Altingiaceae" genus="Liquidambar" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Liquidambar" order="Saxifragales" pageId="220" pageNumber="221" phylum="Tracheophyta" rank="genus">Liquidambar</taxonomicName>
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forest).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="220" pageNumber="221" type="comments">
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<paragraph pageId="220" pageNumber="221">Comments.</paragraph>
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<paragraph pageId="220" pageNumber="221">
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma vexator" order="Hymenoptera" pageId="220" pageNumber="221" phylum="Arthropoda" rank="species" species="vexator">Stenamma vexator</taxonomicName>
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is not a particularly distinctive species, but it should be separable from similar species by the diagnostic character states given above and in the key. It bears the most resemblance to
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma huachucanum" order="Hymenoptera" pageId="220" pageNumber="221" phylum="Arthropoda" rank="species" species="huachucanum">Stenamma huachucanum</taxonomicName>
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, which occurs in sympatry with
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma vexator" order="Hymenoptera" pageId="220" pageNumber="221" phylum="Arthropoda" rank="species" species="vexator">Stenamma vexator</taxonomicName>
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at several sites.
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma vexator" order="Hymenoptera" pageId="220" pageNumber="221" phylum="Arthropoda" rank="species" species="vexator">Stenamma vexator</taxonomicName>
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is usually larger than
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma huachucanum" order="Hymenoptera" pageId="220" pageNumber="221" phylum="Arthropoda" rank="species" species="huachucanum">Stenamma huachucanum</taxonomicName>
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and has a longer, more distinctive petiole. Molecular phylogenetic data tentatively show that
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma vexator" order="Hymenoptera" pageId="220" pageNumber="221" phylum="Arthropoda" rank="species" species="vexator">Stenamma vexator</taxonomicName>
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is sister to the diversum species group, which includes
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma diversum" order="Hymenoptera" pageId="220" pageNumber="221" phylum="Arthropoda" rank="species" species="diversum">Stenamma diversum</taxonomicName>
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and
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma tico" order="Hymenoptera" pageId="220" pageNumber="221" phylum="Arthropoda" rank="species" species="tico">Stenamma tico</taxonomicName>
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(
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<bibRefCitation author="Branstetter, MG" journalOrPublisher="Systematic Entomology" pageId="228" pageNumber="229" pagination="478 - 496" title="Origin and diversification of the cryptic ant genus Stenamma Westwood (Hymenoptera: Formicidae), inferred from multilocus molecular data, biogeography and natural history." url="10.1111/j.1365-3113.2012.00624.x" volume="37" year="2012">Branstetter 2012</bibRefCitation>
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, unpublished data).
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</paragraph>
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<paragraph pageId="220" pageNumber="221">
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Several aberrant populations are worth separating out as variants of
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma vexator" order="Hymenoptera" pageId="220" pageNumber="221" phylum="Arthropoda" rank="species" species="vexator">Stenamma vexator</taxonomicName>
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. Variant 1 (Figure 166
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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) occurs at several sites in Puebla, Mexico. It has the propodeal spines tuberculate, the basal margin of the mandible nearly straight, and the clypeal teeth reduced to blunt nubs. Variant 2 (Figure 166
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<normalizedToken originalValue="D–F">D-F</normalizedToken>
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) is known from a few collections taken at high elevation between Oaxaca and Valle Nacional, in Oaxaca, Mexico. It is larger, darker, and has reduced sculpturing, with the pronotum mostly smooth and shiny. It also has the propodeal spines tuberculate, and the petiolar node pointing more vertically. I have not seen intermediate specimens between this variant and the holotype form of
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma vexator" order="Hymenoptera" pageId="220" pageNumber="221" phylum="Arthropoda" rank="species" species="vexator">Stenamma vexator</taxonomicName>
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, so it is possible that this high elevation variant represents a distinct species. Variant 3 (Figure 166
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<normalizedToken originalValue="G–I">G-I</normalizedToken>
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) is known from the locality Coapilla in Chiapas, Mexico. These specimens are black, have reduced sculpture, and have the propodeal spines almost absent. The petiolar node is also sharper and pointing more vertically. Placement of variant 3 within
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma vexator" order="Hymenoptera" pageId="220" pageNumber="221" phylum="Arthropoda" rank="species" species="vexator">Stenamma vexator</taxonomicName>
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is uncertain and needs to be confirmed with molecular data.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="222" lastPageNumber="223" pageId="220" pageNumber="221" type="material examined">
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<paragraph pageId="220" pageNumber="221">Material examined.</paragraph>
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<paragraph lastPageId="222" lastPageNumber="223" pageId="220" pageNumber="221">
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<normalizedToken originalValue="MÉXICO">MEXICO</normalizedToken>
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: Chiapas: 5km NE Coapilla,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="17.17297">17.17297°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-93.14612">93.14612°W</geoCoordinate>
|
||
, 1850m, 26 May 2008 (M. G. Branstetter); Hidalgo: 18km NE
|
||
<pageBreakToken pageId="221" pageNumber="222" start="start">Jacala</pageBreakToken>
|
||
, nr El Alamo, [ca.
|
||
<geoCoordinate direction="north" orientation="latitude" precision="55" value="21.058">21.058°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="55" value="-99.052">99.052°W</geoCoordinate>
|
||
], 1700m, 10 Jun 1988 (S. & J. Peck); Tlanchinol, 43km SW Huejutla, [ca.
|
||
<geoCoordinate direction="north" orientation="latitude" precision="55" value="20.988">20.988°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="55" value="-98.662">98.662°W</geoCoordinate>
|
||
], 1 Aug 1983 (S. & J. Peck); 43km SW Huejutla, [ca.
|
||
<geoCoordinate direction="north" orientation="latitude" precision="55" value="20.988">20.988°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="55" value="-98.662">98.662°W</geoCoordinate>
|
||
], 1500m, 14 Jun 1983 (S. & J. Peck); 4-6km N Tlanchinol, [ca.
|
||
<geoCoordinate direction="north" orientation="latitude" precision="55" value="21.026">21.026°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="55" value="-98.644">98.644°W</geoCoordinate>
|
||
], 1590m, 6-11 Jul 1973 (A. F. Newton);Oaxaca: 20.6km SW Valle Nacional,
|
||
<geoCoordinate direction="north" orientation="latitude" precision="1" value="17.60404">17.60404°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-96.37786">96.37786°W</geoCoordinate>
|
||
, 1730m, 13 Aug 2009 (M. G. Branstetter); 22.4km SW Valle Nacional,
|
||
<geoCoordinate direction="north" orientation="latitude" precision="1" value="17.59112">17.59112°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-96.39193">96.39193°W</geoCoordinate>
|
||
, 1990m, 13 Aug 2009 (M. G. Branstetter); 23km SW Valle Nacional, Km76, [ca.
|
||
<geoCoordinate direction="north" orientation="latitude" precision="5" value="17.6364">17.6364°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-96.47">96.4700°W</geoCoordinate>
|
||
], 1300m, 26 Jul 1992 (R. S. Anderson); 32km SW Valle Nacional, Km85, [ca.
|
||
<geoCoordinate direction="north" orientation="latitude" precision="5" value="17.5046">17.5046°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-96.3897">96.3897°W</geoCoordinate>
|
||
], 1650m 26 Jul 1992 (R. S. Anderson); Puebla: 2.7km S Apulco, nr Zacapoaxtla, [ca.
|
||
<geoCoordinate direction="north" orientation="latitude" precision="55" value="19.911">19.911°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="55" value="-97.606">97.606°W</geoCoordinate>
|
||
], 1400m, 22 Jul 1987 (R. S. Anderson); 7.6km SW La Cumbre, [ca.
|
||
<geoCoordinate direction="north" orientation="latitude" precision="5" value="20.1416">20.1416°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-98.0217">98.0217°W</geoCoordinate>
|
||
], 1585m, 23 Jul 1987 (R. S. Anderson); Teziutlan, [ca.
|
||
<geoCoordinate direction="north" orientation="latitude" precision="55" value="19.881">19.881°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="55" value="-97.304">97.304°W</geoCoordinate>
|
||
], 1350m, 18 Apr 1946 (F. Bonnet); 2km NE Teziutlan, [ca.
|
||
<geoCoordinate direction="north" orientation="latitude" precision="55" value="19.892">19.892°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="55" value="-97.296">97.296°W</geoCoordinate>
|
||
], 1220m, 19 Jun 1983 (S. & J. Peck); 24km N Xicotepec de Juarez, [ca.
|
||
<geoCoordinate direction="north" orientation="latitude" precision="55" value="20.282">20.282°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="55" value="-97.963">97.963°W</geoCoordinate>
|
||
], 1070m, 17 Jun 1983 (R. S. Anderson);
|
||
<normalizedToken originalValue="Querétaro">Queretaro</normalizedToken>
|
||
: 40km E Landa de Matamoros, [ca.
|
||
<geoCoordinate direction="north" orientation="latitude" precision="55" value="21.297">21.297°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="55" value="-99.09">99.090°W</geoCoordinate>
|
||
], 1520m, 14 Jul 1969 (S. & J. Peck); Veracruz: Cordoba, [ca.
|
||
<geoCoordinate direction="north" orientation="latitude" precision="55" value="18.908">18.908°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="55" value="-96.958">96.958°W</geoCoordinate>
|
||
], 4 Aug 1969 (S. B. Peck); 5km NE Coscomatepec, [ca.
|
||
<geoCoordinate direction="north" orientation="latitude" precision="55" value="19.1">19.100°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="55" value="-97.033">97.033°W</geoCoordinate>
|
||
], 22 Jun 1983 (Peck & Anderson); 1.9km S Huatusco, [ca.
|
||
<geoCoordinate direction="north" orientation="latitude" precision="55" value="19.193">19.193°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="55" value="-96.956">96.956°W</geoCoordinate>
|
||
], 1344m, 2-8 Aug 1969 (S. B. Peck); km38, 3.2km S Huatusco on
|
||
<normalizedToken originalValue="Fortín">Fortin</normalizedToken>
|
||
road, [ca.
|
||
<geoCoordinate direction="north" orientation="latitude" precision="55" value="19.185">19.185°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="55" value="-96.959">96.959°W</geoCoordinate>
|
||
], 1300m, 3 Aug 1965 (Cornell
|
||
<pageBreakToken pageId="222" pageNumber="223" start="start">Univ</pageBreakToken>
|
||
. Mexico Field Party); 7km E Huatusco, [ca.
|
||
<geoCoordinate direction="north" orientation="latitude" precision="55" value="19.207">19.207°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="55" value="-96.914">96.914°W</geoCoordinate>
|
||
], 22 Jun 1983 (Peck & Anderson); 2.7km N Teocelo, [ca.
|
||
<geoCoordinate direction="north" orientation="latitude" precision="555" value="19.4">19.40°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="555" value="-96.98">96.98°W</geoCoordinate>
|
||
], 1130m, 22-24 Jul 1973 (A. F. Newton).
|
||
</paragraph>
|
||
<caption pageId="222" pageNumber="223">
|
||
<paragraph pageId="222" pageNumber="223">
|
||
Figure 168. Distribution map of
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma vexator" order="Hymenoptera" pageId="222" pageNumber="223" phylum="Arthropoda" rank="species" species="vexator">Stenamma vexator</taxonomicName>
|
||
(circles) and
|
||
<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma zelum" order="Hymenoptera" pageId="222" pageNumber="223" phylum="Arthropoda" rank="species" species="zelum">Stenamma zelum</taxonomicName>
|
||
(squares).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |