180 lines
19 KiB
XML
180 lines
19 KiB
XML
<document id="566C6D570AAF03769428EDC5D0C9C3EA" ID-CLB-Dataset="45627" ID-DOI="10.5281/zenodo.211674" ID-GBIF-Dataset="d13a52a2-c0aa-45e9-9c6a-58613b627afb" ID-ISSN="1175-5326" ID-Zenodo-Dep="211674" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460429898246" checkinUser="plazi" docAuthor="Tilbrook, Kevin J." docDate="2012" docId="5C370C4EFFA3FFADFF646FC119F99E9B" docLanguage="en" docName="zt03165p063.pdf" docOrigin="Zootaxa 3165" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Bryopesanser baderae Tilbrook, 2012, n. sp." docType="treatment" docVersion="9" lastPageNumber="59" masterDocId="A00E7436FFB1FFB9FFF369001A099F6F" masterDocTitle="Review of the bryozoan genus Bryopesanser Tilbrook, 2006 (Escharinidae: Cheilostomata) with the description of 11 new species" masterLastPageNumber="63" masterPageNumber="39" pageNumber="57" updateTime="1698331236171" updateUser="ExternalLinkService">
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<mods:mods id="4736BB9E5901C67A829D507B49E77E94" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="ECC6668345E3DB04318026730233CAA3">
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<mods:title id="E5399BDF0374875596A0196FE5FCFCB2">Review of the bryozoan genus Bryopesanser Tilbrook, 2006 (Escharinidae: Cheilostomata) with the description of 11 new species</mods:title>
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</mods:titleInfo>
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<mods:name id="5925F9D6F43B378FD012F981C3A5DEFB" type="personal">
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<mods:roleTerm id="E2AE9FE35DDBFDD9ED38E963C4E9AE4C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6D35A6DA386041D341E6A0D6B87725F4">Tilbrook, Kevin J.</mods:namePart>
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<mods:typeOfResource id="C76B03B080B15788A545957D9792B273">text</mods:typeOfResource>
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<mods:titleInfo id="523F41F78D2B3F1903D460F9BDD93584">
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<mods:title id="BB7C18135ED1C0AFC60B9F3A995F86E6">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="932BBE2F3F60212C47ADC7B1692FDD1A">
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<mods:date id="D1A07D49634E8354B4BBF2BDBE454251">2012</mods:date>
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<mods:detail id="F6EC865519B4B43B273DEA083D7DEE4B" type="volume">
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<mods:number id="AE61514F845AC23A8524ACAFB9EFDD6A">3165</mods:number>
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<mods:start id="18BAD35A3570FA86450C1B552BD9BDAE">39</mods:start>
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<mods:end id="7A6CCD1D21A60B6E303A6A5DC998EB89">63</mods:end>
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<mods:classification id="6844EAC8D18E62825C5DBC792C06B0D9">journal article</mods:classification>
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<mods:identifier id="52514130BB70909E5102A12288DE8FD3" type="CLB-Dataset">45627</mods:identifier>
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<mods:identifier id="D6D5237F5E4BDF8C66079CC0E81C34FE" type="DOI">10.5281/zenodo.211674</mods:identifier>
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<mods:identifier id="57D6AB07C5210505D92FEDF7F7833297" type="GBIF-Dataset">d13a52a2-c0aa-45e9-9c6a-58613b627afb</mods:identifier>
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<mods:identifier id="BBF5B17F7EE39EF53BF26E7DFBB42DBE" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="5FE760E9C0C329A6FAA255E6C9735634" type="Zenodo-Dep">211674</mods:identifier>
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<treatment id="5C370C4EFFA3FFADFF646FC119F99E9B" ID-DOI="http://doi.org/10.5281/zenodo.6493024" ID-GBIF-Taxon="119429752" ID-Zenodo-Dep="6493024" LSID="urn:lsid:plazi:treatment:5C370C4EFFA3FFADFF646FC119F99E9B" httpUri="http://treatment.plazi.org/id/5C370C4EFFA3FFADFF646FC119F99E9B" lastPageId="20" lastPageNumber="59" pageId="18" pageNumber="57">
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<subSubSection id="9C84EED3FFA3FFABFF646FC11BA29993" pageId="18" pageNumber="57" type="nomenclature">
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<paragraph id="D421BD58FFA3FFABFF646FC11BE599B4" blockId="18.[151,492,1729,1788]" box="[151,492,1729,1755]" pageId="18" pageNumber="57">
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<heading id="8F690A34FFA3FFABFF646FC11BE599B4" bold="true" box="[151,492,1729,1755]" fontSize="11" level="1" pageId="18" pageNumber="57" reason="1">
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<emphasis id="E6EA614AFFA3FFABFF646FC11BE599B4" bold="true" box="[151,492,1729,1755]" pageId="18" pageNumber="57">
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<taxonomicName id="139EC6DBFFA3FFABFF646FC11BAA99B4" box="[151,419,1729,1755]" class="Gymnolaemata" family="Escharinidae" genus="Bryopesanser" kingdom="Animalia" order="Cheilostomatida" pageId="18" pageNumber="57" phylum="Bryozoa" rank="species" species="baderae" status="sp. nov.">
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<emphasis id="E6EA614AFFA3FFABFF646FC11BAA99B4" bold="true" box="[151,419,1729,1755]" italics="true" pageId="18" pageNumber="57">Bryopesanser baderae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="FDD9DC31FFA3FFABFE596FC11BE599B4" box="[426,492,1729,1755]" pageId="18" pageNumber="57" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="D421BD58FFA3FFABFF646FE41BA29993" blockId="18.[151,492,1729,1788]" box="[151,427,1764,1788]" pageId="18" pageNumber="57">
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(
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<figureCitation id="4CA5A1DDFFA3FFABFF6D6FE41B4D9993" box="[158,324,1764,1788]" captionStart="FIGURES 46 – 51" captionStartId="19.[151,263,1947,1970]" captionTargetBox="[193,1395,467,1921]" captionTargetId="figure@19.[193,1395,467,1921]" captionTargetPageId="19" captionText="FIGURES 46 – 51, Bryopesanser baderae n. sp. 46 - 48, Holotype specimen, LI 2011 / 152, Great Meteor Bank. 49, Paratype specimen, LI 2011 / 156, Great Meteor Bank. 50, LI 2011 / 154, Great Meteor Bank. 51, LI 2011 / 405, Great Meteor Bank. Scale bars: 46, 1000 µm; 47, 500 µm; 48, 100 µm; 49, 200 µm; 50, 500 µm; 51, 200 µm." httpUri="https://zenodo.org/record/211684/files/figure.png" pageId="18" pageNumber="57">Figures 46–51</figureCitation>
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, Table 1)
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</paragraph>
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</subSubSection>
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<subSubSection id="9C84EED3FFA3FFABFF646E261B0998C5" pageId="18" pageNumber="57" type="materials_examined">
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<paragraph id="D421BD58FFA3FFABFF646E261B0998C5" blockId="18.[151,1436,1830,2035]" pageId="18" pageNumber="57">
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<emphasis id="E6EA614AFFA3FFABFF646E261B0F9850" bold="true" box="[151,262,1830,1855]" pageId="18" pageNumber="57">Material.</emphasis>
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<typeStatus id="0B2503FAFFA3FFABFEFE6E271B7D9850" box="[269,372,1831,1855]" pageId="18" pageNumber="57" type="holotype">
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<emphasis id="E6EA614AFFA3FFABFEFE6E271B7D9850" box="[269,372,1831,1855]" italics="true" pageId="18" pageNumber="57">Holotype</emphasis>
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</typeStatus>
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(here selected): LI 2011/152, M42/3-465, Great Meteor Bank,
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<quantity id="136610BDFFA3FFABFBC56E271E779850" box="[1078,1150,1831,1855]" metricMagnitude="2" metricUnit="m" metricValue="3.02" pageId="18" pageNumber="57" unit="m" value="302.0">302 m</quantity>
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,
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<geoCoordinate id="B1AADB9FFFA3FFABFB796E271F009850" box="[1162,1289,1831,1855]" direction="north" orientation="latitude" pageId="18" pageNumber="57" precision="92" value="29.805">29°48.3´N</geoCoordinate>
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,
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<geoCoordinate id="B1AADB9FFFA3FFABFAE66E271F919850" box="[1301,1432,1831,1855]" direction="west" orientation="longitude" pageId="18" pageNumber="57" precision="92" value="-28.608334">28°36.5´W</geoCoordinate>
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, on coral.
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<typeStatus id="0B2503FAFFA3FFABFEF16E4C1B7D980C" box="[258,372,1868,1891]" pageId="18" pageNumber="57" type="paratype">
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<emphasis id="E6EA614AFFA3FFABFEF16E4C1B7D980C" box="[258,372,1868,1891]" italics="true" pageId="18" pageNumber="57">Paratypes</emphasis>
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</typeStatus>
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(here selected): LI 2011/150, 151 (on coral), LI 2011/157 two colonies on echinoid fragment, same locality as
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<typeStatus id="0B2503FAFFA3FFABFEA56E6F1BB598E8" box="[342,444,1903,1927]" pageId="18" pageNumber="57" type="holotype">holotype</typeStatus>
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.
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<emphasis id="E6EA614AFFA3FFABFE3B6E6F18EC98E8" box="[456,741,1903,1927]" italics="true" pageId="18" pageNumber="57">Other material examined</emphasis>
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: LI 2011/153, 154, 155, 156, 405, on coral, same locality as
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<typeStatus id="0B2503FAFFA3FFABFF646E921AF498C5" box="[151,253,1938,1962]" pageId="18" pageNumber="57" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="9C84EED3FFA3FFAAFF346EB61F149EC3" lastPageId="19" lastPageNumber="58" pageId="18" pageNumber="57" type="description">
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<paragraph id="D421BD58FFA3FFAAFF346EB61F149EC3" blockId="18.[151,1436,1830,2035]" lastBlockId="19.[151,1437,151,428]" lastPageId="19" lastPageNumber="58" pageId="18" pageNumber="57">
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<emphasis id="E6EA614AFFA3FFABFF346EB61B5098A0" bold="true" box="[199,345,1974,1999]" pageId="18" pageNumber="57">Description.</emphasis>
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Autozooids 0.65–0.85 x
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<quantity id="136610BDFFA3FFABFD766EB7193998A0" box="[645,816,1975,1999]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="18" pageNumber="57" unit="mm" value="0.6" valueMax="0.7" valueMin="0.5">0.50–0.70 mm</quantity>
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, irregularly polygonal, distinct, separated by shallow grooves. Frontal shield slightly nodular, slightly convex, evenly perforated with numerous small multiporous pores, areolar septular pores at each lateral and proximal angle. Primary orifice wider than long, ca 0.14 x
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<quantity id="136610BDFFA2FFAAFACA69971F959FC0" box="[1337,1436,151,175]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="19" pageNumber="58" unit="mm" value="0.12">0.12 mm</quantity>
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excluding sinus, anter deeply arched, proximal border straight, condyles smooth, dipping medially towards dropshaped sinus, as wide as long. Oral spines 9, evenly spaced. Proximal peristome hardly developed, a slight median mucro with ontogeny. Avicularia originating lateral to second most proximal pair of spines, rostra medium-sized, with smooth edges, open-ended distally, crossbar complete, slightly distomedially directed, mandibles fan-shaped. Ovicell hyperstomial, producing a slightly thickened proximal edge. Ovicellate zooids with 8 oral spines, the most distal pair incorporated into ovicell. Ancestrula longer than wide, ca 0.40 x
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<quantity id="136610BDFFA2FFAAFC07686F1E559EE8" box="[1012,1116,367,391]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="19" pageNumber="58" unit="mm" value="0.3">0.30 mm</quantity>
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; 10 spines, 5 closely spaced distally, 5 widely spaced proximally; opesia circular, occupying less than one quarter of frontal surface.
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</paragraph>
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</subSubSection>
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<caption id="80E1EDD0FFA2FFAAFF646E9B19E99881" httpUri="https://zenodo.org/record/211684/files/figure.png" pageId="19" pageNumber="58" targetBox="[193,1395,467,1921]" targetPageId="19">
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<paragraph id="D421BD58FFA2FFAAFF646E9B19E99881" blockId="19.[151,1436,1947,2030]" pageId="19" pageNumber="58">
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<emphasis id="E6EA614AFFA2FFAAFF646E9B18DF98DF" bold="true" box="[151,726,1947,1970]" pageId="19" pageNumber="58">
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FIGURES 46–51,
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<taxonomicName id="139EC6DBFFA2FFAAFEAD6E9C184198DE" box="[350,584,1947,1969]" class="Gymnolaemata" family="Escharinidae" genus="Bryopesanser" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="58" phylum="Bryozoa" rank="species" species="baderae" status="sp. nov.">
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<emphasis id="E6EA614AFFA2FFAAFEAD6E9C184198DE" bold="true" box="[350,584,1947,1969]" italics="true" pageId="19" pageNumber="58">Bryopesanser baderae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="FDD9DC31FFA2FFAAFDA26E9C188598DE" box="[593,652,1948,1969]" pageId="19" pageNumber="58" rank="species">n. sp.</taxonomicNameLabel>
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46-48,
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</emphasis>
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Holotype specimen, LI 2011/152, Great Meteor Bank.
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<emphasis id="E6EA614AFFA2FFAAFAE86E9B1F3398DF" bold="true" box="[1307,1338,1947,1968]" pageId="19" pageNumber="58">49,</emphasis>
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Paratype specimen, LI 2011/156, Great Meteor Bank.
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<emphasis id="E6EA614AFFA2FFAAFD936EBA187698A0" bold="true" box="[608,639,1978,1999]" pageId="19" pageNumber="58">50,</emphasis>
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LI 2011/154, Great Meteor Bank.
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<emphasis id="E6EA614AFFA2FFAAFC116EBA1E0B98A0" bold="true" box="[994,1026,1978,1999]" pageId="19" pageNumber="58">51,</emphasis>
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LI 2011/405, Great Meteor Bank. Scale bars:
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<emphasis id="E6EA614AFFA2FFAAFF3D6ED91AEE9881" bold="true" box="[206,231,2009,2030]" pageId="19" pageNumber="58">46</emphasis>
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, 1000 µm;
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<emphasis id="E6EA614AFFA2FFAAFEAE6ED91B7F9881" bold="true" box="[349,374,2009,2030]" pageId="19" pageNumber="58">47</emphasis>
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, 500 µm;
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<emphasis id="E6EA614AFFA2FFAAFE2C6ED91BF19881" bold="true" box="[479,504,2009,2030]" pageId="19" pageNumber="58">48</emphasis>
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, 100 µm;
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<emphasis id="E6EA614AFFA2FFAAFD926ED918739881" bold="true" box="[609,634,2009,2030]" pageId="19" pageNumber="58">49</emphasis>
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, 200 µm;
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<emphasis id="E6EA614AFFA2FFAAFD106ED918F59881" bold="true" box="[739,764,2009,2030]" pageId="19" pageNumber="58">50</emphasis>
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, 500 µm;
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<emphasis id="E6EA614AFFA2FFAAFC976ED919749881" bold="true" box="[868,893,2009,2030]" pageId="19" pageNumber="58">51</emphasis>
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, 200 µm.
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</paragraph>
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</caption>
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<subSubSection id="9C84EED3FFA5FFADFF34699818CC9FBB" pageId="20" pageNumber="59" type="etymology">
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<paragraph id="D421BD58FFA5FFADFF34699818CC9FBB" blockId="20.[151,1436,151,501]" pageId="20" pageNumber="59">
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<emphasis id="E6EA614AFFA5FFADFF3469981B479FDF" bold="true" box="[199,334,152,176]" pageId="20" pageNumber="59">Etymology.</emphasis>
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Named for Dr Beate Bader who collected the specimen, formerly of the Institut für Geowissenschaften, Christian-Albrechts-Universität zu Kiel.
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</paragraph>
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</subSubSection>
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<subSubSection id="9C84EED3FFA5FFADFF3469E01B3A9EE8" pageId="20" pageNumber="59" type="discussion">
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<paragraph id="D421BD58FFA5FFADFF3469E01B3A9EE8" blockId="20.[151,1436,151,501]" pageId="20" pageNumber="59">
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<emphasis id="E6EA614AFFA5FFADFF3469E01B329F97" bold="true" box="[199,315,224,248]" pageId="20" pageNumber="59">Remarks.</emphasis>
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<taxonomicName id="139EC6DBFFA5FFADFEB169E118379F97" box="[322,574,224,248]" class="Gymnolaemata" family="Escharinidae" genus="Bryopesanser" kingdom="Animalia" order="Cheilostomatida" pageId="20" pageNumber="59" phylum="Bryozoa" rank="species" species="baderae" status="sp. nov.">
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<emphasis id="E6EA614AFFA5FFADFEB169E118379F97" box="[322,574,224,248]" italics="true" pageId="20" pageNumber="59">Bryopesanser baderae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="FDD9DC31FFA5FFADFDB569E0188C9F97" box="[582,645,224,248]" pageId="20" pageNumber="59" rank="species">
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<emphasis id="E6EA614AFFA5FFADFDB569E0188C9F97" bold="true" box="[582,645,224,248]" pageId="20" pageNumber="59">n. sp.</emphasis>
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</taxonomicNameLabel>
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is characterised by the details of its primary orifice and the number of oral spines. The species differs from
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<taxonomicName id="139EC6DBFFA5FFADFDCB680518C79E73" box="[568,718,261,284]" class="Gymnolaemata" family="Escharinidae" genus="Escharina" kingdom="Animalia" order="Cheilostomatida" pageId="20" pageNumber="59" phylum="Bryozoa" rank="species" species="pesanseris">
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<emphasis id="E6EA614AFFA5FFADFDCB680518C79E73" box="[568,718,261,284]" italics="true" pageId="20" pageNumber="59">B. pesanseris</emphasis>
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</taxonomicName>
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in the number of oral spines and in this respect most resembles
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<taxonomicName id="139EC6DBFFA5FFADFF6468291B1A9E2F" box="[151,275,296,320]" class="Gymnolaemata" family="Escharinidae" genus="Bryopesanser" kingdom="Animalia" order="Cheilostomatida" pageId="20" pageNumber="59" phylum="Bryozoa" rank="species" species="thricyng" status="sp. nov.">
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<emphasis id="E6EA614AFFA5FFADFF6468291AA19E2F" box="[151,168,297,320]" italics="true" pageId="20" pageNumber="59">B</emphasis>
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.
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<emphasis id="E6EA614AFFA5FFADFF4568281B1A9E2F" box="[182,275,296,320]" italics="true" pageId="20" pageNumber="59">thricyng</emphasis>
|
||
</taxonomicName>
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||
<taxonomicNameLabel id="FDD9DC31FFA5FFADFEE868281B509E2F" box="[283,345,296,320]" pageId="20" pageNumber="59" rank="species">
|
||
<emphasis id="E6EA614AFFA5FFADFEE868281B509E2F" bold="true" box="[283,345,296,320]" pageId="20" pageNumber="59">n. sp.</emphasis>
|
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</taxonomicNameLabel>
|
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and
|
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<taxonomicName id="139EC6DBFFA5FFADFE6268281BED9E50" box="[401,484,295,319]" class="Gymnolaemata" family="Escharinidae" genus="Bryopesanser" kingdom="Animalia" order="Cheilostomatida" pageId="20" pageNumber="59" phylum="Bryozoa" rank="species" species="tiara" status="sp. nov.">B. tiara</taxonomicName>
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<taxonomicNameLabel id="FDD9DC31FFA5FFADFE18682818239E2F" box="[491,554,296,320]" pageId="20" pageNumber="59" rank="species">
|
||
<emphasis id="E6EA614AFFA5FFADFE18682818239E2F" bold="true" box="[491,554,296,320]" pageId="20" pageNumber="59">n. sp.</emphasis>
|
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</taxonomicNameLabel>
|
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which also have nine. However, these two species differ in a number of ways ― shape of primary orifice, extent of orificial peristome, ovicell ornamentation and size, and direction of the avicularian rostra.
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</paragraph>
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</subSubSection>
|
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<subSubSection id="9C84EED3FFA5FFADFF34689319F99E9B" pageId="20" pageNumber="59" type="distribution">
|
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<paragraph id="D421BD58FFA5FFADFF34689319F99E9B" blockId="20.[151,1436,151,501]" pageId="20" pageNumber="59">
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<emphasis id="E6EA614AFFA5FFADFF3468931B699EC3" bold="true" box="[199,352,403,428]" pageId="20" pageNumber="59">Distribution.</emphasis>
|
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<taxonomicName id="139EC6DBFFA5FFADFE9D689518789EC3" box="[366,625,404,428]" class="Gymnolaemata" family="Escharinidae" genus="Bryopesanser" kingdom="Animalia" order="Cheilostomatida" pageId="20" pageNumber="59" phylum="Bryozoa" rank="species" species="baderae" status="sp. nov.">
|
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<emphasis id="E6EA614AFFA5FFADFE9D689518789EC3" box="[366,625,404,428]" italics="true" pageId="20" pageNumber="59">Bryopesanser baderae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FDD9DC31FFA5FFADFD8C689418CD9EC3" box="[639,708,404,428]" pageId="20" pageNumber="59" rank="species">
|
||
<emphasis id="E6EA614AFFA5FFADFD8C689418CD9EC3" bold="true" box="[639,708,404,428]" pageId="20" pageNumber="59">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is found on the Great Meteor Bank, as well as the Hyères (
|
||
<geoCoordinate id="B1AADB9FFFA5FFADFF6C68B71B269EA0" box="[159,303,439,463]" direction="north" orientation="latitude" pageId="20" pageNumber="59" precision="9" value="31.499166">31°29.95´N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="B1AADB9FFFA5FFADFEC868B71BC79EA0" box="[315,462,439,463]" direction="west" orientation="longitude" pageId="20" pageNumber="59" precision="9" value="-28.992332">28°59.54´W</geoCoordinate>
|
||
) and Irving (
|
||
<geoCoordinate id="B1AADB9FFFA5FFADFD9068B718FB9EA0" box="[611,754,439,463]" direction="north" orientation="latitude" pageId="20" pageNumber="59" precision="9" value="31.9965">31°59.79´N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="B1AADB9FFFA5FFADFD0D68B7199B9EA0" box="[766,914,439,463]" direction="west" orientation="longitude" pageId="20" pageNumber="59" precision="9" value="-28.000166">28°00.01´W</geoCoordinate>
|
||
) seamounts (Beate Bader, pers. comm. 2001). The observed depth range is
|
||
<quantity id="136610BDFFA5FFADFE2A68DC18519E9B" box="[473,600,476,500]" metricMagnitude="2" metricUnit="m" metricValue="3.725" metricValueMax="4.55" metricValueMin="2.9" pageId="20" pageNumber="59" unit="m" value="372.5" valueMax="455.0" valueMin="290.0">290–455 m</quantity>
|
||
(Björn Berning, pers. comm. 2011).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |