treatments-xml/data/36/6A/37/366A374BF355FFFAFF47F999FE16EB57.xml
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<document id="66771DEF7D2A6F292BB738BE60C7D259" ID-DOI="10.11646/zootaxa.2796.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5290155" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629714697258" checkinUser="felipe" docAuthor="Tucker, J. K., Tenorio, M. J. &amp; Stahlschmidt, P." docDate="2011" docId="366A374BF355FFFAFF47F999FE16EB57" docLanguage="en" docName="zt02796p014.pdf" docOrigin="Zootaxa 2796 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2796.1.1" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Benthofascis biconica" docType="treatment" docVersion="4" lastPageNumber="5" masterDocId="CA534F33F357FFFEFFD0FFB8FFDDED18" masterDocTitle="The genus Benthofascis (Gastropoda: Conoidea): A revision with descriptions of new species" masterLastPageNumber="14" masterPageNumber="1" pageNumber="3" updateTime="1699100674434" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="40A3EA02B2C97CE28DFE5ADB2753FEF2">The genus Benthofascis (Gastropoda: Conoidea): A revision with descriptions of new species</mods:title>
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<mods:namePart id="DF7AE19EB31982E9BB5EF4DC3F06BDB3">Tucker, J. K.</mods:namePart>
<mods:affiliation id="C12495CA40C53CA16E25C28063AE932A">Illinois Natural History Survey, 8450 Montclaire Avenue, Brighton, Illinois 62012 USA. E-mail: jktucker @ inhs. uiuc. edu</mods:affiliation>
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<mods:namePart id="B59E49B1BFC1BC9E2D2192A5BCCF0F06">Tenorio, M. J.</mods:namePart>
<mods:affiliation id="88ED37493E560BBC725CB89666F2DC06">Facultad de Ciencias, Universidad de Cádiz, 11510 Puerto Real, Cádiz, Spain. E-mail: manuel. tenorio @ uca. es</mods:affiliation>
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<mods:namePart id="146367FC4E49F99ACF7B73C14E4BDE28">Stahlschmidt, P.</mods:namePart>
<mods:affiliation id="68C8E9781D7151418F0B31213A2EA714">Institute for Environmental Science, University Koblenz-Landau, Fortstrasse 7, 76829 Landau, Germany.</mods:affiliation>
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<mods:date id="C79263F345F6982A291894A822B86BAA">2011</mods:date>
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<mods:identifier id="0F00A0C2B2E17E3DF930034CA1FFD981" type="DOI">10.11646/zootaxa.2796.1.1</mods:identifier>
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<mods:identifier id="F59F9FE0E222E15BDEC1126F6278159A" type="Zenodo-Dep">5290155</mods:identifier>
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<treatment id="366A374BF355FFFAFF47F999FE16EB57" ID-DOI="http://doi.org/10.5281/zenodo.5294698" ID-Zenodo-Dep="5294698" LSID="urn:lsid:plazi:treatment:366A374BF355FFFAFF47F999FE16EB57" httpUri="http://treatment.plazi.org/id/366A374BF355FFFAFF47F999FE16EB57" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">
<subSubSection id="F6D9D5D6F355FFFCFF47F999FD86EB22" box="[151,603,1568,1595]" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="BE7C865DF355FFFCFF47F999FD86EB22" blockId="2.[151,603,1568,1628]" box="[151,603,1568,1595]" pageId="2" pageNumber="3">
<heading id="E5343131F355FFFCFF47F999FD86EB22" bold="true" box="[151,603,1568,1595]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<taxonomicName id="79C3FDDEF355FFFCFF47F999FD86EB22" ID-CoL="LHV8" authority="(Hedley, 1903)" baseAuthorityName="Hedley" baseAuthorityYear="1903" box="[151,603,1568,1595]" class="Gastropoda" family="Conorbidae" genus="Benthofascis" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="biconica">
<emphasis id="8CB75A4FF355FFFCFF47F999FD86EB22" bold="true" box="[151,603,1568,1595]" pageId="2" pageNumber="3">
<emphasis id="8CB75A4FF355FFFCFF47F999FE7DEB23" bold="true" box="[151,416,1569,1595]" italics="true" pageId="2" pageNumber="3">Benthofascis biconica</emphasis>
(
<bibRefCitation id="DA52FBACF355FFFCFE60F998FD8EEB22" author="Hedley, C." box="[432,595,1568,1594]" pageId="2" pageNumber="3" pagination="327 - 402" refId="ref9789" refString="Hedley, C. (1903) Scientific results of the trawling expedition of H. M. C. S. &quot; Thetis &quot; off the coast of New South Wales, in February and March, 1896. Mollusca. Part II. Scaphopoda and Gastropoda. Memoirs of the Australian Museum, 4, 327 - 402, pls. 36 - 37." type="journal article" year="1903">Hedley, 1903</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F6D9D5D6F355FFFCFF47F9FCFE7CEB44" box="[151,417,1604,1628]" pageId="2" pageNumber="3" type="description">
<paragraph id="BE7C865DF355FFFCFF47F9FCFE7CEB44" blockId="2.[151,603,1568,1628]" box="[151,417,1604,1628]" pageId="2" pageNumber="3">
<figureCitation id="26F89AD8F355FFFCFF47F9FCFEDDEB44" box="[151,256,1604,1628]" captionStart="FIGURE 1" captionStartId="3.[151,250,1731,1754]" captionTargetBox="[284,1299,195,1716]" captionTargetId="figure-227@3.[284,1299,195,1717]" captionTargetPageId="3" captionText="FIGURE 1. AC. Holotype of Benthofascis biconica (AMS C.016448, 3537 m deep, Shoalhaven Bight, New South Wales, Australia), height 18 mm, width 8 mm, ventral (A) and side (B) views with enlargement of spire (C). DE. Benthofascis biconica (AMS C.468235, 5.7 km E of Mistral Point, Sydney, New South Wales, 33º56.470'S, 151º19.630'E, 7177 m deep), heigth 29.8 mm, width 10.7 mm, ventral view (D) and x-ray (E). F. Benthofascis biconica (AMS C.468235, same locality data as above), height 17.9 mm, width 7.7 mm. GH. Holotype of Benthofascis sarcinula (AMS C.19871, 203 m deep, 20 km miles E of Cape Byron, New South Wales, Australia), height 7 mm, width 4 mm, ventral view (G) and an enlargement of the spire (H). IK. Benthofascis sarcinula (USNM 902892, SSE of Lakes Entrance, Gippsland Lakes, Victoria, Australia), height 16.6 mm, width 7.4 mm, ventral (I) and lateral views (J) with an enlargement of the spire (K). LN. Benthofascis sarcinula (USNM 902893, Victoria, Australia), height 24.6 mm, width 10.7 mm, ventral (L) and lateral (M) views and an x-ray (N). Scale bars represent 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5290157" httpUri="https://zenodo.org/record/5290157/files/figure.png" pageId="2" pageNumber="3">Figures 1</figureCitation>
, AF;
<figureCitation id="26F89AD8F355FFFCFE9EF9FCFE83EB44" box="[334,350,1604,1628]" captionStart="FIGURE 4" captionStartId="11.[151,250,1820,1843]" captionTargetBox="[258,1329,193,1799]" captionTargetId="figure-165@11.[258,1329,193,1799]" captionTargetPageId="11" captionText="FIGURE 4. A, B. Radular teeth extracted from Benthofascis biconica (AMS C.468232, shell length of 15.5 mm), photomicrograph (A) and drawing (B) showing internal structures. C, D. Radular teeth extracted from Benthofascis biconica (AMS C.468234, shell length of 15.2 mm), photomicrograph (C) and drawing (D) showing internal structures. E, F. Radular teeth extracted from Benthofascis pseudobiconica (AMS C.373099, shell length of 17.3 mm), photomicrograph (E) and drawing (F) showing internal structures. Scale bar represents 0.1 mm for AF. G. G. Electron micrographic study of the radular tooth of Benthofascis lozoueti Sysoev &amp; Bouchet, 2001, (MNHN IM200742331, 496 m deep, Norfolk Ridge, 22º50'S, 167º16'E, height 16.4 mm, width 7.0 mm, teeth from the same specimen shown in Fig. 3H). Scale bar dimensions are explained in the figure." figureDoi="http://doi.org/10.5281/zenodo.5290165" httpUri="https://zenodo.org/record/5290165/files/figure.png" pageId="2" pageNumber="3">4</figureCitation>
, AD
</paragraph>
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<subSubSection id="F6D9D5D6F355FFFCFF47F934FDBEEAF0" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="BE7C865DF355FFFCFF47F934FB99EBBB" blockId="2.[151,1436,1675,2024]" box="[151,1092,1675,1700]" pageId="2" pageNumber="3">
<emphasis id="8CB75A4FF355FFFCFF47F934FE98EBBC" bold="true" box="[151,325,1675,1700]" inLineHeading="true" pageId="2" pageNumber="3" reason="1">Type material.</emphasis>
<typeStatus id="617838FFF355FFFCFE9CF934FE69EBBC" box="[332,436,1676,1700]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
is
<collectionCode id="D8D21E98F355FFFCFE04F934FDCCEBBC" box="[468,529,1676,1700]" pageId="2" pageNumber="3">AMS</collectionCode>
C.016448, shell length
<quantity id="793B2BB8F355FFFCFCCCF934FCB0EBBB" box="[796,877,1676,1700]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="3" unit="mm" value="18.0">18 mm</quantity>
, shell width
<quantity id="793B2BB8F355FFFCFC2CF934FB9DEBBB" box="[1020,1088,1676,1700]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="3" unit="mm" value="8.0">8 mm</quantity>
.
</paragraph>
<paragraph id="BE7C865DF355FFFCFF15F908FC2DEBDF" blockId="2.[151,1436,1675,2024]" box="[197,1008,1711,1736]" pageId="2" pageNumber="3">
<materialsCitation id="0EAB8C00F355FFFCFF15F908FC2DEBDF" box="[197,1008,1711,1736]" country="Australia" elevation="36" location="Shoalhaven Bight" pageId="2" pageNumber="3" specimenCount="1" stateProvince="New South Wales" typeStatus="holotype">
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<typeStatus id="617838FFF355FFFCFF15F908FEDDEBD0" box="[197,256,1712,1736]" pageId="2" pageNumber="3">Type</typeStatus>
locality.
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<elevation id="35EE616EF355FFFCFEBAF917FE0FEBDF" box="[362,466,1711,1735]" metricMagnitude="1" metricUnit="m" metricValue="3.6" metricValueMax="3.7" metricValueMin="3.5" pageId="2" pageNumber="3" unit="m" value="36.0" valueMax="37.0" valueMin="35.0">3537 m</elevation>
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,
<location id="BB1CD086F355FFFCFE0DF917FD75EBDF" LSID="urn:lsid:plazi:treatment:366A374BF355FFFAFF47F999FE16EB57:BB1CD086F355FFFCFE0DF917FD75EBDF" box="[477,680,1711,1735]" country="Australia" name="Shoalhaven Bight" pageId="2" pageNumber="3" stateProvince="New South Wales">Shoalhaven Bight</location>
,
<collectingRegion id="7C0748BFF355FFFCFD63F908FCA3EBDF" box="[691,894,1711,1735]" country="Australia" name="New South Wales" pageId="2" pageNumber="3">New South Wales</collectingRegion>
,
<collectingCountry id="C6D4C6CDF355FFFCFC59F917FC2DEBDF" box="[905,1008,1711,1735]" name="Australia" pageId="2" pageNumber="3">Australia</collectingCountry>
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</paragraph>
<paragraph id="BE7C865DF355FFFCFF15F96BFE97EADC" blockId="2.[151,1436,1675,2024]" pageId="2" pageNumber="3">
<emphasis id="8CB75A4FF355FFFCFF15F96BFE49EBF4" bold="true" box="[197,404,1747,1772]" inLineHeading="true" pageId="2" pageNumber="3" reason="1">Other specimens.</emphasis>
<materialsCitation id="0EAB8C00F355FFFCFE4BF96CFDBAEA17" collectionCode="AMS" elevation="74" latitude="-33.941166" location="Sydney" longLatPrecision="1" longitude="151.32716" pageId="2" pageNumber="3" specimenCount="4" stateProvince="New South Wales">
<collectionCode id="D8D21E98F355FFFCFE4BF96CFE05EBF4" box="[411,472,1748,1772]" pageId="2" pageNumber="3">AMS</collectionCode>
C.468235,
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from
<quantity id="793B2BB8F355FFFCFCFBF96CFC53EBF3" box="[811,910,1748,1772]" metricMagnitude="1" metricUnit="m" metricValue="7.4" metricValueMax="7.7" metricValueMin="7.1" pageId="2" pageNumber="3" unit="m" value="74.0" valueMax="77.0" valueMin="71.0">
<elevation id="35EE616EF355FFFCFCFBF96CFC53EBF3" box="[811,910,1748,1772]" metricMagnitude="1" metricUnit="m" metricValue="7.4" metricValueMax="7.7" metricValueMin="7.1" pageId="2" pageNumber="3" unit="m" value="74.0" valueMax="77.0" valueMin="71.0">7177 m</elevation>
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deep,
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<quantity id="793B2BB8F355FFFCFC08F96CFBFAEBF4" box="[984,1063,1748,1772]" metricMagnitude="3" metricUnit="m" metricValue="5.7" pageId="2" pageNumber="3" unit="km" value="5.7">5.7 km</quantity>
E of Mistral Point
</locationDeviation>
,
<location id="BB1CD086F355FFFCFAD6F96CFA86EBF4" LSID="urn:lsid:plazi:treatment:366A374BF355FFFAFF47F999FE16EB57:BB1CD086F355FFFCFAD6F96CFA86EBF4" box="[1286,1371,1748,1772]" latitude="-33.941166" longLatPrecision="1" longitude="151.32716" name="Sydney" pageId="2" pageNumber="3" stateProvince="New South Wales">Sydney</location>
,
<collectingRegion id="7C0748BFF355FFFCFAB7F96CFEF5EA17" country="Australia" name="New South Wales" pageId="2" pageNumber="3">New South Wales</collectingRegion>
,
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,
<geoCoordinate id="DBF7E09AF355FFFCFE1FF94FFDBAEA17" box="[463,615,1783,1807]" degrees="151" direction="east" minutes="19.630" orientation="longitude" pageId="2" pageNumber="3" precision="1" value="151.32716">151º19.630'E</geoCoordinate>
</materialsCitation>
, dredged,
<date id="CA7DA09DF355FFFCFD32F94FFCA0EA17" box="[738,893,1783,1807]" pageId="2" pageNumber="3" value="1972-07-20">20 July 1972</date>
, Station: SBS 2;
<materialsCitation id="0EAB8C00F355FFFCFB98F94FFC94EA2C" collectionCode="AMS" elevation="9" latitude="-37.083332" location="Twofold Bay" longLatPrecision="1183" longitude="149.91667" pageId="2" pageNumber="3" specimenCount="2" stateProvince="New South Wales">
<collectionCode id="D8D21E98F355FFFCFB98F94FFB58EA17" box="[1096,1157,1783,1807]" pageId="2" pageNumber="3">AMS</collectionCode>
C.468237,
<specimenCount id="A8C54DD4F355FFFCFADEF94FFA41EA17" box="[1294,1436,1783,1807]" pageId="2" pageNumber="3" type="generic">2 specimens</specimenCount>
from
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</quantity>
,
<location id="BB1CD086F355FFFCFEC0F8A4FE78EA2C" LSID="urn:lsid:plazi:treatment:366A374BF355FFFAFF47F999FE16EB57:BB1CD086F355FFFCFEC0F8A4FE78EA2C" box="[272,421,1820,1844]" latitude="-37.083332" longLatPrecision="1183" longitude="149.91667" name="Twofold Bay" pageId="2" pageNumber="3" stateProvince="New South Wales">Twofold Bay</location>
,
<collectingRegion id="7C0748BFF355FFFCFE62F8A4FDA3EA2C" box="[434,638,1820,1844]" country="Australia" name="New South Wales" pageId="2" pageNumber="3">New South Wales</collectingRegion>
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,
<geoCoordinate id="DBF7E09AF355FFFCFD31F8A4FC94EA2C" box="[737,841,1820,1844]" degrees="149" direction="east" minutes="55" orientation="longitude" pageId="2" pageNumber="3" precision="925" value="149.91667">149º55'E</geoCoordinate>
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;
<materialsCitation id="0EAB8C00F355FFFCFC86F8A4FCCDEA4F" collectionCode="AMS" elevation="66" latitude="-33.825" location="Sydney" longLatPrecision="119" longitude="151.30167" municipality="North Head" pageId="2" pageNumber="3" specimenCount="2" stateProvince="New South Wales">
<collectionCode id="D8D21E98F355FFFCFC86F8A4FC4FEA2C" box="[854,914,1820,1844]" pageId="2" pageNumber="3">AMS</collectionCode>
C.388783,
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from
<quantity id="793B2BB8F355FFFCFB39F8A4FAF8EA2B" box="[1257,1317,1820,1844]" metricMagnitude="1" metricUnit="m" metricValue="6.6" pageId="2" pageNumber="3" unit="m" value="66.0">
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, off
<collectingMunicipality id="5E181C27F355FFFCFA8BF8A4FF08EA4F" pageId="2" pageNumber="3">North Head</collectingMunicipality>
,
<location id="BB1CD086F355FFFCFF31F887FEEBEA4F" LSID="urn:lsid:plazi:treatment:366A374BF355FFFAFF47F999FE16EB57:BB1CD086F355FFFCFF31F887FEEBEA4F" box="[225,310,1855,1879]" latitude="-33.825" longLatPrecision="119" longitude="151.30167" municipality="North Head" name="Sydney" pageId="2" pageNumber="3" stateProvince="New South Wales">Sydney</location>
,
<collectingRegion id="7C0748BFF355FFFCFE92F8F8FDD3EA4F" box="[322,526,1855,1879]" country="Australia" name="New South Wales" pageId="2" pageNumber="3">New South Wales</collectingRegion>
,
<geoCoordinate id="DBF7E09AF355FFFCFDCAF887FD5AEA4F" box="[538,647,1855,1879]" degrees="33" direction="south" minutes="49.5" orientation="latitude" pageId="2" pageNumber="3" precision="92" value="-33.825">33º49.5'S</geoCoordinate>
,
<geoCoordinate id="DBF7E09AF355FFFCFD43F887FCCDEA4F" box="[659,784,1855,1879]" degrees="151" direction="east" minutes="18.1" orientation="longitude" pageId="2" pageNumber="3" precision="92" value="151.30167">151º18.1'E</geoCoordinate>
</materialsCitation>
;
<materialsCitation id="0EAB8C00F355FFFCFCCDF887FC35EA64" collectionCode="AMS" elevation="74" latitude="-33.941166" location="Sydney" longLatPrecision="1" longitude="151.32716" pageId="2" pageNumber="3" specimenCount="1" stateProvince="New South Wales">
<collectionCode id="D8D21E98F355FFFCFCCDF887FC84EA4F" box="[797,857,1855,1879]" pageId="2" pageNumber="3">AMS</collectionCode>
C.468232,
<specimenCount id="A8C54DD4F355FFFCFC0EF887FB81EA4F" box="[990,1116,1855,1879]" pageId="2" pageNumber="3" type="generic">1 specimen</specimenCount>
from
<quantity id="793B2BB8F355FFFCFB73F887FAD4EA4F" box="[1187,1289,1855,1879]" metricMagnitude="1" metricUnit="m" metricValue="7.4" metricValueMax="7.7" metricValueMin="7.1" pageId="2" pageNumber="3" unit="m" value="74.0" valueMax="77.0" valueMin="71.0">
<elevation id="35EE616EF355FFFCFB73F887FAD4EA4F" box="[1187,1289,1855,1879]" metricMagnitude="1" metricUnit="m" metricValue="7.4" metricValueMax="7.7" metricValueMin="7.1" pageId="2" pageNumber="3" unit="m" value="74.0" valueMax="77.0" valueMin="71.0">7177 m</elevation>
</quantity>
,
<locationDeviation id="AFCE9F7FF355FFFCFAC5F887FEF3EA64" location="Mistral Point" pageId="2" pageNumber="3">
<quantity id="793B2BB8F355FFFCFAC5F887FAB9EA4F" box="[1301,1380,1855,1879]" metricMagnitude="3" metricUnit="m" metricValue="5.7" pageId="2" pageNumber="3" unit="km" value="5.7">5.7 km</quantity>
E of Mistral Point
</locationDeviation>
,
<location id="BB1CD086F355FFFCFEEBF8DCFE4DEA64" LSID="urn:lsid:plazi:treatment:366A374BF355FFFAFF47F999FE16EB57:BB1CD086F355FFFCFEEBF8DCFE4DEA64" box="[315,400,1892,1916]" latitude="-33.941166" longLatPrecision="1" longitude="151.32716" name="Sydney" pageId="2" pageNumber="3" stateProvince="New South Wales">Sydney</location>
,
<collectingRegion id="7C0748BFF355FFFCFE4CF8DCFDB4EA64" box="[412,617,1892,1916]" country="Australia" name="New South Wales" pageId="2" pageNumber="3">New South Wales</collectingRegion>
,
<geoCoordinate id="DBF7E09AF355FFFCFDA7F8DCFD22EA64" box="[631,767,1892,1916]" degrees="33" direction="south" minutes="56.470" orientation="latitude" pageId="2" pageNumber="3" precision="1" value="-33.941166">33º56.470'S</geoCoordinate>
,
<geoCoordinate id="DBF7E09AF355FFFCFCDCF8DCFC7FEA64" box="[780,930,1892,1916]" degrees="151" direction="east" minutes="19.630" orientation="longitude" pageId="2" pageNumber="3" precision="1" value="151.32716">151º19.630'E</geoCoordinate>
(wet)
</materialsCitation>
;
<materialsCitation id="0EAB8C00F355FFFCFC26F8DCFBCBEA87" collectionCode="AMS" elevation="77" latitude="-33.969166" location="Sydney" longLatPrecision="1" longitude="151.30034" pageId="2" pageNumber="3" specimenCount="1" stateProvince="New South Wales">
<collectionCode id="D8D21E98F355FFFCFC26F8DCFBEFEA64" box="[1014,1074,1892,1916]" pageId="2" pageNumber="3">AMS</collectionCode>
C.468233,
<specimenCount id="A8C54DD4F355FFFCFB68F8DCFAE5EA64" box="[1208,1336,1892,1916]" pageId="2" pageNumber="3" type="generic">1 specimen</specimenCount>
from
<quantity id="793B2BB8F355FFFCFA50F8DCFF72EA87" metricMagnitude="1" metricUnit="m" metricValue="7.7" pageId="2" pageNumber="3" unit="m" value="77.0">
<elevation id="35EE616EF355FFFCFA50F8DCFF72EA87" metricMagnitude="1" metricUnit="m" metricValue="7.7" pageId="2" pageNumber="3" unit="m" value="77.0">77 m</elevation>
</quantity>
,
<locationDeviation id="AFCE9F7FF355FFFCFF6BF83FFE78EAB8" box="[187,421,1927,1952]" location="Long Bay" pageId="2" pageNumber="3">
<quantity id="793B2BB8F355FFFCFF6BF83FFF28EA87" box="[187,245,1927,1951]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="3" unit="km" value="5.0">5 km</quantity>
E of Long Bay
</locationDeviation>
,
<location id="BB1CD086F355FFFCFE61F83FFDDBEA87" LSID="urn:lsid:plazi:treatment:366A374BF355FFFAFF47F999FE16EB57:BB1CD086F355FFFCFE61F83FFDDBEA87" box="[433,518,1927,1951]" latitude="-33.969166" longLatPrecision="1" longitude="151.30034" name="Sydney" pageId="2" pageNumber="3" stateProvince="New South Wales">Sydney</location>
,
<collectingRegion id="7C0748BFF355FFFCFDC2F830FD00EA87" box="[530,733,1927,1951]" country="Australia" name="New South Wales" pageId="2" pageNumber="3">New South Wales</collectingRegion>
,
<geoCoordinate id="DBF7E09AF355FFFCFD39F83FFCAFEA87" box="[745,882,1927,1951]" degrees="33" direction="south" minutes="58.150" orientation="latitude" pageId="2" pageNumber="3" precision="1" value="-33.969166">33º58.150'S</geoCoordinate>
,
<geoCoordinate id="DBF7E09AF355FFFCFCAFF83FFBCBEA87" box="[895,1046,1927,1951]" degrees="151" direction="east" minutes="18.020" orientation="longitude" pageId="2" pageNumber="3" precision="1" value="151.30034">151º18.020'E</geoCoordinate>
</materialsCitation>
, dredged 20 February, 1973, Shelf Benthic Survey.
</paragraph>
<paragraph id="BE7C865DF355FFFCFF15F868FDBEEAF0" blockId="2.[151,1436,1675,2024]" box="[197,611,2000,2024]" pageId="2" pageNumber="3">
<emphasis id="8CB75A4FF355FFFCFF15F868FEC5EAF0" bold="true" box="[197,280,2000,2024]" inLineHeading="true" pageId="2" pageNumber="3" reason="1">Range.</emphasis>
<collectingRegion id="7C0748BFF355FFFCFECFF869FE34EAF0" box="[287,489,2000,2024]" country="Australia" name="New South Wales" pageId="2" pageNumber="3">New South Wales</collectingRegion>
,
<collectingCountry id="C6D4C6CDF355FFFCFE25F868FDBDEAF0" box="[501,608,2000,2024]" name="Australia" pageId="2" pageNumber="3">Australia</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="EABCD6D5F354FFFDFF47F97BFEE6EAF5" ID-DOI="http://doi.org/10.5281/zenodo.5290157" ID-Zenodo-Dep="5290157" httpUri="https://zenodo.org/record/5290157/files/figure.png" pageId="3" pageNumber="4" startId="3.[151,250,1731,1754]" targetBox="[284,1299,195,1716]" targetPageId="3">
<paragraph id="BE7C865DF354FFFDFF47F97BFEE6EAF5" blockId="3.[151,1436,1731,2029]" pageId="3" pageNumber="4">
<emphasis id="8CB75A4FF354FFFDFF47F97BFE91EBC2" bold="true" box="[151,332,1731,1754]" pageId="3" pageNumber="4">FIGURE 1. AC</emphasis>
. Holotype of
<taxonomicName id="79C3FDDEF354FFFDFE0CF97BFD61EBC1" baseAuthorityName="Hedley" baseAuthorityYear="1903" box="[476,700,1731,1753]" class="Gastropoda" family="Conorbidae" genus="Benthofascis" kingdom="Animalia" order="Neogastropoda" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="biconica">
<emphasis id="8CB75A4FF354FFFDFE0CF97BFD61EBC1" box="[476,700,1731,1753]" italics="true" pageId="3" pageNumber="4">Benthofascis biconica</emphasis>
</taxonomicName>
(AMS C.016448, 3537 m deep, Shoalhaven Bight, New South Wales, Australia), height 18 mm, width 8 mm, ventral (
<emphasis id="8CB75A4FF354FFFDFDACF959FD53EBE0" bold="true" box="[636,654,1761,1784]" pageId="3" pageNumber="4">A</emphasis>
) and side (
<emphasis id="8CB75A4FF354FFFDFD2FF95AFCCDEBEF" bold="true" box="[767,784,1762,1783]" pageId="3" pageNumber="4">B</emphasis>
) views with enlargement of spire (
<emphasis id="8CB75A4FF354FFFDFBA0F959FB5FEBE0" bold="true" box="[1136,1154,1761,1784]" pageId="3" pageNumber="4">C</emphasis>
).
<emphasis id="8CB75A4FF354FFFDFB47F95AFB15EBEF" bold="true" box="[1175,1224,1762,1783]" pageId="3" pageNumber="4">DE</emphasis>
.
<taxonomicName id="79C3FDDEF354FFFDFB04F95AFF6BEA0F" baseAuthorityName="Hedley" baseAuthorityYear="1903" class="Gastropoda" family="Conorbidae" genus="Benthofascis" kingdom="Animalia" order="Neogastropoda" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="biconica">
<emphasis id="8CB75A4FF354FFFDFB04F95AFF6BEA0F" italics="true" pageId="3" pageNumber="4">Benthofascis biconica</emphasis>
</taxonomicName>
(AMS C.468235, 5.7 km E of Mistral Point, Sydney, New South Wales, 33º56.470'S, 151º19.630'E, 7177 m deep), heigth 29.8 mm, width 10.7 mm, ventral view (
<emphasis id="8CB75A4FF354FFFDFD92F898FD89EA2D" bold="true" box="[578,596,1824,1845]" pageId="3" pageNumber="4">D</emphasis>
) and x-ray (
<emphasis id="8CB75A4FF354FFFDFD09F898FD37EA2D" bold="true" box="[729,746,1824,1845]" pageId="3" pageNumber="4">E</emphasis>
).
<emphasis id="8CB75A4FF354FFFDFCD2F898FCD2EA2D" bold="true" box="[770,783,1824,1845]" pageId="3" pageNumber="4">F</emphasis>
.
<taxonomicName id="79C3FDDEF354FFFDFCCEF8A7FC22EA2D" baseAuthorityName="Hedley" baseAuthorityYear="1903" box="[798,1023,1823,1845]" class="Gastropoda" family="Conorbidae" genus="Benthofascis" kingdom="Animalia" order="Neogastropoda" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="biconica">
<emphasis id="8CB75A4FF354FFFDFCCEF8A7FC22EA2D" box="[798,1023,1823,1845]" italics="true" pageId="3" pageNumber="4">Benthofascis biconica</emphasis>
</taxonomicName>
(AMS C.468235, same locality data as above), height 17.9 mm, width 7.7 mm.
<emphasis id="8CB75A4FF354FFFDFDFFF886FDBEEA4D" bold="true" box="[559,611,1854,1877]" pageId="3" pageNumber="4">GH</emphasis>
. Holotype of
<taxonomicName id="79C3FDDEF354FFFDFD20F886FC0AEA4C" baseAuthorityName="Hedley" baseAuthorityYear="1905" box="[752,983,1854,1876]" class="Gastropoda" family="Conorbidae" genus="Benthofascis" kingdom="Animalia" order="Neogastropoda" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="sarcinula">
<emphasis id="8CB75A4FF354FFFDFD20F886FC0AEA4C" box="[752,983,1854,1876]" italics="true" pageId="3" pageNumber="4">Benthofascis sarcinula</emphasis>
</taxonomicName>
(AMS C.19871, 203 m deep, 20 km miles E of Cape Byron, New South Wales, Australia), height 7 mm, width 4 mm, ventral view (
<emphasis id="8CB75A4FF354FFFDFBDBF8E4FBC2EA6B" bold="true" box="[1035,1055,1884,1907]" pageId="3" pageNumber="4">G</emphasis>
) and an enlargement of the spire (
<emphasis id="8CB75A4FF354FFFDFAA9F8E5FA50EA6A" bold="true" box="[1401,1421,1885,1906]" pageId="3" pageNumber="4">H</emphasis>
).
<emphasis id="8CB75A4FF354FFFDFF47F8C4FF1CEA89" bold="true" box="[151,193,1916,1937]" pageId="3" pageNumber="4">IK</emphasis>
.
<taxonomicName id="79C3FDDEF354FFFDFF1EF8C3FE6BEA89" baseAuthorityName="Hedley" baseAuthorityYear="1905" box="[206,438,1915,1937]" class="Gastropoda" family="Conorbidae" genus="Benthofascis" kingdom="Animalia" order="Neogastropoda" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="sarcinula">
<emphasis id="8CB75A4FF354FFFDFF1EF8C3FE6BEA89" box="[206,438,1915,1937]" italics="true" pageId="3" pageNumber="4">Benthofascis sarcinula</emphasis>
</taxonomicName>
(USNM 902892, SSE of Lakes Entrance, Gippsland Lakes, Victoria, Australia), height 16.6 mm, width 7.4 mm, ventral (
<emphasis id="8CB75A4FF354FFFDFE40F822FE47EAB7" bold="true" box="[400,410,1946,1967]" pageId="3" pageNumber="4">I</emphasis>
) and lateral views (
<emphasis id="8CB75A4FF354FFFDFDBCF822FDA4EAB7" bold="true" box="[620,633,1946,1967]" pageId="3" pageNumber="4">J</emphasis>
) with an enlargement of the spire (
<emphasis id="8CB75A4FF354FFFDFC3EF822FBDFEAB7" bold="true" box="[1006,1026,1946,1967]" pageId="3" pageNumber="4">K</emphasis>
).
<emphasis id="8CB75A4FF354FFFDFBCAF822FB94EAB7" bold="true" box="[1050,1097,1946,1967]" pageId="3" pageNumber="4">LN</emphasis>
.
<taxonomicName id="79C3FDDEF354FFFDFB89F822FA9CEAA8" baseAuthorityName="Hedley" baseAuthorityYear="1905" box="[1113,1345,1946,1968]" class="Gastropoda" family="Conorbidae" genus="Benthofascis" kingdom="Animalia" order="Neogastropoda" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="sarcinula">
<emphasis id="8CB75A4FF354FFFDFB89F822FA9CEAA8" box="[1113,1345,1946,1968]" italics="true" pageId="3" pageNumber="4">Benthofascis sarcinula</emphasis>
</taxonomicName>
(USNM 902893, Victoria, Australia), height 24.6 mm, width 10.7 mm, ventral (
<emphasis id="8CB75A4FF354FFFDFCA3F801FC59EAD6" bold="true" box="[883,900,1977,1998]" pageId="3" pageNumber="4">L</emphasis>
) and lateral (
<emphasis id="8CB75A4FF354FFFDFBDFF801FBFAEAD6" bold="true" box="[1039,1063,1977,1998]" pageId="3" pageNumber="4">M</emphasis>
) views and an x-ray (
<emphasis id="8CB75A4FF354FFFDFADBF801FAC0EAD6" bold="true" box="[1291,1309,1977,1998]" pageId="3" pageNumber="4">N</emphasis>
). Scale bars represent 1 mm.
</paragraph>
</caption>
<subSubSection id="F6D9D5D6F353FFFAFF15FF2FFC93EFBF" pageId="4" pageNumber="5" type="description">
<paragraph id="BE7C865DF353FFFAFF15FF2FFC93EFBF" blockId="4.[151,1437,151,1615]" pageId="4" pageNumber="5">
<emphasis id="8CB75A4FF353FFFAFF15FF2FFE8AEDA8" bold="true" box="[197,343,151,176]" inLineHeading="true" pageId="4" pageNumber="5" reason="1">Description.</emphasis>
Shell solid, biconical, slightly angled at the shoulder, of more than six whorls. Color apparently flesh-tint. Sculpture: below the shoulder the shell is furrowed by numerous fine spiral grooves, crossed by arcuate growth lines, above the furrows are broader and fewer. Aperture narrow, sinus sutural and deep, outer lip appears to have curved far forward, columella angled in center, spreading broadly and with a small anterior plication. Length,
<quantity id="793B2BB8F353FFFAFF47FE9FFF34EC27" box="[151,233,295,319]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="4" pageNumber="5" unit="mm" value="18.0">18 mm</quantity>
; breadth,
<quantity id="793B2BB8F353FFFAFE86FE9FFE48EC27" box="[342,405,295,319]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="5" unit="mm" value="8.0">8 mm</quantity>
(quoted from
<bibRefCitation id="DA52FBACF353FFFAFDE7FE9FFD15EC27" author="Hedley, C." box="[567,712,295,319]" pageId="4" pageNumber="5" pagination="327 - 402" refId="ref9789" refString="Hedley, C. (1903) Scientific results of the trawling expedition of H. M. C. S. &quot; Thetis &quot; off the coast of New South Wales, in February and March, 1896. Mollusca. Part II. Scaphopoda and Gastropoda. Memoirs of the Australian Museum, 4, 327 - 402, pls. 36 - 37." type="journal article" year="1903">Hedley 1903</bibRefCitation>
, p. 385386). Inner shell walls are resorbed (
<figureCitation id="26F89AD8F353FFFAFB15FE9FFAC7EC27" box="[1221,1306,295,319]" captionStart="FIGURE 1" captionStartId="3.[151,250,1731,1754]" captionTargetBox="[284,1299,195,1716]" captionTargetId="figure-227@3.[284,1299,195,1717]" captionTargetPageId="3" captionText="FIGURE 1. AC. Holotype of Benthofascis biconica (AMS C.016448, 3537 m deep, Shoalhaven Bight, New South Wales, Australia), height 18 mm, width 8 mm, ventral (A) and side (B) views with enlargement of spire (C). DE. Benthofascis biconica (AMS C.468235, 5.7 km E of Mistral Point, Sydney, New South Wales, 33º56.470'S, 151º19.630'E, 7177 m deep), heigth 29.8 mm, width 10.7 mm, ventral view (D) and x-ray (E). F. Benthofascis biconica (AMS C.468235, same locality data as above), height 17.9 mm, width 7.7 mm. GH. Holotype of Benthofascis sarcinula (AMS C.19871, 203 m deep, 20 km miles E of Cape Byron, New South Wales, Australia), height 7 mm, width 4 mm, ventral view (G) and an enlargement of the spire (H). IK. Benthofascis sarcinula (USNM 902892, SSE of Lakes Entrance, Gippsland Lakes, Victoria, Australia), height 16.6 mm, width 7.4 mm, ventral (I) and lateral views (J) with an enlargement of the spire (K). LN. Benthofascis sarcinula (USNM 902893, Victoria, Australia), height 24.6 mm, width 10.7 mm, ventral (L) and lateral (M) views and an x-ray (N). Scale bars represent 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5290157" httpUri="https://zenodo.org/record/5290157/files/figure.png" pageId="4" pageNumber="5">Fig. 1E</figureCitation>
). The operculum is leaf-shaped with a terminal nucleus. It is fairly large and covers most of the apertural opening. The operculum of a specimen (AMS C.468234) with a shell length of
<quantity id="793B2BB8F353FFFAFC98FED7FC71EC9F" box="[840,940,367,391]" metricMagnitude="-2" metricUnit="m" metricValue="1.52" pageId="4" pageNumber="5" unit="mm" value="15.2">15.2 mm</quantity>
measured
<quantity id="793B2BB8F353FFFAFBF6FED7FB57EC9F" box="[1062,1162,367,391]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" pageId="4" pageNumber="5" unit="mm" value="3.25">3.25 mm</quantity>
in length. We examined a total of
<specimenCount id="A8C54DD4F353FFFAFED1FE2CFE45ECB4" box="[257,408,404,428]" pageId="4" pageNumber="5" type="generic">28 specimens</specimenCount>
that could be accurately measured. Shell length for these averaged
<quantity id="793B2BB8F353FFFAFB5DFE2CFB2DECB3" box="[1165,1264,404,428]" metricMagnitude="-2" metricUnit="m" metricValue="2.6399999999999997" pageId="4" pageNumber="5" unit="mm" value="26.4">26.4 mm</quantity>
(range =
<quantity id="793B2BB8F353FFFAFA8DFE2CFEDDECD7" metricMagnitude="-2" metricUnit="m" metricValue="2.5949999999999998" metricValueMax="4.18" metricValueMin="1.01" pageId="4" pageNumber="5" unit="mm" value="25.95" valueMax="41.8" valueMin="10.1">10.1 41.8 mm</quantity>
). Shell width averaged
<quantity id="793B2BB8F353FFFAFDDCFE0FFDADECD7" box="[524,624,439,463]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="4" pageNumber="5" unit="mm" value="10.7">10.7 mm</quantity>
(range =
<quantity id="793B2BB8F353FFFAFD31FE0FFCA1ECD7" box="[737,892,439,463]" metricMagnitude="-2" metricUnit="m" metricValue="1.11" metricValueMax="1.76" metricValueMin="0.45999999999999996" pageId="4" pageNumber="5" unit="mm" value="11.1" valueMax="17.6" valueMin="4.6">4.617.6 mm</quantity>
). The radula has a barb and blade at its anterior end (
<figureCitation id="26F89AD8F353FFFAFF1EFE64FE8BECEC" box="[206,342,476,500]" captionStart="FIGURE 4" captionStartId="11.[151,250,1820,1843]" captionTargetBox="[258,1329,193,1799]" captionTargetId="figure-165@11.[258,1329,193,1799]" captionTargetPageId="11" captionText="FIGURE 4. A, B. Radular teeth extracted from Benthofascis biconica (AMS C.468232, shell length of 15.5 mm), photomicrograph (A) and drawing (B) showing internal structures. C, D. Radular teeth extracted from Benthofascis biconica (AMS C.468234, shell length of 15.2 mm), photomicrograph (C) and drawing (D) showing internal structures. E, F. Radular teeth extracted from Benthofascis pseudobiconica (AMS C.373099, shell length of 17.3 mm), photomicrograph (E) and drawing (F) showing internal structures. Scale bar represents 0.1 mm for AF. G. G. Electron micrographic study of the radular tooth of Benthofascis lozoueti Sysoev &amp; Bouchet, 2001, (MNHN IM200742331, 496 m deep, Norfolk Ridge, 22º50'S, 167º16'E, height 16.4 mm, width 7.0 mm, teeth from the same specimen shown in Fig. 3H). Scale bar dimensions are explained in the figure." figureDoi="http://doi.org/10.5281/zenodo.5290165" httpUri="https://zenodo.org/record/5290165/files/figure.png" pageId="4" pageNumber="5">Figs. 4AD</figureCitation>
). The barb is sharply tipped and this distal tip is curved away from the shaft of the tooth. The barb is about one-third as long as the blade. The distal end of the blade slopes towards the shaft of the tooth and is not much elevated above the shaft of the tooth. There is no anterior fold but there is a C-fold. This fold demonstrates that the tooth is enrolled a minimum of 360 degrees. There are no serrations and no terminating cusp. There is a fold on the shaft of the tooth that begins posterior to the distal end of the blade. This fold extends to the waist of the tooth (
<figureCitation id="26F89AD8F353FFFAFF30FD37FEA6EFBF" box="[224,379,655,679]" captionStart="FIGURE 4" captionStartId="11.[151,250,1820,1843]" captionTargetBox="[258,1329,193,1799]" captionTargetId="figure-165@11.[258,1329,193,1799]" captionTargetPageId="11" captionText="FIGURE 4. A, B. Radular teeth extracted from Benthofascis biconica (AMS C.468232, shell length of 15.5 mm), photomicrograph (A) and drawing (B) showing internal structures. C, D. Radular teeth extracted from Benthofascis biconica (AMS C.468234, shell length of 15.2 mm), photomicrograph (C) and drawing (D) showing internal structures. E, F. Radular teeth extracted from Benthofascis pseudobiconica (AMS C.373099, shell length of 17.3 mm), photomicrograph (E) and drawing (F) showing internal structures. Scale bar represents 0.1 mm for AF. G. G. Electron micrographic study of the radular tooth of Benthofascis lozoueti Sysoev &amp; Bouchet, 2001, (MNHN IM200742331, 496 m deep, Norfolk Ridge, 22º50'S, 167º16'E, height 16.4 mm, width 7.0 mm, teeth from the same specimen shown in Fig. 3H). Scale bar dimensions are explained in the figure." figureDoi="http://doi.org/10.5281/zenodo.5290165" httpUri="https://zenodo.org/record/5290165/files/figure.png" pageId="4" pageNumber="5">Figs. 4A &amp; C</figureCitation>
). The tooth has a slight but distinct waist.
</paragraph>
</subSubSection>
<subSubSection id="F6D9D5D6F353FFFAFF15FD0BFE16EB57" pageId="4" pageNumber="5" type="discussion">
<paragraph id="BE7C865DF353FFFAFF15FD0BFB2BEE67" blockId="4.[151,1437,151,1615]" pageId="4" pageNumber="5">
<emphasis id="8CB75A4FF353FFFAFF15FD0BFE97EFD4" bold="true" box="[197,330,691,716]" inLineHeading="true" pageId="4" pageNumber="5" reason="1">Discussion.</emphasis>
<bibRefCitation id="DA52FBACF353FFFAFE83FD0CFE2AEFD4" author="Hedley, C." box="[339,503,692,716]" pageId="4" pageNumber="5" pagination="327 - 402" refId="ref9789" refString="Hedley, C. (1903) Scientific results of the trawling expedition of H. M. C. S. &quot; Thetis &quot; off the coast of New South Wales, in February and March, 1896. Mollusca. Part II. Scaphopoda and Gastropoda. Memoirs of the Australian Museum, 4, 327 - 402, pls. 36 - 37." type="journal article" year="1903">Hedley (1903)</bibRefCitation>
aptly described the
<typeStatus id="617838FFF353FFFAFD32FD0CFC99EFD4" box="[738,836,692,716]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
of this species (
<figureCitation id="26F89AD8F353FFFAFBD2FD0CFB56EFD4" box="[1026,1163,692,716]" captionStart="FIGURE 1" captionStartId="3.[151,250,1731,1754]" captionTargetBox="[284,1299,195,1716]" captionTargetId="figure-227@3.[284,1299,195,1717]" captionTargetPageId="3" captionText="FIGURE 1. AC. Holotype of Benthofascis biconica (AMS C.016448, 3537 m deep, Shoalhaven Bight, New South Wales, Australia), height 18 mm, width 8 mm, ventral (A) and side (B) views with enlargement of spire (C). DE. Benthofascis biconica (AMS C.468235, 5.7 km E of Mistral Point, Sydney, New South Wales, 33º56.470'S, 151º19.630'E, 7177 m deep), heigth 29.8 mm, width 10.7 mm, ventral view (D) and x-ray (E). F. Benthofascis biconica (AMS C.468235, same locality data as above), height 17.9 mm, width 7.7 mm. GH. Holotype of Benthofascis sarcinula (AMS C.19871, 203 m deep, 20 km miles E of Cape Byron, New South Wales, Australia), height 7 mm, width 4 mm, ventral view (G) and an enlargement of the spire (H). IK. Benthofascis sarcinula (USNM 902892, SSE of Lakes Entrance, Gippsland Lakes, Victoria, Australia), height 16.6 mm, width 7.4 mm, ventral (I) and lateral views (J) with an enlargement of the spire (K). LN. Benthofascis sarcinula (USNM 902893, Victoria, Australia), height 24.6 mm, width 10.7 mm, ventral (L) and lateral (M) views and an x-ray (N). Scale bars represent 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5290157" httpUri="https://zenodo.org/record/5290157/files/figure.png" pageId="4" pageNumber="5">Figs. 1AC</figureCitation>
).
<bibRefCitation id="DA52FBACF353FFFAFB70FD0CFA93EFD4" author="Laseron, C. F." box="[1184,1358,692,716]" pageId="4" pageNumber="5" refId="ref10053" refString="Laseron, C. F. (1954) Revision of the New South Wales Turridae. Royal Zoological Society of New South Wales Australian Zoological Handbook. Royal Zoological Society of New South Wales, Sydney, 56 pp., 12 pls." type="book" year="1954">Laseron (1954)</bibRefCitation>
reproduced Hedley's figure and correctly identified the species. However, some subsequent authors have misidentified specimens of
<taxonomicName id="79C3FDDEF353FFFAFEEAFD44FD59EE0C" authorityName="Tucker &amp; Tenorio &amp; Stahlschmidt" authorityYear="2011" box="[314,644,764,788]" class="Gastropoda" family="Conorbidae" genus="Benthofascis" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="pseudobiconica">
<emphasis id="8CB75A4FF353FFFAFEEAFD44FD59EE0C" box="[314,644,764,788]" italics="true" pageId="4" pageNumber="5">Benthofascis pseudobiconica</emphasis>
</taxonomicName>
as
<taxonomicName id="79C3FDDEF353FFFAFD62FD45FCEBEE0C" baseAuthorityName="Hedley" baseAuthorityYear="1903" box="[690,822,764,788]" class="Gastropoda" family="Conorbidae" genus="Benthofascis" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="biconica">
<emphasis id="8CB75A4FF353FFFAFD62FD45FCEBEE0C" box="[690,822,764,788]" italics="true" pageId="4" pageNumber="5">B. biconica</emphasis>
</taxonomicName>
(e.g.
<bibRefCitation id="DA52FBACF353FFFAFCAFFD44FBCEEE0C" author="Powell, A. W. B." box="[895,1043,764,788]" pageId="4" pageNumber="5" pagination="1 - 184" refId="ref10226" refString="Powell, A. W. B. (1966) The molluscan families Speightiidae and Turridae an evaluation of the valid taxa, both Recent and fossil, with lists of characteristic species. Bulletin of the Aukland Institute and Museum, 5, 1 - 184, 23 pls." type="journal article" year="1966">Powell 1966</bibRefCitation>
;
<bibRefCitation id="DA52FBACF353FFFAFBF4FD44FB67EE0C" author="Wilson, B." box="[1060,1210,764,788]" pageId="4" pageNumber="5" refId="ref10912" refString="Wilson, B. (1994) Australian Marine Shells. Vol. 2 (Prosobranch Gastropods Part Two (Neogastropods )). Odyssey Publishing, Kallaroo, Western Australia, 370 pp." type="book" year="1994">Wilson 1994</bibRefCitation>
;
<bibRefCitation id="DA52FBACF353FFFAFB1AFD44FF09EE2F" author="Tucker, J. K. &amp; Tenorio, M. J." pageId="4" pageNumber="5" refId="ref10740" refString="Tucker, J. K. &amp; Tenorio, M. J. (2009) Systematic Classification of Recent and Fossil Conoidean Gastropods, with Keys to the Genera of Cone Shells. ConchBooks, Hackenheim, Germany, 295 pp." type="book" year="2009">Tucker &amp; Tenorio 2009</bibRefCitation>
). These two species are similar to each other but differ in shell shape. The shoulders of
<taxonomicName id="79C3FDDEF353FFFAFB6EFC99FAE2EE20" baseAuthorityName="Hedley" baseAuthorityYear="1903" box="[1214,1343,800,824]" class="Gastropoda" family="Conorbidae" genus="Benthofascis" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="biconica">
<emphasis id="8CB75A4FF353FFFAFB6EFC99FAE2EE20" box="[1214,1343,800,824]" italics="true" pageId="4" pageNumber="5">B. biconica</emphasis>
</taxonomicName>
are distinctly angular (
<figureCitation id="26F89AD8F353FFFAFE99FCFCFE12EE43" box="[329,463,836,860]" captionStart="FIGURE 1" captionStartId="3.[151,250,1731,1754]" captionTargetBox="[284,1299,195,1716]" captionTargetId="figure-227@3.[284,1299,195,1717]" captionTargetPageId="3" captionText="FIGURE 1. AC. Holotype of Benthofascis biconica (AMS C.016448, 3537 m deep, Shoalhaven Bight, New South Wales, Australia), height 18 mm, width 8 mm, ventral (A) and side (B) views with enlargement of spire (C). DE. Benthofascis biconica (AMS C.468235, 5.7 km E of Mistral Point, Sydney, New South Wales, 33º56.470'S, 151º19.630'E, 7177 m deep), heigth 29.8 mm, width 10.7 mm, ventral view (D) and x-ray (E). F. Benthofascis biconica (AMS C.468235, same locality data as above), height 17.9 mm, width 7.7 mm. GH. Holotype of Benthofascis sarcinula (AMS C.19871, 203 m deep, 20 km miles E of Cape Byron, New South Wales, Australia), height 7 mm, width 4 mm, ventral view (G) and an enlargement of the spire (H). IK. Benthofascis sarcinula (USNM 902892, SSE of Lakes Entrance, Gippsland Lakes, Victoria, Australia), height 16.6 mm, width 7.4 mm, ventral (I) and lateral views (J) with an enlargement of the spire (K). LN. Benthofascis sarcinula (USNM 902893, Victoria, Australia), height 24.6 mm, width 10.7 mm, ventral (L) and lateral (M) views and an x-ray (N). Scale bars represent 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5290157" httpUri="https://zenodo.org/record/5290157/files/figure.png" pageId="4" pageNumber="5">Figs. 1A, D</figureCitation>
, &amp; F), whereas those of
<taxonomicName id="79C3FDDEF353FFFAFD32FCFDFC6CEE44" authorityName="Tucker &amp; Tenorio &amp; Stahlschmidt" authorityYear="2011" box="[738,945,836,860]" class="Gastropoda" family="Conorbidae" genus="Benthofascis" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="pseudobiconica">
<emphasis id="8CB75A4FF353FFFAFD32FCFDFC6CEE44" box="[738,945,836,860]" italics="true" pageId="4" pageNumber="5">B. pseudobiconica</emphasis>
</taxonomicName>
are not (
<figureCitation id="26F89AD8F353FFFAFBC5FCFCFB48EE44" box="[1045,1173,836,860]" captionStart="FIGURE 2" captionStartId="7.[151,250,1794,1817]" captionTargetBox="[266,1308,193,1756]" captionTargetId="figure-171@7.[266,1308,193,1756]" captionTargetPageId="7" captionText="FIGURE 2. AD. Holotype of Benthofascis angularis new species (MNHN 23067, 30 m deep, off Cape le Grande, Esperance, southwest Western Australia), height 27 mm, width 11.1 mm, ventral (A) and lateral (B) views with enlargement of spire (C) and an x-ray (D). E, F. Holotype of Benthofascis pseudobiconica new species (MNHN 23069, off Cape Moreton, Queensland, Australia), height 39.8mm, width 12.8 mm, ventral (E) and lateral (F) views. G.Paratype of Benthofascis pseudobiconica new species (ANSP 424978, 183 m deep, off Cape Moreton, S. Queensland, Australia), height 35.3 mm, width 11.8 mm, ventral view. HK. Paratype of Benthofascis pseudobiconica new species (USNM 902891, South Australia, Australia) height 36.4 mm, width 12.4 mm, ventral (H) and lateral (I) views with an enlargement of the spire (J) and an x-ray (K). Scale bars represent 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5290161" httpUri="https://zenodo.org/record/5290161/files/figure.png" pageId="4" pageNumber="5">Figs. 2E, G</figureCitation>
, &amp; H). The more angular shoulders give
<taxonomicName id="79C3FDDEF353FFFAFEB5FCD1FE38EE98" baseAuthorityName="Hedley" baseAuthorityYear="1903" box="[357,485,872,896]" class="Gastropoda" family="Conorbidae" genus="Benthofascis" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="biconica">
<emphasis id="8CB75A4FF353FFFAFEB5FCD1FE38EE98" box="[357,485,872,896]" italics="true" pageId="4" pageNumber="5">B. biconica</emphasis>
</taxonomicName>
a stouter appearance compared to the long-bodied
<taxonomicName id="79C3FDDEF353FFFAFBF1FCD1FB32EE98" authorityName="Tucker &amp; Tenorio &amp; Stahlschmidt" authorityYear="2011" box="[1057,1263,872,896]" class="Gastropoda" family="Conorbidae" genus="Benthofascis" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="pseudobiconica">
<emphasis id="8CB75A4FF353FFFAFBF1FCD1FB32EE98" box="[1057,1263,872,896]" italics="true" pageId="4" pageNumber="5">B. pseudobiconica</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="BE7C865DF353FFFAFF15FC34FACEE92C" blockId="4.[151,1437,151,1615]" pageId="4" pageNumber="5">
Two other species have subangular shoulders, namely
<taxonomicName id="79C3FDDEF353FFFAFCF5FC35FC72EEBC" authorityName="Tucker &amp; Tenorio &amp; Stahlschmidt" authorityYear="2011" box="[805,943,908,932]" class="Gastropoda" family="Conorbidae" genus="Benthofascis" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="angularis">
<emphasis id="8CB75A4FF353FFFAFCF5FC35FCEBEEBC" box="[805,822,909,932]" italics="true" pageId="4" pageNumber="5">B</emphasis>
.
<emphasis id="8CB75A4FF353FFFAFC94FC34FC72EEBC" box="[836,943,908,932]" italics="true" pageId="4" pageNumber="5">angularis</emphasis>
</taxonomicName>
(
<figureCitation id="26F89AD8F353FFFAFC6EFC34FB8AEEBC" box="[958,1111,908,932]" captionStart="FIGURE 2" captionStartId="7.[151,250,1794,1817]" captionTargetBox="[266,1308,193,1756]" captionTargetId="figure-171@7.[266,1308,193,1756]" captionTargetPageId="7" captionText="FIGURE 2. AD. Holotype of Benthofascis angularis new species (MNHN 23067, 30 m deep, off Cape le Grande, Esperance, southwest Western Australia), height 27 mm, width 11.1 mm, ventral (A) and lateral (B) views with enlargement of spire (C) and an x-ray (D). E, F. Holotype of Benthofascis pseudobiconica new species (MNHN 23069, off Cape Moreton, Queensland, Australia), height 39.8mm, width 12.8 mm, ventral (E) and lateral (F) views. G.Paratype of Benthofascis pseudobiconica new species (ANSP 424978, 183 m deep, off Cape Moreton, S. Queensland, Australia), height 35.3 mm, width 11.8 mm, ventral view. HK. Paratype of Benthofascis pseudobiconica new species (USNM 902891, South Australia, Australia) height 36.4 mm, width 12.4 mm, ventral (H) and lateral (I) views with an enlargement of the spire (J) and an x-ray (K). Scale bars represent 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5290161" httpUri="https://zenodo.org/record/5290161/files/figure.png" pageId="4" pageNumber="5">Figs. 2A &amp; C</figureCitation>
) and
<taxonomicName id="79C3FDDEF353FFFAFB45FC35FAD0EEBC" authorityName="Sysoev &amp; Bouchet" authorityYear="2001" box="[1173,1293,908,932]" class="Gastropoda" family="Conorbidae" genus="Benthofacies" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="lozoueti">
<emphasis id="8CB75A4FF353FFFAFB45FC35FAD0EEBC" box="[1173,1293,908,932]" italics="true" pageId="4" pageNumber="5">B. lozoueti</emphasis>
</taxonomicName>
(
<figureCitation id="26F89AD8F353FFFAFACCFC34FA4AEEBB" box="[1308,1431,908,932]" captionStart="FIGURE 3" captionStartId="10.[151,249,1608,1631]" captionTargetBox="[319,1260,177,1588]" captionTargetId="figure-275@10.[319,1260,177,1588]" captionTargetPageId="10" captionText="FIGURE 3. AC. Holotype of Benthofascis conorbioides new species (MNHN 23068, trawled 80120 m, off Mooloolaba, S. Queensland, Australia), height 38.7 mm, width 15.4 mm, ventral view (A), with an enlargement of the spire (B) and an x-ray (C). D. Paratype of Benthofascis conorbioides new species (SBMNH 424097, trawled on sand and rubble, 80120 m, off Mooloolaba, S. Queensland, Australia) height 41.8 mm, width 17.0 mm, ventral view. E. Holotype of Benthofascis atractoides (SAMA T750A, Kent Town Bore, Clayey greensands, Janjukian?, L. Miocene), height 16.5 mm, width 6.5 mm (reproduced from Long, 1981, Pl. 6, Fig. 14, with permission), ventral view. F. Holotype of Benthofascis otwayensis (NMVM P42959, Point Flinders, Cape Otway, Victoria, Australia. Glen Aire Clay, U. Aldingan, L. Oligocene), height 19.75 mm, width 8.75 mm (reproduced from Long, 1981, Pl. 6, Fig. 15, with permission), ventral view. G. Paratype of Benthofascis lozoueti (NMSA L5679/T1867), South New Caledonia, N/O &quot;Alis&quot; campaign, Stn. DW 721, 22º54'S, 167º17'E, height 16.5 mm, width 7.2 mm, ventral view. H. Benthofascis lozoueti (MNHN IM200742331, 496 m deep, Norfolk Ridge, 22º50'S, 167º16'E), length 16.4 mm, width 7.0 mm, x-ray. IK. Holotype of Benthofascis lozoueti (MNHN 3053, 530 m deep, Norfolk Ridge, 22º58'S, 167º20'E), height 21.5 mm, width 15 mm, ventral (I) and lateral (J) views with an enlargement of the spire (K). L. Paratype of Benthofascis lozoueti (AMS C.415308), 525574 m deep, New Caledonia, 22º54'S, 167º17'E, height 17.5 mm, width 8.0 mm, ventral view. Scale bars represent 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5290163" httpUri="https://zenodo.org/record/5290163/files/figure.png" pageId="4" pageNumber="5">Figs. 3I, K</figureCitation>
, &amp; L). The latter species has a nearly smooth protoconch with more than 2 whorls (
<figureCitation id="26F89AD8F353FFFAFBE0FC17FB5DEEDF" box="[1072,1152,943,967]" captionStart="FIGURE 3" captionStartId="10.[151,249,1608,1631]" captionTargetBox="[319,1260,177,1588]" captionTargetId="figure-275@10.[319,1260,177,1588]" captionTargetPageId="10" captionText="FIGURE 3. AC. Holotype of Benthofascis conorbioides new species (MNHN 23068, trawled 80120 m, off Mooloolaba, S. Queensland, Australia), height 38.7 mm, width 15.4 mm, ventral view (A), with an enlargement of the spire (B) and an x-ray (C). D. Paratype of Benthofascis conorbioides new species (SBMNH 424097, trawled on sand and rubble, 80120 m, off Mooloolaba, S. Queensland, Australia) height 41.8 mm, width 17.0 mm, ventral view. E. Holotype of Benthofascis atractoides (SAMA T750A, Kent Town Bore, Clayey greensands, Janjukian?, L. Miocene), height 16.5 mm, width 6.5 mm (reproduced from Long, 1981, Pl. 6, Fig. 14, with permission), ventral view. F. Holotype of Benthofascis otwayensis (NMVM P42959, Point Flinders, Cape Otway, Victoria, Australia. Glen Aire Clay, U. Aldingan, L. Oligocene), height 19.75 mm, width 8.75 mm (reproduced from Long, 1981, Pl. 6, Fig. 15, with permission), ventral view. G. Paratype of Benthofascis lozoueti (NMSA L5679/T1867), South New Caledonia, N/O &quot;Alis&quot; campaign, Stn. DW 721, 22º54'S, 167º17'E, height 16.5 mm, width 7.2 mm, ventral view. H. Benthofascis lozoueti (MNHN IM200742331, 496 m deep, Norfolk Ridge, 22º50'S, 167º16'E), length 16.4 mm, width 7.0 mm, x-ray. IK. Holotype of Benthofascis lozoueti (MNHN 3053, 530 m deep, Norfolk Ridge, 22º58'S, 167º20'E), height 21.5 mm, width 15 mm, ventral (I) and lateral (J) views with an enlargement of the spire (K). L. Paratype of Benthofascis lozoueti (AMS C.415308), 525574 m deep, New Caledonia, 22º54'S, 167º17'E, height 17.5 mm, width 8.0 mm, ventral view. Scale bars represent 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5290163" httpUri="https://zenodo.org/record/5290163/files/figure.png" pageId="4" pageNumber="5">Fig. 3J</figureCitation>
), a feature unique among the
<taxonomicName id="79C3FDDEF353FFFAFF10FC6CFE72EEF4" box="[192,431,980,1004]" class="Gastropoda" family="Conorbidae" genus="Benthofascis" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="8CB75A4FF353FFFAFF10FC6CFE92EEF4" box="[192,335,980,1004]" italics="true" pageId="4" pageNumber="5">Benthofascis</emphasis>
species.
</taxonomicName>
Besides that
<taxonomicName id="79C3FDDEF353FFFAFD94FC6DFD61EEF4" authorityName="Sysoev &amp; Bouchet" authorityYear="2001" box="[580,700,980,1004]" class="Gastropoda" family="Conorbidae" genus="Benthofacies" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="lozoueti">
<emphasis id="8CB75A4FF353FFFAFD94FC6DFD61EEF4" box="[580,700,980,1004]" italics="true" pageId="4" pageNumber="5">B. lozoueti</emphasis>
</taxonomicName>
is unique among
<taxonomicName id="79C3FDDEF353FFFAFC53FC6CFBCFEEF4" authorityName="Iredale" authorityYear="1936" box="[899,1042,980,1004]" class="Gastropoda" family="Conorbidae" genus="Benthofascis" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="5" phylum="Mollusca" rank="genus">
<emphasis id="8CB75A4FF353FFFAFC53FC6CFBCFEEF4" box="[899,1042,980,1004]" italics="true" pageId="4" pageNumber="5">Benthofascis</emphasis>
</taxonomicName>
in that the inner shell walls are not remodeled (
<figureCitation id="26F89AD8F353FFFAFECEFC4FFEA7E917" box="[286,378,1015,1039]" captionStart="FIGURE 3" captionStartId="10.[151,249,1608,1631]" captionTargetBox="[319,1260,177,1588]" captionTargetId="figure-275@10.[319,1260,177,1588]" captionTargetPageId="10" captionText="FIGURE 3. AC. Holotype of Benthofascis conorbioides new species (MNHN 23068, trawled 80120 m, off Mooloolaba, S. Queensland, Australia), height 38.7 mm, width 15.4 mm, ventral view (A), with an enlargement of the spire (B) and an x-ray (C). D. Paratype of Benthofascis conorbioides new species (SBMNH 424097, trawled on sand and rubble, 80120 m, off Mooloolaba, S. Queensland, Australia) height 41.8 mm, width 17.0 mm, ventral view. E. Holotype of Benthofascis atractoides (SAMA T750A, Kent Town Bore, Clayey greensands, Janjukian?, L. Miocene), height 16.5 mm, width 6.5 mm (reproduced from Long, 1981, Pl. 6, Fig. 14, with permission), ventral view. F. Holotype of Benthofascis otwayensis (NMVM P42959, Point Flinders, Cape Otway, Victoria, Australia. Glen Aire Clay, U. Aldingan, L. Oligocene), height 19.75 mm, width 8.75 mm (reproduced from Long, 1981, Pl. 6, Fig. 15, with permission), ventral view. G. Paratype of Benthofascis lozoueti (NMSA L5679/T1867), South New Caledonia, N/O &quot;Alis&quot; campaign, Stn. DW 721, 22º54'S, 167º17'E, height 16.5 mm, width 7.2 mm, ventral view. H. Benthofascis lozoueti (MNHN IM200742331, 496 m deep, Norfolk Ridge, 22º50'S, 167º16'E), length 16.4 mm, width 7.0 mm, x-ray. IK. Holotype of Benthofascis lozoueti (MNHN 3053, 530 m deep, Norfolk Ridge, 22º58'S, 167º20'E), height 21.5 mm, width 15 mm, ventral (I) and lateral (J) views with an enlargement of the spire (K). L. Paratype of Benthofascis lozoueti (AMS C.415308), 525574 m deep, New Caledonia, 22º54'S, 167º17'E, height 17.5 mm, width 8.0 mm, ventral view. Scale bars represent 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5290163" httpUri="https://zenodo.org/record/5290163/files/figure.png" pageId="4" pageNumber="5">Fig. 3H</figureCitation>
). We were able to confirm via x-ray that all other species of
<taxonomicName id="79C3FDDEF353FFFAFBEAFC40FB14E908" authorityName="Iredale" authorityYear="1936" box="[1082,1225,1016,1040]" class="Gastropoda" family="Conorbidae" genus="Benthofascis" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="5" phylum="Mollusca" rank="genus">
<emphasis id="8CB75A4FF353FFFAFBEAFC40FB14E908" box="[1082,1225,1016,1040]" italics="true" pageId="4" pageNumber="5">Benthofascis</emphasis>
</taxonomicName>
remodel the inner shell walls.
<taxonomicName id="79C3FDDEF353FFFAFECCFBA5FE7BE92C" authorityName="Tucker &amp; Tenorio &amp; Stahlschmidt" authorityYear="2011" box="[284,422,1052,1076]" class="Gastropoda" family="Conorbidae" genus="Benthofascis" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="angularis">
<emphasis id="8CB75A4FF353FFFAFECCFBA5FEF0E92C" box="[284,301,1053,1076]" italics="true" pageId="4" pageNumber="5">B</emphasis>
.
<emphasis id="8CB75A4FF353FFFAFEEBFBA4FE7BE92C" box="[315,422,1052,1076]" italics="true" pageId="4" pageNumber="5">angularis</emphasis>
</taxonomicName>
does not have the punctate sutural ramps similar to those that
<taxonomicName id="79C3FDDEF353FFFAFBB0FBA5FB3DE92C" baseAuthorityName="Hedley" baseAuthorityYear="1903" box="[1120,1248,1052,1076]" class="Gastropoda" family="Conorbidae" genus="Benthofascis" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="biconica">
<emphasis id="8CB75A4FF353FFFAFBB0FBA5FBACE92C" box="[1120,1137,1053,1076]" italics="true" pageId="4" pageNumber="5">B</emphasis>
.
<emphasis id="8CB75A4FF353FFFAFB50FBA4FB3DE92C" box="[1152,1248,1052,1076]" italics="true" pageId="4" pageNumber="5">biconica</emphasis>
</taxonomicName>
has.
</paragraph>
<paragraph id="BE7C865DF353FFFAFF15FBF8FB41E9DC" blockId="4.[151,1437,151,1615]" pageId="4" pageNumber="5">
The body whorl ornamentation of
<taxonomicName id="79C3FDDEF353FFFAFD9CFBF9FD22E940" authorityName="Tucker &amp; Tenorio &amp; Stahlschmidt" authorityYear="2011" box="[588,767,1088,1112]" class="Gastropoda" family="Conorbidae" genus="Benthofascis" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="conorbioides">
<emphasis id="8CB75A4FF353FFFAFD9CFBF9FD22E940" box="[588,767,1088,1112]" italics="true" pageId="4" pageNumber="5">B. conorbioides</emphasis>
</taxonomicName>
differs from that of
<taxonomicName id="79C3FDDEF353FFFAFC39FBF9FBB7E940" baseAuthorityName="Hedley" baseAuthorityYear="1903" box="[1001,1130,1088,1112]" class="Gastropoda" family="Conorbidae" genus="Benthofascis" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="biconica">
<emphasis id="8CB75A4FF353FFFAFC39FBF9FBB7E940" box="[1001,1130,1088,1112]" italics="true" pageId="4" pageNumber="5">B. biconica</emphasis>
</taxonomicName>
. The former species (
<figureCitation id="26F89AD8F353FFFAFAB4FBF8FF01E963" captionStart="FIGURE 3" captionStartId="10.[151,249,1608,1631]" captionTargetBox="[319,1260,177,1588]" captionTargetId="figure-275@10.[319,1260,177,1588]" captionTargetPageId="10" captionText="FIGURE 3. AC. Holotype of Benthofascis conorbioides new species (MNHN 23068, trawled 80120 m, off Mooloolaba, S. Queensland, Australia), height 38.7 mm, width 15.4 mm, ventral view (A), with an enlargement of the spire (B) and an x-ray (C). D. Paratype of Benthofascis conorbioides new species (SBMNH 424097, trawled on sand and rubble, 80120 m, off Mooloolaba, S. Queensland, Australia) height 41.8 mm, width 17.0 mm, ventral view. E. Holotype of Benthofascis atractoides (SAMA T750A, Kent Town Bore, Clayey greensands, Janjukian?, L. Miocene), height 16.5 mm, width 6.5 mm (reproduced from Long, 1981, Pl. 6, Fig. 14, with permission), ventral view. F. Holotype of Benthofascis otwayensis (NMVM P42959, Point Flinders, Cape Otway, Victoria, Australia. Glen Aire Clay, U. Aldingan, L. Oligocene), height 19.75 mm, width 8.75 mm (reproduced from Long, 1981, Pl. 6, Fig. 15, with permission), ventral view. G. Paratype of Benthofascis lozoueti (NMSA L5679/T1867), South New Caledonia, N/O &quot;Alis&quot; campaign, Stn. DW 721, 22º54'S, 167º17'E, height 16.5 mm, width 7.2 mm, ventral view. H. Benthofascis lozoueti (MNHN IM200742331, 496 m deep, Norfolk Ridge, 22º50'S, 167º16'E), length 16.4 mm, width 7.0 mm, x-ray. IK. Holotype of Benthofascis lozoueti (MNHN 3053, 530 m deep, Norfolk Ridge, 22º58'S, 167º20'E), height 21.5 mm, width 15 mm, ventral (I) and lateral (J) views with an enlargement of the spire (K). L. Paratype of Benthofascis lozoueti (AMS C.415308), 525574 m deep, New Caledonia, 22º54'S, 167º17'E, height 17.5 mm, width 8.0 mm, ventral view. Scale bars represent 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5290163" httpUri="https://zenodo.org/record/5290163/files/figure.png" pageId="4" pageNumber="5">Figs. 3A, B</figureCitation>
, &amp; D) and the two fossil species (
<figureCitation id="26F89AD8F353FFFAFD85FBDCFD3AE964" box="[597,743,1124,1148]" captionStart="FIGURE 3" captionStartId="10.[151,249,1608,1631]" captionTargetBox="[319,1260,177,1588]" captionTargetId="figure-275@10.[319,1260,177,1588]" captionTargetPageId="10" captionText="FIGURE 3. AC. Holotype of Benthofascis conorbioides new species (MNHN 23068, trawled 80120 m, off Mooloolaba, S. Queensland, Australia), height 38.7 mm, width 15.4 mm, ventral view (A), with an enlargement of the spire (B) and an x-ray (C). D. Paratype of Benthofascis conorbioides new species (SBMNH 424097, trawled on sand and rubble, 80120 m, off Mooloolaba, S. Queensland, Australia) height 41.8 mm, width 17.0 mm, ventral view. E. Holotype of Benthofascis atractoides (SAMA T750A, Kent Town Bore, Clayey greensands, Janjukian?, L. Miocene), height 16.5 mm, width 6.5 mm (reproduced from Long, 1981, Pl. 6, Fig. 14, with permission), ventral view. F. Holotype of Benthofascis otwayensis (NMVM P42959, Point Flinders, Cape Otway, Victoria, Australia. Glen Aire Clay, U. Aldingan, L. Oligocene), height 19.75 mm, width 8.75 mm (reproduced from Long, 1981, Pl. 6, Fig. 15, with permission), ventral view. G. Paratype of Benthofascis lozoueti (NMSA L5679/T1867), South New Caledonia, N/O &quot;Alis&quot; campaign, Stn. DW 721, 22º54'S, 167º17'E, height 16.5 mm, width 7.2 mm, ventral view. H. Benthofascis lozoueti (MNHN IM200742331, 496 m deep, Norfolk Ridge, 22º50'S, 167º16'E), length 16.4 mm, width 7.0 mm, x-ray. IK. Holotype of Benthofascis lozoueti (MNHN 3053, 530 m deep, Norfolk Ridge, 22º58'S, 167º20'E), height 21.5 mm, width 15 mm, ventral (I) and lateral (J) views with an enlargement of the spire (K). L. Paratype of Benthofascis lozoueti (AMS C.415308), 525574 m deep, New Caledonia, 22º54'S, 167º17'E, height 17.5 mm, width 8.0 mm, ventral view. Scale bars represent 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5290163" httpUri="https://zenodo.org/record/5290163/files/figure.png" pageId="4" pageNumber="5">Figs. 3E &amp; F</figureCitation>
) have spiral grooves that are widely spaced and that cover the entire body whorl rather than the more closely spaced grooves of
<taxonomicName id="79C3FDDEF353FFFAFC40FB31FBCFE9B8" baseAuthorityName="Hedley" baseAuthorityYear="1903" box="[912,1042,1160,1184]" class="Gastropoda" family="Conorbidae" genus="Benthofascis" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="biconica">
<emphasis id="8CB75A4FF353FFFAFC40FB31FBCFE9B8" box="[912,1042,1160,1184]" italics="true" pageId="4" pageNumber="5">B. biconica</emphasis>
</taxonomicName>
and
<taxonomicName id="79C3FDDEF353FFFAFB9DFB31FAC2E9B8" authorityName="Tucker &amp; Tenorio &amp; Stahlschmidt" authorityYear="2011" box="[1101,1311,1160,1184]" class="Gastropoda" family="Conorbidae" genus="Benthofascis" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="pseudobiconica">
<emphasis id="8CB75A4FF353FFFAFB9DFB31FAC2E9B8" box="[1101,1311,1160,1184]" italics="true" pageId="4" pageNumber="5">B. pseudobiconica</emphasis>
</taxonomicName>
. The body whorls of
<taxonomicName id="79C3FDDEF353FFFAFED8FB15FE5DE9DC" authorityName="Sysoev &amp; Bouchet" authorityYear="2001" box="[264,384,1196,1220]" class="Gastropoda" family="Conorbidae" genus="Benthofacies" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="lozoueti">
<emphasis id="8CB75A4FF353FFFAFED8FB15FE5DE9DC" box="[264,384,1196,1220]" italics="true" pageId="4" pageNumber="5">B. lozoueti</emphasis>
</taxonomicName>
(
<figureCitation id="26F89AD8F353FFFAFE5FFB14FDD6E9DB" box="[399,523,1196,1220]" captionStart="FIGURE 3" captionStartId="10.[151,249,1608,1631]" captionTargetBox="[319,1260,177,1588]" captionTargetId="figure-275@10.[319,1260,177,1588]" captionTargetPageId="10" captionText="FIGURE 3. AC. Holotype of Benthofascis conorbioides new species (MNHN 23068, trawled 80120 m, off Mooloolaba, S. Queensland, Australia), height 38.7 mm, width 15.4 mm, ventral view (A), with an enlargement of the spire (B) and an x-ray (C). D. Paratype of Benthofascis conorbioides new species (SBMNH 424097, trawled on sand and rubble, 80120 m, off Mooloolaba, S. Queensland, Australia) height 41.8 mm, width 17.0 mm, ventral view. E. Holotype of Benthofascis atractoides (SAMA T750A, Kent Town Bore, Clayey greensands, Janjukian?, L. Miocene), height 16.5 mm, width 6.5 mm (reproduced from Long, 1981, Pl. 6, Fig. 14, with permission), ventral view. F. Holotype of Benthofascis otwayensis (NMVM P42959, Point Flinders, Cape Otway, Victoria, Australia. Glen Aire Clay, U. Aldingan, L. Oligocene), height 19.75 mm, width 8.75 mm (reproduced from Long, 1981, Pl. 6, Fig. 15, with permission), ventral view. G. Paratype of Benthofascis lozoueti (NMSA L5679/T1867), South New Caledonia, N/O &quot;Alis&quot; campaign, Stn. DW 721, 22º54'S, 167º17'E, height 16.5 mm, width 7.2 mm, ventral view. H. Benthofascis lozoueti (MNHN IM200742331, 496 m deep, Norfolk Ridge, 22º50'S, 167º16'E), length 16.4 mm, width 7.0 mm, x-ray. IK. Holotype of Benthofascis lozoueti (MNHN 3053, 530 m deep, Norfolk Ridge, 22º58'S, 167º20'E), height 21.5 mm, width 15 mm, ventral (I) and lateral (J) views with an enlargement of the spire (K). L. Paratype of Benthofascis lozoueti (AMS C.415308), 525574 m deep, New Caledonia, 22º54'S, 167º17'E, height 17.5 mm, width 8.0 mm, ventral view. Scale bars represent 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5290163" httpUri="https://zenodo.org/record/5290163/files/figure.png" pageId="4" pageNumber="5">Figs. 3I, K</figureCitation>
, &amp; L) and
<taxonomicName id="79C3FDDEF353FFFAFD54FB14FCD2E9DC" authorityName="Tucker &amp; Tenorio &amp; Stahlschmidt" authorityYear="2011" box="[644,783,1196,1220]" class="Gastropoda" family="Conorbidae" genus="Benthofascis" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="13" phylum="Mollusca" rank="species" species="angularis">
B.
<emphasis id="8CB75A4FF353FFFAFD74FB14FCD2E9DC" box="[676,783,1196,1220]" italics="true" pageId="4" pageNumber="5">angularis</emphasis>
</taxonomicName>
(
<figureCitation id="26F89AD8F353FFFAFCCFFB14FC64E9DC" box="[799,953,1196,1220]" captionStart="FIGURE 2" captionStartId="7.[151,250,1794,1817]" captionTargetBox="[266,1308,193,1756]" captionTargetId="figure-171@7.[266,1308,193,1756]" captionTargetPageId="7" captionText="FIGURE 2. AD. Holotype of Benthofascis angularis new species (MNHN 23067, 30 m deep, off Cape le Grande, Esperance, southwest Western Australia), height 27 mm, width 11.1 mm, ventral (A) and lateral (B) views with enlargement of spire (C) and an x-ray (D). E, F. Holotype of Benthofascis pseudobiconica new species (MNHN 23069, off Cape Moreton, Queensland, Australia), height 39.8mm, width 12.8 mm, ventral (E) and lateral (F) views. G.Paratype of Benthofascis pseudobiconica new species (ANSP 424978, 183 m deep, off Cape Moreton, S. Queensland, Australia), height 35.3 mm, width 11.8 mm, ventral view. HK. Paratype of Benthofascis pseudobiconica new species (USNM 902891, South Australia, Australia) height 36.4 mm, width 12.4 mm, ventral (H) and lateral (I) views with an enlargement of the spire (J) and an x-ray (K). Scale bars represent 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5290161" httpUri="https://zenodo.org/record/5290161/files/figure.png" pageId="4" pageNumber="5">Figs. 2A &amp; C</figureCitation>
) are almost smooth.
</paragraph>
<paragraph id="BE7C865DF353FFFAFF15FB68FE16EB57" blockId="4.[151,1437,151,1615]" pageId="4" pageNumber="5">
We illustrate the radulae of two of these species with micrographs and with drawings that show the internal structure of the teeth for the first time. Previous drawings (e.g.,
<bibRefCitation id="DA52FBACF353FFFAFCBDFB4CFBDEE814" author="Powell, A. W. B." box="[877,1027,1268,1292]" pageId="4" pageNumber="5" pagination="1 - 184" refId="ref10226" refString="Powell, A. W. B. (1966) The molluscan families Speightiidae and Turridae an evaluation of the valid taxa, both Recent and fossil, with lists of characteristic species. Bulletin of the Aukland Institute and Museum, 5, 1 - 184, 23 pls." type="journal article" year="1966">Powell, 1966</bibRefCitation>
) were misleading as they suggested that the teeth were simple hollow tubes with no internal structure (see
<bibRefCitation id="DA52FBACF353FFFAFC6DFAA0FB0BE828" author="Tucker, J. K. &amp; Tenorio, M. J." box="[957,1238,1304,1328]" pageId="4" pageNumber="5" refId="ref10740" refString="Tucker, J. K. &amp; Tenorio, M. J. (2009) Systematic Classification of Recent and Fossil Conoidean Gastropods, with Keys to the Genera of Cone Shells. ConchBooks, Hackenheim, Germany, 295 pp." type="book" year="2009">Tucker &amp; Tenorio, 2009</bibRefCitation>
). Electron micrographs available for
<taxonomicName id="79C3FDDEF353FFFAFEAFFA85FCF0E84C" authority="(Sysoev &amp; Bouchet, 2001)" baseAuthorityName="Sysoev &amp; Bouchet" baseAuthorityYear="2001" box="[383,813,1340,1364]" class="Gastropoda" family="Conorbidae" genus="Benthofacies" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="lozoueti">
<emphasis id="8CB75A4FF353FFFAFEAFFA85FE25E84C" box="[383,504,1340,1364]" italics="true" pageId="4" pageNumber="5">B. lozoueti</emphasis>
(
<bibRefCitation id="DA52FBACF353FFFAFDD7FA84FCF8E84C" author="Sysoev, A. V. &amp; Bouchet, P." box="[519,805,1340,1364]" pageId="4" pageNumber="5" pagination="271 - 320" refId="ref10507" refString="Sysoev, A. V. &amp; Bouchet, P. (2001) New and uncommon turriform gastropods (Gastropoda: Conoidea) from the South-West Pacific. Memoires du Museum National d'Histoire Naturelle, Paris, serie A, Zoologie, 185, 271 - 320." type="journal article" year="2001">Sysoev &amp; Bouchet, 2001</bibRefCitation>
)
</taxonomicName>
also do not show internal structures. The radulae of
<taxonomicName id="79C3FDDEF353FFFAFA54FA85FF2AE860" baseAuthorityName="Hedley" baseAuthorityYear="1903" class="Gastropoda" family="Conorbidae" genus="Benthofascis" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="biconica">
<emphasis id="8CB75A4FF353FFFAFA54FA85FF2AE860" italics="true" pageId="4" pageNumber="5">B. biconica</emphasis>
</taxonomicName>
and
<taxonomicName id="79C3FDDEF353FFFAFEFDFAD9FE26E860" authorityName="Tucker &amp; Tenorio &amp; Stahlschmidt" authorityYear="2011" box="[301,507,1376,1400]" class="Gastropoda" family="Conorbidae" genus="Benthofascis" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="pseudobiconica">
<emphasis id="8CB75A4FF353FFFAFEFDFAD9FE26E860" box="[301,507,1376,1400]" italics="true" pageId="4" pageNumber="5">B. pseudobiconica</emphasis>
</taxonomicName>
that we were able to examine had unexpected structures. There is a definite C-fold and a slight waist both features that were not obvious in previously published drawings of radulae from
<taxonomicName id="79C3FDDEF353FFFAFAF3FA3CFF6BE8A7" authorityName="Iredale" authorityYear="1936" class="Gastropoda" family="Conorbidae" genus="Benthofascis" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="5" phylum="Mollusca" rank="genus">
<emphasis id="8CB75A4FF353FFFAFAF3FA3CFF6BE8A7" italics="true" pageId="4" pageNumber="5">Benthofascis</emphasis>
</taxonomicName>
. This is important because it demonstrates that the tooth is enrolled more than 360 degrees. Moreover a fold is obvious along the shaft of the tooth in all three species. This fold may be homologous to the shaft fold that characterizes the
<taxonomicName id="79C3FDDEF353FFFAFEC2FA48FD11EB10" authority="(Tucker &amp; Tenorio, 2009)" baseAuthorityName="Tucker &amp; Tenorio" baseAuthorityYear="2009" box="[274,716,1520,1544]" class="Gastropoda" family="Conilithidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="5" phylum="Mollusca" rank="family">
Conilithidae (
<bibRefCitation id="DA52FBACF353FFFAFE7BFA48FD19EB10" author="Tucker, J. K. &amp; Tenorio, M. J." box="[427,708,1520,1544]" pageId="4" pageNumber="5" refId="ref10740" refString="Tucker, J. K. &amp; Tenorio, M. J. (2009) Systematic Classification of Recent and Fossil Conoidean Gastropods, with Keys to the Genera of Cone Shells. ConchBooks, Hackenheim, Germany, 295 pp." type="book" year="2009">Tucker &amp; Tenorio, 2009</bibRefCitation>
)
</taxonomicName>
. However, this fold is not a shaft fold because it appears that in
<taxonomicName id="79C3FDDEF353FFFAFF48F9ACFEFAEB34" authorityName="Iredale" authorityYear="1936" box="[152,295,1556,1580]" class="Gastropoda" family="Conorbidae" genus="Benthofascis" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="5" phylum="Mollusca" rank="genus">
<emphasis id="8CB75A4FF353FFFAFF48F9ACFEFAEB34" box="[152,295,1556,1580]" italics="true" pageId="4" pageNumber="5">Benthofascis</emphasis>
</taxonomicName>
this fold is an internal one associated with the enrollment of the tooth. The shaft fold of the
<taxonomicName id="79C3FDDEF353FFFAFAE0F9ACFF63EB57" baseAuthorityName="Tucker &amp; Tenorio" baseAuthorityYear="2009" class="Gastropoda" family="Conilithidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="5" phylum="Mollusca" rank="family">Conilithidae</taxonomicName>
is an external structure.
</paragraph>
</subSubSection>
</treatment>
</document>