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<document id="851D73AF2424F8964F35A71657F8775E" ID-DOI="10.11646/zootaxa.3734.5.1" ID-GBIF-Dataset="51673b52-5717-4920-846b-f1c560357734" ID-ISSN="1175-5326" ID-Zenodo-Dep="219026" ID-ZooBank="2C4411BF-233F-48F2-BA6F-423A9F8EE8B1" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460696948925" checkinUser="plazi" docAuthor="Vagalinski, Boyan, Lazányi, Eszter &amp; Golovatch, Sergei" docDate="2013" docId="323D6308FFBBF304FF2FFD4C5CA7FF43" docLanguage="en" docName="zt03734p520.pdf" docOrigin="Zootaxa 3734 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Megaphyllum (Parancistrum) serratum Verhoeff 1898" docType="treatment" docVersion="10" lastPageNumber="510" masterDocId="CE041B70FFBCF30DFFB8FFFF5865FFBB" masterDocTitle="Redescription of the subgenus Parancistrum Verhoeff, 1943, an eastern Mediterranean lineage of the millipede genus Megaphyllum Verhoeff, 1894 (Diplopoda: Julida: Julidae: Brachyiulini)" masterLastPageNumber="520" masterPageNumber="501" pageNumber="508" updateTime="1698353915693" updateUser="plazi">
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<mods:title id="BA6322BE1E523B4F17A2AE136BDEE829">Redescription of the subgenus Parancistrum Verhoeff, 1943, an eastern Mediterranean lineage of the millipede genus Megaphyllum Verhoeff, 1894 (Diplopoda: Julida: Julidae: Brachyiulini)</mods:title>
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<mods:namePart id="22A798F8ABE3F656A0DE29CFD30A58CA">Vagalinski, Boyan</mods:namePart>
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<treatment id="323D6308FFBBF304FF2FFD4C5CA7FF43" ID-DOI="http://doi.org/10.5281/zenodo.5672398" ID-GBIF-Taxon="119581922" ID-Zenodo-Dep="5672398" LSID="urn:lsid:plazi:treatment:323D6308FFBBF304FF2FFD4C5CA7FF43" httpUri="http://treatment.plazi.org/id/323D6308FFBBF304FF2FFD4C5CA7FF43" lastPageId="9" lastPageNumber="510" pageId="7" pageNumber="508">
<subSubSection id="F28E8195FFBBF30AFF2FFD4C5A26FCE8" pageId="7" pageNumber="508" type="nomenclature">
<paragraph id="BA2BD21EFFBBF30AFF2FFD4C5B26FD75" blockId="7.[151,835,691,751]" box="[151,835,691,718]" pageId="7" pageNumber="508">
<heading id="E1636572FFBBF30AFF2FFD4C5B26FD75" bold="true" box="[151,835,691,718]" fontSize="11" level="1" pageId="7" pageNumber="508" reason="1">
<taxonomicName id="7D94A99DFFBBF30AFF2FFD4C5B26FD75" ID-CoL="8KR6D" authority="Verhoeff, 1898" authorityName="Verhoeff" authorityYear="1898" box="[151,835,691,718]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="serratum" subGenus="Parancistrum">
<emphasis id="88E00E0CFFBBF30AFF2FFD4C5B26FD75" bold="true" box="[151,835,691,718]" pageId="7" pageNumber="508">
<emphasis id="88E00E0CFFBBF30AFF2FFD4C5A11FD76" bold="true" box="[151,628,691,717]" italics="true" pageId="7" pageNumber="508">Megaphyllum (Parancistrum) serratum</emphasis>
(Verhoeff, 1898)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="BA2BD21EFFBBF30AFF2FFD295971FD54" blockId="7.[151,835,691,751]" box="[151,276,726,751]" pageId="7" pageNumber="508">
<figureCitation id="22AFCE9BFFBBF30AFF2FFD295971FD54" box="[151,276,726,751]" captionStart="FIGURES 19 22" captionStartId="8.[151,269,1756,1779]" captionTargetBox="[174,1410,203,1724]" captionTargetId="figure@8.[170,1417,195,1732]" captionTargetPageId="8" captionText="FIGURES 19 22. Megaphyllum serratum syntypes, male (19 21) and female (22): (19) right gonopods, mesal, slightly anterior view; (20) right promere, posterior view; (21) right opisthomere, meso-anterior view; (22) right vulva, lateroposterior view (setae omitted, receptaculum seen by transparency, but drawn with solid lines for better emphasis). Abbreviations: ap: anterior process, asp: anterior protrusion of the solenomere, at: apodematic tube, ch: central hump, ct: connecting tube, fc: flagellum channel, fl: flagellum, g: median groove, lh: lateral hump, mh: mesal hump, P: promere, pa: posterior ampulla, ph: posterior hump, pp: posterior process, r: median ridge, s: solenomere, sf: spine-like filaments. Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/219030/files/figure.png" pageId="7" pageNumber="508">Figs 1922</figureCitation>
</paragraph>
<paragraph id="BA2BD21EFFBBF30AFF2FFCE15A26FCE8" blockId="7.[151,789,798,943]" pageId="7" pageNumber="508">
<taxonomicName id="7D94A99DFFBBF30AFF2FFCE15A59FC8F" authority="Verhoeff, 1898: 297" authorityName="Verhoeff" authorityPageNumber="297" authorityYear="1898" box="[151,572,798,821]" class="Diplopoda" family="Julidae" genus="Brachyiulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="serratus">
<emphasis id="88E00E0CFFBBF30AFF2FFCE1590FFC8E" box="[151,362,798,821]" italics="true" pageId="7" pageNumber="508">Brachyiulus serratus</emphasis>
Verhoeff, 1898: 297
</taxonomicName>
298, fig. 7.
<taxonomicName id="7D94A99DFFBBF30AFF2FFCC25A5AFCE8" authority="Verhoeff 1923: 131" authorityName="Verhoeff" authorityPageNumber="131" authorityYear="1923" box="[151,575,829,851]" class="Diplopoda" family="Julidae" genus="Brachyiulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="serratus">
<emphasis id="88E00E0CFFBBF30AFF2FFCC2590FFCE8" box="[151,362,829,851]" italics="true" pageId="7" pageNumber="508">Brachyiulus serratus</emphasis>
: Verhoeff 1923: 131
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="F28E8195FFBBF30AFF2FFCA35A33FC14" pageId="7" pageNumber="508" type="description">
<paragraph id="BA2BD21EFFBBF30AFF2FFCA35A33FC14" blockId="7.[151,789,798,943]" pageId="7" pageNumber="508">
<taxonomicName id="7D94A99DFFBBF30AFF2FFCA35A2EFCC9" authority="Attems 1927: 229" authorityName="Attems" authorityPageNumber="229" authorityYear="1927" box="[151,587,860,882]" class="Diplopoda" family="Julidae" genus="Chromatoiulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="serratus">
<emphasis id="88E00E0CFFBBF30AFF2FFCA359E2FCC9" box="[151,391,860,882]" italics="true" pageId="7" pageNumber="508">Chromatoiulus serratus</emphasis>
: Attems 1927: 229
</taxonomicName>
230, figs 301302.
<taxonomicName id="7D94A99DFFBBF30AFF2FFC855ABBFC2B" authority="Attems 1940: 307" authorityName="Attems" authorityPageNumber="307" authorityYear="1940" box="[151,734,890,912]" class="Diplopoda" family="Julidae" genus="Chromatoiulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="serratus" subGenus="Phauloiulus">
<emphasis id="88E00E0CFFBBF30AFF2FFC855A7DFC2B" box="[151,536,890,912]" italics="true" pageId="7" pageNumber="508">Chromatoiulus (Phauloiulus) serratus</emphasis>
: Attems 1940: 307
</taxonomicName>
.
<taxonomicName id="7D94A99DFFBBF30AFF2FFC665A34FC14" authority="Enghoff 2006: 183" authorityName="Enghoff" authorityPageNumber="183" authorityYear="2006" box="[151,593,921,943]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="serratum">
<emphasis id="88E00E0CFFBBF30AFF2FFC6659E6FC14" box="[151,387,921,943]" italics="true" pageId="7" pageNumber="508">Megaphyllum serratum</emphasis>
: Enghoff 2006: 183
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="F28E8195FFBBF30AFF2FFC235945FBA3" pageId="7" pageNumber="508" type="materials_examined">
<paragraph id="BA2BD21EFFBBF30AFF2FFC235945FBA3" blockId="7.[151,1436,988,2021]" pageId="7" pageNumber="508">
<emphasis id="88E00E0CFFBBF30AFF2FFC2359E5FC4E" bold="true" box="[151,384,988,1013]" pageId="7" pageNumber="508">Material examined.</emphasis>
MNKB: Cat. No. 2265,
<typeStatus id="652F6CBCFFBBF30AFD2CFC225A93FC4E" box="[660,758,989,1013]" pageId="7" pageNumber="508" type="syntype">syntypes</typeStatus>
<specimenCount id="AC921997FFBBF30AFD45FC235B43FC48" box="[765,806,988,1012]" pageId="7" pageNumber="508" type="generic">1 ♂</specimenCount>
, 1 ♀,
<specimenCount id="AC921997FFBBF30AFCD2FC235BC9FC4E" box="[874,940,988,1013]" pageId="7" pageNumber="508" type="generic">1 juv.</specimenCount>
; Verhoeff
<typeStatus id="652F6CBCFFBBF30AFBA7FC225C13FC4E" box="[1055,1142,989,1013]" pageId="7" pageNumber="508" type="syntype">syntype</typeStatus>
slide No. 1201: gonopods + leg-pair 2.
</paragraph>
</subSubSection>
<subSubSection id="F28E8195FFBBF30AFF7FFBDB5A80F85E" pageId="7" pageNumber="508" type="diagnosis">
<paragraph id="BA2BD21EFFBBF30AFF7FFBDB5AE1FB13" blockId="7.[151,1436,988,2021]" pageId="7" pageNumber="508">
<emphasis id="88E00E0CFFBBF30AFF7FFBDB5924FB86" bold="true" box="[199,321,1060,1085]" pageId="7" pageNumber="508">Diagnosis.</emphasis>
Differs from
<taxonomicName id="7D94A99DFFBBF30AFE5DFBDA5A11FB87" box="[485,628,1060,1084]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="curvifolii">
<emphasis id="88E00E0CFFBBF30AFE5DFBDA5A11FB87" box="[485,628,1060,1084]" italics="true" pageId="7" pageNumber="508">M. curvifolii</emphasis>
</taxonomicName>
,
<taxonomicName id="7D94A99DFFBBF30AFD3EFBDA5B02FB87" box="[646,871,1060,1084]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="genezarethanum">
<emphasis id="88E00E0CFFBBF30AFD3EFBDA5B02FB87" box="[646,871,1060,1084]" italics="true" pageId="7" pageNumber="508">M. genezarethanum</emphasis>
</taxonomicName>
,
<taxonomicName id="7D94A99DFFBBF30AFCC1FBDA5C41FB87" box="[889,1060,1060,1084]" class="Diplopoda" family="Julidae" genus="Chromatoiulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="tenenbaumi">
<emphasis id="88E00E0CFFBBF30AFCC1FBDA5C41FB87" box="[889,1060,1060,1084]" italics="true" pageId="7" pageNumber="508">M. tenenbaumi</emphasis>
</taxonomicName>
, and
<taxonomicName id="7D94A99DFFBBF30AFBD1FBDA5C99FB87" box="[1129,1276,1061,1084]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="arcuatum" status="sp. nov.">
<emphasis id="88E00E0CFFBBF30AFBD1FBDA5C99FB87" box="[1129,1276,1061,1084]" italics="true" pageId="7" pageNumber="508">M. arcuatum</emphasis>
</taxonomicName>
<emphasis id="88E00E0CFFBBF30AFABEFBDB5D2CFB86" bold="true" box="[1286,1353,1060,1085]" pageId="7" pageNumber="508">
<taxonomicNameLabel id="93D3B377FFBBF30AFABEFBDB5D2CFB86" box="[1286,1353,1060,1085]" pageId="7" pageNumber="508" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the serrated, rather than smooth, anterior opisthomere process; and from
<taxonomicName id="7D94A99DFFBBF30AFC59FBB65C14FBE4" box="[993,1137,1096,1120]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="turcicum">
<emphasis id="88E00E0CFFBBF30AFC59FBB65C14FBE4" box="[993,1137,1096,1120]" italics="true" pageId="7" pageNumber="508">M. turcicum</emphasis>
</taxonomicName>
by the narrow posterior opisthomere process which is slightly shorter than solenomere, as opposed to the same process being very wide, higher than solenomere in the latter species.
</paragraph>
<paragraph id="BA2BD21EFFBBF30AFF7FFB4B5970F9E3" blockId="7.[151,1436,988,2021]" pageId="7" pageNumber="508">
<emphasis id="88E00E0CFFBBF30AFF7FFB4B59FFFB76" bold="true" box="[199,410,1204,1229]" pageId="7" pageNumber="508">Descriptive notes.</emphasis>
According to Verhoeffs (1898) original description, this species seems to be comparable in size with
<taxonomicName id="7D94A99DFFBBF30AFEB4FB265988FB4B" box="[268,493,1240,1264]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="transsylvanicum">
<emphasis id="88E00E0CFFBBF30AFEB4FB265988FB4B" box="[268,493,1240,1264]" italics="true" pageId="7" pageNumber="508">M. transsylvanicum</emphasis>
</taxonomicName>
. However, the examined specimens proved to be considerably smaller: male in developmental stadium XII, 52+1+T, body length
<quantity id="7D6C7FFBFFBBF30AFD6FFB035B4EFAAF" box="[727,811,1276,1300]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" pageId="7" pageNumber="508" unit="mm" value="29.0">29 mm</quantity>
, height
<quantity id="7D6C7FFBFFBBF30AFC30FB035B86FAAF" box="[904,995,1276,1300]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="7" pageNumber="508" unit="mm" value="1.7">1.7 mm</quantity>
; female in stadium XIIXIII, 48+1+T, length
<quantity id="7D6C7FFBFFBBF30AFF5FFADF595EFA8C" box="[231,315,1312,1336]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="7" pageNumber="508" unit="mm" value="33.0">33 mm</quantity>
, height
<quantity id="7D6C7FFBFFBBF30AFE21FADF5990FA8C" box="[409,501,1312,1336]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="7" pageNumber="508" unit="mm" value="2.2">2.2 mm</quantity>
. 20 labral setae. Collum with 45 shallow striae near posterolateral corner; male mandibular stipites fairly large, protruding mostly ventrad, with a rounded infero-anterior corner. Ozopores placed just behind suture between pro- and metazona, slightly sinuous in anterior body rings; male pleurotergum 7 almost unmodified, only its ventral metazonal edge slightly convex. Metazona deeply striated, with 10 striae in a square with sides equal to metazonal length just below ozopore level; metazonal hind margins densely pilose. Pre-anal process directed slightly ventrad, rather short, strongly robust, surpassed by longest setae on anal valves. Subanal scale triangular, pointed, protruding by ca 1/3 of its length behind anal valves rear contour in the male, blunt, not protruding in the female. Anal valves densely setose, with a distinct row of short, stiff setae along caudal margin of each valve.
</paragraph>
<paragraph id="BA2BD21EFFBBF30AFF7FF99B5CBAF8EE" blockId="7.[151,1436,988,2021]" pageId="7" pageNumber="508">
Gonopods: Promere (
<figureCitation id="22AFCE9BFFBBF30AFE01F99B5A6FF9C7" box="[441,522,1636,1661]" captionStart="FIGURES 19 22" captionStartId="8.[151,269,1756,1779]" captionTargetBox="[174,1410,203,1724]" captionTargetId="figure@8.[170,1417,195,1732]" captionTargetPageId="8" captionText="FIGURES 19 22. Megaphyllum serratum syntypes, male (19 21) and female (22): (19) right gonopods, mesal, slightly anterior view; (20) right promere, posterior view; (21) right opisthomere, meso-anterior view; (22) right vulva, lateroposterior view (setae omitted, receptaculum seen by transparency, but drawn with solid lines for better emphasis). Abbreviations: ap: anterior process, asp: anterior protrusion of the solenomere, at: apodematic tube, ch: central hump, ct: connecting tube, fc: flagellum channel, fl: flagellum, g: median groove, lh: lateral hump, mh: mesal hump, P: promere, pa: posterior ampulla, ph: posterior hump, pp: posterior process, r: median ridge, s: solenomere, sf: spine-like filaments. Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/219030/files/figure.png" pageId="7" pageNumber="508">Fig. 20</figureCitation>
and
<emphasis id="88E00E0CFFBBF30AFDFBF99B5A31F9C0" bold="true" box="[579,596,1636,1659]" italics="true" pageId="7" pageNumber="508">P</emphasis>
in
<figureCitation id="22AFCE9BFFBBF30AFDC2F99B5AAEF9C7" box="[634,715,1636,1661]" captionStart="FIGURES 19 22" captionStartId="8.[151,269,1756,1779]" captionTargetBox="[174,1410,203,1724]" captionTargetId="figure@8.[170,1417,195,1732]" captionTargetPageId="8" captionText="FIGURES 19 22. Megaphyllum serratum syntypes, male (19 21) and female (22): (19) right gonopods, mesal, slightly anterior view; (20) right promere, posterior view; (21) right opisthomere, meso-anterior view; (22) right vulva, lateroposterior view (setae omitted, receptaculum seen by transparency, but drawn with solid lines for better emphasis). Abbreviations: ap: anterior process, asp: anterior protrusion of the solenomere, at: apodematic tube, ch: central hump, ct: connecting tube, fc: flagellum channel, fl: flagellum, g: median groove, lh: lateral hump, mh: mesal hump, P: promere, pa: posterior ampulla, ph: posterior hump, pp: posterior process, r: median ridge, s: solenomere, sf: spine-like filaments. Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/219030/files/figure.png" pageId="7" pageNumber="508">Fig. 19</figureCitation>
) ca 2 times longer than broad, with a rounded incision at distolateral margin; median groove (
<emphasis id="88E00E0CFFBBF30AFDABF9775A44F924" bold="true" box="[531,545,1672,1695]" italics="true" pageId="7" pageNumber="508">g</emphasis>
) very deep and narrow, tightly subtending opisthomere
<emphasis id="88E00E0CFFBBF30AFB72F9775D73F924" box="[1226,1302,1672,1695]" italics="true" pageId="7" pageNumber="508">in situ</emphasis>
. Flagellum considerably longer than promere. Opisthomere (
<figureCitation id="22AFCE9BFFBBF30AFD72F9535B2AF97F" box="[714,847,1708,1733]" captionStart="FIGURES 19 22" captionStartId="8.[151,269,1756,1779]" captionTargetBox="[174,1410,203,1724]" captionTargetId="figure@8.[170,1417,195,1732]" captionTargetPageId="8" captionText="FIGURES 19 22. Megaphyllum serratum syntypes, male (19 21) and female (22): (19) right gonopods, mesal, slightly anterior view; (20) right promere, posterior view; (21) right opisthomere, meso-anterior view; (22) right vulva, lateroposterior view (setae omitted, receptaculum seen by transparency, but drawn with solid lines for better emphasis). Abbreviations: ap: anterior process, asp: anterior protrusion of the solenomere, at: apodematic tube, ch: central hump, ct: connecting tube, fc: flagellum channel, fl: flagellum, g: median groove, lh: lateral hump, mh: mesal hump, P: promere, pa: posterior ampulla, ph: posterior hump, pp: posterior process, r: median ridge, s: solenomere, sf: spine-like filaments. Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/219030/files/figure.png" pageId="7" pageNumber="508">Figs 19, 21</figureCitation>
) with a narrow, slightly tapering, posterior process (
<emphasis id="88E00E0CFFBBF30AFF18F92F58D9F95C" bold="true" box="[160,188,1744,1767]" italics="true" pageId="7" pageNumber="508">pp</emphasis>
), with several blunt teeth at distomesal margin; a flattened, mesad deviating, anterior process (
<emphasis id="88E00E0CFFBBF30AFA99F92F5D58F95C" bold="true" box="[1313,1341,1744,1767]" italics="true" pageId="7" pageNumber="508">ap</emphasis>
) with a serrated apical part; and a slender, tubular solenomere (
<emphasis id="88E00E0CFFBBF30AFCA8F90B5B7EF8B0" bold="true" box="[784,795,1780,1803]" italics="true" pageId="7" pageNumber="508">s</emphasis>
) with a finger-like anterior protrusion (
<emphasis id="88E00E0CFFBBF30AFB67F90B5D63F8B0" bold="true" box="[1247,1286,1780,1803]" italics="true" pageId="7" pageNumber="508">asp</emphasis>
). Promere
<emphasis id="88E00E0CFFBBF30AFA3EF90A58A5F894" italics="true" pageId="7" pageNumber="508">in situ</emphasis>
considerably shorter than opisthomere; promere-opisthomere connection exceptionally strong due to a chitinous knot at their bases, similar to those normally observed between promeres of most julids.
</paragraph>
<paragraph id="BA2BD21EFFBBF30AFF7FF8A05A80F85E" blockId="7.[151,1436,988,2021]" pageId="7" pageNumber="508">
Vulvae (
<figureCitation id="22AFCE9BFFBBF30AFE9FF8A0591EF8C3" box="[295,379,1887,1912]" captionStart="FIGURES 19 22" captionStartId="8.[151,269,1756,1779]" captionTargetBox="[174,1410,203,1724]" captionTargetId="figure@8.[170,1417,195,1732]" captionTargetPageId="8" captionText="FIGURES 19 22. Megaphyllum serratum syntypes, male (19 21) and female (22): (19) right gonopods, mesal, slightly anterior view; (20) right promere, posterior view; (21) right opisthomere, meso-anterior view; (22) right vulva, lateroposterior view (setae omitted, receptaculum seen by transparency, but drawn with solid lines for better emphasis). Abbreviations: ap: anterior process, asp: anterior protrusion of the solenomere, at: apodematic tube, ch: central hump, ct: connecting tube, fc: flagellum channel, fl: flagellum, g: median groove, lh: lateral hump, mh: mesal hump, P: promere, pa: posterior ampulla, ph: posterior hump, pp: posterior process, r: median ridge, s: solenomere, sf: spine-like filaments. Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/219030/files/figure.png" pageId="7" pageNumber="508">Fig. 22</figureCitation>
) strongly asymmetric: lateral hump of bursa (
<emphasis id="88E00E0CFFBBF30AFC35F8A05BC1F8C3" bold="true" box="[909,932,1887,1912]" italics="true" pageId="7" pageNumber="508">lh</emphasis>
) protruding apically high above both mesal hump (
<emphasis id="88E00E0CFFBBF30AFF5EF87C596EF827" bold="true" box="[230,267,1923,1948]" italics="true" pageId="7" pageNumber="508">mh</emphasis>
) and operculum. Short, rounded hyaline lamellae present apically on both bursa and operculum. Setation dense all over vulva. Shape of receptaculum (
<emphasis id="88E00E0CFFBBF30AFD18F8575AD3F804" bold="true" box="[672,694,1960,1983]" italics="true" pageId="7" pageNumber="508">at</emphasis>
,
<emphasis id="88E00E0CFFBBF30AFD7DF8575A84F804" bold="true" box="[709,737,1960,1983]" italics="true" pageId="7" pageNumber="508">pa</emphasis>
) typical of the subgenus, but smaller in relation to vulva size compared to condition observed in the other species.
</paragraph>
</subSubSection>
<caption id="EEEB8296FFB4F305FF2FF923590EF808" httpUri="https://zenodo.org/record/219030/files/figure.png" pageId="8" pageNumber="509" targetBox="[174,1410,203,1724]" targetPageId="8">
<paragraph id="BA2BD21EFFB4F305FF2FF923590EF808" blockId="8.[151,1437,1755,1971]" pageId="8" pageNumber="509">
<emphasis id="88E00E0CFFB4F305FF2FF923593BF948" bold="true" box="[151,350,1756,1779]" pageId="8" pageNumber="509">FIGURES 1922.</emphasis>
<taxonomicName id="7D94A99DFFB4F305FEDCF9245A3BF948" box="[356,606,1755,1779]" class="Diplopoda" family="Julidae" genus="Megaphyllum" kingdom="Animalia" order="Julida" pageId="8" pageNumber="509" phylum="Arthropoda" rank="species" species="serratum">
<emphasis id="88E00E0CFFB4F305FEDCF9245A3BF948" box="[356,606,1755,1779]" italics="true" pageId="8" pageNumber="509">Megaphyllum serratum</emphasis>
</taxonomicName>
syntypes, male (1921) and female (22):
<emphasis id="88E00E0CFFB4F305FBA6F9235C2FF948" bold="true" box="[1054,1098,1756,1779]" pageId="8" pageNumber="509">(19)</emphasis>
right gonopods, mesal, slightly anterior view;
<emphasis id="88E00E0CFFB4F305FE8EF9035906F8A8" bold="true" box="[310,355,1788,1811]" pageId="8" pageNumber="509">(20)</emphasis>
right promere, posterior view;
<emphasis id="88E00E0CFFB4F305FD01F9035A80F8A8" bold="true" box="[697,741,1788,1811]" pageId="8" pageNumber="509">(21)</emphasis>
right opisthomere, meso-anterior view;
<emphasis id="88E00E0CFFB4F305FB24F9035CADF8A8" bold="true" box="[1180,1224,1788,1811]" pageId="8" pageNumber="509">(22)</emphasis>
right vulva, lateroposterior view (setae omitted, receptaculum seen by transparency, but drawn with solid lines for better emphasis). Abbreviations:
<emphasis id="88E00E0CFFB4F305FE83F8C45933F8E8" bold="true" box="[315,342,1851,1875]" italics="true" pageId="8" pageNumber="509">ap</emphasis>
: anterior process,
<emphasis id="88E00E0CFFB4F305FDA1F8C45A5BF8E8" bold="true" box="[537,574,1851,1875]" italics="true" pageId="8" pageNumber="509">asp</emphasis>
: anterior protrusion of the solenomere,
<emphasis id="88E00E0CFFB4F305FC5BF8C45B9DF8E8" bold="true" box="[995,1016,1851,1875]" italics="true" pageId="8" pageNumber="509">at</emphasis>
: apodematic tube,
<emphasis id="88E00E0CFFB4F305FB79F8C45CB9F8E8" bold="true" box="[1217,1244,1851,1875]" italics="true" pageId="8" pageNumber="509">ch</emphasis>
: central hump,
<emphasis id="88E00E0CFFB4F305FA39F8C45DF1F8E8" bold="true" box="[1409,1428,1851,1875]" italics="true" pageId="8" pageNumber="509">ct</emphasis>
: connecting tube,
<emphasis id="88E00E0CFFB4F305FEF7F8A45901F8C8" bold="true" box="[335,356,1883,1907]" italics="true" pageId="8" pageNumber="509">fc</emphasis>
: flagellum channel,
<emphasis id="88E00E0CFFB4F305FDFAF8A45A37F8C8" bold="true" box="[578,594,1883,1907]" italics="true" pageId="8" pageNumber="509">fl</emphasis>
: flagellum,
<emphasis id="88E00E0CFFB4F305FD6DF8A45A87F8C8" bold="true" box="[725,738,1883,1907]" italics="true" pageId="8" pageNumber="509">g</emphasis>
: median groove,
<emphasis id="88E00E0CFFB4F305FC18F8A45BD3F8C8" bold="true" box="[928,950,1883,1907]" italics="true" pageId="8" pageNumber="509">lh</emphasis>
: lateral hump,
<emphasis id="88E00E0CFFB4F305FBE2F8A45C18F8C8" bold="true" box="[1114,1149,1883,1907]" italics="true" pageId="8" pageNumber="509">mh</emphasis>
: mesal hump,
<emphasis id="88E00E0CFFB4F305FAA5F8A45D48F8C8" bold="true" box="[1309,1325,1883,1907]" italics="true" pageId="8" pageNumber="509">P</emphasis>
: promere,
<emphasis id="88E00E0CFFB4F305FF2FF88458D7F828" bold="true" box="[151,178,1915,1939]" italics="true" pageId="8" pageNumber="509">pa</emphasis>
: posterior ampulla,
<emphasis id="88E00E0CFFB4F305FE30F88459C1F828" bold="true" box="[392,420,1915,1939]" italics="true" pageId="8" pageNumber="509">ph</emphasis>
: posterior hump,
<emphasis id="88E00E0CFFB4F305FDDAF8845A18F828" bold="true" box="[610,637,1915,1939]" italics="true" pageId="8" pageNumber="509">pp</emphasis>
: posterior process,
<emphasis id="88E00E0CFFB4F305FCF6F8845B3DF828" bold="true" box="[846,856,1915,1939]" italics="true" pageId="8" pageNumber="509">r</emphasis>
: median ridge,
<emphasis id="88E00E0CFFB4F305FBB8F8845C6FF828" bold="true" box="[1024,1034,1915,1939]" italics="true" pageId="8" pageNumber="509">s</emphasis>
: solenomere,
<emphasis id="88E00E0CFFB4F305FB19F8845CD1F828" bold="true" box="[1185,1204,1915,1939]" italics="true" pageId="8" pageNumber="509">sf</emphasis>
: spine-like filaments. Scale bars: 0.2 mm.
</paragraph>
</caption>
<subSubSection id="F28E8195FFB5F304FF7FFF685C0BFF0B" box="[199,1134,151,176]" pageId="9" pageNumber="510" type="distribution">
<paragraph id="BA2BD21EFFB5F304FF7FFF685C0BFF0B" blockId="9.[151,1436,151,248]" box="[199,1134,151,176]" pageId="9" pageNumber="510">
<emphasis id="88E00E0CFFB5F304FF7FFF685905FF0B" bold="true" box="[199,352,151,176]" pageId="9" pageNumber="510">Distribution.</emphasis>
<collectingCountry id="C283928EFFB5F304FEDFFF6859DDFF0B" box="[359,440,151,176]" name="Turkey" pageId="9" pageNumber="510">Turkey</collectingCountry>
, Pernata, Inevi (
<typeStatus id="652F6CBCFFB5F304FDD5FF675AFAFF0B" box="[621,671,152,176]" pageId="9" pageNumber="510">type</typeStatus>
locality) (Verhoeff 1898; Enghoff 2006).
</paragraph>
</subSubSection>
<subSubSection id="F28E8195FFB5F304FF7FFF435CA7FF43" pageId="9" pageNumber="510" type="discussion">
<paragraph id="BA2BD21EFFB5F304FF7FFF435CA7FF43" blockId="9.[151,1436,151,248]" pageId="9" pageNumber="510">
<emphasis id="88E00E0CFFB5F304FF7FFF43595EFF6E" bold="true" box="[199,315,188,213]" pageId="9" pageNumber="510">Remarks.</emphasis>
At the present the examined individuals are all strongly faded, but, according to Verhoeff (1898), only the males have yellow-reddish dorsolateral sides which might suggest sexual dimorphism.
</paragraph>
</subSubSection>
</treatment>
</document>