treatments-xml/data/0C/12/05/0C12051D2343FFCBFECCF9FECF91B42D.xml

<document id="D5D1F51D658462321C1412B6090D28F0" ID-CLB-Dataset="5480" ID-DOI="10.11646/zootaxa.230.1.1" ID-GBIF-Dataset="a880f550-16bc-4733-9407-d09212861392" ID-ISSN="1175­5334" ID-Zenodo-Dep="5019111" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624465299970" checkinUser="felipe" docAuthor="Ponder, Winston F." docDate="2003" docId="0C12051D2343FFCBFECCF9FECF91B42D" docLanguage="en" docName="zt00230.pdf" docOrigin="Zootaxa 230 (1)" docSource="http://biotaxa.org/Zootaxa/article/view/zootaxa.230.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Gabbia davisi Ponder 2003, n. sp." docType="treatment" docVersion="8" lastPageNumber="49" masterDocId="F02B7D65236DFFFBFFC4FFFECC32B02B" masterDocTitle="Monograph of the Australian Bithyniidae (Caenogastropoda: Rissooidea)" masterLastPageNumber="126" masterPageNumber="1" pageNumber="47" updateTime="1699020163718" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="EB0338D8702AAA7CC559FC6A1665D3EC">Monograph of the Australian Bithyniidae (Caenogastropoda: Rissooidea)</mods:title>
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<paragraph id="8404B40B2343FFD5FECCF9FECDC5B631" blockId="46.[264,503,1536,1562]" box="[264,503,1536,1562]" pageId="46" pageNumber="47">
<heading id="DF4C03672343FFD5FECCF9FECDC5B631" bold="true" box="[264,503,1536,1562]" fontSize="11" level="1" pageId="46" pageNumber="47" reason="1">
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<taxonomicName id="43BBCF882343FFD5FECCF9FECD9FB631" ID-CoL="6K5FZ" authority="Ponder, 2003" authorityName="Ponder" authorityYear="2003" box="[264,429,1536,1562]" class="Gastropoda" family="Bithyniidae" genus="Gabbia" kingdom="Animalia" order="Littorinimorpha" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="davisi" status="sp. nov.">
<emphasis id="B6CF68192343FFD5FECCF9FECD9FB631" bold="true" box="[264,429,1536,1562]" italics="true" pageId="46" pageNumber="47">Gabbia davisi</emphasis>
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<taxonomicNameLabel id="ADFCD5622343FFD5FE71F9FECDC5B631" box="[437,503,1536,1562]" pageId="46" pageNumber="47" rank="species">n. sp.</taxonomicNameLabel>
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<emphasis id="B6CF68192343FFD5FECCF9B8CDB7B674" box="[264,389,1606,1631]" italics="true" pageId="46" pageNumber="47">Etymology</emphasis>
</paragraph>
<paragraph id="8404B40B2343FFD5FEFCF990CFEAB69B" blockId="46.[264,1323,1606,1712]" pageId="46" pageNumber="47">
Named for Assoc. Prof. Andrew Davis, University of Wollongong, NSW, 
<collectingCountry id="FCACF49B2343FFD5FB5EF990C93AB6A3" box="[1178,1288,1646,1672]" name="Australia" pageId="46" pageNumber="47">Australia</collectingCountry>
, in appreciation of his companionship and assistance in the field.
</paragraph>
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<typeStatus id="5B000AA92343FFD5FECCF925CD0DB6DF" box="[264,319,1755,1780]" pageId="46" pageNumber="47">Type</typeStatus>
material
</emphasis>
</paragraph>
<paragraph id="8404B40B2343FFD5FEFCF8FDC86CB746" blockId="46.[264,1325,1755,1901]" pageId="46" pageNumber="47">
<materialsCitation id="34D3BE562343FFD5FEFCF8FDC869B746" ID-GBIF-Occurrence="3321535499" collectingDate="2002-10-04" collectionCode="AMS" collectorName="W. F. Ponder &amp; A. Davis" latitude="-22.261" location="Lake Huffer Stn" longLatPrecision="1" longitude="145.3405" municipality="Qld" pageId="46" pageNumber="47" specimenCount="2" typeStatus="holotype">
<location id="8164E2D02343FFD5FEFCF8FDCDC5B736" LSID="urn:lsid:plazi:treatment:0C12051D2343FFCBFECCF9FECF91B42D:8164E2D02343FFD5FEFCF8FDCDC5B736" box="[312,503,1795,1821]" latitude="-22.261" longLatPrecision="1" longitude="145.3405" municipality="Qld" name="Lake Huffer Stn" pageId="46" pageNumber="47">Lake Huffer Stn</location>
, large spring near lake, pool at base of spring, 
<collectingMunicipality id="64602E712343FFD5FBD1F8FDC876B736" box="[1045,1092,1795,1821]" pageId="46" pageNumber="47">Qld</collectingMunicipality>
, 
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, 
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, in mud, 
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<collectingDate id="E0416B232343FFD5FE30F8D5CEABB76E" box="[500,665,1835,1861]" pageId="46" pageNumber="47" value="2002-10-04">4 OCT 2002</collectingDate>
</date>
, 
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&amp; 
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(
<typeStatus id="5B000AA92343FFD5FC1BF8D5C863B76E" box="[991,1105,1835,1861]" pageId="46" pageNumber="47" type="holotype">Holotype</typeStatus>
, 
<collectionCode id="E2AA2CCE2343FFD5FBA7F8D5C890B76E" box="[1123,1186,1835,1861]" pageId="46" pageNumber="47">AMS</collectionCode>
C.417668; 
<typeStatus id="5B000AA92343FFD5FECCF8ADCD4AB746" box="[264,376,1875,1901]" pageId="46" pageNumber="47" type="paratype">paratypes</typeStatus>
AMS C.417617, 414 wet, 5 on SEM stub; QM MO71711, 10)
</materialsCitation>
.
</paragraph>
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<emphasis id="B6CF68192342FFD4FECCFEE2CDA0B11E" box="[264,402,284,309]" italics="true" pageId="47" pageNumber="48">Description</emphasis>
</paragraph>
<paragraph id="8404B40B2342FFD4FEFCFEBAC8E1B265" blockId="47.[264,1323,284,670]" pageId="47" pageNumber="48">
Shell (
<figureCitation id="1C80A88E2342FFD4FE41FEBACDDEB175" box="[389,492,324,350]" captionStart="FIGURE 13" captionStartId="41.[264,368,1426,1450]" captionTargetBox="[311,1270,315,1412]" captionTargetId="figure-166@41.[271,1331,305,1424]" captionTargetPageId="41" captionText="FIGURE 13. Shells: A, B, Gabbia campicola, AMS, C.318702; C, D, Gabbia pallidula, C, paratype, AMS, C.308007, D, holotype AMS, C.417667; E, Gabbia davisi, holotype, AMS, C.417668; F, G, Gabbia napierensis, F, paratype, AMS, C.337372, G, holotype, S10978; H, I, Gabbia cf. rotunda, AMS, C.385439; Scales: A – D, F – I, 500 m, E, 1mm." figureDoi="http://doi.org/10.5281/zenodo.5019136" httpUri="https://zenodo.org/record/5019136/files/figure.png" pageId="47" pageNumber="48">Fig. 13E</figureCitation>
) medium (up to 
<quantity id="434319EE2342FFD4FD76FEBACF36B175" box="[690,772,324,350]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="47" pageNumber="48" unit="mm" value="6.7">6.7mm</quantity>
in length), broadly­ovate­conic, up to 4.6 convex whorls. Protoconch of about 1.3­1.5 almost smooth whorls, with traces of extremely fine spiral striae, last half whorl with weak axial threads. Teleoconch sculptured with fine collabral growth lines only; base evenly convex; umbilicus narrowly open, wider in juveniles. Aperture broadly ovate; peristome thin; inner lip slightly thickened, outer lip thin, prosocline. Colour: shell opaque to semi­translucent; periostracum pale yellowish to yellowish­brown. A few thin, rather indistinct, pale brown varices sometimes present.
</paragraph>
<paragraph id="8404B40B2342FFD4FEFCFDA2CEDAB2B5" blockId="47.[264,1323,284,670]" pageId="47" pageNumber="48">
Dimensions. See 
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for dimensions of 
<typeStatus id="5B000AA92342FFD4FCF1FDA2CFA9B25D" box="[821,923,604,630]" pageId="47" pageNumber="48" type="holotype">holotype</typeStatus>
and Appendix, 
<tableCitation id="C93981B02342FFD4FB90FDA2C88EB25D" box="[1108,1212,604,630]" captionStart="TABLE 29" captionStartId="118.[264,352,348,372]" captionTargetPageId="118" captionText="TABLE 29. Shell measurements and teleoconch whorl counts. The figures given are the minimum ­ maximum (mean) and standard deviation. The number of specimens measured is given in parentheses in the second column and in a few cases a smaller number of specimens is noted for the whorl counts. All material is AMS unless otherwise noted." httpUri="http://table.plazi.org/id/D0C4E483231BFF8DFECCFEA2CF4BB1F8" pageId="47" pageNumber="48" tableUuid="D0C4E483231BFF8DFECCFEA2CF4BB1F8">Table 29</tableCitation>
, for summary shell dimensions and whorl counts.
</paragraph>
<caption id="D0C4E4832342FFD4FECCFD20CF71B2DE" box="[264,835,734,758]" pageId="47" pageNumber="48" startId="47.[264,352,734,758]" targetBox="[278,1306,795,898]" targetIsTable="true" targetPageId="47">
<paragraph id="8404B40B2342FFD4FECCFD20CF71B2DE" blockId="47.[264,835,734,758]" box="[264,835,734,758]" pageId="47" pageNumber="48">
<emphasis id="B6CF68192342FFD4FECCFD20CD46B2DD" bold="true" box="[264,372,734,758]" pageId="47" pageNumber="48">TABLE 9</emphasis>
. Dimensions of holotype of 
<taxonomicName id="43BBCF882342FFD4FD61FD21CF0EB2DD" authorityName="Ponder" authorityYear="2003" box="[677,828,735,758]" class="Gastropoda" family="Bithyniidae" genus="Gabbia" kingdom="Animalia" order="Littorinimorpha" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="davisi">
<emphasis id="B6CF68192342FFD4FD61FD21CF0EB2DD" box="[677,828,735,758]" italics="true" pageId="47" pageNumber="48">Gabbia davisi</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8404B40B2342FFD4FE0EFCE5C8C2B3A9" pageId="47" pageNumber="48">
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<th id="795ADF3523420004FE0EFCE5CE24B319" box="[458,534,795,818]" gridcol="1" gridrow="0" pageId="47" pageNumber="48">Length</th>
<th id="795ADF3523420004FD92FCE5CEAAB319" box="[598,664,795,818]" gridcol="2" gridrow="0" pageId="47" pageNumber="48">Width</th>
<th id="795ADF3523420004FD1EFCE5CF0BB319" box="[730,825,795,818]" gridcol="3" gridrow="0" pageId="47" pageNumber="48">Aperture</th>
<th id="795ADF3523420004FCB3FCE5CFE4B319" box="[887,982,795,818]" gridcol="4" gridrow="0" pageId="47" pageNumber="48">Aperture</th>
<th id="795ADF3523420004FBCAFCE5C84AB319" box="[1038,1144,795,818]" gridcol="5" gridrow="0" pageId="47" pageNumber="48">Length of</th>
<th id="795ADF3523420004FB62FCE5C928B319" box="[1190,1306,795,818]" gridcol="6" gridrow="0" pageId="47" pageNumber="48">Number of</th>
</tr>
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<td id="795ADF3523420004FD1EFCC3CF0BB37F" box="[730,825,829,852]" gridcol="3" gridrow="1" pageId="47" pageNumber="48">length</td>
<td id="795ADF3523420004FCB3FCC3CFE4B37F" box="[887,982,829,852]" gridcol="4" gridrow="1" pageId="47" pageNumber="48">width</td>
<td id="795ADF3523420004FBCAFCC3C84AB37F" box="[1038,1144,829,852]" gridcol="5" gridrow="1" pageId="47" pageNumber="48">last whorl</td>
<td id="795ADF3523420004FB62FCC3C928B37F" box="[1190,1306,829,852]" gridcol="6" gridrow="1" pageId="47" pageNumber="48">whorls</td>
</tr>
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<th id="795ADF3523420004FED2FC95CD4BB3A9" box="[278,377,875,898]" gridcol="0" gridrow="2" pageId="47" pageNumber="48">Holotype</th>
<td id="795ADF3523420004FE0EFC95CE24B3A9" box="[458,534,875,898]" gridcol="1" gridrow="2" pageId="47" pageNumber="48">5.07</td>
<td id="795ADF3523420004FD92FC95CEAAB3A9" box="[598,664,875,898]" gridcol="2" gridrow="2" pageId="47" pageNumber="48">3.89</td>
<td id="795ADF3523420004FD1EFC95CF0BB3A9" box="[730,825,875,898]" gridcol="3" gridrow="2" pageId="47" pageNumber="48">2.75</td>
<td id="795ADF3523420004FCB3FC95CFE4B3A9" box="[887,982,875,898]" gridcol="4" gridrow="2" pageId="47" pageNumber="48">2.54</td>
<td id="795ADF3523420004FBCAFC95C84AB3A9" box="[1038,1144,875,898]" gridcol="5" gridrow="2" pageId="47" pageNumber="48">4.23</td>
<td id="795ADF3523420004FB62FC95C928B3A9" box="[1190,1306,875,898]" gridcol="6" gridrow="2" pageId="47" pageNumber="48">4.0</td>
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</paragraph>
<paragraph id="8404B40B2342FFD4FEFCFC1DCDC2B466" blockId="47.[264,1324,995,1901]" pageId="47" pageNumber="48">
Operculum (
<figureCitation id="1C80A88E2342FFD4FE0AFC1DCE7CB3D6" box="[462,590,995,1021]" captionStart="FIGURE 15" captionStartId="44.[264,368,1766,1790]" captionTargetBox="[357,1260,401,1755]" captionTargetId="figure-142@44.[342,1277,393,1756]" captionTargetPageId="44" captionText="FIGURE 15. Opercula: A, B, Gabbia campicola, AMS, C.318702; C, D, Gabbia pallidula, paratype, AMS, C.308007; E, F, Gabbia davisi, paratype, AMS, C.417617; G ­ I, Gabbia napierensis, paratype, WAM, 1006 ­76; J, K, Gabbia cf. rotunda, AMS, C.385439; L, M, Gabbia rotunda, paratype, AMS, C.400134; A, C, E, G, I, J, L outer side, B, D, F, H, K, M inner side. Scales: 500 m." figureDoi="http://doi.org/10.5281/zenodo.5019142" httpUri="https://zenodo.org/record/5019142/files/figure.png" pageId="47" pageNumber="48">Fig. 15E,F</figureCitation>
) typical of genus. Ovate, white; with several distinct concentric growth ridges; inner surface with muscle scar occupying most of surface, sculptured with weak pustules.
</paragraph>
<paragraph id="8404B40B2342FFD4FEFCFBA5CF26B62E" blockId="47.[264,1324,995,1901]" pageId="47" pageNumber="48">
Radula (Appendix 
<tableCitation id="C93981B02342FFD4FDD3FBA5CEB2B45E" box="[535,640,1115,1141]" captionStart="TABLE 30" captionStartId="123.[264,352,284,308]" captionTargetPageId="123" captionText="TABLE 30. Radular cusp counts." httpUri="http://table.plazi.org/id/D0C4E4832316FF80FECCFEE2CE5DB118" pageId="47" pageNumber="48" tableUuid="D0C4E4832316FF80FECCFEE2CE5DB118">Table 30</tableCitation>
; 
<figureCitation id="1C80A88E2342FFD4FD48FBA5CF38B45E" box="[652,778,1115,1141]" captionStart="FIGURE 16" captionStartId="45.[264,368,1540,1564]" captionTargetBox="[345,1242,313,1530]" captionTargetId="figure-178@45.[341,1245,305,1532]" captionTargetPageId="45" captionText="FIGURE 16. Radulae: A, Gabbia pallidula, paratype, AMS, C.308007; B, C, Gabbia campicola, AMS, C.318702; D, Gabbia fontana, AMS, C.307906; E, F, Gabbia davisi, paratype, AMS, C.417617; G – J, Gabbia napierensis, paratype, –AMS, C.337372; K, L, Gabbia cf. rotunda, AMS, C.385439. A, B, E, H, I, L, central teeth, C, half row of teeth, D, central and lateral teeth, F, J, lateral teeth, G, marginal teeth, K, lateral and marginal teeth; Scales: 20 m." figureDoi="http://doi.org/10.5281/zenodo.5019146" httpUri="https://zenodo.org/record/5019146/files/figure.png" pageId="47" pageNumber="48">Fig. 16E,F</figureCitation>
) typical of genus. Central teeth with 4­5 cusps on either side of median cusp which is up to about 1.5 length of adjacent cusps and its base up to about 1.5 width; median cusp tapering and pointed. Face of central tooth with 1 pair of prominent cusps about third to nearly half of tooth height; lie inside lateral margin with weakly denticulate ridge representing up to 4 additional rudimentary basal denticles; lateral margins straight to slightly concave, at about 50­60º; basal tongue rather long, narrow and blunt. Lateral teeth with cusp formula 3 + 1 + 3­4; cutting edge slightly less than half length of lateral part of tooth; median cusp about 1.5 length of adjacent cusps, tapering with blunt to pointed end; inner side with excavation at base, upper edge of lateral part of tooth at about 70º to cutting edge, lateral edge strongly concave. Inner marginal teeth with 15­21 cusps, outer marginals with 8­9 cusps.
</paragraph>
<paragraph id="8404B40B2342FFD4FEFCF9EDC8DAB656" blockId="47.[264,1324,995,1901]" pageId="47" pageNumber="48">Head­foot grey to black except for tentacles distal to eyes. Tentacles unpigmented except for grey stripe in middle. Mantle roof black with small to large, close white spots and patches giving it netted appearance. Visceral coil unpigmented to black dorsally.</paragraph>
<paragraph id="8404B40B2342FFD4FEFCF975CDEFB746" blockId="47.[264,1324,995,1901]" pageId="47" pageNumber="48">
Anatomy. Gill with apices at about third gill width from right; 55­61 filaments (n=3). Osphradium slightly posterior to middle of gill or at middle. Penis (
<figureCitation id="1C80A88E2342FFD4FBD9F94DC845B6E6" box="[1053,1143,1715,1741]" captionStart="FIGURE 7" captionStartId="23.[264,368,1693,1717]" captionTargetId="figure-16@24.[272,1314,321,1892]" captionTargetPageId="24" captionText="FIGURE 7. Penis and accessory gland. A, D – Gabbia campicola, A, ventral view, C.332829, D, dorsal view, USNM, 803665; B – Gabbia rotunda, ventral view, AMS, C.400134; C – Gabbia davisi, ventral view, AMS, C.417617; E – Gabbia beecheyi, ventral view, AMS, C.332843; F, G – Gabbia fontana, dorsal view, AMS, C.380863; H – Gabbia spiralis, dorsal view, AMS, C.331849; I, J – Gabbia vertiginosa, dorsal view, AMS, C.203392, I, enlargement of distal end of accessory lobe; K – Gabbia adusta, dorsal view, AMS, C.203394. Scales: 500 m." figureDoi="http://doi.org/10.5281/zenodo.5097805" httpUri="https://zenodo.org/record/5097805/files/figure.png" pageId="47" pageNumber="48">Fig. 7C</figureCitation>
) and accessory lobe about equal in length to penial lobe or slightly longer, accessory lobe expanded to markedly expanded distally; accessory gland very long and tightly coiled. Pallial oviduct similar to 
<emphasis id="B6CF68192342FFD4FE4CF8D2CE1BB76E" box="[392,553,1836,1861]" italics="true" pageId="47" pageNumber="48">
<taxonomicName id="43BBCF882342FFD4FE4CF8D2CE16B76E" box="[392,548,1836,1861]" class="Gastropoda" family="Bithyniidae" genus="Gabbia" kingdom="Animalia" order="Littorinimorpha" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="vertiginos">G. vertiginos</taxonomicName>
,
</emphasis>
with narrow bursa copulatrix reaching edge of albumen gland (AMS C.417617).
</paragraph>
<paragraph id="8404B40B235DFFCBFEFCFEE2CD88B185" blockId="48.[264,1323,284,430]" pageId="48" pageNumber="49">
Distribution (
<figureCitation id="1C80A88E235DFFCBFE12FEE2CE29B11D" box="[470,539,284,310]" captionStart="FIGURE 9" captionStartId="26.[264,368,1373,1397]" captionTargetBox="[276,1298,569,1360]" captionTargetId="figure-233@26.[227,1311,552,1369]" captionTargetPageId="26" captionText="FIGURE 9. Distribution of Gabbia vertiginosa (o), G. iredalei (), G. cf. iredalei (), G. campicola (), G. fontana (), G. pallidula (☆), G. davisi () and G. rotunda (" figureDoi="http://doi.org/10.5281/zenodo.5019128" httpUri="https://zenodo.org/record/5019128/files/figure.png" pageId="48" pageNumber="49">Fig. 9</figureCitation>
) and habitat. Known only from a single spring on Lake Huffer Station in western 
<collectingRegion id="467F7AE9235DFFCBFE00FEBACE7DB175" box="[452,591,324,350]" country="Australia" name="Queensland" pageId="48" pageNumber="49">Queensland</collectingRegion>
where it was found in a small, shallow muddy pool near the head of a large spring and, abundantly, in a large, shallow muddy pool at the base of the spring outflow.
</paragraph>
<paragraph id="8404B40B235DFFCBFECCFE1ACD5CB1D6" blockId="48.[264,1323,484,1030]" box="[264,366,484,509]" pageId="48" pageNumber="49">
<emphasis id="B6CF6819235DFFCBFECCFE1ACD5CB1D6" box="[264,366,484,509]" italics="true" pageId="48" pageNumber="49">Remarks</emphasis>
</paragraph>
<paragraph id="8404B40B235DFFCBFEFCFDF2CF1BB3A5" blockId="48.[264,1323,484,1030]" pageId="48" pageNumber="49">
This species closely resembles 
<taxonomicName id="43BBCF88235DFFCBFD61FDF2CF0DB20E" box="[677,831,524,549]" class="Gastropoda" family="Bithyniidae" genus="Gabbia" kingdom="Animalia" order="Littorinimorpha" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="campicola">
<emphasis id="B6CF6819235DFFCBFD61FDF2CF0DB20E" box="[677,831,524,549]" italics="true" pageId="48" pageNumber="49">G. campicola</emphasis>
</taxonomicName>
in shell characters and measurements and in radular characters. Its whorls tend to be more convex and it is much more distinctly umbilicate, especially in juveniles which are rather widely umbilicate. 
<taxonomicName id="43BBCF88235DFFCBFB95FDA2C8CAB25E" authorityName="Ponder" authorityYear="2003" box="[1105,1272,604,629]" class="Gastropoda" family="Bithyniidae" genus="Gabbia" kingdom="Animalia" order="Littorinimorpha" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="davisi">
<emphasis id="B6CF6819235DFFCBFB95FDA2C8CAB25E" box="[1105,1272,604,629]" italics="true" pageId="48" pageNumber="49">Gabbia davisi</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="43BBCF88235DFFCBFEB8FD7ACE25B2B6" box="[380,535,644,669]" class="Gastropoda" family="Bithyniidae" genus="Gabbia" kingdom="Animalia" order="Littorinimorpha" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="campicola">
<emphasis id="B6CF6819235DFFCBFEB8FD7ACE25B2B6" box="[380,535,644,669]" italics="true" pageId="48" pageNumber="49">G. campicola</emphasis>
</taxonomicName>
in the gill apices being at about a third of the gill width (compared with on or close to the right edge). It also differs from 
<taxonomicName id="43BBCF88235DFFCBFC53FD52C800B2EE" box="[919,1074,684,709]" class="Gastropoda" family="Bithyniidae" genus="Gabbia" kingdom="Animalia" order="Littorinimorpha" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="campicola">
<emphasis id="B6CF6819235DFFCBFC53FD52C800B2EE" box="[919,1074,684,709]" italics="true" pageId="48" pageNumber="49">G. campicola</emphasis>
</taxonomicName>
and all other similar species in the accessory prostate being unusually very long, narrow and tangled. The pigmented mantle, long accessory gland and umbilicus separate this species from 
<emphasis id="B6CF6819235DFFCBFB7EFD02CDD7B316" italics="true" pageId="48" pageNumber="49">
<taxonomicName id="43BBCF88235DFFCBFB7EFD02CD03B316" authorityName="Ponder" authorityYear="2003" class="Gastropoda" family="Bithyniidae" genus="Gabbia" kingdom="Animalia" order="Littorinimorpha" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="pallidula">G. pallidula</taxonomicName>
. 
<taxonomicName id="43BBCF88235DFFCBFEFAFCDACDD7B316" authorityName="Ponder" authorityYear="2003" box="[318,485,804,829]" class="Gastropoda" family="Bithyniidae" genus="Gabbia" kingdom="Animalia" order="Littorinimorpha" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="davisi">Gabbia davisi</taxonomicName>
</emphasis>
is found in a spring very near the 
<typeStatus id="5B000AA9235DFFCBFC4CFCDACF88B315" box="[904,954,804,830]" pageId="48" pageNumber="49">type</typeStatus>
locality of 
<taxonomicName id="43BBCF88235DFFCBFB83FCDAC8E1B316" authorityName="Ponder" authorityYear="2003" box="[1095,1235,804,829]" class="Gastropoda" family="Bithyniidae" genus="Gabbia" kingdom="Animalia" order="Littorinimorpha" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="pallidula">
<emphasis id="B6CF6819235DFFCBFB83FCDAC8E1B316" box="[1095,1235,804,829]" italics="true" pageId="48" pageNumber="49">G. pallidula</emphasis>
</taxonomicName>
, and is contrasted in the remarks under that species. 
<taxonomicName id="43BBCF88235DFFCBFC82FCB2C821B34E" authority="Ponder, 2003" authorityName="Ponder" authorityYear="2003" box="[838,1043,844,869]" class="Gastropoda" family="Bithyniidae" genus="Gabbia" kingdom="Animalia" order="Littorinimorpha" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="fontana" status="sp. nov.">
<emphasis id="B6CF6819235DFFCBFC82FCB2C821B34E" box="[838,1043,844,869]" italics="true" pageId="48" pageNumber="49">Gabbia fontana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADFCD562235DFFCBFBE5FCB2C85AB34D" box="[1057,1128,844,870]" pageId="48" pageNumber="49" rank="species">n. sp.</taxonomicNameLabel>
, another spring endemic, is contrasted below under that taxon.
</paragraph>
<paragraph id="8404B40B235DFFCBFEFCFC62CF91B42D" blockId="48.[264,1323,484,1030]" pageId="48" pageNumber="49">
Despite the apparent lack of shell differences, a disciminant analysis including all similar species from NE 
<collectingRegion id="467F7AE9235DFFCBFDC4FC3ACEB8B3F5" box="[512,650,964,990]" country="Australia" name="Queensland" pageId="48" pageNumber="49">Queensland</collectingRegion>
separates this species reasonably well from 
<taxonomicName id="43BBCF88235DFFCBFB55FC3AC918B3F6" box="[1169,1322,964,989]" class="Gastropoda" family="Bithyniidae" genus="Gabbia" kingdom="Animalia" order="Littorinimorpha" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="campicola">
<emphasis id="B6CF6819235DFFCBFB55FC3AC918B3F6" box="[1169,1322,964,989]" italics="true" pageId="48" pageNumber="49">G. campicola</emphasis>
</taxonomicName>
and other similar taxa (
<figureCitation id="1C80A88E235DFFCBFDD2FC12CEB4B42D" box="[534,646,1004,1030]" captionStart="FIGURE 10" captionStartId="33.[264,368,1428,1452]" captionTargetBox="[287,1305,329,1414]" captionTargetId="figure-211@33.[280,1306,321,1424]" captionTargetPageId="33" captionText="FIGURE 10. A ­ Plot of the shell width/shell length (SWSH) against length/number of teleoconch whorls (SHTW) for two geographic sets of samples of G. vertiginosa. x – Sydney area; O – New England area. B ­ D ­ Plots of the canonical scores obtained from a discriminant function analysis using all shell measurements and whorl counts. B, x – G. iredalei; O – G. campicola; + ­ G. vertiginosa. C, x – G. aff. vertiginosa; O – G. campicola; + ­ G. iredalei; ­ G. aff. iredalei; ­ G. vertiginosa. D, x – G. davisi; O – G. campicola; + ­ G. fontana; ­ G. pallidula; ­ G. rotunda;" figureDoi="http://doi.org/10.5281/zenodo.5019130" httpUri="https://zenodo.org/record/5019130/files/figure.png" pageId="48" pageNumber="49">Figs 10D</figureCitation>
, 
<figureCitation id="1C80A88E235DFFCBFD56FC12CEDBB42D" box="[658,745,1004,1030]" captionStart="FIGURE 17" captionStartId="46.[264,368,1315,1339]" captionTargetBox="[309,1284,320,1293]" captionTargetId="figure-186@46.[302,1285,313,1305]" captionText="FIGURE 17. A ­ D, ­ Plots of the canonical scores obtained from a discriminant function analysis using all shell measurements and whorl counts. A, x – G. fontana; O – G. davisi; + ­ G. pallidula. B, x – G. obesa; O – G. smithii; + ­ G. tumida. C, x – G. fontana; O – G. davisi; + ­ G. obesa; ­ G. pallidula; ­ G. tumida. D, x – G. smithii; O – G. napierensis; + ­ G. tumida." figureDoi="http://doi.org/10.5281/zenodo.5097817" httpUri="https://zenodo.org/record/5097817/files/figure.png" pageId="48" pageNumber="49">17A, C</figureCitation>
; 
<tableCitation id="C93981B0235DFFCBFD33FC12CF58B42D" box="[759,874,1004,1030]" captionStart="TABLE 10" captionStartId="48.[264,352,1090,1114]" captionTargetPageId="48" captionText="TABLE 10. Classification matrix (cases in row categories classified into columns) resulting from a discriminant function analysis using shell dimensions and whorl counts for G. campicola, G. davisi, G. fontana, G. pallidula, G. rotunda and G. cf. rotunda." httpUri="http://table.plazi.org/id/D0C4E483235DFFCBFECCFBBCCF7FB488" pageId="48" pageNumber="49" tableUuid="D0C4E483235DFFCBFECCFBBCCF7FB488">Tables 10</tableCitation>
, 11).
</paragraph>
</subSubSection>
</treatment>
</document>